485 lines
58 KiB
XML
485 lines
58 KiB
XML
<document id="E9E0D670AE44FF42FAD8700F00D00422" ID-DOI="10.11646/zootaxa.3986.2.4" ID-GBIF-Dataset="99ab0619-d0df-4186-a2be-5198484ca988" ID-ISSN="1175-5326" ID-Zenodo-Dep="245930" ID-ZooBank="02CCFD53-C564-4DEF-86B5-21B0951768EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461262132840" checkinUser="plazi" docAuthor="Hoese, Douglass F. & Allen, Gerald R." docDate="2015" docId="03C28793FFCFFFB1FF4CFF4145A6F86A" docLanguage="en" docName="zt03986p216.pdf" docOrigin="Zootaxa 3986 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glossogobius macrocephalus Hoese & Allen, 2015, n. sp." docType="treatment" docUuid="27059EAC-D85F-4A0E-BFE8-AB42F47C8BA3" docUuidSource="ZooBank" docVersion="9" lastPageNumber="207" masterDocId="FFFBFFEBFFCDFFB7FFDBFFD64417FF8C" masterDocTitle="Descriptions of three new species of Glossogobius (Teleostei: Gobiidae) from New Guinea" masterLastPageNumber="216" masterPageNumber="201" pageNumber="203" updateTime="1698609731161" updateUser="plazi">
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<mods:title id="F437EBEE4C06D481894A34DF9F3079B4">Descriptions of three new species of Glossogobius (Teleostei: Gobiidae) from New Guinea</mods:title>
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<mods:namePart id="561A244EC2812E53DB498C627CEED133">Hoese, Douglass F.</mods:namePart>
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<mods:namePart id="209B44E64B43D3EC8AF6F0AFE590B312">Allen, Gerald R.</mods:namePart>
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<mods:title id="22D79E5AB456C0193BF0F4348C9FAF8C">Zootaxa</mods:title>
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<mods:part id="9B6A585C38883324DC70C5E406E08884">
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<mods:date id="2CD902A2E9736DB5AE39198DC5AAFBBE">2015</mods:date>
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<mods:number id="71BF82C0B6B419C1D92561863C1AB629">3986</mods:number>
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<mods:classification id="CFF3734D228F90C2AB5CBF86009837BB">journal article</mods:classification>
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<mods:identifier id="BD54E7DD5811166C9151ECBE665A1313" type="DOI">10.11646/zootaxa.3986.2.4</mods:identifier>
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<treatment id="03C28793FFCFFFB1FF4CFF4145A6F86A" ID-DOI="http://doi.org/10.5281/zenodo.6093993" ID-GBIF-Taxon="119637134" ID-Zenodo-Dep="6093993" LSID="urn:lsid:zoobank.org:act:27059EAC-D85F-4A0E-BFE8-AB42F47C8BA3" httpUri="http://treatment.plazi.org/id/03C28793FFCFFFB1FF4CFF4145A6F86A" lastPageId="6" lastPageNumber="207" pageId="2" pageNumber="203">
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<subSubSection id="C371650EFFCFFFB5FF4CFF4147B2FEB0" pageId="2" pageNumber="203" type="nomenclature">
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<paragraph id="8BD43685FFCFFFB5FF4CFF414656FF3E" blockId="2.[151,577,151,247]" box="[151,577,151,178]" pageId="2" pageNumber="203">
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<heading id="D09C81E9FFCFFFB5FF4CFF414656FF3E" bold="true" box="[151,577,151,178]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
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<emphasis id="B91FEA97FFCFFFB5FF4CFF414656FF3E" bold="true" box="[151,577,151,178]" pageId="2" pageNumber="203">
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<taxonomicName id="4C6B4D06FFCFFFB5FF4CFF4145E0FF3D" box="[151,503,151,177]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="macrocephalus" status="sp. nov.">
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<emphasis id="B91FEA97FFCFFFB5FF4CFF4145E0FF3D" bold="true" box="[151,503,151,177]" italics="true" pageId="2" pageNumber="203">Glossogobius macrocephalus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22C57ECFFCFFFB5FE25FF4E4656FF3E" box="[510,577,152,178]" pageId="2" pageNumber="203" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD43685FFCFFFB5FF4CFF6C452FFF7B" blockId="2.[151,577,151,247]" pageId="2" pageNumber="203">
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<figureCitation id="13502A00FFCFFFB5FF4CFF6C44F3FF5F" box="[151,228,186,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1954,1976]" captionTargetBox="[309,1256,221,1906]" captionTargetId="figure@4.[294,1292,194,1933]" captionTargetPageId="4" captionText="FIGURE 2. Sensory papillae of three species of Glossogobius: a. Glossogobius macrocephalus, based on holotype WAMP. 26972 – 003 and paratype WAMP. 26972 – 003; b. Glossogobius multipapillus, composite based on holotype and several Paratypes; c. Glossogobius sentaniensis, based largely on holotype, WAMP. 27852 – 004." httpUri="https://zenodo.org/record/245932/files/figure.png" pageId="2" pageNumber="203">Figs. 2</figureCitation>
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a, 3, 4,
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<tableCitation id="C6E9033EFFCFFFB5FEEEFF6C45B8FF5F" box="[309,431,186,211]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStartId-0="2.[151,239,1621,1644]" captionStartId-1="3.[151,239,152,175]" captionStartId-2="3.[151,239,538,561]" captionStartId-3="3.[151,239,925,948]" captionStartId-4="3.[151,239,1312,1335]" captionStartId-5="3.[151,239,1699,1722]" captionText-0="TABLE 1. Second dorsal fin-ray counts of 3 species of Glossogobius." captionText-1="TABLE 2. Anal fin-ray counts of 3 species of Glossogobius." captionText-2="TABLE 3. Pectoral fin-ray counts of 3 species of Glossogobius." captionText-3="TABLE 4. Predorsal scale counts of 3 species of Glossogobius." captionText-4="TABLE 5. Longitudinal scale count for three species of Glossogobius." captionText-5="TABLE 6. Transvers scale count for three species of Glossogobius." pageId="2" pageNumber="203">Tables 1–6</tableCitation>
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Bighead Goby
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</paragraph>
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||
<paragraph id="8BD43685FFCFFFB5FF4CFEF047B2FEB0" blockId="2.[151,933,294,316]" box="[151,933,294,316]" pageId="2" pageNumber="203">
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<treatmentCitationGroup id="AB7B11ABFFCFFFB5FF4CFEF047B2FEB0" box="[151,933,294,316]" pageId="2" pageNumber="203">
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<taxonomicName id="4C6B4D06FFCFFFB5FF4CFEF04536FEB0" ID-CoL="4P99" box="[151,289,294,316]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="genus">
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<emphasis id="B91FEA97FFCFFFB5FF4CFEF04536FEB0" box="[151,289,294,316]" italics="true" pageId="2" pageNumber="203">Glossogobius</emphasis>
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</taxonomicName>
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sp. 13. —
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<treatmentCitation id="0ACA1094FFCFFFB5FE4BFEF0462BFEB0" author="Allen" box="[400,572,294,316]" page="187" pageId="2" pageNumber="203" year="1991">
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<bibRefCitation id="EFFA4B74FFCFFFB5FE4BFEF0461DFEB0" author="Allen" box="[400,522,294,316]" pageId="2" pageNumber="203" refString="Allen, G. R. (1991) Field Guide to the Freshwater Fishes of New Guinea. Publication No. 9. Christensen Research Institute, Madang, 268 pp." type="book" year="1991">Allen, 1991</bibRefCitation>
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: 187
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</treatmentCitation>
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, pl. 15, fig. 3 (
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<collectingCountry id="F37C7615FFCFFFB5FD0BFEF0478DFEB0" box="[720,922,294,316]" name="Papua New Guinea" pageId="2" pageNumber="203">Papua New Guinea</collectingCountry>
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).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C371650EFFCFFFB5FF4CFEBF44DFFD9D" pageId="2" pageNumber="203" type="materials_examined">
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<paragraph id="8BD43685FFCFFFB5FF4CFEBF45A7FE2A" blockId="2.[151,1437,361,1574]" pageId="2" pageNumber="203">
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<emphasis id="B91FEA97FFCFFFB5FF4CFEBF451CFE0D" bold="true" box="[151,267,361,385]" pageId="2" pageNumber="203">
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||
<typeStatus id="54D08827FFCFFFB5FF4CFEBF4511FE0D" box="[151,262,361,385]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED7AAE40FFCFFFB5FEC9FEBF4540FE0D" LSID="urn:lsid:biocol.org:col:34981" box="[274,343,361,385]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="203">WAM</collectionCode>
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P.26972–004,
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<quantity id="4C939B60FFCFFFB5FE25FEBF4659FE0D" box="[510,590,361,385]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="203" unit="mm" value="73.0">73 mm</quantity>
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SL male, creek
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<quantity id="4C939B60FFCFFFB5FCDDFEBF4757FE0E" box="[774,832,361,386]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="203" unit="km" value="4.0">4 km</quantity>
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E of Lake Tebera (
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||
<geoCoordinate id="EE5F5042FFCFFFB5FBC1FEBF4070FE0E" box="[1050,1127,361,386]" direction="south" orientation="latitude" pageId="2" pageNumber="203" precision="925" value="-6.75">6°45'S</geoCoordinate>
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,
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||
<geoCoordinate id="EE5F5042FFCFFFB5FBA8FEBF40CAFE0D" box="[1139,1245,361,385]" direction="east" orientation="longitude" pageId="2" pageNumber="203" precision="925" value="144.65">144°39'E</geoCoordinate>
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),
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||
<date id="FFD51045FFCFFFB5FB29FEBF4145FE0D" box="[1266,1362,361,385]" pageId="2" pageNumber="203" value="1924-09">24 Sept.</date>
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||
, 1980, G. Allen & B. Parkinson.
|
||
</paragraph>
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||
<paragraph id="8BD43685FFCFFFB5FF1CFE6744DFFD9D" blockId="2.[151,1437,361,1574]" pageId="2" pageNumber="203">
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||
<emphasis id="B91FEA97FFCFFFB5FF1CFE674551FE45" bold="true" box="[199,326,433,457]" pageId="2" pageNumber="203">
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<typeStatus id="54D08827FFCFFFB5FF1CFE674556FE45" box="[199,321,433,457]" pageId="2" pageNumber="203" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED7AAE40FFCFFFB5FE8EFE67458DFE45" LSID="urn:lsid:biocol.org:col:34981" box="[341,410,433,457]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="203">WAM</collectionCode>
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P.26972–003, 2(59–66), taken with
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<typeStatus id="54D08827FFCFFFB5FC85FE6747D3FE46" box="[862,964,433,458]" pageId="2" pageNumber="203" type="holotype">holotype</typeStatus>
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||
;
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<collectionCode id="ED7AAE40FFCFFFB5FC03FE67400BFE45" LSID="urn:lsid:biocol.org:col:34981" box="[984,1052,433,457]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="203">WAM</collectionCode>
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P.30383–003, 2(23–112), small tributary creek and Lake Tebera,
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<date id="FFD51045FFCFFFB5FDCBFE0046B1FE62" box="[528,678,470,494]" pageId="2" pageNumber="203" value="1991-10-19">19 Oct. 1991</date>
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, G. Allen; AMS I.46502–001, 1(86), same data as
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<collectionCode id="ED7AAE40FFCFFFB5FB33FE00413AFE62" LSID="urn:lsid:biocol.org:col:34981" box="[1256,1325,470,494]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="203">WAM</collectionCode>
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P.30383– 0 0 3.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C371650EFFCFFFB5FF1CFDCA471BFA1A" pageId="2" pageNumber="203" type="diagnosis">
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<paragraph id="8BD43685FFCFFFB5FF1CFDCA4799FC4E" blockId="2.[151,1437,361,1574]" pageId="2" pageNumber="203">
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<emphasis id="B91FEA97FFCFFFB5FF1CFDCA452DFDB9" bold="true" box="[199,314,540,565]" pageId="2" pageNumber="203">Diagnosis</emphasis>
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. Preoperculum without a spine, but with small flat projection adjacent to lower preopercular pore. Mental fraenum low with short lateral lobes. Mouth small; reaching to below anterior margin of eyes; jaws forming an angle of 40–50° with body axis; upper margin of upper jaw in line with lower margin of pupil. Head robust and large, cheeks slightly bulbose and tapering. Interorbital narrow, less than eye length. Predorsal area scaled forward to just behind eye, with a narrow naked patch between and behind eyes. Cheek naked. Operculum naked. Pectoral base with 1–3 vertical rows of scales. Prepelvic area about three-quarters scaled in 7–8 rows, naked anteriorly. Belly largely scaled, with a very small naked patch behind pelvic insertion. First dorsal fin spine 3–5 extending beyond other spines when fin depressed. First dorsal fin with black spot posteriorly on membrane surrounding sixth dorsal spine, spot faint in female. Pelvic fin thin, length much greater than width, reaching anus. Second dorsal rays I,9–10; anal rays I,8; pectoral rays 17–18; longitudinal scale count 30–31; predorsal scale count 15–18; transverse scale count (TRB) 9.5–10.5; gill rakers on outer face of first arch 1+1+6–8=8–10; lower gill rakers on outer face of second arch 6–7; branched caudal rays 14–15; vertebrae 12+16=28.
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</paragraph>
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<paragraph id="8BD43685FFCFFFB5FF1CFC1B471BFA1A" blockId="2.[151,1437,361,1574]" pageId="2" pageNumber="203">
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<emphasis id="B91FEA97FFCFFFB5FF1CFC1B4531FC69" box="[199,294,973,997]" italics="true" pageId="2" pageNumber="203">Papillae</emphasis>
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: Line 1 (before nasal pore) composed of 2 rows of papillae (sometimes incomplete). Line 2 (between nasal pores) composed of a single row of papillae usually continuous across snout. Line 5 (suborbital) composed of single row of papillae (double in one specimens on one side of head below infraorbital pore), joining line 7 below posterior nostril. Line 6 (suborbital branch) absent. Lines 7, 9, 10 (LL cheek rows) composed of a single row of papillae, but irregular anteriorly appearing double, reaching to posterior preopercular margin. Lines 8 and 11 (LT row) reaching just short of posterior preopercular margin. Line 12 (Outer POP–mandibular) composed of single row of papillae, appearing double along lips, without gap at end of lips. Line 13 (Inner POP–mandibular) composed of 2–3 rows of papillae. Line 20 (opercular VT row) composed of single row of papillae dorsally, and double ventrally. Line 21 (Upper opercular OT) composed of single row of papillae, strongly curved ventrally meeting or almost meeting line 22. Line 22 (Lower opercular OT) composed of single row of papillae, curving upward along posterior opercular margin. Several vertical papillae rows on belly. A single curved line anteriorly on most body scales (often obscure dorsally and posteriorly); chin with an oval patch of papillae behind mental fraenum, with a notch posteromedially without papillae.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C371650EFFCFFFB1FF1CFA7740BCFB89" lastPageId="6" lastPageNumber="207" pageId="2" pageNumber="203" type="description">
|
||
<paragraph id="8BD43685FFCFFFB5FF1CFA774050F9AA" blockId="2.[151,1437,361,1574]" pageId="2" pageNumber="203">
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||
<emphasis id="B91FEA97FFCFFFB5FF1CFA774546FA36" bold="true" box="[199,337,1441,1466]" pageId="2" pageNumber="203">Description</emphasis>
|
||
. Based on
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<specimenCount id="9D6DFD0CFFCFFFB5FE09FA774649FA36" box="[466,606,1441,1466]" pageId="2" pageNumber="203" type="generic">6 specimens</specimenCount>
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||
<quantity id="4C939B60FFCFFFB5FDBDFA7446FAFA35" box="[614,749,1442,1466]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="11.2" metricValueMin="2.3" pageId="2" pageNumber="203" unit="mm" value="67.5" valueMax="112.0" valueMin="23.0">23–112 mm</quantity>
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SL, data not taken on smallest. Count of
|
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<typeStatus id="54D08827FFCFFFB5FB10FA77413AFA36" box="[1227,1325,1441,1466]" pageId="2" pageNumber="203" type="holotype">holotype</typeStatus>
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indicated with an asterisk, where a range of counts exists. First dorsal spines 6(5*); gill rakers on outer face of first arch 0+1+7(1), 1+1+6(1). 1+1+7(2), 1+1+8(1); segmented caudal rays 9/8(5*); branched caudal rays 7/7 (2), 7/8(1), 8/ 7(2*); vertebrae 12+16(3*). Counts of other meristic characters given in
|
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<tableCitation id="C6E9033EFFCFFFB5FC1CF9DB4054F9AA" box="[967,1091,1549,1574]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStartId-0="2.[151,239,1621,1644]" captionStartId-1="3.[151,239,152,175]" captionStartId-2="3.[151,239,538,561]" captionStartId-3="3.[151,239,925,948]" captionStartId-4="3.[151,239,1312,1335]" captionStartId-5="3.[151,239,1699,1722]" captionText-0="TABLE 1. Second dorsal fin-ray counts of 3 species of Glossogobius." captionText-1="TABLE 2. Anal fin-ray counts of 3 species of Glossogobius." captionText-2="TABLE 3. Pectoral fin-ray counts of 3 species of Glossogobius." captionText-3="TABLE 4. Predorsal scale counts of 3 species of Glossogobius." captionText-4="TABLE 5. Longitudinal scale count for three species of Glossogobius." captionText-5="TABLE 6. Transvers scale count for three species of Glossogobius." pageId="2" pageNumber="203">Tables 1–6</tableCitation>
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||
.
|
||
</paragraph>
|
||
<caption id="DF14660DFFCFFFB5FF4CF9834795F9E0" box="[151,898,1620,1644]" pageId="2" pageNumber="203">
|
||
<paragraph id="8BD43685FFCFFFB5FF4CF9834795F9E0" blockId="2.[151,898,1620,1644]" box="[151,898,1620,1644]" pageId="2" pageNumber="203">
|
||
<emphasis id="B91FEA97FFCFFFB5FF4CF983451DF9E0" bold="true" box="[151,266,1621,1644]" pageId="2" pageNumber="203">TABLE 1.</emphasis>
|
||
Second dorsal fin-ray counts of 3 species of
|
||
<taxonomicName id="4C6B4D06FFCFFFB5FD31F982476CF9E0" box="[746,891,1620,1644]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCFFFB5FD31F982476CF9E0" box="[746,891,1620,1644]" italics="true" pageId="2" pageNumber="203">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCFFFB5FF44F95E4784F912" blockId="2.[159,1307,1672,1943]" box="[159,915,1672,1694]" pageId="2" pageNumber="203">Species Locality Second dorsal fin</paragraph>
|
||
<paragraph id="8BD43685FFCFFFB4FD39F964473FFF6C" blockId="2.[159,1307,1672,1943]" lastBlockId="3.[159,1294,202,473]" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
|
||
I,
|
||
<date id="FFD51045FFCFFFB5FD2BF964476BF94B" box="[752,892,1713,1735]" pageId="2" pageNumber="203">7 I</date>
|
||
,
|
||
<date id="FFD51045FFCFFFB5FC5AF967405BF94B" box="[897,1100,1713,1735]" pageId="2" pageNumber="203">8 I</date>
|
||
,
|
||
<date id="FFD51045FFCFFFB5FB8BF96740ECF94B" box="[1104,1275,1713,1735]" pageId="2" pageNumber="203">9 I</date>
|
||
,10
|
||
<taxonomicName id="4C6B4D06FFCFFFB5FF44F90D4542F97C" box="[159,341,1754,1776]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCFFFB5FF44F90D4542F97C" box="[159,341,1754,1776]" italics="true" pageId="2" pageNumber="203">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- - 2 3*
|
||
<taxonomicName id="4C6B4D06FFCFFFB5FF44F8D3463CF896" authority="Safia" authorityName="Safia" box="[159,555,1796,1818]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCFFFB5FF44F8D34556F896" box="[159,321,1796,1818]" italics="true" pageId="2" pageNumber="203">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
1 27 - - Popondetta 1 22* 4 - Kokoda - 12 - -
|
||
<taxonomicName id="4C6B4D06FFCFFFB5FF44F8544520F81B" box="[159,311,1922,1943]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCFFFB5FF44F8544520F81B" box="[159,311,1922,1943]" italics="true" pageId="2" pageNumber="203">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- 27* - - Species Locality Analfin
|
||
</paragraph>
|
||
<caption id="DF14660DFFCEFFB4FF4CFF4E4737FF23" box="[151,800,151,175]" pageId="3" pageNumber="204">
|
||
<paragraph id="8BD43685FFCEFFB4FF4CFF4E4737FF23" blockId="3.[151,800,151,175]" box="[151,800,151,175]" pageId="3" pageNumber="204">
|
||
<emphasis id="B91FEA97FFCEFFB4FF4CFF4E451EFF23" bold="true" box="[151,265,152,175]" pageId="3" pageNumber="204">TABLE 2.</emphasis>
|
||
Anal fin-ray counts of 3 species of
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FD53FF41470EFF23" box="[648,793,151,175]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCEFFB4FD53FF41470EFF23" box="[648,793,151,175]" italics="true" pageId="3" pageNumber="204">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCEFFB4FD0FFF2240BCFDEF" blockId="3.[159,1294,202,473]" lastBlockId="3.[159,1321,589,860]" pageId="3" pageNumber="204">
|
||
I.
|
||
<date id="FFD51045FFCEFFB4FD39FF224784FE86" box="[738,915,244,266]" pageId="3" pageNumber="204">6 I</date>
|
||
,
|
||
<date id="FFD51045FFCEFFB4FC4CFF234050FE86" box="[919,1095,244,266]" pageId="3" pageNumber="204">7 I</date>
|
||
,
|
||
<date id="FFD51045FFCEFFB4FB97FF2240EBFE86" box="[1100,1276,244,266]" pageId="3" pageNumber="204">8 I</date>
|
||
,9
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FEC84542FEBF" box="[159,341,285,307]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FEC84542FEBF" box="[159,341,285,307]" italics="true" pageId="3" pageNumber="204">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- - 5* -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FE91460FFED0" authority="Safia" authorityName="Safia" box="[159,536,326,348]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FE914556FED0" box="[159,321,326,348]" italics="true" pageId="3" pageNumber="204">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
<date id="FFD51045FFCEFFB4FD0FFE90405DFED0" box="[724,1098,326,348]" pageId="3" pageNumber="204" value="1928-01-01">1 28 1</date>
|
||
- Popondetta - 25* 2 - Kokoda 1 11 -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FE124520FE55" box="[159,311,452,473]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FE124520FE55" box="[159,311,452,473]" italics="true" pageId="3" pageNumber="204">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- - 27* 2 Species Locality Pectoral ray counts
|
||
</paragraph>
|
||
<caption id="DF14660DFFCEFFB4FF4CFDCC4754FDBD" box="[151,835,537,561]" pageId="3" pageNumber="204">
|
||
<paragraph id="8BD43685FFCEFFB4FF4CFDCC4754FDBD" blockId="3.[151,835,537,561]" box="[151,835,537,561]" pageId="3" pageNumber="204">
|
||
<emphasis id="B91FEA97FFCEFFB4FF4CFDCC451EFDBD" bold="true" box="[151,265,538,561]" pageId="3" pageNumber="204">TABLE 3.</emphasis>
|
||
Pectoral fin-ray counts of 3 species of
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FD70FDCF472BFDBD" box="[683,828,537,561]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCEFFB4FD70FDCF472BFDBD" box="[683,828,537,561]" italics="true" pageId="3" pageNumber="204">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCEFFB4FD19FDA046DEFC6A" blockId="3.[159,1321,589,860]" lastBlockId="3.[159,1406,976,1247]" pageId="3" pageNumber="204">
|
||
16 17 18 19 20
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FD774542FD3A" box="[159,341,672,694]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FD774542FD3A" box="[159,341,672,694]" italics="true" pageId="3" pageNumber="204">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- 4* 1 - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FD1C4626FD53" authority="Safia" authorityName="Safia" box="[159,561,713,735]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FD1C4556FD53" box="[159,321,713,735]" italics="true" pageId="3" pageNumber="204">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
<date id="FFD51045FFCEFFB4FD19FD1F47E1FD53" box="[706,1014,713,735]" pageId="3" pageNumber="204" value="1921-08-01">8 21 1</date>
|
||
- - Popondetta - 1 19* 6 - Kokoda - 2 9 1 -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FC914520FCD0" box="[159,311,839,860]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FC914520FCD0" box="[159,311,839,860]" italics="true" pageId="3" pageNumber="204">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- - 3 20* 5 Species Locality Predorsal scales
|
||
</paragraph>
|
||
<caption id="DF14660DFFCEFFB4FF4CFC4B472AFC38" box="[151,829,924,948]" pageId="3" pageNumber="204">
|
||
<paragraph id="8BD43685FFCEFFB4FF4CFC4B472AFC38" blockId="3.[151,829,924,948]" box="[151,829,924,948]" pageId="3" pageNumber="204">
|
||
<emphasis id="B91FEA97FFCEFFB4FF4CFC4B451EFC38" bold="true" box="[151,265,925,948]" pageId="3" pageNumber="204">TABLE 4.</emphasis>
|
||
Predorsal scale counts of 3 species of
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FD7EFC4A4721FC38" box="[677,822,924,948]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCEFFB4FD7EFC4A4721FC38" box="[677,822,924,948]" italics="true" pageId="3" pageNumber="204">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCEFFB4FDFCFC2F4071FAE5" blockId="3.[159,1406,976,1247]" lastBlockId="3.[159,1387,1363,1634]" pageId="3" pageNumber="204">
|
||
<date id="FFD51045FFCEFFB4FDFCFC2F46D7FB83" box="[551,704,1017,1039]" pageId="3" pageNumber="204" value="2014-12-13">12 13 14</date>
|
||
15 16 17 18 25 26 27 28 29 30 31
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FBF54542FBB5" box="[159,341,1059,1081]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FBF54542FBB5" box="[159,341,1059,1081]" italics="true" pageId="3" pageNumber="204">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- - - 1 1 2* 1 - - - - - - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FB9B45A1FBEE" authority="Safia" authorityName="Safia" box="[159,438,1100,1122]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FB9B4556FBEE" box="[159,321,1100,1122]" italics="true" pageId="3" pageNumber="204">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
4 3 6 2 - - - - - - - - - - Popondetta 1 3 5 2 - - - - - - - - - - Kokoda 1 4 3 4 1 - - - - - - - - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FB1C4520FB53" box="[159,311,1226,1247]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FB1C4520FB53" box="[159,311,1226,1247]" italics="true" pageId="3" pageNumber="204">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- - - - - - - 2 - 3 8* 2 5 2 Species Locality Longitudinal Scale Count
|
||
</paragraph>
|
||
<caption id="DF14660DFFCEFFB4FF4CFAF6479EFABB" box="[151,905,1311,1335]" pageId="3" pageNumber="204">
|
||
<paragraph id="8BD43685FFCEFFB4FF4CFAF6479EFABB" blockId="3.[151,905,1311,1335]" box="[151,905,1311,1335]" pageId="3" pageNumber="204">
|
||
<emphasis id="B91FEA97FFCEFFB4FF4CFAF6451DFABB" bold="true" box="[151,266,1312,1335]" pageId="3" pageNumber="204">TABLE 5.</emphasis>
|
||
Longitudinal scale count for three species of
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FD2AFAC94795FABB" box="[753,898,1311,1335]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCEFFB4FD2AFAC94795FABB" box="[753,898,1311,1335]" italics="true" pageId="3" pageNumber="204">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCEFFB4FDCCFAAA4050F967" blockId="3.[159,1387,1363,1634]" lastBlockId="3.[159,1331,1749,2021]" pageId="3" pageNumber="204">
|
||
25 26 27 28 29 30 31 32 33 34 35
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FA704542FA37" box="[159,341,1445,1467]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FA704542FA37" box="[159,341,1445,1467]" italics="true" pageId="3" pageNumber="204">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- - - - 1 3* 1 - - - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44FA0645ADFA69" authority="Safia" authorityName="Safia" box="[159,442,1487,1509]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44FA064556FA69" box="[159,321,1487,1509]" italics="true" pageId="3" pageNumber="204">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
1 5 6 - - - - - - - Popondetta -
|
||
<date id="FFD51045FFCEFFB4FDB1FA2E470BF982" box="[618,796,1528,1550]" pageId="3" pageNumber="204" value="2010-03-02">3 10 2</date>
|
||
- - - - - - - Kokoda 1 3 7 1 - - - - - - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44F99A4520F9EE" box="[159,311,1612,1634]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44F99A4520F9EE" box="[159,311,1612,1634]" italics="true" pageId="3" pageNumber="204">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- - - - - - 4 6 5 3* 1 TransverseScaleCount
|
||
</paragraph>
|
||
<caption id="DF14660DFFCEFFB4FF4CF9754770F936" box="[151,871,1698,1722]" pageId="3" pageNumber="204">
|
||
<paragraph id="8BD43685FFCEFFB4FF4CF9754770F936" blockId="3.[151,871,1698,1722]" box="[151,871,1698,1722]" pageId="3" pageNumber="204">
|
||
<emphasis id="B91FEA97FFCEFFB4FF4CF975451DF936" bold="true" box="[151,266,1699,1722]" pageId="3" pageNumber="204">TABLE 6.</emphasis>
|
||
Transvers scale count for three species of
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FD14F9744777F936" box="[719,864,1698,1722]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCEFFB4FD14F9744777F936" box="[719,864,1698,1722]" italics="true" pageId="3" pageNumber="204">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFCEFFB2FF44F92945A0F8C2" blockId="3.[159,1331,1749,2021]" lastBlockId="5.[151,1437,1269,2014]" lastPageId="5" lastPageNumber="206" pageId="3" pageNumber="204">
|
||
Species Locality 8.5 9.5 10.5 11.5 12.5
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44F8FF4542F8B2" box="[159,341,1832,1854]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44F8FF4542F8B2" box="[159,341,1832,1854]" italics="true" pageId="3" pageNumber="204">G. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
- 3* 2 - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44F88445CAF8E4" authority="Safia" authorityName="Safia" box="[159,477,1874,1896]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44F8844556F8E4" box="[159,321,1874,1896]" italics="true" pageId="3" pageNumber="204">G. multipapillus</emphasis>
|
||
Safia
|
||
</taxonomicName>
|
||
15 - - - - Popondetta 7* 3 - - - Kokoda - - - - -
|
||
<taxonomicName id="4C6B4D06FFCEFFB4FF44F8194520F869" box="[159,311,1999,2021]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="sentaniensis">
|
||
<emphasis id="B91FEA97FFCEFFB4FF44F8194520F869" box="[159,311,1999,2021]" italics="true" pageId="3" pageNumber="204">G. sentaniensis</emphasis>
|
||
</taxonomicName>
|
||
- - - 13* 9 Head large, slightly depressed, length 32.9–34.9% SL, depth 17.3–18.3% SL, width 18.7–21.2% SL. Snout rounded in dorsal view, straight in side view; 9.4–11.7% SL. Eye about
|
||
<quantity id="4C939B60FFC8FFB2FC6EFACF4032FABE" box="[949,1061,1305,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="5" pageNumber="206" unit="in" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 in</quantity>
|
||
snout, 6.2–8% SL (6.2–7% SL in specimens over
|
||
<quantity id="4C939B60FFC8FFB2FE8AFAE845B4FADA" box="[337,419,1342,1366]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="206" unit="mm" value="70.0">70 mm</quantity>
|
||
SL). Small bump below anterior nostril present. Anterior nostril at end of short tube, 1 nostril diameter above upper lip. Posterior nostril a pore, midway between eye and upper lip, 3–4 nostril diameters from eye and 1–2 from anterior nostril. Preoperculum short, distance from end of eye to upper posterior preopercular margin slightly less than eye length. Postorbital long, subequal to distance from tip of snout to posterior half of eye. Gill opening reaching to below posterior preopercular margin. Teeth in upper jaw: outer row of teeth conical, slightly enlarged and wideset, 3 inner rows of smaller depressible, inwardly directed teeth, innermost row larger than middle rows. Teeth in lower jaw: teeth in outer row conical, slightly enlarged and wideset anteriorly, 2 inner rows of smaller depressible teeth, teeth in innermost row larger than teeth in middle row. Tongue tip bilobed. Gill rakers on outer face of first arch short and broad, one–third to one–quarter filament length. Rakers on inner face of first arch and other arches short and denticulate. Body covered mostly with large ctenoid scales, cycloid predorsal area, pectoral base, prepelvic area and midline of belly. First dorsal fin round to triangular shaped. Second dorsal fin subequal in height to first dorsal fin. Anal fin slightly lower than dorsal fins. Pectoral fin with pointed margin, reaching to above a point just before anus to anal origin. Pelvic disc thin, length greater than width, reaching about ¾ of way to anus or just reaching anus; fifth ray with 10 terminal tips of rays at posterior margin of fin in
|
||
<typeStatus id="54D08827FFC8FFB2FE95F8E345A3F8C2" box="[334,436,1845,1870]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF14660DFFC9FFB3FF4CF8744006F87A" httpUri="https://zenodo.org/record/245932/files/figure.png" pageId="4" pageNumber="205" targetBox="[309,1256,221,1906]" targetPageId="4">
|
||
<paragraph id="8BD43685FFC9FFB3FF4CF8744006F87A" blockId="4.[151,1436,1954,2038]" pageId="4" pageNumber="205">
|
||
<emphasis id="B91FEA97FFC9FFB3FF4CF874450CF83B" bold="true" box="[151,283,1954,1976]" pageId="4" pageNumber="205">FIGURE 2.</emphasis>
|
||
Sensory papillae of three species of
|
||
<taxonomicName id="4C6B4D06FFC9FFB3FD10F8744742F834" box="[715,853,1954,1976]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFC9FFB3FD10F8744742F834" box="[715,853,1954,1976]" italics="true" pageId="4" pageNumber="205">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
: a.
|
||
<taxonomicName id="4C6B4D06FFC9FFB3FC50F87440A9F834" box="[907,1214,1954,1976]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFC9FFB3FC50F87440A9F834" box="[907,1214,1954,1976]" italics="true" pageId="4" pageNumber="205">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
, based on holotype WAMP.26972–003 and paratype WAMP.26972–003; b.
|
||
<taxonomicName id="4C6B4D06FFC9FFB3FD0FF81747FCF85B" box="[724,1003,1985,2007]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="multipapillus">
|
||
<emphasis id="B91FEA97FFC9FFB3FD0FF81747FCF85B" box="[724,1003,1985,2007]" italics="true" pageId="4" pageNumber="205">Glossogobius multipapillus</emphasis>
|
||
</taxonomicName>
|
||
, composite based on holotype and several Paratypes; c.
|
||
<emphasis id="B91FEA97FFC9FFB3FEC5F8364626F87A" box="[286,561,2016,2038]" italics="true" pageId="4" pageNumber="205">
|
||
<taxonomicName id="4C6B4D06FFC9FFB3FEC5F836463AF87A" box="[286,557,2016,2038]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="sentaniensis">Glossogobius sentaniensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
based largely on holotype, WAMP.27852–004.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF14660DFFC8FFB2FF4CFD57405AFD1B" box="[151,1101,641,663]" httpUri="https://zenodo.org/record/245933/files/figure.png" pageId="5" pageNumber="206" targetBox="[151,1436,193,619]" targetPageId="5">
|
||
<paragraph id="8BD43685FFC8FFB2FF4CFD57405AFD1B" blockId="5.[151,1101,641,663]" box="[151,1101,641,663]" pageId="5" pageNumber="206">
|
||
<emphasis id="B91FEA97FFC8FFB2FF4CFD574505FD1A" bold="true" box="[151,274,641,663]" pageId="5" pageNumber="206">FIGURE 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C6B4D06FFC8FFB2FE42FD5746D4FD1B" box="[409,707,641,663]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFC8FFB2FE42FD5746D4FD1B" box="[409,707,641,663]" italics="true" pageId="5" pageNumber="206">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
, WAM P.26972–004, 73 mm SL male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF14660DFFC8FFB2FF4CFB6E4125FB42" box="[151,1330,1208,1230]" httpUri="https://zenodo.org/record/245934/files/figure.png" pageId="5" pageNumber="206" targetBox="[151,1436,712,1186]" targetPageId="5">
|
||
<paragraph id="8BD43685FFC8FFB2FF4CFB6E4125FB42" blockId="5.[151,1330,1208,1230]" box="[151,1330,1208,1230]" pageId="5" pageNumber="206">
|
||
<emphasis id="B91FEA97FFC8FFB2FF4CFB6E4505FB41" bold="true" box="[151,274,1208,1230]" pageId="5" pageNumber="206">FIGURE 4.</emphasis>
|
||
Photo of freshly collected specimen of
|
||
<taxonomicName id="4C6B4D06FFC8FFB2FD7EFB6E47D8FB42" box="[677,975,1208,1230]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFC8FFB2FD7EFB6E47D8FB42" box="[677,975,1208,1230]" italics="true" pageId="5" pageNumber="206">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
from Lake Tebera, photo G. Allen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD43685FFC8FFB1FF1CF88C44E5FF58" blockId="5.[151,1437,1269,2014]" lastBlockId="6.[151,1437,151,2022]" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206">
|
||
<emphasis id="B91FEA97FFC8FFB2FF1CF88C4558F8FD" box="[199,335,1882,1906]" italics="true" pageId="5" pageNumber="206">Head pores</emphasis>
|
||
: nasal pore before and medial posterior nostril; anterior interorbital pore present posterior interorbital pore present; postorbital pore behind eye present; infraorbital pore below postorbital present; lateral canal pore above preoperculum present; lateral canal pore above posterior preopercular margin absent; terminal lateral canal pore above anterior operculum present; short tube above operculum, with pore at each end present or represented by open trench; 3 preopercular pores, upper in line with lower margin of eye; widely separated from lower 2.
|
||
</paragraph>
|
||
<paragraph id="8BD43685FFCBFFB1FF1CFF364727FD65" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CFF3645AEFF74" box="[199,441,224,248]" italics="true" pageId="6" pageNumber="207">Coloration in alcohol</emphasis>
|
||
: Head and body brown. A dark brown bar from anteroventral margin of eye extending to upper lip; posterior end of lips whitish; an oval brown blotch at posteroventral margin of eye, followed by a small irregularly shaped dark brown blotch; an irregular dark brown blotch on cheek below anterior half of eye; sometimes a series of small dark brown spots forming a horizontal line on cheek; chin pale to faint grey, lighter than top of head; operculum with dark brown mottling. Pectoral base with an upper dark brown spot followed by a curved brown bar on bases of pectoral rays; a second dark brown spot ventrally. Body with faint brown mottling on back forming indistinct saddles; midsides with series of vertically elongate spots, usually occurring in groups of 2 spots; first group below front of first dorsal fin, second below posterior end of first dorsal fin, third below beginning of second dorsal fin, fourth below second dorsal fin origin, fifth below middle of second dorsal fin, sixth below posterior end of second dorsal fin, seventh on middle of caudal peduncle, eighth at posterior end of caudal peduncle, extending onto base of caudal fin, a triangular (apex forward) brown spot on base of caudal fin, just touching last caudal peduncle spot; individual spots in each group coalesced into single large spot in largest specimen (
|
||
<quantity id="4C939B60FFCBFFB1FED4FD5C457BFD2D" box="[271,364,650,674]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="207" unit="mm" value="110.0">110 mm</quantity>
|
||
SL). All fins grey. First dorsal fin with mottling; a black spot on membrane surrounding middle of sixth dorsal spine. Second dorsal fin with small dark brown spots forming rows. Caudal fin grey with brown spots forming 3–5 short oblique rows, fin darkest ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BD43685FFCBFFB1FF1CFD22471AFC32" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CFD224654FC80" box="[199,579,756,780]" italics="true" pageId="6" pageNumber="207">Coloration when freshly collected</emphasis>
|
||
(
|
||
<figureCitation id="13502A00FFCBFFB1FD88FD224681FC80" box="[595,662,756,781]" captionStart="FIGURE 4" captionStartId="5.[151,250,1208,1230]" captionTargetBox="[151,1436,712,1186]" captionTargetId="figure@5.[151,1436,712,1186]" captionTargetPageId="5" captionText="FIGURE 4. Photo of freshly collected specimen of Glossogobius macrocephalus from Lake Tebera, photo G. Allen." httpUri="https://zenodo.org/record/245934/files/figure.png" pageId="6" pageNumber="207">Fig. 4</figureCitation>
|
||
): Head and body yellowish tan; dark brown to blackish markings on head and pectoral-fin base similar to those described for preseserved specimens; fins translucent to slightly yellowish; first dorsal fin with mottling and reddish-yellow margin; a black spot on membrane surrounding middle of sixth dorsal spine. Second dorsal fin with brownish outer margin and small dark brown spots forming rows; caudal fin dusky yellowish brown to dark grey along posterior and ventral margins and with dark-brown spots forming 3–5 short oblique rows in central portion of fin.
|
||
</paragraph>
|
||
<paragraph id="8BD43685FFCBFFB1FF1CFC1E40BCFB89" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CFC1E4522FC6D" bold="true" box="[199,309,968,993]" pageId="6" pageNumber="207">Variation</emphasis>
|
||
: Specimens collected from Lake Tebera differ slightly from specimens in the creek in being darker, with the spotting pattern obscured and chin dark to attachment of branchiostegal membranes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371650EFFCBFFB1FF1CFBD94603F813" pageId="6" pageNumber="207" type="distribution">
|
||
<paragraph id="8BD43685FFCBFFB1FF1CFBD94543FA79" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CFBD9454EFBA4" bold="true" box="[199,345,1039,1064]" pageId="6" pageNumber="207">Distribution</emphasis>
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FEBBFBC646BEFBA4" box="[352,681,1040,1064]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCBFFB1FEBBFBC646BEFBA4" box="[352,681,1040,1064]" italics="true" pageId="6" pageNumber="207">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
is known only from the Lake Tebera basin, including the main lake and in-flowing tributaries, located in the Southern Highlands of
|
||
<collectingCountry id="F37C7615FFCBFFB1FCB3FBE24052FBC0" box="[872,1093,1075,1100]" name="Papua New Guinea" pageId="6" pageNumber="207">Papua New Guinea</collectingCountry>
|
||
. The
|
||
<typeStatus id="54D08827FFCBFFB1FB59FBE240A5FBC0" box="[1154,1202,1076,1100]" pageId="6" pageNumber="207">type</typeStatus>
|
||
locality consisted of a small stream flowing into Lake Tabera about
|
||
<quantity id="4C939B60FFCBFFB1FD1EFB814715FBE3" box="[709,770,1110,1135]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="207" unit="km" value="4.0">4 km</quantity>
|
||
east of the lake. The water was exceptionally clear with respective temperature and pH values of 21°C and 7.8 recorded at the time of collection. The main lake and stream lie at an elevation of approximately
|
||
<specimenCount id="9D6DFD0CFFCBFFB1FDEAFB4B4697FB39" box="[561,640,1181,1206]" pageId="6" pageNumber="207" type="generic">
|
||
<quantity id="4C939B60FFCBFFB1FDEAFB4B466BFB39" box="[561,636,1181,1206]" metricMagnitude="2" metricUnit="m" metricValue="7.95" pageId="6" pageNumber="207" unit="m" value="795.0">795 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Lake Tebera is situated in a narrow, elongate basin, about
|
||
<quantity id="4C939B60FFCBFFB1FAC0FB484174FB3A" box="[1307,1379,1181,1206]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="207" unit="km" value="10.0">10 km</quantity>
|
||
long and
|
||
<quantity id="4C939B60FFCBFFB1FF1CFB174501FB56" box="[199,278,1217,1242]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="207" unit="km" value="1.5">1.5 km</quantity>
|
||
wide. The basin is surrounded by steep mountains that rise to about
|
||
<quantity id="4C939B60FFCBFFB1FBFAFB174060FB55" box="[1057,1143,1217,1241]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="207" unit="m" value="1000.0">1000 m</quantity>
|
||
elevation. Apparently the drainage is isolated from the Erave-Purari system, lying just a few km to the east, although it is probably connected via subterranean flow. The lake contains numerous floating rafts of aquatic vegetation and the shoreline is swampy and lined with tall grass. Grassland and swamp habitat transition to dense rainforest on the eastern end of the basin, and the
|
||
<typeStatus id="54D08827FFCBFFB1FF29FA864535FAEB" box="[242,290,1360,1383]" pageId="6" pageNumber="207">type</typeStatus>
|
||
locality is situated in the latter environment. The stream was about
|
||
<quantity id="4C939B60FFCBFFB1FBFCFA984078FAEB" box="[1063,1135,1358,1383]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="207" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</quantity>
|
||
in width with a maximum depth of one m. The new species was relatively common on the mainly gravel/rock bottom. The melanotaeniid
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FF4CFA40466AFA22" authority="Allen, 1981" authorityName="Allen" authorityYear="1981" box="[151,637,1429,1454]" class="Actinopterygii" family="Melanotaeniidae" genus="Melanotaenia" kingdom="Animalia" order="Atheriniformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="herbertaxelrodi">
|
||
<emphasis id="B91FEA97FFCBFFB1FF4CFA4045FCFA22" box="[151,491,1430,1454]" italics="true" pageId="6" pageNumber="207">Melanotaenia herbertaxelrodi</emphasis>
|
||
<bibRefCitation id="EFFA4B74FFCBFFB1FE2EFA43466AFA22" author="Allen" box="[501,637,1429,1454]" pageId="6" pageNumber="207" refString="Allen, G. R. (1981) Central highland rainbows from Papua New Guinea, with descriptions of two new species (Melanotaeniidae). Tropical Fish Hobbyist, 29 (5), 20 - 33, 68 - 81." type="journal article" year="1981">Allen, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
was the only other fish that was observed. Underwater observations revealed the new species was also common in the main lake during a visit to the Tebera Basin by the second author in
|
||
<date id="FFD51045FFCBFFB1FF6FFA0A4547FA79" box="[180,336,1500,1525]" pageId="6" pageNumber="207" value="1991-10">October 1991</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD43685FFCBFFB1FF1CFA294603F813" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CFA29461CF994" bold="true" box="[199,523,1535,1560]" pageId="6" pageNumber="207">Similarity to other species.</emphasis>
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FDCFF9D64748F994" box="[532,863,1536,1560]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCBFFB1FDCFF9D64748F994" box="[532,863,1536,1560]" italics="true" pageId="6" pageNumber="207">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
differs from the lowland species of
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FADFF9D6418BF994" box="[1284,1436,1536,1560]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91FEA97FFCBFFB1FADFF9D6418BF994" box="[1284,1436,1536,1560]" italics="true" pageId="6" pageNumber="207">Glossogobius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B91FEA97FFCBFFB1FF7BF9F245A0F9B0" box="[160,439,1572,1596]" italics="true" pageId="6" pageNumber="207">
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FF7BF9F244E8F9B0" box="[160,255,1572,1596]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="celebius">celebius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FED1F9F24572F9B0" box="[266,357,1572,1596]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="clitellus">clitellus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FEAAF9F245A0F9B0" box="[369,439,1572,1596]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="illimis">illimis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FE36F9F2462AF9B7" box="[493,573,1572,1595]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="munroi">
|
||
<emphasis id="B91FEA97FFCBFFB1FE36F9F2462AF9B7" box="[493,573,1572,1595]" italics="true" pageId="6" pageNumber="207">munroi</emphasis>
|
||
</taxonomicName>
|
||
) treated by
|
||
<bibRefCitation id="EFFA4B74FFCBFFB1FD1FF9F247A9F9B0" author="Hoese" box="[708,958,1571,1596]" pageId="6" pageNumber="207" refString="Hoese, D. F. & Allen, G. R. (2012) A review of the amphidromous species of the Glossogobius celebius complex, with description of three new species. Cybium, 35 (4), 269 - 284." type="journal article" year="2012">Hoese & Allen (2012)</bibRefCitation>
|
||
in lacking opercular scales. It differs from
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FF4CF99145ADF9D3" box="[151,442,1607,1631]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="bicirrhosus">
|
||
<emphasis id="B91FEA97FFCBFFB1FF4CF99145ADF9D3" box="[151,442,1607,1631]" italics="true" pageId="6" pageNumber="207">Glossogobius bicirrhosus</emphasis>
|
||
</taxonomicName>
|
||
in lacking barbels extending from the mental fraenum. It differs from the highland species
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FF2AF9BD45FEF90F" box="[241,489,1643,1667]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="bulmeri">
|
||
<emphasis id="B91FEA97FFCBFFB1FF2AF9BD45FEF90F" box="[241,489,1643,1667]" italics="true" pageId="6" pageNumber="207">Glossogobius bulmeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FDF8F9BD46B4F90E" box="[547,675,1643,1666]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="torrentis">
|
||
<emphasis id="B91FEA97FFCBFFB1FDF8F9BD46B4F90E" box="[547,675,1643,1666]" italics="true" pageId="6" pageNumber="207">G. torrentis</emphasis>
|
||
</taxonomicName>
|
||
in having an elongate pelvic disc and predorsal scales reaching to near eye (versus rounded pelvic fin and predorsal scales reduced, not reaching before posterior preopercular margin). It differs from
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FE75F9674688F945" box="[430,671,1713,1737]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="coatesi">
|
||
<emphasis id="B91FEA97FFCBFFB1FE75F9674688F945" box="[430,671,1713,1737]" italics="true" pageId="6" pageNumber="207">Glossogobius coatesi</emphasis>
|
||
</taxonomicName>
|
||
from the Sepik River system in having more dorsal rays (I, 9–10 versus I,8). It differs from
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FE16F90346A1F961" box="[461,694,1749,1773]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="hoesei">
|
||
<emphasis id="B91FEA97FFCBFFB1FE16F90346A1F961" box="[461,694,1749,1773]" italics="true" pageId="6" pageNumber="207">Glossogobius hoesei</emphasis>
|
||
</taxonomicName>
|
||
from the Vogelkop Peninsula in western New
|
||
<collectingCountry id="F37C7615FFCBFFB1FB0DF9034130F962" box="[1238,1319,1749,1774]" name="Guinea-Bissau" pageId="6" pageNumber="207">Guinea</collectingCountry>
|
||
in having fewer anal rays (I,8 versus I,9–10).
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FDFEF92E4778F89C" box="[549,879,1784,1808]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="B91FEA97FFCBFFB1FDFEF92E4778F89C" box="[549,879,1784,1808]" italics="true" pageId="6" pageNumber="207">Glossogobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C6B4D06FFCBFFB1FBE4F92E4122F89C" box="[1087,1333,1784,1808]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="bulmeri">
|
||
<emphasis id="B91FEA97FFCBFFB1FBE4F92E4122F89C" box="[1087,1333,1784,1808]" italics="true" pageId="6" pageNumber="207">Glossogobius bulmeri</emphasis>
|
||
</taxonomicName>
|
||
from the highland rivers of
|
||
<collectingCountry id="F37C7615FFCBFFB1FEB8F8CB462DF8B9" box="[355,570,1820,1845]" name="Papua New Guinea" pageId="6" pageNumber="207">Papua New Guinea</collectingCountry>
|
||
in fin ray counts, sensory papillae and other morphological features. It differs from that species in having a slightly larger head (length 32.9–34.9% SL versus 28.5–32.4 %SL, head depth 17.3– 18.3% SL, versus 25.2–27.4 %SL), longer snout (9.4–11.7% SL versus 8.4–10.4% SL), longer pelvic fin (reaching anus versus ending short of anus).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371650EFFCBFFB1FF1CF87C45A6F86A" pageId="6" pageNumber="207" type="etymology">
|
||
<paragraph id="8BD43685FFCBFFB1FF1CF87C45A6F86A" blockId="6.[151,1437,151,2022]" pageId="6" pageNumber="207">
|
||
<emphasis id="B91FEA97FFCBFFB1FF1CF87C4559F84E" bold="true" box="[199,334,1962,1986]" pageId="6" pageNumber="207">Etymology.</emphasis>
|
||
From the Greek
|
||
<emphasis id="B91FEA97FFCBFFB1FDD7F87C464AF84E" box="[524,605,1962,1986]" italics="true" pageId="6" pageNumber="207">makros</emphasis>
|
||
= large and
|
||
<emphasis id="B91FEA97FFCBFFB1FD33F87C4728F84E" box="[744,831,1962,1986]" italics="true" pageId="6" pageNumber="207">kephale</emphasis>
|
||
= head, treated as a noun in apposition, relating to the large head of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |