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<document id="9DAC97E204D58790824787943606B3F9" ID-DOI="10.5852/ejt.2024.925.2467" ID-ISSN="2118-9773" ID-Zenodo-Dep="10837829" ID-ZooBank="B09375C0-6E28-4E2D-BB7C-26E1753171E2" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1710844381467" checkinUser="plazi" docAuthor="Elm, Laura-Marie Von &amp; Hoffman, Leon" docDate="2024" docId="03BAA406FFCEFFA7B814D230BBF3FA07" docLanguage="en" docName="EJT.2024.925.268-291.pdf" docOrigin="European Journal of Taxonomy 925" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2467/10969" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Granulina ronaldi Elm &amp; Hoffman 2024, sp. nov." docType="treatment" docUuid="38316ACE-B8E8-4743-8F4D-59785DA8AED2" docUuidSource="ZooBank" docVersion="3" lastPageNumber="282" masterDocId="FF83DC7EFFC2FFA9BA6ED11FB921FFCC" masterDocTitle="Revision of deep-water species in Granulina (Gastropoda: Granulinidae) from Mauritania and Western Sahara" masterLastPageNumber="291" masterPageNumber="268" pageNumber="280" updateTime="1710942960895" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FDB6412B44A8A5E2D67129C2E2A3C975">Revision of deep-water species in Granulina (Gastropoda: Granulinidae) from Mauritania and Western Sahara</mods:title>
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<mods:namePart id="5E79299E8DAEBE9BEB24110AC4D27882">Elm, Laura-Marie Von</mods:namePart>
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<mods:affiliation id="4442D48532BDC0C0B6BDD024B1668110">Thüringerstrasse 3, 26180 Rastede, Germany.</mods:affiliation>
<mods:nameIdentifier id="0510F6329B30FCCE7C721339092FD6F2" type="email">laura-marievonelm@kabelmail.de</mods:nameIdentifier>
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<mods:namePart id="75A934A588D03AB4FE648E5CB7186392">Hoffman, Leon</mods:namePart>
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<mods:affiliation id="6A109DE531101F2B3F2A9CE787BA5371">Senckenberg am Meer, Südstrand 40, 26382 Wilhelmshaven, Germany.</mods:affiliation>
<mods:nameIdentifier id="C4A1FEF636D35C9AEBCE447585FC5B47" type="email">leon.hoffman@senckenberg.de</mods:nameIdentifier>
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<mods:title id="3EEE198B457FCE913653163806A14BE5">European Journal of Taxonomy</mods:title>
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<mods:date id="48FF28B895A886513E38BFA876593BFD">2024</mods:date>
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<mods:number id="618C1270F3848C96B282EB16BABB8F18">2024-03-18</mods:number>
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<mods:identifier id="72575CA854A802B5C57E3B5C389ABBE7" type="DOI">10.5852/ejt.2024.925.2467</mods:identifier>
<mods:identifier id="C7F52B023405FA2A8C080322B11245EB" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="AB6AE64E55BB9EB9F07017EAB84E7FC1" type="Zenodo-Dep">10837829</mods:identifier>
<mods:identifier id="C4BB68A30DE7BB59E752BA304224F5E7" type="ZooBank">B09375C0-6E28-4E2D-BB7C-26E1753171E2</mods:identifier>
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<treatment id="03BAA406FFCEFFA7B814D230BBF3FA07" LSID="urn:lsid:zoobank.org:act:38316ACE-B8E8-4743-8F4D-59785DA8AED2" httpUri="http://treatment.plazi.org/id/03BAA406FFCEFFA7B814D230BBF3FA07" lastPageId="14" lastPageNumber="282" pageId="12" pageNumber="280">
<subSubSection id="C309469BFFCEFFA5B814D230BA9BFC86" box="[634,954,815,842]" pageId="12" pageNumber="280" type="nomenclature">
<paragraph id="8BAC1510FFCEFFA5B814D230BA9BFC86" blockId="12.[363,1225,815,914]" box="[634,954,815,842]" pageId="12" pageNumber="280">
<heading id="D0E4A27CFFCEFFA5B814D230BA9BFC86" box="[634,954,815,842]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="280" reason="2">
<taxonomicName id="4C136E93FFCEFFA5B814D230BA79FC86" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[634,856,815,842]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="280" phylum="Mollusca" rank="species" species="ronaldi" status="sp. nov.">
<emphasis id="B967C902FFCEFFA5B814D230BA79FC86" bold="true" box="[634,856,815,842]" italics="true" pageId="12" pageNumber="280">Granulina ronaldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCEFFA5B90ED22FBA9BFC86" box="[864,954,816,842]" pageId="12" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309469BFFCEFFA5BB05D24CBA6CFC5E" pageId="12" pageNumber="280" type="description">
<paragraph id="8BAC1510FFCEFFA5BB05D24CBDE8FCA2" blockId="12.[363,1225,815,914]" box="[363,1225,851,878]" pageId="12" pageNumber="280">
<uri id="FF821912FFCEFFA5BB05D24CBDE8FCA2" box="[363,1225,851,878]" pageId="12" pageNumber="280">
urn:lsid:zoobank.org:act:
<uuid id="FFB52FC5FFCEFFA5B8E6D24CBDE8FCA2" box="[648,1225,851,878]" pageId="12" pageNumber="280">38316ACE-B8E8-4743-8F4D-59785DA8AED2</uuid>
</uri>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5B889D268BA6CFC5E" blockId="12.[363,1225,815,914]" box="[743,845,887,914]" pageId="12" pageNumber="280">
<figureCitation id="13280995FFCEFFA5B889D268BA10FC5E" box="[743,817,887,914]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[280,1266,441,1796]" captionTargetId="figure-101@10.[216,1372,404,1821]" captionTargetPageId="10" captionText="Fig. 5. Location map of the new species in Granulina Jousseaume, 1888 off Mauritania and Western Sahara. White circles show all investigated stations; colour symbols show locations of shells from new species. Granulina reginae sp. nov. is presented by yellow squares, G. sigridae sp. nov. as red diamonds, G. sandrae sp. nov. as blue triangles, and G. ronaldi sp. nov. as green circles. Bathymetric data from GEBCO; contours 500 m" figureDoi="http://doi.org/10.5281/zenodo.10837843" httpUri="https://zenodo.org/record/10837843/files/figure.png" pageId="12" pageNumber="280">Figs 5</figureCitation>
,
<figureCitation id="13280995FFCEFFA5B950D267BA6CFC5E" box="[830,845,888,914]" captionStart="Fig" captionStartId="13.[189,232,1787,1813]" captionTargetBox="[228,1360,556,1742]" captionTargetId="figure-181@13.[228,1360,556,1742]" captionTargetPageId="13" captionText="Fig. 7. Granulina ronaldi sp. nov., Mauritania, off Banc dArguin, CANCAP/3.154. AD. Holotype (SMF359026). AB. Ventral and side views, height 2.9 mm, width 1.7 mm, tumidity 1.4 mm. C. Columellar folds. D. Micro-sculpture above second columellar fold, see arrow in C. EF. Paratype (SMF359020), ventral and side views, height 2.5 mm, width 1.5 mm, tumidity 1.3 mm. GH. Paratype (SMF359027), central and side views, height 2.5 mm, width 1.4 mm, tumidity 1.3 mm, callus line indicated by white dots. IJ. Paratype (SMF359029), ventral and side views, height 2.5 mm, width 1.5 mm, tumidity 1.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10837847" httpUri="https://zenodo.org/record/10837847/files/figure.png" pageId="12" pageNumber="280">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309469BFFCEFFA5BAD3D2A6B869FBA2" pageId="12" pageNumber="280" type="diagnosis">
<paragraph id="8BAC1510FFCEFFA5BAD3D2A6B817FC1F" blockId="12.[189,310,953,979]" box="[189,310,953,979]" pageId="12" pageNumber="280">
<heading id="D0E4A27CFFCEFFA5BAD3D2A6B817FC1F" bold="true" box="[189,310,953,979]" fontSize="11" level="3" pageId="12" pageNumber="280" reason="3">
<emphasis id="B967C902FFCEFFA5BAD3D2A6B817FC1F" bold="true" box="[189,310,953,979]" pageId="12" pageNumber="280">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BAD3D2F6B869FBA2" blockId="12.[189,1398,1000,1134]" pageId="12" pageNumber="280">
Small shell, height maximum
<quantity id="4CEBB8F5FFCEFFA5B840D2F6BBAEFBCF" box="[558,655,1001,1027]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="280" unit="mm" value="2.9">2.9 mm</quantity>
, width maximum
<quantity id="4CEBB8F5FFCEFFA5B901D2F6BAECFBCF" box="[879,973,1001,1027]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="280" unit="mm" value="1.7">1.7 mm</quantity>
(width/height ratio 0.59), tumidity maximum
<quantity id="4CEBB8F5FFCEFFA5BB55D512B8B9FBEB" box="[315,408,1037,1063]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="280" unit="mm" value="1.4">1.4 mm</quantity>
, ovoid-angular shape, rather narrow shape; no labial denticles on interior of external lip, tapered and weakly rostrated apically and abapically, widely channelled abapically, colour white, translucent.
</paragraph>
</subSubSection>
<subSubSection id="C309469BFFCEFFA5BAD3D589B85EFA87" pageId="12" pageNumber="280" type="etymology">
<paragraph id="8BAC1510FFCEFFA5BAD3D589B865FB7C" blockId="12.[189,324,1174,1200]" box="[189,324,1174,1200]" pageId="12" pageNumber="280">
<heading id="D0E4A27CFFCEFFA5BAD3D589B865FB7C" bold="true" box="[189,324,1174,1200]" fontSize="11" level="3" pageId="12" pageNumber="280" reason="3">
<emphasis id="B967C902FFCEFFA5BAD3D589B865FB7C" bold="true" box="[189,324,1174,1200]" pageId="12" pageNumber="280">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BAD3D5DAB85EFA87" blockId="12.[189,1399,1221,1355]" pageId="12" pageNumber="280">
The name (masculine) of the species acknowledges Ronald Janssen, retired curator of the Mollusca collection in the Senckenberg Museum Frankfurt who salvaged the
<emphasis id="B967C902FFCEFFA5B9B2D5F5BD10FACF" box="[988,1073,1258,1283]" italics="true" pageId="12" pageNumber="280">Meteor</emphasis>
material when it was about to be discarded and who initiated the preservation and sorting process. He also allowed us to study the
<emphasis id="B967C902FFCEFFA5BAD0D42DB832FA87" box="[190,275,1330,1355]" italics="true" pageId="12" pageNumber="280">Meteor</emphasis>
samples.
</paragraph>
</subSubSection>
<subSubSection id="C309469BFFCEFFA4BAD3D46CBA67FE41" lastPageId="13" lastPageNumber="281" pageId="12" pageNumber="280" type="materials_examined">
<paragraph id="8BAC1510FFCEFFA5BAD3D46CB84DFA41" blockId="12.[189,364,1395,1421]" box="[189,364,1395,1421]" pageId="12" pageNumber="280">
<heading id="D0E4A27CFFCEFFA5BAD3D46CB84DFA41" bold="true" box="[189,364,1395,1421]" fontSize="11" level="3" pageId="12" pageNumber="280" reason="3">
<emphasis id="B967C902FFCEFFA5BAD3D46CB84DFA41" bold="true" box="[189,364,1395,1421]" pageId="12" pageNumber="280">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BA84D4BDB87DFA70" blockId="12.[187,1399,1442,1540]" box="[234,348,1442,1468]" pageId="12" pageNumber="280">
<emphasis id="B967C902FFCEFFA5BA84D4BDB87DFA70" bold="true" box="[234,348,1442,1468]" pageId="12" pageNumber="280">
<typeStatus id="54A8ABB2FFCEFFA5BA84D4BDB87DFA70" box="[234,348,1442,1468]" pageId="12" pageNumber="280" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BAD3D4D9B8F0F9C8" blockId="12.[187,1399,1442,1540]" pageId="12" pageNumber="280">
<materialsCitation id="3B7B1F4DFFCEFFA5BAD3D4D9B8F0F9C8" collectingDate="1978-10-31" collectionCode="SMF" country="Mauritania" elevation="200" latitude="19.35" location="Banc d'Arguin" longLatPrecision="555" longitude="-16.9333" pageId="12" pageNumber="280" specimenCode="SMF359026" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F3045580FFCEFFA5BAD3D4D9B857FA2C" box="[189,374,1478,1504]" name="Mauritania" pageId="12" pageNumber="280">MAURITANIA</collectingCountry>
<emphasis id="B967C902FFCEFFA5BBE3D4D9BB67FA13" bold="true" box="[397,582,1477,1504]" pageId="12" pageNumber="280">
<location id="8ECC43CBFFCEFFA5BBE3D4D9BB67FA13" LSID="urn:lsid:plazi:treatment:03BAA406FFCEFFA7B814D230BBF3FA07:8ECC43CBFFCEFFA5BBE3D4D9BB67FA13" box="[397,582,1477,1504]" country="Mauritania" latitude="19.35" longLatPrecision="555" longitude="-16.9333" name="Banc d'Arguin" pageId="12" pageNumber="280">Banc dArguin</location>
</emphasis>
• 1 shell;
<geoCoordinate id="EE2773D7FFCEFFA5B8D8D4D9BA00FA2C" box="[694,801,1478,1505]" degrees="19.35" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="555" value="19.35">19.35° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5B945D4D9BA98FA2C" box="[811,953,1478,1504]" degrees="16.9333" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="5" value="-16.9333">16.9333° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCEFFA5BE67D4D9BD75FA2C" box="[1033,1108,1478,1504]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="280" unit="m" value="200.0">
<elevation id="003EF223FFCEFFA5BE67D4D9BD75FA2C" box="[1033,1108,1478,1504]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="280" unit="m" value="200.0">200 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCEFFA5BE31D4D9BDD7FA2C" box="[1119,1270,1478,1504]" pageId="12" pageNumber="280" value="1978-10-31">
<collectingDate id="EFE9CA38FFCEFFA5BE31D4D9BDD7FA2C" box="[1119,1270,1478,1504]" pageId="12" pageNumber="280" value="1978-10-31">31 Oct. 1978</collectingDate>
</date>
; CANCAP stn 3.154;
<specimenCode id="DBB5BD6BFFCEFFA5BB58D4F6B8ECF9C8" box="[310,461,1513,1540]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">SMF359026</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BA84D733B8CBF98A" blockId="12.[187,1398,1580,1786]" box="[234,490,1580,1606]" pageId="12" pageNumber="280">
<emphasis id="B967C902FFCEFFA5BA84D733B846F98A" bold="true" box="[234,359,1580,1606]" pageId="12" pageNumber="280">
<typeStatus id="54A8ABB2FFCEFFA5BA84D733B846F98A" box="[234,359,1580,1606]" pageId="12" pageNumber="280" type="paratype">Paratypes</typeStatus>
</emphasis>
(40 shells)
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BAD3D74FBBEFF936" blockId="12.[187,1398,1580,1786]" pageId="12" pageNumber="280">
<materialsCitation id="3B7B1F4DFFCEFFA5BAD3D74FB8D4F942" collectingDate="1978-10-31" collectionCode="SMF" country="Mauritania" elevation="200" latitude="19.35" location="Banc d'Arguin" longLatPrecision="555" longitude="-16.9333" pageId="12" pageNumber="280" specimenCode="SMF359020, SMF359027, SMF359029" specimenCount="1" stateProvince="Banc d'Arguin" typeStatus="paratype">
<collectingCountry id="F3045580FFCEFFA5BAD3D74FB857F9A6" box="[189,374,1616,1642]" name="Mauritania" pageId="12" pageNumber="280">MAURITANIA</collectingCountry>
<emphasis id="B967C902FFCEFFA5BBCFD74FBB43F9A5" bold="true" box="[417,610,1615,1642]" pageId="12" pageNumber="280">
<collectingRegion id="49D7DBF2FFCEFFA5BBCFD74FBB43F9A5" box="[417,610,1615,1642]" pageId="12" pageNumber="280">Banc dArguin</collectingRegion>
</emphasis>
• 3 shells; same collection data as for
<typeStatus id="54A8ABB2FFCEFFA5BE0ED750BDEBF9A5" box="[1120,1226,1615,1641]" pageId="12" pageNumber="280">holotype</typeStatus>
;
<specimenCode id="DBB5BD6BFFCEFFA5BEB2D750BC53F9A6" box="[1244,1394,1615,1642]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">
<collectionCode id="ED028DD5FFCEFFA5BEB2D750BC36F9A6" box="[1244,1303,1615,1642]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="12" pageNumber="280">SMF</collectionCode>
359020
</specimenCode>
,
<specimenCode id="DBB5BD6BFFCEFFA5BAD3D76CB872F942" box="[189,339,1651,1678]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">
<collectionCode id="ED028DD5FFCEFFA5BAD3D76CB9D8F942" box="[189,249,1651,1678]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="12" pageNumber="280">SMF</collectionCode>
359027
</specimenCode>
,
<specimenCode id="DBB5BD6BFFCEFFA5BB0CD76CB8D4F942" box="[354,501,1651,1678]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">
<collectionCode id="ED028DD5FFCEFFA5BB0CD76CB8BDF942" box="[354,412,1651,1678]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="12" pageNumber="280">SMF</collectionCode>
359029
</specimenCode>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCEFFA5B87BD76CBC40F942" box="[533,1377,1651,1678]" collectingDate="1978-10-31" collectionCode="RMNH" country="Mauritania" elevation="200" latitude="19.35" location="Banc d'Arguin" longLatPrecision="555" longitude="-16.9333" pageId="12" pageNumber="280" specimenCount="1" stateProvince="Banc d'Arguin" typeStatus="paratype">
25 shells; same collection data as for
<typeStatus id="54A8ABB2FFCEFFA5B989D76CBD70F941" box="[999,1105,1651,1677]" pageId="12" pageNumber="280">holotype</typeStatus>
;
<collectionCode id="ED028DD5FFCEFFA5BE0FD76CBD9DF942" box="[1121,1212,1651,1678]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="280" type="Museum">RMNH</collectionCode>
.MOL.351810
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCEFFA5BAD3D788BC57F97E" box="[189,1398,1687,1714]" collectingDate="1988-05-22" collectionCode="SMF" country="Mauritania" elevation="151" latitude="19.94" location="Tyro Mauritania" longLatPrecision="762" longitude="-17.48" pageId="12" pageNumber="280" specimenCode="SMF359028" specimenCount="1" stateProvince="Banc d'Arguin" typeStatus="paratype">
8 shells;
<geoCoordinate id="EE2773D7FFCEFFA5BB4ED788B8AAF97E" box="[288,395,1687,1714]" degrees="19.94" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="555" value="19.94">19.94° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5BBFAD788BB25F97E" box="[404,516,1687,1714]" degrees="17.48" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="555" value="-17.48">17.48° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCEFFA5B83AD788BBBEF97E" box="[596,671,1687,1714]" metricMagnitude="2" metricUnit="m" metricValue="1.51" pageId="12" pageNumber="280" unit="m" value="151.0">
<elevation id="003EF223FFCEFFA5B83AD788BBBEF97E" box="[596,671,1687,1714]" metricMagnitude="2" metricUnit="m" metricValue="1.51" pageId="12" pageNumber="280" unit="m" value="151.0">151 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCEFFA5B8C7D788BA65F97D" box="[681,836,1687,1714]" pageId="12" pageNumber="280" value="1988-05-22">
<collectingDate id="EFE9CA38FFCEFFA5B8C7D788BA65F97D" box="[681,836,1687,1714]" pageId="12" pageNumber="280" value="1988-05-22">22 May 1988</collectingDate>
</date>
;
<location id="8ECC43CBFFCEFFA5B923D787BD2BF97D" LSID="urn:lsid:plazi:treatment:03BAA406FFCEFFA7B814D230BBF3FA07:8ECC43CBFFCEFFA5B923D787BD2BF97D" box="[845,1034,1687,1714]" country="Mauritania" latitude="19.94" longLatPrecision="762" longitude="-17.48" name="Tyro Mauritania" pageId="12" pageNumber="280" stateProvince="Banc d'Arguin">
Tyro
<collectingCountry id="F3045580FFCEFFA5B9E7D788BD2BF97D" box="[905,1034,1687,1713]" name="Mauritania" pageId="12" pageNumber="280">Mauritania</collectingCountry>
</location>
I stn B5; box core;
<specimenCode id="DBB5BD6BFFCEFFA5BE8AD788BC57F97E" box="[1252,1398,1687,1714]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">
<collectionCode id="ED028DD5FFCEFFA5BE8AD788BC3EF97E" box="[1252,1311,1687,1714]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="12" pageNumber="280">SMF</collectionCode>
359028
</specimenCode>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCEFFA5BAA3D7A4BBEBF936" collectedFrom="sandy clay with shells" collectingDate="1978-10-30" collectingMethod="van Veen grab" collectionCode="RMNH" country="Mauritania" elevation="203" latitude="18.9833" longLatPrecision="7" longitude="-16.8333" pageId="12" pageNumber="280" specimenCode="RMNH.MOL.351811" specimenCount="4" stateProvince="Banc d'Arguin" typeStatus="paratype">
<specimenCount id="9D15DE99FFCEFFA5BAA3D7A4B80BF919" box="[205,298,1723,1749]" pageId="12" pageNumber="280">4 shells</specimenCount>
;
<geoCoordinate id="EE2773D7FFCEFFA5BB56D7A4B8E2F919" box="[312,451,1723,1749]" degrees="18.9833" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="5" value="18.9833">18.9833° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5BBBED7A4BB42F919" box="[464,611,1723,1749]" degrees="16.8333" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="5" value="-16.8333">16.8333° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCEFFA5B8D4D7A4BA29F919" box="[698,776,1723,1749]" metricMagnitude="2" metricUnit="m" metricValue="2.03" pageId="12" pageNumber="280" unit="m" value="203.0">
<elevation id="003EF223FFCEFFA5B8D4D7A4BA29F919" box="[698,776,1723,1749]" metricMagnitude="2" metricUnit="m" metricValue="2.03" pageId="12" pageNumber="280" unit="m" value="203.0">203 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCEFFA5B978D7A4BA95F919" box="[790,948,1723,1749]" pageId="12" pageNumber="280" value="1978-10-30">
<collectingDate id="EFE9CA38FFCEFFA5B978D7A4BA95F919" box="[790,948,1723,1749]" pageId="12" pageNumber="280" value="1978-10-30">30 Oct. 1978</collectingDate>
</date>
; CANCAP stn 3.140;
<collectedFrom id="380F56EDFFCEFFA5BED2D7A4B825F935" pageId="12" pageNumber="280">sandy clay with shells</collectedFrom>
;
<collectingMethod id="52526D07FFCEFFA5BB7ED7C0B89AF935" box="[272,443,1759,1785]" pageId="12" pageNumber="280">van Veen grab</collectingMethod>
;
<specimenCode id="DBB5BD6BFFCEFFA5BBA9D7C0BBEBF936" box="[455,714,1759,1786]" pageId="12" pageNumber="280">RMNH.MOL.351811</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BA84D63EBB96F8F7" blockId="12.[189,1398,1825,1923]" box="[234,695,1825,1852]" pageId="12" pageNumber="280">
<emphasis id="B967C902FFCEFFA5BA84D63EBB07F8F7" bold="true" box="[234,550,1825,1851]" pageId="12" pageNumber="280">Other material examined</emphasis>
(159 shells)
</paragraph>
<paragraph id="8BAC1510FFCEFFA5BAD3D65ABA73F84F" blockId="12.[189,1398,1825,1923]" pageId="12" pageNumber="280">
<materialsCitation id="3B7B1F4DFFCEFFA5BAD3D65ABA6FF84F" collectingDate="2010-10-24" collectionCode="SaM" country="Mauritania" elevation="246" latitude="20.3651" location="Banc d'Arguin" longLatPrecision="7" longitude="-17.6834" pageId="12" pageNumber="280" specimenCount="1" stateProvince="Banc d'Arguin">
<collectingCountry id="F3045580FFCEFFA5BAD3D65AB852F893" box="[189,371,1861,1887]" name="Mauritania" pageId="12" pageNumber="280">MAURITANIA</collectingCountry>
<emphasis id="B967C902FFCEFFA5BBF9D65ABB72F892" bold="true" box="[407,595,1860,1887]" pageId="12" pageNumber="280">
<collectingRegion id="49D7DBF2FFCEFFA5BBF9D65ABB72F892" box="[407,595,1860,1887]" pageId="12" pageNumber="280">Banc dArguin</collectingRegion>
</emphasis>
• 12 shells;
<geoCoordinate id="EE2773D7FFCEFFA5B883D65BBA58F893" box="[749,889,1860,1887]" degrees="20.3651" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="5" value="20.3651">20.3651° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5B9E6D65BBD38F893" box="[904,1049,1860,1887]" degrees="17.6834" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="5" value="-17.6834">17.6834° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCEFFA5BE1CD65BBDE3F893" box="[1138,1218,1860,1887]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="280" unit="m" value="246.0">
<elevation id="003EF223FFCEFFA5BE1CD65BBDE3F893" box="[1138,1218,1860,1887]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="280" unit="m" value="246.0">246 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCEFFA5BEBCD65ABC52F893" box="[1234,1395,1861,1887]" pageId="12" pageNumber="280" value="2010-10-24">
<collectingDate id="EFE9CA38FFCEFFA5BEBCD65ABC52F893" box="[1234,1395,1861,1887]" pageId="12" pageNumber="280" value="2010-10-24">24 Oct. 2010</collectingDate>
</date>
; MSM163 stn GeoB14704; box core in muddy silt;
<collectionCode id="ED028DD5FFCEFFA5B97AD676BA6FF84F" box="[788,846,1897,1923]" pageId="12" pageNumber="280">SaM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC1510FFCEFFA4BAD3D6B4BA67FE41" blockId="12.[189,1399,1962,2025]" lastBlockId="13.[189,1398,265,397]" lastPageId="13" lastPageNumber="281" pageId="12" pageNumber="280">
<materialsCitation id="3B7B1F4DFFCEFFA5BAD3D6B4BA57F824" collectingDate="1977-02-25" collectionCode="SMF" country="Western Sahara" elevation="81" latitude="21.325" location="North of Dakhla" longLatPrecision="520" longitude="-17.37" pageId="12" pageNumber="280" specimenCount="1">
<collectingCountry id="F3045580FFCEFFA5BAD3D6B4B8E9F809" box="[189,456,1963,1989]" name="Western Sahara" pageId="12" pageNumber="280">WESTERN SAHARA</collectingCountry>
<emphasis id="B967C902FFCEFFA5BB82D6B5BBBFF809" bold="true" box="[492,670,1962,1989]" pageId="12" pageNumber="280">off Cap Blanc</emphasis>
• 6 shells;
<geoCoordinate id="EE2773D7FFCEFFA5B948D6B5BA85F809" box="[806,932,1962,1989]" degrees="21.325" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="55" value="21.325">21.325° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5B9DCD6B4BD06F809" box="[946,1063,1963,1989]" degrees="17.37" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="555" value="-17.37">17.37° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCEFFA5BEEED6B4BDE0F809" box="[1152,1217,1963,1989]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="12" pageNumber="280" unit="m" value="81.0">
<elevation id="003EF223FFCEFFA5BEEED6B4BDE0F809" box="[1152,1217,1963,1989]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="12" pageNumber="280" unit="m" value="81.0">81 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCEFFA5BEBED6B5BC53F809" box="[1232,1394,1962,1989]" pageId="12" pageNumber="280" value="1977-02-25">
<collectingDate id="EFE9CA38FFCEFFA5BEBED6B5BC53F809" box="[1232,1394,1962,1989]" pageId="12" pageNumber="280" value="1977-02-25">25 Feb. 1977</collectingDate>
</date>
; M44 stn 235KG649; box core;
<collectionCode id="ED028DD5FFCEFFA5B828D6D1BBA3F824" box="[582,642,1998,2024]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="280">SMF</collectionCode>
.
<emphasis id="B967C902FFCEFFA5B8CBD6D1BA57F824" bold="true" box="[677,886,1998,2024]" pageId="12" pageNumber="280">
<location id="8ECC43CBFFCEFFA5B8CBD6D1BA57F824" LSID="urn:lsid:plazi:treatment:03BAA406FFCEFFA7B814D230BBF3FA07:8ECC43CBFFCEFFA5B8CBD6D1BA57F824" box="[677,886,1998,2024]" country="Western Sahara" latitude="21.325" longLatPrecision="520" longitude="-17.37" name="North of Dakhla" pageId="12" pageNumber="280">North of Dakhla</location>
</emphasis>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCEFFA4B9FFD6D1BA56FEEF" collectingDate="1982-02-05" collectionCode="SMF" elevation="411" lastPageId="13" lastPageNumber="281" latitude="25.3333" longLatPrecision="7" longitude="-16.1533" pageId="12" pageNumber="280" specimenCount="2">
<specimenCount id="9D15DE99FFCEFFA5B9FFD6D1BACFF824" box="[913,1006,1998,2024]" pageId="12" pageNumber="280">2 shells</specimenCount>
;
<geoCoordinate id="EE2773D7FFCEFFA5B992D6D1BDA9F825" box="[1020,1160,1998,2025]" degrees="25.3333" direction="north" orientation="latitude" pageId="12" pageNumber="280" precision="5" value="25.3333">25.3333° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCEFFA5BEF8D6D1BC09F825" box="[1174,1320,1998,2025]" degrees="16.1533" direction="west" orientation="longitude" pageId="12" pageNumber="280" precision="5" value="-16.1533">16.1533° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCFFFA4BAD3D016B82DFEEF" box="[189,268,265,291]" metricMagnitude="2" metricUnit="m" metricValue="4.11" pageId="13" pageNumber="281" unit="m" value="411.0">
<elevation id="003EF223FFCFFFA4BAD3D016B82DFEEF" box="[189,268,265,291]" metricMagnitude="2" metricUnit="m" metricValue="4.11" pageId="13" pageNumber="281" unit="m" value="411.0">411 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCFFFA4BB75D016B891FEEF" box="[283,432,265,292]" pageId="13" pageNumber="281" value="1982-02-05">
<collectingDate id="EFE9CA38FFCFFFA4BB75D016B891FEEF" box="[283,432,265,292]" pageId="13" pageNumber="281" value="1982-02-05">5 Feb. 1982</collectingDate>
</date>
; M60 stn 49KG921, box core;
<collectionCode id="ED028DD5FFCFFFA4B952D016BA56FEEF" box="[828,887,265,291]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="281">SMF</collectionCode>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCFFFA4B9FBD016BA48FE8A" collectingDate="1982-02-05" collectionCode="SMF" elevation="407" latitude="25.3333" longLatPrecision="7" longitude="-16.1533" pageId="13" pageNumber="281" specimenCount="1">
<specimenCount id="9D15DE99FFCFFFA4B9FBD016BAC8FEEF" box="[917,1001,265,291]" pageId="13" pageNumber="281">1 shell</specimenCount>
;
<geoCoordinate id="EE2773D7FFCFFFA4B996D016BDA4FEEF" box="[1016,1157,265,292]" degrees="25.3333" direction="north" orientation="latitude" pageId="13" pageNumber="281" precision="5" value="25.3333">25.3333° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCFFFA4BEFAD016BC07FEEF" box="[1172,1318,265,292]" degrees="16.1533" direction="west" orientation="longitude" pageId="13" pageNumber="281" precision="5" value="-16.1533">16.1533° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCFFFA4BAD3D033B82DFE8B" box="[189,268,300,327]" metricMagnitude="2" metricUnit="m" metricValue="4.07" pageId="13" pageNumber="281" unit="m" value="407.0">
<elevation id="003EF223FFCFFFA4BAD3D033B82DFE8B" box="[189,268,300,327]" metricMagnitude="2" metricUnit="m" metricValue="4.07" pageId="13" pageNumber="281" unit="m" value="407.0">407 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCFFFA4BB77D033B88AFE8A" box="[281,427,300,327]" pageId="13" pageNumber="281" value="1982-02-05">
<collectingDate id="EFE9CA38FFCFFFA4BB77D033B88AFE8A" box="[281,427,300,327]" pageId="13" pageNumber="281" value="1982-02-05">5 Feb. 1982</collectingDate>
</date>
; M60 stn 43KG922; box core;
<collectionCode id="ED028DD5FFCFFFA4B940D033BA48FE8A" box="[814,873,300,326]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="281">SMF</collectionCode>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCFFFA4B9EAD033BA75FEA5" collectingDate="1975-02-22" collectionCode="SMF" elevation="87" latitude="25.1833" longLatPrecision="7" longitude="-15.8033" pageId="13" pageNumber="281" specimenCount="10">
<specimenCount id="9D15DE99FFCFFFA4B9EAD033BAD1FE8A" box="[900,1008,300,326]" pageId="13" pageNumber="281">10 shells</specimenCount>
;
<geoCoordinate id="EE2773D7FFCFFFA4B990D033BDABFE8B" box="[1022,1162,300,327]" degrees="25.1833" direction="north" orientation="latitude" pageId="13" pageNumber="281" precision="5" value="25.1833">25.1833° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCFFFA4BEF9D033BC09FE8B" box="[1175,1320,300,327]" degrees="15.8033" direction="west" orientation="longitude" pageId="13" pageNumber="281" precision="5" value="-15.8033">15.8033° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCFFFA4BAD0D050B9DEFEA6" box="[190,255,335,362]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="281" unit="m" value="87.0">
<elevation id="003EF223FFCFFFA4BAD0D050B9DEFEA6" box="[190,255,335,362]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="281" unit="m" value="87.0">87 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCFFFA4BB63D050B88EFEA6" box="[269,431,335,362]" pageId="13" pageNumber="281" value="1975-02-22">
<collectingDate id="EFE9CA38FFCFFFA4BB63D050B88EFEA6" box="[269,431,335,362]" pageId="13" pageNumber="281" value="1975-02-22">22 Feb. 1975</collectingDate>
</date>
; M36 stn 95KG6; box core;
<collectionCode id="ED028DD5FFCFFFA4B977D050BA75FEA5" box="[793,852,335,361]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="281">SMF</collectionCode>
</materialsCitation>
<materialsCitation id="3B7B1F4DFFCFFFA4B91FD050BA63FE41" collectingDate="1975-02-21" collectionCode="SMF" elevation="53" latitude="25.0" longLatPrecision="7" longitude="-15.5167" pageId="13" pageNumber="281" specimenCount="128">
<specimenCount id="9D15DE99FFCFFFA4B91FD050BACDFEA5" box="[881,1004,335,361]" pageId="13" pageNumber="281">128 shells</specimenCount>
;
<geoCoordinate id="EE2773D7FFCFFFA4B995D050BDA6FEA6" box="[1019,1159,335,362]" degrees="25.0000" direction="north" orientation="latitude" pageId="13" pageNumber="281" precision="5" value="25.0">25.0000° N</geoCoordinate>
,
<geoCoordinate id="EE2773D7FFCFFFA4BEFBD050BC06FEA6" box="[1173,1319,335,362]" degrees="15.5167" direction="west" orientation="longitude" pageId="13" pageNumber="281" precision="5" value="-15.5167">15.5167° W</geoCoordinate>
; depth
<quantity id="4CEBB8F5FFCFFFA4BAD1D06DB9DFFE41" box="[191,254,370,397]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="13" pageNumber="281" unit="m" value="53.0">
<elevation id="003EF223FFCFFFA4BAD1D06DB9DFFE41" box="[191,254,370,397]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="13" pageNumber="281" unit="m" value="53.0">53 m</elevation>
</quantity>
;
<date id="FFAD33D0FFCFFFA4BB64D06CB886FE41" box="[266,423,370,397]" pageId="13" pageNumber="281" value="1975-02-21">
<collectingDate id="EFE9CA38FFCFFFA4BB64D06CB886FE41" box="[266,423,370,397]" pageId="13" pageNumber="281" value="1975-02-21">21 Feb. 1975</collectingDate>
</date>
; M36 stn 94KG2; box core;
<collectionCode id="ED028DD5FFCFFFA4B968D06CBA63FE41" box="[774,834,371,397]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="281">SMF</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6C4598FFCFFFA4BAD3D7E4B8D0F825" ID-DOI="http://doi.org/10.5281/zenodo.10837847" ID-Zenodo-Dep="10837847" httpUri="https://zenodo.org/record/10837847/files/figure.png" pageId="13" pageNumber="281" startId="13.[189,232,1787,1813]" targetBox="[228,1360,556,1742]" targetPageId="13">
<paragraph id="8BAC1510FFCFFFA4BAD3D7E4B8D0F825" blockId="13.[189,1399,1787,2025]" pageId="13" pageNumber="281">
<emphasis id="B967C902FFCFFFA4BAD3D7E4B82FF8D9" bold="true" box="[189,270,1787,1813]" pageId="13" pageNumber="281">Fig. 7.</emphasis>
<taxonomicName id="4C136E93FFCFFFA4BB76D7E4B8D3F8D9" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[280,498,1787,1813]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="281" phylum="Mollusca" rank="species" species="ronaldi" status="sp. nov.">
<emphasis id="B967C902FFCFFFA4BB76D7E4B8D3F8D9" box="[280,498,1787,1813]" italics="true" pageId="13" pageNumber="281">Granulina ronaldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCFFFA4BB92D7E3BB7DF8DA" box="[508,604,1788,1814]" pageId="13" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
, Mauritania, off Banc dArguin, CANCAP/3.154.
<emphasis id="B967C902FFCFFFA4BED1D7E4BDD9F8D9" bold="true" box="[1215,1272,1787,1814]" pageId="13" pageNumber="281">AD</emphasis>
. Holotype (SMF359026).
<emphasis id="B967C902FFCFFFA4BB14D601B893F8F4" bold="true" box="[378,434,1822,1849]" pageId="13" pageNumber="281">AB</emphasis>
. Ventral and side views, height 2.9 mm, width 1.7 mm, tumidity 1.4 mm.
<emphasis id="B967C902FFCFFFA4BAD3D65EB9F3F897" bold="true" box="[189,210,1857,1883]" pageId="13" pageNumber="281">C</emphasis>
. Columellar folds.
<emphasis id="B967C902FFCFFFA4BBD7D65DB8ECF890" bold="true" box="[441,461,1858,1884]" pageId="13" pageNumber="281">D</emphasis>
. Micro-sculpture above second columellar fold, see arrow in C.
<emphasis id="B967C902FFCFFFA4BEA3D65DBC21F890" bold="true" box="[1229,1280,1858,1884]" pageId="13" pageNumber="281">EF</emphasis>
. Paratype (SMF359020), ventral and side views, height 2.5 mm, width 1.5 mm, tumidity 1.3 mm.
<emphasis id="B967C902FFCFFFA4BEABD67BBC20F8B3" bold="true" box="[1221,1281,1892,1919]" pageId="13" pageNumber="281">GH</emphasis>
. Paratype (SMF359027), central and side views, height 2.5 mm, width 1.4 mm, tumidity 1.3 mm, callus line indicated by white dots.
<emphasis id="B967C902FFCFFFA4BB84D6B4BB32F809" bold="true" box="[490,531,1963,1990]" pageId="13" pageNumber="281">IJ</emphasis>
. Paratype (SMF359029), ventral and side views, height 2.5 mm, width 1.5 mm, tumidity 1.2 mm.
</paragraph>
</caption>
<subSubSection id="C309469BFFCCFFA7BAD3D017BA4AFD83" pageId="14" pageNumber="282" type="description">
<paragraph id="8BAC1510FFCCFFA7BAD3D017B86FFEEE" blockId="14.[189,334,264,290]" box="[189,334,264,290]" pageId="14" pageNumber="282">
<heading id="D0E4A27CFFCCFFA7BAD3D017B86FFEEE" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="14" pageNumber="282" reason="3">
<emphasis id="B967C902FFCCFFA7BAD3D017B86FFEEE" bold="true" box="[189,334,264,290]" pageId="14" pageNumber="282">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC1510FFCCFFA7BAD3D027BA4AFD83" blockId="14.[187,1400,312,592]" pageId="14" pageNumber="282">Apex with highly convex, blunt, narrowing channel. External lip smoothly rounded, strongly thickened from apex to base, maximum convexity slightly above periphery, reclining towards apex and base, widely channelled at highly convex base, pointed at base columella, internally smooth (without labial denticles). Four discrete oblique folds at base of straight columella, with round edges, third fold longest. Parietal area widely convex and smooth. Macroscopic sculpture smooth with microscopic irregular growth lines; microscopic sculpture of flakes oriented towards exterior on callus of external lip, columella, columellar folds, base and apex, callus reclining in the middle part; internally smooth. Aperture curved, elongated, wide at base. Protoconch invisible, convolved inside shell.</paragraph>
</subSubSection>
<subSubSection id="C309469BFFCCFFA7BAD3D367BBF3FA07" pageId="14" pageNumber="282" type="discussion">
<paragraph id="8BAC1510FFCCFFA7BAD3D367B80EFD5E" blockId="14.[189,303,632,658]" box="[189,303,632,658]" pageId="14" pageNumber="282">
<heading id="D0E4A27CFFCCFFA7BAD3D367B80EFD5E" bold="true" box="[189,303,632,658]" fontSize="11" level="3" pageId="14" pageNumber="282" reason="3">
<emphasis id="B967C902FFCCFFA7BAD3D367B80EFD5E" bold="true" box="[189,303,632,658]" pageId="14" pageNumber="282">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC1510FFCCFFA7BAD3D3B7BDA0FC9E" blockId="14.[188,1400,679,850]" pageId="14" pageNumber="282">
Because of the missing labial denticles, this species cannot be mistaken for any hitherto known species found north of
<collectingCountry id="F3045580FFCCFFA7BB17D3D4B8FBFD29" box="[377,474,715,741]" name="Senegal" pageId="14" pageNumber="282">Senegal</collectingCountry>
. The morphology is similar to
<taxonomicName id="4C136E93FFCCFFA7B938D3D4BD10FD29" baseAuthorityName="Monterosato" baseAuthorityYear="1869" box="[854,1073,715,741]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="occulta">
<emphasis id="B967C902FFCCFFA7B938D3D4BD10FD29" box="[854,1073,715,741]" italics="true" pageId="14" pageNumber="282">Granulina occulta</emphasis>
</taxonomicName>
but that species has labial denticles. The outline of the present species is not as broad (width/height ratio 0.59) as in
<taxonomicName id="4C136E93FFCCFFA7BE90D3F0B83DFCE1" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B967C902FFCCFFA7BE90D3F0B83DFCE1" italics="true" pageId="14" pageNumber="282">Granulina sandrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCCFFA7BB4CD20BB85DFCE2" box="[290,380,788,814]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
(width/height ratio 0.69) and it has a narrow aperture.
<taxonomicName id="4C136E93FFCCFFA7B994D20CBDF4FCE1" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[1018,1237,787,813]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="reginae" status="sp. nov.">
<emphasis id="B967C902FFCCFFA7B994D20CBDF4FCE1" box="[1018,1237,787,813]" italics="true" pageId="14" pageNumber="282">Granulina reginae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCCFFA7BEB4D20BBC14FCE2" box="[1242,1333,788,814]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
has a wide, notched apical channel and its last whorl is widely rounded at the periphery.
</paragraph>
<paragraph id="8BAC1510FFCCFFA7BAD3D265B864FB85" blockId="14.[189,1401,890,1097]" pageId="14" pageNumber="282">
<taxonomicName id="4C136E93FFCCFFA7BAD3D265B8B7FC58" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[189,406,890,916]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="ronaldi" status="sp. nov.">
<emphasis id="B967C902FFCCFFA7BAD3D265B8B7FC58" box="[189,406,890,916]" italics="true" pageId="14" pageNumber="282">Granulina ronaldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCCFFA7BBCDD264BB20FC59" box="[419,513,891,917]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
could be mistaken for
<taxonomicName id="4C136E93FFCCFFA7B94ED264BA87FC58" authority="Elm &amp; Hoffman, 2024" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[800,934,890,917]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="sigridae" status="sp. nov.">
<emphasis id="B967C902FFCCFFA7B94ED264BA87FC58" box="[800,934,890,917]" italics="true" pageId="14" pageNumber="282">G. sigridae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2547479FFCCFFA7B9DCD264BD31FC59" box="[946,1040,891,917]" pageId="14" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
but that species has different columellar folds; the two upper ones are often coalescent and the third one is not as long as the fourth one. The micro-sculpture in the present species covers a wider area at the base of the callus and apex and is not as retracted in the parietal area as in
<taxonomicName id="4C136E93FFCCFFA7B8C5D2F8BA57FBCD" box="[683,886,999,1025]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="subSpecies" species="sigridae" subSpecies="either">
<emphasis id="B967C902FFCCFFA7B8C5D2F8BA57FBCD" box="[683,886,999,1025]" italics="true" pageId="14" pageNumber="282">G. sigridae either</emphasis>
</taxonomicName>
. The outer lip of the present species is more thickened, with weaker convexity and slightly angular. The (ab-) apical outlines are more convex than in
<taxonomicName id="4C136E93FFCCFFA7BAD0D530B81CFB85" authorityName="Elm &amp; Hoffman" authorityYear="2024" box="[190,317,1071,1097]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="sigridae">
<emphasis id="B967C902FFCCFFA7BAD0D530B81CFB85" box="[190,317,1071,1097]" italics="true" pageId="14" pageNumber="282">G. sigridae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAC1510FFCCFFA7BAD3D56DBBACFA8C" blockId="14.[189,1400,1137,1345]" pageId="14" pageNumber="282">
<bibRefCitation id="EF8268E1FFCCFFA7BAD3D56DB89EFB47" author="La Perna R. &amp; Landau B. &amp; Marquet R." box="[189,447,1137,1164]" pageId="14" pageNumber="282" pagination="111 - 120" refId="ref12002" refString="La Perna R., Landau B. &amp; Marquet R. 2001. Granulina (Gastropoda, Marginellidae) from the Pliocene of Malaga (southern Spain) with descriptions of four new species. Cainozoic Research 1 (1 - 2): 111 - 120. Available from http: // natuurtijdschriften. nl / download? type = document &amp; docid = 541677 [accessed 5 Mar. 2024]." type="journal article" year="2001">
La Perna
<emphasis id="B967C902FFCCFFA7BB41D56DB843FB47" box="[303,354,1137,1163]" italics="true" pageId="14" pageNumber="282">et al</emphasis>
. (2001)
</bibRefCitation>
described the similar
<taxonomicName id="4C136E93FFCCFFA7B8ABD56EBC17FB47" authority="La Perna, Landau &amp; Marquet, 2001" authorityName="La Perna, Landau &amp; Marquet" authorityYear="2001" box="[709,1334,1137,1164]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="longa">
<emphasis id="B967C902FFCCFFA7B8ABD56EBAABFB47" box="[709,906,1137,1163]" italics="true" pageId="14" pageNumber="282">Granulina longa</emphasis>
<bibRefCitation id="EF8268E1FFCCFFA7B9FFD56DBC17FB47" author="La Perna R. &amp; Landau B. &amp; Marquet R." box="[913,1334,1137,1164]" pageId="14" pageNumber="282" pagination="111 - 120" refId="ref12002" refString="La Perna R., Landau B. &amp; Marquet R. 2001. Granulina (Gastropoda, Marginellidae) from the Pliocene of Malaga (southern Spain) with descriptions of four new species. Cainozoic Research 1 (1 - 2): 111 - 120. Available from http: // natuurtijdschriften. nl / download? type = document &amp; docid = 541677 [accessed 5 Mar. 2024]." type="journal article" year="2001">La Perna, Landau &amp; Marquet, 2001</bibRefCitation>
</taxonomicName>
from the Pliocene near Estepona (western Mediterranean,
<collectingCountry id="F3045580FFCCFFA7B95CD58ABA5BFB63" box="[818,890,1173,1199]" name="Spain" pageId="14" pageNumber="282">Spain</collectingCountry>
).
<taxonomicName id="4C136E93FFCCFFA7B9FED58ABD76FB63" authorityName="La Perna, Landau &amp; Marquet" authorityYear="2001" box="[912,1111,1173,1199]" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="longa">
<emphasis id="B967C902FFCCFFA7B9FED58ABD76FB63" box="[912,1111,1173,1199]" italics="true" pageId="14" pageNumber="282">Granulina longa</emphasis>
</taxonomicName>
differs from the present species by a well-rounded oval outline, a wide aperture and a thin lip with fine labial denticles.
<bibRefCitation id="EF8268E1FFCCFFA7BF37D5A5B8B9FB34" author="La Perna R. &amp; Landau B. &amp; da Silva C. M." pageId="14" pageNumber="282" pagination="36 - 42" refId="ref12077" refString="La Perna R., Landau B. &amp; da Silva C. M. 2003. The genus Granulina (Gastropoda, Marginellidae) from the Atlantic Iberian Pliocene with description of a new species from Portugal. Iberus 21 (1): 36 - 42. Available from https: // zenodo. org / records / 4515821 [accessed 26 Feb. 2024]." type="journal article" year="2003">
La Perna
<emphasis id="B967C902FFCCFFA7BB69D5C0B81AFB34" box="[263,315,1246,1272]" italics="true" pageId="14" pageNumber="282">et al</emphasis>
. (2003)
</bibRefCitation>
described another similar species:
<taxonomicName id="4C136E93FFCCFFA7B958D5C1B9D9FAD0" authority="La Perna, Landau &amp; da Silva, 2003" authorityName="La Perna, Landau &amp; da Silva" authorityYear="2003" class="Gastropoda" family="Granulinidae" genus="Granulina" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="282" phylum="Mollusca" rank="species" species="choffati">
<emphasis id="B967C902FFCCFFA7B958D5C1BD33FB34" box="[822,1042,1246,1272]" italics="true" pageId="14" pageNumber="282">Granulina choffati</emphasis>
<bibRefCitation id="EF8268E1FFCCFFA7BE77D5C1B9D9FAD0" author="La Perna R. &amp; Landau B. &amp; da Silva C. M." pageId="14" pageNumber="282" pagination="36 - 42" refId="ref12077" refString="La Perna R., Landau B. &amp; da Silva C. M. 2003. The genus Granulina (Gastropoda, Marginellidae) from the Atlantic Iberian Pliocene with description of a new species from Portugal. Iberus 21 (1): 36 - 42. Available from https: // zenodo. org / records / 4515821 [accessed 26 Feb. 2024]." type="journal article" year="2003">La Perna, Landau &amp; da Silva, 2003</bibRefCitation>
</taxonomicName>
from the Atlantic Pliocene in
<collectingCountry id="F3045580FFCCFFA7B80CD41DBBE8FAD0" box="[610,713,1282,1308]" name="Portugal" pageId="14" pageNumber="282">Portugal</collectingCountry>
. It is differentiated from the present species by fine labial denticulations, and a more oval outline.
</paragraph>
<paragraph id="8BAC1510FFCCFFA7BAD3D477BBF3FA07" blockId="14.[189,1400,1384,1483]" pageId="14" pageNumber="282">
The bathymetric range is
<quantity id="4CEBB8F5FFCCFFA7BB81D476BB49FA4F" box="[495,616,1385,1411]" metricMagnitude="2" metricUnit="m" metricValue="2.46" metricValueMax="4.11" metricValueMin="0.8099999999999999" pageId="14" pageNumber="282" unit="m" value="246.0" valueMax="411.0" valueMin="81.0">
81
<specimenCount id="9D15DE99FFCCFFA7B877D476BB49FA4F" box="[537,616,1385,1411]" count="411" pageId="14" pageNumber="282" type="generic">411 m</specimenCount>
</quantity>
. The latitudinal range is 18.9
<geoCoordinate id="EE2773D7FFCCFFA7B9BAD477BD12FA4F" box="[980,1075,1384,1411]" degrees="25.4" direction="north" orientation="latitude" pageId="14" pageNumber="282" precision="5555" value="25.4">25.4° N</geoCoordinate>
, which is the northernmost occurrence of the new species. Some shells were found in sandy or muddy silt occasionally with other shells. The habitat of the species is uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>