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<document id="03A1FF942B78192EC67E4C4A75EBEE4D" ID-DOI="10.5281/zenodo.205436" ID-GBIF-Dataset="d9857903-c1ca-4729-ac92-dfed41c53827" ID-ISSN="1175-5326" ID-Zenodo-Dep="205436" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460338987085" checkinUser="plazi" docAuthor="Nieto, Carolina &amp; Emmerich, Daniel" docDate="2011" docId="03ADC05DFFCEFFF2FF330C50D95AFB68" docLanguage="en" docName="zt02996p065.pdf" docOrigin="Zootaxa 2996" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cloeodes dialutoi Nieto &amp; Emmerich, 2011, sp. nov." docType="treatment" docVersion="11" lastPageNumber="61" masterDocId="FF94B825FFCCFFF6FFA40F7ADC3EFFAC" masterDocTitle="Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay" masterLastPageNumber="65" masterPageNumber="57" pageNumber="59" updateTime="1698260647811" updateUser="plazi">
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<mods:title id="D0CD15F0A622AC32ED7CE297859F2DA1">Three new species of the genus Cloeodes Traver (Ephemeroptera: Baetidae) from Uruguay</mods:title>
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<mods:namePart id="6EC17E5E69A471559AAD6BB4BD870938">Nieto, Carolina</mods:namePart>
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<mods:namePart id="106F0A29C2618A5C5F797E9FCFB1CCAE">Emmerich, Daniel</mods:namePart>
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<mods:title id="6A8631D6FF899F676302D648EA193127">Zootaxa</mods:title>
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<mods:date id="F69F079974A5809ACDB52D8B1F49FA45">2011</mods:date>
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<mods:number id="81A3E5D4FB56268585957C1B545E8B03">2996</mods:number>
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<mods:identifier id="E9AF99EB485A4DDDF25B295579043E75" type="DOI">10.5281/zenodo.205436</mods:identifier>
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<mods:identifier id="BDC9D9D785CA7ACB7CE2E19D2F3CAD14" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03ADC05DFFCEFFF2FF330C50D95AFB68" ID-DOI="http://doi.org/10.5281/zenodo.6183251" ID-GBIF-Taxon="119405918" ID-Zenodo-Dep="6183251" LSID="urn:lsid:plazi:treatment:03ADC05DFFCEFFF2FF330C50D95AFB68" httpUri="http://treatment.plazi.org/id/03ADC05DFFCEFFF2FF330C50D95AFB68" lastPageId="4" lastPageNumber="61" pageId="2" pageNumber="59">
<subSubSection id="C31E22C0FFCEFFF2FF330C50DDACFC9B" lastPageId="4" lastPageNumber="61" pageId="2" pageNumber="59" type="nomenclature">
<paragraph id="8BBB714BFFCEFFF4FF330C50DD81FCEF" blockId="2.[151,447,810,870]" box="[151,447,810,835]" pageId="2" pageNumber="59">
<heading id="D0F3C627FFCEFFF4FF330C50DD81FCEF" bold="true" box="[151,447,810,835]" fontSize="10" level="2" pageId="2" pageNumber="59" reason="2">
<emphasis id="B970AD59FFCEFFF4FF330C50DD81FCEF" bold="true" box="[151,447,810,835]" pageId="2" pageNumber="59">
<taxonomicName id="4C040AC8FFCEFFF4FF330C50DD61FCEF" ID-CoL="5ZBTV" box="[151,351,810,835]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="dialutoi" status="sp. nov.">
<emphasis id="B970AD59FFCEFFF4FF330C50DD61FCEF" bold="true" box="[151,351,810,835]" italics="true" pageId="2" pageNumber="59">Cloeodes dialutoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFCEFFF4FEC20C51DD81FCEF" box="[358,447,811,835]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB714BFFCEFFF4FF330C34DD02FCCA" blockId="2.[151,447,810,870]" box="[151,316,846,870]" pageId="2" pageNumber="59">
(
<figureCitation id="133F6DCEFFCEFFF4FF3B0C34DCD3FCCA" box="[159,237,846,870]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1916,1939]" captionTargetBox="[161,1425,926,1890]" captionTargetId="figure@2.[161,1425,926,1890]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov." httpUri="https://zenodo.org/record/205437/files/figure.png" pageId="2" pageNumber="59">Figs. 1</figureCitation>
,
<figureCitation id="133F6DCEFFCEFFF4FF5E0C34DD0DFCCA" box="[250,307,846,870]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="2" pageNumber="59">4–14</figureCitation>
)
</paragraph>
<caption id="DF7B21C3FFCEFFF4FF330806DCFDF818" httpUri="https://zenodo.org/record/205437/files/figure.png" pageId="2" pageNumber="59" targetBox="[161,1425,926,1890]" targetPageId="2">
<paragraph id="8BBB714BFFCEFFF4FF330806DCFDF818" blockId="2.[151,1435,1916,1972]" pageId="2" pageNumber="59">
<emphasis id="B970AD59FFCEFFF4FF330806DD02F83E" bold="true" box="[151,316,1916,1939]" pageId="2" pageNumber="59">FIGURES 1–3.</emphasis>
Nymphs, dorsal habitus. 1,
<taxonomicName id="4C040AC8FFCEFFF4FDFA0807DF2DF83F" box="[606,787,1917,1939]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="dialutoi" status="sp. nov.">
<emphasis id="B970AD59FFCEFFF4FDFA0807DF2DF83F" box="[606,787,1917,1939]" italics="true" pageId="2" pageNumber="59">Cloeodes dialutoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFCEFFF4FCBF0807DF52F83E" box="[795,876,1917,1938]" pageId="2" pageNumber="59" rank="species">
<emphasis id="B970AD59FFCEFFF4FCBF0807DF52F83E" bold="true" box="[795,876,1917,1938]" pageId="2" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
2,
<taxonomicName id="4C040AC8FFCEFFF4FC2B0807D800F83F" box="[911,1086,1917,1939]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="guenoa" status="sp. nov.">
<emphasis id="B970AD59FFCEFFF4FC2B0807D800F83F" box="[911,1086,1917,1939]" italics="true" pageId="2" pageNumber="59">Cloeodes guenoa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFCEFFF4FBE20807D8A6F83E" box="[1094,1176,1917,1938]" pageId="2" pageNumber="59" rank="species">
<emphasis id="B970AD59FFCEFFF4FBE20807D8A6F83E" bold="true" box="[1094,1176,1917,1938]" pageId="2" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
3,
<taxonomicName id="4C040AC8FFCEFFF4FB1F0807D948F83F" box="[1211,1398,1917,1939]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="vaimaca" status="sp. nov.">
<emphasis id="B970AD59FFCEFFF4FB1F0807D948F83F" box="[1211,1398,1917,1939]" italics="true" pageId="2" pageNumber="59">Cloeodes vaimaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFCEFFF4FAD90807DCFDF818" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF7B21C3FFCFFFF5FF3308EAD8BEF847" httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="3" pageNumber="60" targetBox="[242,1339,215,1904]" targetPageId="3">
<paragraph id="8BBB714BFFCFFFF5FF3308EAD8BEF847" blockId="3.[151,1436,1935,2027]" pageId="3" pageNumber="60">
<emphasis id="B970AD59FFCFFFF5FF3308EADD77F809" bold="true" box="[151,329,1936,1959]" pageId="3" pageNumber="60">FIGURES 4–14.</emphasis>
<taxonomicName id="4C040AC8FFCFFFF5FEF408EADE38F80A" box="[336,518,1936,1958]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="dialutoi" status="sp. nov.">
<emphasis id="B970AD59FFCFFFF5FEF408EADE38F80A" box="[336,518,1936,1958]" italics="true" pageId="3" pageNumber="60">Cloeodes dialutoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFCFFFF5FDAA08EBDE5EF80A" box="[526,608,1937,1958]" pageId="3" pageNumber="60" rank="species">
<emphasis id="B970AD59FFCFFFF5FDAA08EBDE5EF80A" bold="true" box="[526,608,1937,1958]" pageId="3" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
Nymph. Mouthparts (Figs. 4–9): 4, labrum, left d.v., right v.v.; 5, left mandible v.v.; 6, right mandible v.v.; 7, hypopharynx v.v.; 8, maxilla v.v.; 9, labium, left d.v., right v.v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14a, cercus and terminal filament; 14b, cercus, detail of setae.
</paragraph>
</caption>
<paragraph id="8BBB714BFFC8FFF2FF330FEDD813FE2B" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
Nymph (
<figureCitation id="133F6DCEFFC8FFF2FF5F0FEDDD03FF03" box="[251,317,151,175]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1916,1939]" captionTargetBox="[161,1425,926,1890]" captionTargetId="figure@2.[161,1425,926,1890]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov." httpUri="https://zenodo.org/record/205437/files/figure.png" pageId="4" pageNumber="61">Fig. 1</figureCitation>
). Length: body,
<quantity id="4CFCDCAEFFC8FFF2FE5E0FEDDEBBFF03" box="[506,645,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="4" pageNumber="61" unit="mm" value="4.65" valueMax="4.7" valueMin="4.6">4.6–4.7 mm</quantity>
; cerci, 1.9–2.0 mm; terminal filament,
<quantity id="4CFCDCAEFFC8FFF2FB9A0FEDD8F7FF03" box="[1086,1225,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="61" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
. Antennae broken. Head yellowish brown, longer than wide, compound eyes reddish brown. Antennae pale yellow. Mouthparts (
<figureCitation id="133F6DCEFFC8FFF2FAC70FC6DCFBFF5B" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Figs. 4–9</figureCitation>
): labrum (
<figureCitation id="133F6DCEFFC8FFF2FE9E0FA5DD40FF5B" box="[314,382,223,247]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 4</figureCitation>
) with one subapical seta centrally, anterior margin with bifid setae (similar to
<figureCitation id="133F6DCEFFC8FFF2FB530FA5D976FF5B" box="[1271,1352,223,247]" captionStart="FIGURES 15 – 25" captionStartId="5.[151,264,1941,1964]" captionTargetBox="[222,1343,207,1894]" captionTargetId="figure@5.[212,1378,179,1930]" captionTargetPageId="5" captionText="FIGURES 15 – 25. Cloeodes guenoa sp. nov. Nymph. Mouthparts (Figs. 15 – 20): 15 a, labrum, left d. v., right v. v.; 15 b, labrum: apical setae; 16, left mandible v. v.; 17, right mandible v. v.; 18, hypopharynx v. v.; 19, maxilla v. v.; 20, labium, left d. v., right v. v. 21, leg I. 22, posterior margin of tergum IV. 23, paraproct. 24, gill IV. 25, cercus and terminal filament." httpUri="https://zenodo.org/record/205439/files/figure.png" pageId="4" pageNumber="61">Fig. 15</figureCitation>
b). Left mandible (
<figureCitation id="133F6DCEFFC8FFF2FEB40E7EDD6BFEB0" box="[272,341,260,284]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 5</figureCitation>
) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible (
<figureCitation id="133F6DCEFFC8FFF2FEFD0E5DDDA1FE93" box="[345,415,295,319]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 6</figureCitation>
) without setae between prostheca and mola. Hypopharynx (
<figureCitation id="133F6DCEFFC8FFF2FBF10E5DD8A0FE93" box="[1109,1182,295,319]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 7</figureCitation>
): lingua with rounded projection and subequal in length to superlinguae. Maxillae (
<figureCitation id="133F6DCEFFC8FFF2FCE40E36DFBAFEC8" box="[832,900,332,356]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 8</figureCitation>
), palpi as long as galea-lacinia, two segmented, segment II with a constriction. Labium (
<figureCitation id="133F6DCEFFC8FFF2FDFD0E15DEA3FE2B" box="[601,669,367,391]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 9</figureCitation>
) with segment III of palpi rounded.
</paragraph>
<paragraph id="8BBB714BFFC8FFF2FF610EEEDF11FDBB" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
Thorax yellowish brown. Fore wing pads yellowish. Pleura brownish, sterna pale yellow. Legs pale yellow, femora with median brownish bands, apical margin of tibiae and basal margin of tarsi brownish. Dorsal margin of femora (
<figureCitation id="133F6DCEFFC8FFF2FF510EA6DD78FE58" box="[245,326,476,500]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 10</figureCitation>
) with a row of blunt spines, apically with a subquadrangular projection and two blunt spines. Tarsal claws 0.3 times the length of tarsi.
<collectingCountry id="F31331DBFFC8FFF2FDBA0E85DE68FDBB" box="[542,598,511,535]" name="India" pageId="4" pageNumber="61">Hind</collectingCountry>
wing pads present.
</paragraph>
<paragraph id="8BBB714BFFC8FFF2FF610D5EDD32FCB8" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
Abdomen as in
<figureCitation id="133F6DCEFFC8FFF2FED80D5EDDFEFD90" box="[380,448,548,572]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1916,1939]" captionTargetBox="[161,1425,926,1890]" captionTargetId="figure@2.[161,1425,926,1890]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov." httpUri="https://zenodo.org/record/205437/files/figure.png" pageId="4" pageNumber="61">Fig. 1</figureCitation>
, segments I, IV, VIII with reddish markings, segments II-III, VI-VII, IX-X brownish, segment V pale brown, segments I-VI with two pale spots near midline. Abdominal terga with covers and some scales (following
<bibRefCitation id="EF950CBAFFC8FFF2FEFA0D16DDD5FD28" author="Kluge" box="[350,491,620,644]" pageId="4" pageNumber="61" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. Pages 527 - 535 in Landolt P; Sartori M. Ephemeroptera &amp; Plecoptera. Biology-Ecology-Systematics. Mauron + Tinguely &amp; Lachat, SA. Fribourg." type="book chapter" year="1997">Kluge, 1997</bibRefCitation>
), posterior margin of terga with spines as in
<figureCitation id="133F6DCEFFC8FFF2FC780D16D813FD28" box="[988,1069,620,644]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 11</figureCitation>
. Paraprocts (
<figureCitation id="133F6DCEFFC8FFF2FB190D16D930FD28" box="[1213,1294,620,644]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 12</figureCitation>
) with 12–13 spines apically. Gills (
<figureCitation id="133F6DCEFFC8FFF2FE2B0DF5DDDEFD0B" box="[399,480,655,679]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 13</figureCitation>
) translucent, trachea pigmented, 2 times the length of each tergum. Caudal filaments pale yellow, except apically brownish, with flattened setae basally sclerotized (
<figureCitation id="133F6DCEFFC8FFF2FB840DCED84BFD60" box="[1056,1141,692,716]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 14</figureCitation>
b). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline every two segments (
<figureCitation id="133F6DCEFFC8FFF2FF3B0D86DCCCFCB8" box="[159,242,764,788]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 14</figureCitation>
a).
</paragraph>
<paragraph id="8BBB714BFFC8FFF2FF630C65DDACFC9B" blockId="4.[151,1437,151,1220]" box="[199,402,799,824]" pageId="4" pageNumber="61">
<emphasis id="B970AD59FFC8FFF2FF630C65DD23FC94" bold="true" box="[199,285,799,824]" pageId="4" pageNumber="61">Adults.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="C31E22C0FFC8FFF2FF630C3EDFEDFCD3" pageId="4" pageNumber="61" type="etymology">
<paragraph id="8BBB714BFFC8FFF2FF630C3EDFEDFCD3" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
<emphasis id="B970AD59FFC8FFF2FF630C3EDD70FCF0" bold="true" box="[199,334,836,860]" pageId="4" pageNumber="61">Etymology.</emphasis>
This species is dedicated to William Dialuto, because of him the richness of Rocha's ecosystems is known. His commitment to preservation of the environment is invaluable.
</paragraph>
</subSubSection>
<subSubSection id="C31E22C0FFC8FFF2FF630CF1DE91FB98" pageId="4" pageNumber="61" type="diagnosis">
<paragraph id="8BBB714BFFC8FFF2FF630CF1DE91FB98" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
<emphasis id="B970AD59FFC8FFF2FF630CF1DD7FFC08" bold="true" box="[199,321,907,932]" pageId="4" pageNumber="61">Diagnosis.</emphasis>
<taxonomicName id="4C040AC8FFC8FFF2FEED0CF6DE2EFC08" box="[329,528,908,932]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="dialutoi" status="sp. nov.">
<emphasis id="B970AD59FFC8FFF2FEED0CF6DE2EFC08" box="[329,528,908,932]" italics="true" pageId="4" pageNumber="61">Cloeodes dialutoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2431022FFC8FFF2FDB30CF6DE4CFC08" box="[535,626,908,932]" pageId="4" pageNumber="61" rank="species">
<emphasis id="B970AD59FFC8FFF2FDB30CF6DE4CFC08" bold="true" box="[535,626,908,932]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from the other species of the genus by the following combination of characters: 1) hind wing pads present; 2) labium with segment III of palpi rounded (
<figureCitation id="133F6DCEFFC8FFF2FA8D0CD5D950FC6B" box="[1321,1390,943,967]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 9</figureCitation>
); 3) dorsal margin of femora with a row of blunt spines, apically with a subquadrangular projection and two blunt spines (
<figureCitation id="133F6DCEFFC8FFF2FF480C8DDD7EFBA3" box="[236,320,1015,1039]" captionStart="FIGURES 4 – 14" captionStartId="3.[151,264,1936,1959]" captionTargetBox="[242,1339,215,1904]" captionTargetId="figure@3.[196,1396,193,1910]" captionTargetPageId="3" captionText="FIGURES 4 – 14. Cloeodes dialutoi sp. nov. Nymph. Mouthparts (Figs. 4 – 9): 4, labrum, left d. v., right v. v.; 5, left mandible v. v.; 6, right mandible v. v.; 7, hypopharynx v. v.; 8, maxilla v. v.; 9, labium, left d. v., right v. v. 10, leg I. 11, posterior margin of tergum IV. 12, paraproct. 13, gill IV. 14 a, cercus and terminal filament; 14 b, cercus, detail of setae." httpUri="https://zenodo.org/record/205438/files/figure.png" pageId="4" pageNumber="61">Fig. 10</figureCitation>
); 4) abdominal color pattern in
<figureCitation id="133F6DCEFFC8FFF2FD000C8DDED8FBA3" box="[676,742,1015,1039]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1916,1939]" captionTargetBox="[161,1425,926,1890]" captionTargetId="figure@2.[161,1425,926,1890]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Nymphs, dorsal habitus. 1, Cloeodes dialutoi sp. nov. 2, Cloeodes guenoa sp. nov. 3, Cloeodes vaimaca sp. nov." httpUri="https://zenodo.org/record/205437/files/figure.png" pageId="4" pageNumber="61">Fig. 1</figureCitation>
, with segments I, IV, VIII with reddish spots, segments II-III, VI-VII, IX-X brownish, segment V pale brown.
</paragraph>
</subSubSection>
<subSubSection id="C31E22C0FFC8FFF2FF630B45D95AFB68" pageId="4" pageNumber="61" type="materials_examined">
<paragraph id="8BBB714BFFC8FFF2FF630B45D95AFB68" blockId="4.[151,1437,151,1220]" pageId="4" pageNumber="61">
<emphasis id="B970AD59FFC8FFF2FF630B45DD08FBF4" bold="true" box="[199,310,1087,1112]" pageId="4" pageNumber="61">Material.</emphasis>
<typeStatus id="54BFCFE9FFC8FFF2FE9A0B3ADD98FBF4" box="[318,422,1088,1112]" pageId="4" pageNumber="61" type="holotype">Holotype</typeStatus>
male mature nymph:
<collectingCountry id="F31331DBFFC8FFF2FD070B3ADF12FBF4" box="[675,812,1088,1112]" name="Uruguay" pageId="4" pageNumber="61">URUGUAY</collectingCountry>
, Rocha, Paraje. El Oratorio, Finca Sr. Cabrera, Arroyo Sauce Chico de los Méndez,
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,
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,
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,
<date id="FFBA578BFFC8FFF2FC340B1ED82CFBD0" box="[912,1042,1124,1148]" pageId="4" pageNumber="61" value="2010-03-24">24/ 3/ 2010</date>
, Emmerich &amp; Molineri colls.
<typeStatus id="54BFCFE9FFC8FFF2FAC50B1FDCE8FB0D" pageId="4" pageNumber="61" type="paratype">Paratypes</typeStatus>
: six nymphs (one dissected) same data as
<typeStatus id="54BFCFE9FFC8FFF2FD740BF2DF08FB0C" box="[720,822,1160,1184]" pageId="4" pageNumber="61" type="holotype">holotype</typeStatus>
. Two nymphs: Maldonado, R 109, Cerro Cordillera, Arroyo sin nombre,
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,
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,
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.
</specimenCount>
<date id="FFBA578BFFC8FFF2FCF20BD6DFE7FB68" box="[854,985,1196,1220]" pageId="4" pageNumber="61" value="2010-03-23">23/ 3/ 2010</date>
. D. Emmerich &amp; C. Molineri colls.
</paragraph>
</subSubSection>
</treatment>
</document>
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