treatments-xml/data/03/95/D5/0395D56BF67FFFCF858BFDDBFD641DBF.xml

<document id="EEE283B43F0D1940695A8A29DB995F9F" ID-DOI="10.11646/zootaxa.5346.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8354610" ID-ZooBank="5C8322AF-F2F8-48A3-8684-93971F78D817" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695025223290" checkinUser="plazi" docAuthor="Vishnudattan, N. K., Rubal, Marcos &amp; Nandan, S. Bijoy" docDate="2023" docId="0395D56BF67FFFCF858BFDDBFD641DBF" docLanguage="en" docName="zootaxa.5346.2.4.pdf" docOrigin="Zootaxa 5346 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5346.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batillipes kalami Vishnudattan, Rubal &amp; Bijoy Nandan 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="169" masterDocId="FFACAD13F67DFFC9851CFF9CFFD71A21" masterDocTitle="A new species of Batillipes (Arthrotardigrada: Batillipedidae) from the mid littoral zone of the southeast coast of India" masterLastPageNumber="172" masterPageNumber="163" pageNumber="165" updateTime="1695025635594" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="527D23362399C6992E1CD9CF2D720840">A new species of Batillipes (Arthrotardigrada: Batillipedidae) from the mid littoral zone of the southeast coast of India</mods:title>
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<mods:namePart id="DEC264A9E79284D6C0545527249C2155">Vishnudattan, N. K.</mods:namePart>
<mods:affiliation id="0F29526195BC36D6FE78C23CE233FE42">Department of Marine Biology, Microbiology &amp; Biochemistry, School of Marine Sciences, Cochin University of Science &amp; Technology, Fine Arts Avenue, Cochin- 682016, Kerala, India.</mods:affiliation>
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<mods:namePart id="69C4F968F3304751EEFEA0608562438F">Rubal, Marcos</mods:namePart>
<mods:affiliation id="0EC9D5DC07C5E622CFD65F6E10B87FD5">Centre of Molecular and Environmental Biology (CBMA / ARNET), Department of Biology, University of Minho, 4710 - 057 Braga, Portugal.</mods:affiliation>
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<mods:namePart id="5957EB9226A8DC355D337BE7BD4195DB">Nandan, S. Bijoy</mods:namePart>
<mods:affiliation id="A8D555C784C0AEB8F3DF837AEFD9C877">Department of Marine Biology, Microbiology &amp; Biochemistry, School of Marine Sciences, Cochin University of Science &amp; Technology, Fine Arts Avenue, Cochin- 682016, Kerala, India.</mods:affiliation>
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<taxonomicName id="4C3C1FFEF67FFFCB858BFDDBFC931840" authority="Vishnudattan, Rubal &amp; Bijoy Nandan" authorityName="Vishnudattan, Rubal &amp; Bijoy Nandan" authorityYear="2023" box="[151,836,583,610]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="165" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67FFFCB858BFDDBFEB31843" bold="true" box="[151,356,583,610]" italics="true" pageId="2" pageNumber="165">Batillipes kalami</emphasis>
Vishnudattan, Rubal &amp; Bijoy Nandan
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(
<figureCitation id="130778F8F67FFFCB8583FDF7FEDC18A7" box="[159,267,619,646]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,2000,2024]" captionStartId-1="4.[151,250,1490,1514]" captionStartId-2="6.[151,250,989,1013]" captionStartId-3="7.[151,250,1723,1747]" captionTargetBox-0="[291,1295,270,1973]" captionTargetBox-1="[151,1435,179,1463]" captionTargetBox-2="[151,1436,179,964]" captionTargetBox-3="[181,1406,179,1698]" captionTargetId-0="figure-55@3.[291,1295,270,1973]" captionTargetId-1="figure-23@4.[151,1435,179,1463]" captionTargetId-2="figure-23@6.[151,1436,179,964]" captionTargetId-3="figure-17@7.[181,1406,179,1698]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Batillipes kalami sp. nov., dorsal drawing of holotype female [MB/SBN/VD9]." captionText-1="FIGURE 3. Batillipes kalami sp. nov., (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection." captionText-2="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." captionText-3="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi-0="http://doi.org/10.5281/zenodo.8354616" figureDoi-1="http://doi.org/10.5281/zenodo.8354618" figureDoi-2="http://doi.org/10.5281/zenodo.8354622" figureDoi-3="http://doi.org/10.5281/zenodo.8354624" httpUri-0="https://zenodo.org/record/8354616/files/figure.png" httpUri-1="https://zenodo.org/record/8354618/files/figure.png" httpUri-2="https://zenodo.org/record/8354622/files/figure.png" httpUri-3="https://zenodo.org/record/8354624/files/figure.png" pageId="2" pageNumber="165">Figs. 2–5</figureCitation>
, 
<tableCitation id="C6BE51C6F67FFFCB840AFDF7FEB018A4" box="[278,359,619,645]" captionStart="TABLE 1" captionStartId="5.[151,239,548,572]" captionText="TABLE 1. Morphometrics (μm) of Batillipes kalami sp. nov., n, number of specimens observed." pageId="2" pageNumber="165">Table 1</tableCitation>
)
</paragraph>
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<subSubSection id="C32637F6F67FFFCB858BFD2FFE2B1E30" pageId="2" pageNumber="165" type="diagnosis">
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<emphasis id="B948B86FF67FFFCB858BFD2FFEC518EC" bold="true" box="[151,274,691,717]" pageId="2" pageNumber="165">Diagnosis.</emphasis>
Medium sized 
<taxonomicName id="4C3C1FFEF67FFFCB84CEFD2FFDEA18EC" authorityName="Richters" authorityYear="1909" box="[466,573,691,717]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="165" phylum="Tardigrada" rank="genus">
<emphasis id="B948B86FF67FFFCB84CEFD2FFDEA18EC" box="[466,573,691,717]" italics="true" pageId="2" pageNumber="165">Batillipes</emphasis>
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with cephalic region comprising a pair of tubular unconstricted-smooth primary clavae and indistinct secondary clavae. Cephalic cirri with sharp distal tips. External cirri are separated from the cirrophore of the primary clavae and lateral cirri by an indentation. Head is distinct and separated from the body by a neck constriction followed by a semi-circular lateral body projection. Sensory spines found on all legs. Leg IV sensory organ is short and finger like with van der Land’s organ at the mid region and a slightly swollen terminal portion. Single pair of lateral body projection between leg I–II, while two pairs of lateral body projection found between leg II and III. Lateral body projection between leg III and IV is longer and blunt ended. Body cuticle including lateral processes are finely punctated. Female gonopore is rosette-like while it is circular in males. Medium sized, terminally sharpened caudal spike attached on a cup shaped caudal base. Toes 3 and 4 of leg IV are shortest and are equal in length.
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<emphasis id="B948B86FF67FFFCB85DBFB87FE6B1E14" bold="true" box="[199,444,1051,1077]" pageId="2" pageNumber="165">Materials examined.</emphasis>
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/SBN/VD9: adult female mounted in glycerol.
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<typeStatus id="5487DADFF67FFFCB85DBFBDCFEEE1E78" box="[199,313,1088,1113]" pageId="2" pageNumber="165">Paratypes</typeStatus>
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: 
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/SBN/VD10-11, VD13, VD15, and VD18: 
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females mounted in glycerol
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; 
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/SBN/ VD12, VD14, VD16-17: 
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males mounted in glycerol
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.
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<emphasis id="B948B86FF67FFFCB85DBFB14FE8A1E80" box="[199,349,1159,1185]" italics="true" pageId="2" pageNumber="165">Type locality</emphasis>
: 
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<taxonomicName id="4C3C1FFEF67FFFCB846DFB1BFBA51E80" authority="Coral Beach, Mandapam, Maraikayar Pattinam, TamilNadu" authorityName="Coral Beach, Mandapam, Maraikayar Pattinam, TamilNadu" box="[369,1138,1159,1185]" class="Amphibia" family="Microhylidae" genus="Mini" kingdom="Animalia" order="Anura" pageId="2" pageNumber="165" phylum="Chordata" rank="genus">Mini Coral Beach, Mandapam, Maraikayar Pattinam, TamilNadu</taxonomicName>
, southeast coast of 
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(
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, 
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.
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<emphasis id="B948B86FF67FFFCB85DBFB4CFE5C1EC8" box="[199,395,1231,1257]" italics="true" pageId="2" pageNumber="165">Type depositories</emphasis>
: 
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Ten specimens containing slides mounted in glycerol were submitted in the Marine Biology Museum (voucher no: 
<collectionCode id="ED2DFCB8F67FFFCB848BFB6FFE131F2C" box="[407,452,1267,1293]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="2" pageNumber="165">MB</collectionCode>
/SBN/VD9-18), of the 
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and Technology (
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, Cochin-682016, 
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, 
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<paragraph id="8B83647DF67FFFCB85DBFAA7FCEB1D6C" blockId="2.[151,1437,691,2014]" pageId="2" pageNumber="165">
<emphasis id="B948B86FF67FFFCB85DBFAA7FE3C1F74" bold="true" box="[199,491,1339,1365]" pageId="2" pageNumber="165">
Description of 
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.
</emphasis>
Adult female with a body length of 170.2 μm (from the anterior rim of cephalic region to the posterior end of body excluding caudal spike) and a width of 51.5 μm between third and fourth leg (
<figureCitation id="130778F8F67FFFCB8583FA1FFF381FBC" box="[159,239,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,250,2000,2024]" captionTargetBox="[291,1295,270,1973]" captionTargetId="figure-55@3.[291,1295,270,1973]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes kalami sp. nov., dorsal drawing of holotype female [MB/SBN/VD9]." figureDoi="http://doi.org/10.5281/zenodo.8354616" httpUri="https://zenodo.org/record/8354616/files/figure.png" pageId="2" pageNumber="165">Figs. 2</figureCitation>
, 
<figureCitation id="130778F8F67FFFCB85E0FA1FFEDB1FBC" box="[252,268,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[151,1435,179,1463]" captionTargetId="figure-23@4.[151,1435,179,1463]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes kalami sp. nov., (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection." figureDoi="http://doi.org/10.5281/zenodo.8354618" httpUri="https://zenodo.org/record/8354618/files/figure.png" pageId="2" pageNumber="165">3</figureCitation>
; 
<tableCitation id="C6BE51C6F67FFFCB8405FA1FFEBC1FBC" box="[281,363,1411,1437]" captionStart="TABLE 1" captionStartId="5.[151,239,548,572]" captionText="TABLE 1. Morphometrics (μm) of Batillipes kalami sp. nov., n, number of specimens observed." pageId="2" pageNumber="165">Table 1</tableCitation>
). Body width slightly increasing from cephalic to caudal region. Translucent body cuticle with fine punctations. Punctation is more evident in the ventral cuticle. Head is trapezoid in shape with a width of 45.4 μm bearing a set of cephalic appendages and separated from the body by a neck constriction. All the cephalic cirri are sharp tipped at the terminal portion. Internal cirri of 12.7 μm length bearing cirrophores (2.3 μm) are attached dorsally on the anterior rim of cephalic region (
<figureCitation id="130778F8F67FFFCB8781F98FFD211C0F" box="[669,758,1555,1582]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="2" pageNumber="165">Fig. 5A</figureCitation>
). Unpaired median cirrus (9.8 μm) attached to cirrophore (2.6 μm) is 4 μm distant from the anterior rim of mid cephalic region (
<figureCitation id="130778F8F67FFFCB8673F9ABFC1E1C70" box="[879,969,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="2" pageNumber="165">Fig. 4A</figureCitation>
). External cirri with indistinct cirrophores is 12.1 μm long (
<figureCitation id="130778F8F67FFFCB8444F9C7FE641C54" box="[344,435,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="2" pageNumber="165">Fig. 4A</figureCitation>
). Primary clava (11.7 μm) is smooth and unconstricted tubular structure with a blunt end (
<figureCitation id="130778F8F67FFFCB8583F9E3FF2E1CB8" box="[159,249,1663,1689]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="2" pageNumber="165">Fig. 4A</figureCitation>
). Lateral cirri, 29.3 μm long is found very proximal to primary clava which arise from the common pedestal (
<figureCitation id="130778F8F67FFFCB8583F93FFF2C1C9F" box="[159,251,1699,1726]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="2" pageNumber="165">Fig. 5A</figureCitation>
). Van der Land’s organ is present at the base of each primary clava (
<figureCitation id="130778F8F67FFFCB811BF93FFBB51C9C" box="[1031,1122,1699,1725]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="2" pageNumber="165">Fig. 4A</figureCitation>
). External cephalic cirri are separated from the cirrophore of the primary clavae and lateral cirri by an indentation (
<figureCitation id="130778F8F67FFFCB8142F95BFB6F1CC3" box="[1118,1208,1735,1762]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="2" pageNumber="165">Fig. 5A</figureCitation>
). Secondary clava is indistinct. Eye spots not visible. Ventral conical shaped mouth is 15.4 μm wide (
<figureCitation id="130778F8F67FFFCB8104F977FBA71D27" box="[1048,1136,1771,1798]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="2" pageNumber="165">Fig. 5B</figureCitation>
). Circular pharyngeal bulb with 21.5 μm diameter is 15.4 μm posterior to the anterior rim of cephalic region. Buccal tube, stylet and placoids were not clearly visible after mounting slide in the 
<typeStatus id="5487DADFF67FFFCB87CFF8AFFCEE1D6C" box="[723,825,1843,1869]" pageId="2" pageNumber="165">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B83647DF67FFFCA85DBF8CBFE541AD8" blockId="2.[151,1437,691,2014]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="166" pageId="2" pageNumber="165">
Five pairs of dorso-lateral body projections are present: a pair of semi-circular lateral projections, 9.9 μm long are found between the head and the first pair of legs. A blunt lateral body projection of 9.4 μm long are present between first and second leg pairs. Two pairs of blunt lateral projections are found between the second and third leg pairs. Among them first pair (11.5 μm) is attached more dorsally while the other pair (9.7 μm) is oriented a little more ventrally. Long blunt ended (18.4 μm), finely punctated-transparent lateral projections are present between third and fourth leg pairs. All lateral body projections are lined by finely punctated and thin transparent sheet at its margin (
<figureCitation id="130778F8F67EFFCA85E9FF43FE921AD8" box="[245,325,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,2000,2024]" captionTargetBox="[291,1295,270,1973]" captionTargetId="figure-55@3.[291,1295,270,1973]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes kalami sp. nov., dorsal drawing of holotype female [MB/SBN/VD9]." figureDoi="http://doi.org/10.5281/zenodo.8354616" httpUri="https://zenodo.org/record/8354616/files/figure.png" pageId="3" pageNumber="166">Figs. 2</figureCitation>
, 
<figureCitation id="130778F8F67EFFCA844DFF43FEA11AD8" box="[337,374,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[151,1435,179,1463]" captionTargetId="figure-23@4.[151,1435,179,1463]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes kalami sp. nov., (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection." figureDoi="http://doi.org/10.5281/zenodo.8354618" httpUri="https://zenodo.org/record/8354618/files/figure.png" pageId="3" pageNumber="166">3A</figureCitation>
).
</paragraph>
<caption id="DF4334F5F67EFFCA858BF84CFBEA1DC9" ID-DOI="http://doi.org/10.5281/zenodo.8354616" ID-Zenodo-Dep="8354616" box="[151,1085,2000,2024]" httpUri="https://zenodo.org/record/8354616/files/figure.png" pageId="3" pageNumber="166" startId="3.[151,250,2000,2024]" targetBox="[291,1295,270,1973]" targetPageId="3" targetType="figure">
<paragraph id="8B83647DF67EFFCA858BF84CFBEA1DC9" blockId="3.[151,1085,2000,2024]" box="[151,1085,2000,2024]" pageId="3" pageNumber="166">
<emphasis id="B948B86FF67EFFCA858BF84CFEC41DC9" bold="true" box="[151,275,2000,2024]" pageId="3" pageNumber="166">FIGURE 2.</emphasis>
<taxonomicName id="4C3C1FFEF67EFFCA8405F84CFE111DC9" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[281,454,2000,2024]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="166" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67EFFCA8405F84CFE111DC9" box="[281,454,2000,2024]" italics="true" pageId="3" pageNumber="166">Batillipes kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67EFFCA84D0F84CFDCA1DC9" bold="true" box="[460,541,2000,2024]" pageId="3" pageNumber="166">
<taxonomicNameLabel id="A27B0514F67EFFCA84D0F84CFDCA1DC9" box="[460,541,2000,2024]" pageId="3" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal drawing of holotype female [MB/SBN/VD9].
</paragraph>
</caption>
<caption id="DF4334F5F679FFCD858BFA4EFED21C13" ID-DOI="http://doi.org/10.5281/zenodo.8354618" ID-Zenodo-Dep="8354618" httpUri="https://zenodo.org/record/8354618/files/figure.png" pageId="4" pageNumber="167" startId="4.[151,250,1490,1514]" targetBox="[151,1435,179,1463]" targetPageId="4" targetType="figure">
<paragraph id="8B83647DF679FFCD858BFA4EFED21C13" blockId="4.[151,1437,1490,1586]" pageId="4" pageNumber="167">
<emphasis id="B948B86FF679FFCD858BFA4EFEC11FCB" bold="true" box="[151,278,1490,1514]" pageId="4" pageNumber="167">FIGURE 3.</emphasis>
<taxonomicName id="4C3C1FFEF679FFCD8403FA4EFE191FCB" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[287,462,1490,1514]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="167" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF679FFCD8403FA4EFE191FCB" box="[287,462,1490,1514]" italics="true" pageId="4" pageNumber="167">Batillipes kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF679FFCD84C4FA4EFDFC1FCB" bold="true" box="[472,555,1490,1514]" pageId="4" pageNumber="167">
<taxonomicNameLabel id="A27B0514F679FFCD84C4FA4EFDFC1FCB" box="[472,555,1490,1514]" pageId="4" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection.
</paragraph>
</caption>
<paragraph id="8B83647DF679FFCD85DBF9C5FED01D20" blockId="4.[151,1437,1625,2042]" pageId="4" pageNumber="167">
All the pair of legs possess sensory spines of varying length (9.5–12.8 μm) with sharp tip at its distal end except for leg IV sensory organ. Leg IV sensory organ is a finger like structure terminated by a slightly swollen tip (
<figureCitation id="130778F8F679FFCD8073F9E0FF6C1C9B" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="4" pageNumber="167">Fig. 4C</figureCitation>
), 11.7 μm long with a proximal portion and a distal portion separated by van der Land’s organ (
<figureCitation id="130778F8F679FFCD81C6F93CFAE41C9B" box="[1242,1331,1696,1722]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="4" pageNumber="167">Fig. 4C</figureCitation>
). Sharply pointed and long Cirri E (22.5 μm) are attached dorsolaterally between caudal lateral body projection and leg IV (
<figureCitation id="130778F8F679FFCD8583F97BFF2D1D20" box="[159,250,1767,1793]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="4" pageNumber="167">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8B83647DF679FFCD85DBF896FD461DDB" blockId="4.[151,1437,1625,2042]" pageId="4" pageNumber="167">
Telescopic legs with an average length of 27.8 μm and toes of varying length. Among the first three pair of legs, toe 2 and 4 are the shortest (6.1, 6 μm) and are of same length. Toe 1,3,6 (8.7–11.8 μm) are medium sized while the 5 
<superScript id="7C49C935F679FFCD85B9F8CDFF651D7E" attach="left" box="[165,178,1873,1887]" fontSize="6" pageId="4" pageNumber="167">th</superScript>
toe is the longest (17.1 μm). On the fourth pair of legs, toe 3 and 4 are almost sessile, of same length (5.6, 5.5 μm) and the shortest. Toe 1 (7.4 μm), toe 2 (11.9 μm) and toe 6 (7.3 μm) are longer than toes 3 and 4, while toe 5 (12.9 μm) is the longest (
<figureCitation id="130778F8F679FFCD84B2F804FE2A1D93" box="[430,509,1944,1970]" captionStart="FIGURE 2" captionStartId="3.[151,250,2000,2024]" captionTargetBox="[291,1295,270,1973]" captionTargetId="figure-55@3.[291,1295,270,1973]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes kalami sp. nov., dorsal drawing of holotype female [MB/SBN/VD9]." figureDoi="http://doi.org/10.5281/zenodo.8354616" httpUri="https://zenodo.org/record/8354616/files/figure.png" pageId="4" pageNumber="167">Figs. 2</figureCitation>
, 
<figureCitation id="130778F8F679FFCD8714F804FDFC1D93" box="[520,555,1944,1970]" captionStart="FIGURE 3" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[151,1435,179,1463]" captionTargetId="figure-23@4.[151,1435,179,1463]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes kalami sp. nov., (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection." figureDoi="http://doi.org/10.5281/zenodo.8354618" httpUri="https://zenodo.org/record/8354618/files/figure.png" pageId="4" pageNumber="167">3B</figureCitation>
, 
<figureCitation id="130778F8F679FFCD872BF804FD8C1D92" box="[567,603,1944,1971]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="4" pageNumber="167">5C</figureCitation>
). The orientation of the toes of leg IV corresponds to the group A proposed by 
<bibRefCitation id="EFAD198CF679FFCD85A7F820FE431DF7" author="Santos, E. &amp; Rubal, M. &amp; Veiga, P. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." box="[187,404,1979,2006]" pageId="4" pageNumber="167" pagination="1 - 32" refId="ref4574" refString="Santos, E., Rubal, M., Veiga, P., Da Rocha, C. M. C. &amp; Fontoura, P. (2018) Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species. European Journal of Taxonomy, 425, 1 - 32. https: // doi. org / 10.5852 / ejt. 2018.425" type="journal article" year="2018">Santos et al (2018)</bibRefCitation>
. Toes possess distally enlarged stalk with terminally attached ovoid and transparent suction discs (6.2 μm long x 5.2 μm wide on leg IV).
</paragraph>
<paragraph id="8B83647DF678FFCC85DBFF0BFCC21B3C" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="168">
Rosette-shaped female gonopore with 6.1 μm width is present 6.6 μm anterior to the anus and 29.1 μm above the caudal base (
<figureCitation id="130778F8F678FFCC8448FF27FE7A1AF4" box="[340,429,186,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1723,1747]" captionTargetBox="[181,1406,179,1698]" captionTargetId="figure-17@7.[181,1406,179,1698]" captionTargetPageId="7" captionText="FIGURE 5. Batillipes kalami sp. nov., LM images of (A) cephalic region showing head indentation; (B) ventral view of cephalic region showing mouth opening and toe pattern of first pair of legs; (C) ventral view of caudal region showing female gonopore and toe pattern of fourth pair of legs. ic, internal cirrus; lc, lateral cirrus; i, indentation; mo, mouth opening; go, gonopore." figureDoi="http://doi.org/10.5281/zenodo.8354624" httpUri="https://zenodo.org/record/8354624/files/figure.png" pageId="5" pageNumber="168">Fig. 5C</figureCitation>
). Caudal apparatus (22 μm long) consists of a cup shaped caudal base (10.1 μm long and 16.2 μm wide), where the base of caudal spike is attached. Caudal spike is 12 μm long with a sharp tip at its distal end lined by transparent sheet at its margin (
<figureCitation id="130778F8F678FFCC879BFE9FFD001B3C" box="[647,727,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,2000,2024]" captionTargetBox="[291,1295,270,1973]" captionTargetId="figure-55@3.[291,1295,270,1973]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes kalami sp. nov., dorsal drawing of holotype female [MB/SBN/VD9]." figureDoi="http://doi.org/10.5281/zenodo.8354616" httpUri="https://zenodo.org/record/8354616/files/figure.png" pageId="5" pageNumber="168">Figs. 2</figureCitation>
, 
<figureCitation id="130778F8F678FFCC87FFFE9FFCDF1B3C" box="[739,776,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[151,1435,179,1463]" captionTargetId="figure-23@4.[151,1435,179,1463]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes kalami sp. nov., (A) LM image of dorsal view of holotype female [MB/SBN/VD9] showing lateral body projections; (B) DIC image of dorsal view of paratype male [MB/SBN/VD14] showing toe patterns of leg I–IV; Bp, body projection." figureDoi="http://doi.org/10.5281/zenodo.8354618" httpUri="https://zenodo.org/record/8354618/files/figure.png" pageId="5" pageNumber="168">3A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32637F6F678FFCC85DBFEBBFD661BA8" pageId="5" pageNumber="168" type="discussion">
<paragraph id="8B83647DF678FFCC85DBFEBBFD661BA8" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC85DBFEBBFEEC1B60" bold="true" box="[199,315,295,321]" pageId="5" pageNumber="168">Remarks.</emphasis>
No sexual dimorphism was found in this species, where both sexes are morphologically and metrically much similar, with an only exception of having a circular gonopore in males. Buccal tube, stylet and placoids were clearly visible in few of the 
<typeStatus id="5487DADFF678FFCC84CDFEF3FDEB1BA8" box="[465,572,367,393]" pageId="5" pageNumber="168">paratypes</typeStatus>
(
<figureCitation id="130778F8F678FFCC8750FEF3FD731BA8" box="[588,676,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,989,1013]" captionTargetBox="[151,1436,179,964]" captionTargetId="figure-23@6.[151,1436,179,964]" captionTargetPageId="6" captionText="FIGURE 4. Batillipes kalami sp. nov., (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E." figureDoi="http://doi.org/10.5281/zenodo.8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="5" pageNumber="168">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32637F6F678FFCC85DBFE0FFE911BD4" pageId="5" pageNumber="168" type="etymology">
<paragraph id="8B83647DF678FFCC85DBFE0FFE911BD4" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC85DBFE0FFE9F1B8C" bold="true" box="[199,328,403,429]" pageId="5" pageNumber="168">Etymology</emphasis>
. The specific epithet, kalami refers to Dr. A. P. J. Abdul Kalam, Indian aerospace scientist and eleventh president of 
<collectingCountry id="F32B24EDF678FFCC8493FE2BFE1E1BF0" box="[399,457,439,465]" name="India" pageId="5" pageNumber="168">India</collectingCountry>
also known as the “Missile Man of 
<collectingCountry id="F32B24EDF678FFCC8672FE2BFC7D1BF0" box="[878,938,439,465]" name="India" pageId="5" pageNumber="168">India</collectingCountry>
” who was native to the 
<typeStatus id="5487DADFF678FFCC81DFFE2BFB241BF0" box="[1219,1267,439,465]" pageId="5" pageNumber="168">type</typeStatus>
locality of the current species.
</paragraph>
</subSubSection>
<caption id="DF4334F5F678FFCC858BFDB8FB77181D" box="[151,1184,548,573]" pageId="5" pageNumber="168" startId="5.[151,239,548,572]" targetType="table">
<paragraph id="8B83647DF678FFCC858BFDB8FB77181D" blockId="5.[151,1184,548,573]" box="[151,1184,548,573]" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC858BFDB8FEDD181D" bold="true" box="[151,266,548,572]" pageId="5" pageNumber="168">TABLE 1.</emphasis>
Morphometrics (μm) of 
<taxonomicName id="4C3C1FFEF678FFCC8708FDB8FD1D181D" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[532,714,548,572]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="168" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF678FFCC8708FDB8FD1D181D" box="[532,714,548,572]" italics="true" pageId="5" pageNumber="168">Batillipes kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF678FFCC87CCFDB9FCF1181C" bold="true" box="[720,806,549,573]" pageId="5" pageNumber="168">
<taxonomicNameLabel id="A27B0514F678FFCC87CCFDB9FCF1181C" box="[720,806,549,573]" pageId="5" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="B948B86FF678FFCC862FFDB9FC97181D" box="[819,832,549,572]" italics="true" pageId="5" pageNumber="168">n</emphasis>
, number of specimens observed.
</paragraph>
</caption>
<paragraph id="8B83647DF678FFCC85BFFDD0FAF11D93" pageId="5" pageNumber="168">
<table id="F93C96DDF678003685BFFDD0FA8E1D93" box="[163,1369,588,1970]" gridcols="4" gridrows="33" pageId="5" pageNumber="168">
<tr id="350C663FF678003685BFFDD0FA8E18A3" box="[163,1369,588,642]" gridrow="0" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFDD0FE5318A3" box="[163,388,588,642]" gridcol="0" gridrow="0" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC85BFFDD0FECB1845" bold="true" box="[163,284,588,612]" pageId="5" pageNumber="168">Characters</emphasis>
</th>
<th id="76DD0F43F67800368754FDD0FD7C18A3" box="[584,683,588,642]" gridcol="1" gridrow="0" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC8754FDD0FD7C1845" bold="true" box="[584,683,588,612]" pageId="5" pageNumber="168">Holotype</emphasis>
female
</th>
<th id="76DD0F43F678003687E1FDD0FBDA18A3" box="[765,1037,588,642]" gridcol="2" gridrow="0" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC867AFDD0FBDA18A3" bold="true" pageId="5" pageNumber="168">
Paratypes average females, 
<emphasis id="B948B86FF678FFCC86F8FDF6FC2518A0" bold="true" box="[996,1010,618,641]" italics="true" pageId="5" pageNumber="168">n</emphasis>
=5
</emphasis>
</th>
<th id="76DD0F43F67800368197FDD0FA8E18A3" box="[1163,1369,588,642]" gridcol="3" gridrow="0" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC81A0FDD0FA8E18A0" bold="true" pageId="5" pageNumber="168">
Paratypes average males, 
<emphasis id="B948B86FF678FFCC802CFDF6FAE918A0" bold="true" box="[1328,1342,618,641]" italics="true" pageId="5" pageNumber="168">n</emphasis>
=4
</emphasis>
</th>
</tr>
<tr id="350C663FF678003685BFFD0EFA8E188B" box="[163,1369,658,682]" gridrow="1" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFD0EFE53188B" box="[163,388,658,682]" gridcol="0" gridrow="1" pageId="5" pageNumber="168">
<emphasis id="B948B86FF678FFCC85BFFD0EFEFE188B" bold="true" box="[163,297,658,682]" pageId="5" pageNumber="168">Voucher No.</emphasis>
</th>
<td id="76DD0F43F67800368754FD0EFD7C188B" box="[584,683,658,682]" gridcol="1" gridrow="1" pageId="5" pageNumber="168">VD9</td>
<td id="76DD0F43F678003687E1FD0EFBDA188B" box="[765,1037,658,682]" gridcol="2" gridrow="1" pageId="5" pageNumber="168">VD10, 11, 13, 15, 18</td>
<td id="76DD0F43F67800368197FD0EFA8E188B" box="[1163,1369,658,682]" gridcol="3" gridrow="1" pageId="5" pageNumber="168">VD12, 14, 16, 17</td>
</tr>
<tr id="350C663FF678003685BFFD7EFA8E18DB" box="[163,1369,738,762]" gridrow="2" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFD7EFE5318DB" box="[163,388,738,762]" gridcol="0" gridrow="2" pageId="5" pageNumber="168">Body length</th>
<td id="76DD0F43F67800368754FD7EFD7C18DB" box="[584,683,738,762]" gridcol="1" gridrow="2" pageId="5" pageNumber="168">170.2</td>
<td id="76DD0F43F678003687E1FD7EFBDA18DB" box="[765,1037,738,762]" gridcol="2" gridrow="2" pageId="5" pageNumber="168">164.2 ± 4.9</td>
<td id="76DD0F43F67800368197FD7EFA8E18DB" box="[1163,1369,738,762]" gridcol="3" gridrow="2" pageId="5" pageNumber="168">162.5 ± 5.1</td>
</tr>
<tr id="350C663FF678003685BFFC96FA8E1902" box="[163,1369,778,803]" gridrow="3" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFC96FE531902" box="[163,388,778,803]" gridcol="0" gridrow="3" pageId="5" pageNumber="168">Body width</th>
<td id="76DD0F43F67800368754FC96FD7C1902" box="[584,683,778,803]" gridcol="1" gridrow="3" pageId="5" pageNumber="168">51.5</td>
<td id="76DD0F43F678003687E1FC96FBDA1902" box="[765,1037,778,803]" gridcol="2" gridrow="3" pageId="5" pageNumber="168">49.5 ± 2.8</td>
<td id="76DD0F43F67800368197FC96FA8E1902" box="[1163,1369,778,803]" gridcol="3" gridrow="3" pageId="5" pageNumber="168">49.8 ± 2.7</td>
</tr>
<tr id="350C663FF678003685BFFCAFFA8E196A" box="[163,1369,819,843]" gridrow="4" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFCAFFE53196A" box="[163,388,819,843]" gridcol="0" gridrow="4" pageId="5" pageNumber="168">Cephalic region width</th>
<td id="76DD0F43F67800368754FCAFFD7C196A" box="[584,683,819,843]" gridcol="1" gridrow="4" pageId="5" pageNumber="168">45.4</td>
<td id="76DD0F43F678003687E1FCAFFBDA196A" box="[765,1037,819,843]" gridcol="2" gridrow="4" pageId="5" pageNumber="168">44.9 ± 2.1</td>
<td id="76DD0F43F67800368197FCAFFA8E196A" box="[1163,1369,819,843]" gridcol="3" gridrow="4" pageId="5" pageNumber="168">45.2 ± 2.3</td>
</tr>
<tr id="350C663FF678003685BFFCC7FA8E1952" box="[163,1369,859,883]" gridrow="5" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFCC7FE531952" box="[163,388,859,883]" gridcol="0" gridrow="5" pageId="5" pageNumber="168">Median cirrus</th>
<td id="76DD0F43F67800368754FCC7FD7C1952" box="[584,683,859,883]" gridcol="1" gridrow="5" pageId="5" pageNumber="168">9.8</td>
<td id="76DD0F43F678003687E1FCC7FBDA1952" box="[765,1037,859,883]" gridcol="2" gridrow="5" pageId="5" pageNumber="168">9.7 ± 0.7</td>
<td id="76DD0F43F67800368197FCC7FA8E1952" box="[1163,1369,859,883]" gridcol="3" gridrow="5" pageId="5" pageNumber="168">9.8 ± 0.9</td>
</tr>
<tr id="350C663FF678003685BFFC1FFA8E19BA" box="[163,1369,899,923]" gridrow="6" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFC1FFE5319BA" box="[163,388,899,923]" gridcol="0" gridrow="6" pageId="5" pageNumber="168">Internal cirri</th>
<td id="76DD0F43F67800368754FC1FFD7C19BA" box="[584,683,899,923]" gridcol="1" gridrow="6" pageId="5" pageNumber="168">12.7</td>
<td id="76DD0F43F678003687E1FC1FFBDA19BA" box="[765,1037,899,923]" gridcol="2" gridrow="6" pageId="5" pageNumber="168">12.5 ± 1.2</td>
<td id="76DD0F43F67800368197FC1FFA8E19BA" box="[1163,1369,899,923]" gridcol="3" gridrow="6" pageId="5" pageNumber="168">12.2 ± 0.9</td>
</tr>
<tr id="350C663FF678003685BFFC37FA8E19E5" box="[163,1369,939,964]" gridrow="7" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFC37FE5319E5" box="[163,388,939,964]" gridcol="0" gridrow="7" pageId="5" pageNumber="168">External cirri</th>
<td id="76DD0F43F67800368754FC37FD7C19E5" box="[584,683,939,964]" gridcol="1" gridrow="7" pageId="5" pageNumber="168">12.1</td>
<td id="76DD0F43F678003687E1FC37FBDA19E5" box="[765,1037,939,964]" gridcol="2" gridrow="7" pageId="5" pageNumber="168">11.8 ± 1.5</td>
<td id="76DD0F43F67800368197FC37FA8E19E5" box="[1163,1369,939,964]" gridcol="3" gridrow="7" pageId="5" pageNumber="168">11.9 ± 1.2</td>
</tr>
<tr id="350C663FF678003685BFFC48FA8E19CD" box="[163,1369,980,1004]" gridrow="8" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFC48FE5319CD" box="[163,388,980,1004]" gridcol="0" gridrow="8" pageId="5" pageNumber="168">Lateral cirri</th>
<td id="76DD0F43F67800368754FC48FD7C19CD" box="[584,683,980,1004]" gridcol="1" gridrow="8" pageId="5" pageNumber="168">29.3</td>
<td id="76DD0F43F678003687E1FC48FBDA19CD" box="[765,1037,980,1004]" gridcol="2" gridrow="8" pageId="5" pageNumber="168">27.9 ± 0.9</td>
<td id="76DD0F43F67800368197FC48FA8E19CD" box="[1163,1369,980,1004]" gridcol="3" gridrow="8" pageId="5" pageNumber="168">28.0 ± 1.1</td>
</tr>
<tr id="350C663FF678003685BFFC60FA8E1E35" box="[163,1369,1020,1044]" gridrow="9" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFC60FE531E35" box="[163,388,1020,1044]" gridcol="0" gridrow="9" pageId="5" pageNumber="168">Primary clavae</th>
<td id="76DD0F43F67800368754FC60FD7C1E35" box="[584,683,1020,1044]" gridcol="1" gridrow="9" pageId="5" pageNumber="168">11.7</td>
<td id="76DD0F43F678003687E1FC60FBDA1E35" box="[765,1037,1020,1044]" gridcol="2" gridrow="9" pageId="5" pageNumber="168">10.9 ± 0.9</td>
<td id="76DD0F43F67800368197FC60FA8E1E35" box="[1163,1369,1020,1044]" gridcol="3" gridrow="9" pageId="5" pageNumber="168">11.4 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFFBB8FA8E1E1D" box="[163,1369,1060,1084]" gridrow="10" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFBB8FE531E1D" box="[163,388,1060,1084]" gridcol="0" gridrow="10" pageId="5" pageNumber="168">Cirri E</th>
<td id="76DD0F43F67800368754FBB8FD7C1E1D" box="[584,683,1060,1084]" gridcol="1" gridrow="10" pageId="5" pageNumber="168">22.5</td>
<td id="76DD0F43F678003687E1FBB8FBDA1E1D" box="[765,1037,1060,1084]" gridcol="2" gridrow="10" pageId="5" pageNumber="168">20.9 ± 1.5</td>
<td id="76DD0F43F67800368197FBB8FA8E1E1D" box="[1163,1369,1060,1084]" gridcol="3" gridrow="10" pageId="5" pageNumber="168">21.9 ± 1.2</td>
</tr>
<tr id="350C663FF678003685BFFBD0FA8E1E44" box="[163,1369,1100,1125]" gridrow="11" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFBD0FE531E44" box="[163,388,1100,1125]" gridcol="0" gridrow="11" pageId="5" pageNumber="168">Caudal apparatus</th>
<td id="76DD0F43F67800368754FBD0FD7C1E44" box="[584,683,1100,1125]" gridcol="1" gridrow="11" pageId="5" pageNumber="168">23.4</td>
<td id="76DD0F43F678003687E1FBD0FBDA1E44" box="[765,1037,1100,1125]" gridcol="2" gridrow="11" pageId="5" pageNumber="168">22.5 ± 3.9</td>
<td id="76DD0F43F67800368197FBD0FA8E1E44" box="[1163,1369,1100,1125]" gridcol="3" gridrow="11" pageId="5" pageNumber="168">22.8 ± 4.2</td>
</tr>
<tr id="350C663FF678003685BFFBE9FA8E1EAC" box="[163,1369,1141,1165]" gridrow="12" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFBE9FE531EAC" box="[163,388,1141,1165]" gridcol="0" gridrow="12" pageId="5" pageNumber="168">Body projection 1</th>
<td id="76DD0F43F67800368754FBE9FD7C1EAC" box="[584,683,1141,1165]" gridcol="1" gridrow="12" pageId="5" pageNumber="168">9.9</td>
<td id="76DD0F43F678003687E1FBE9FBDA1EAC" box="[765,1037,1141,1165]" gridcol="2" gridrow="12" pageId="5" pageNumber="168">9.6 ± 1.1</td>
<td id="76DD0F43F67800368197FBE9FA8E1EAC" box="[1163,1369,1141,1165]" gridcol="3" gridrow="12" pageId="5" pageNumber="168">9.6 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFFB01FA8E1E94" box="[163,1369,1181,1205]" gridrow="13" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFB01FE531E94" box="[163,388,1181,1205]" gridcol="0" gridrow="13" pageId="5" pageNumber="168">Body projection 2</th>
<td id="76DD0F43F67800368754FB01FD7C1E94" box="[584,683,1181,1205]" gridcol="1" gridrow="13" pageId="5" pageNumber="168">9.4</td>
<td id="76DD0F43F678003687E1FB01FBDA1E94" box="[765,1037,1181,1205]" gridcol="2" gridrow="13" pageId="5" pageNumber="168">9.2 ± 1.5</td>
<td id="76DD0F43F67800368197FB01FA8E1E94" box="[1163,1369,1181,1205]" gridcol="3" gridrow="13" pageId="5" pageNumber="168">9.2 ± 1.3</td>
</tr>
<tr id="350C663FF678003685BFFB59FA8E1EFC" box="[163,1369,1221,1245]" gridrow="14" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFB59FE531EFC" box="[163,388,1221,1245]" gridcol="0" gridrow="14" pageId="5" pageNumber="168">Body projection 3</th>
<td id="76DD0F43F67800368754FB59FD7C1EFC" box="[584,683,1221,1245]" gridcol="1" gridrow="14" pageId="5" pageNumber="168">11.5</td>
<td id="76DD0F43F678003687E1FB59FBDA1EFC" box="[765,1037,1221,1245]" gridcol="2" gridrow="14" pageId="5" pageNumber="168">11.6 ± 1.3</td>
<td id="76DD0F43F67800368197FB59FA8E1EFC" box="[1163,1369,1221,1245]" gridcol="3" gridrow="14" pageId="5" pageNumber="168">11.8 ± 1.4</td>
</tr>
<tr id="350C663FF678003685BFFB71FA8E1F27" box="[163,1369,1261,1286]" gridrow="15" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFB71FE531F27" box="[163,388,1261,1286]" gridcol="0" gridrow="15" pageId="5" pageNumber="168">Body projection 4</th>
<td id="76DD0F43F67800368754FB71FD7C1F27" box="[584,683,1261,1286]" gridcol="1" gridrow="15" pageId="5" pageNumber="168">9.7</td>
<td id="76DD0F43F678003687E1FB71FBDA1F27" box="[765,1037,1261,1286]" gridcol="2" gridrow="15" pageId="5" pageNumber="168">9.6 ± 0.8</td>
<td id="76DD0F43F67800368197FB71FA8E1F27" box="[1163,1369,1261,1286]" gridcol="3" gridrow="15" pageId="5" pageNumber="168">9.5 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFFA8AFA8E1F0F" box="[163,1369,1302,1326]" gridrow="16" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFA8AFE531F0F" box="[163,388,1302,1326]" gridcol="0" gridrow="16" pageId="5" pageNumber="168">Body projection 5</th>
<td id="76DD0F43F67800368754FA8AFD7C1F0F" box="[584,683,1302,1326]" gridcol="1" gridrow="16" pageId="5" pageNumber="168">18.4</td>
<td id="76DD0F43F678003687E1FA8AFBDA1F0F" box="[765,1037,1302,1326]" gridcol="2" gridrow="16" pageId="5" pageNumber="168">18.2 ± 1.4</td>
<td id="76DD0F43F67800368197FA8AFA8E1F0F" box="[1163,1369,1302,1326]" gridcol="3" gridrow="16" pageId="5" pageNumber="168">18.7 ± 2.0</td>
</tr>
<tr id="350C663FF678003685BFFAA2FA8E1F77" box="[163,1369,1342,1366]" gridrow="17" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFAA2FE531F77" box="[163,388,1342,1366]" gridcol="0" gridrow="17" pageId="5" pageNumber="168">Leg I–III: Toe 1</th>
<td id="76DD0F43F67800368754FAA2FD7C1F77" box="[584,683,1342,1366]" gridcol="1" gridrow="17" pageId="5" pageNumber="168">8.7</td>
<td id="76DD0F43F678003687E1FAA2FBDA1F77" box="[765,1037,1342,1366]" gridcol="2" gridrow="17" pageId="5" pageNumber="168">7.9 ± 0.9</td>
<td id="76DD0F43F67800368197FAA2FA8E1F77" box="[1163,1369,1342,1366]" gridcol="3" gridrow="17" pageId="5" pageNumber="168">8.0 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFFAFAFA8E1F5F" box="[163,1369,1382,1406]" gridrow="18" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFAFAFE531F5F" box="[163,388,1382,1406]" gridcol="0" gridrow="18" pageId="5" pageNumber="168">Toe 2</th>
<td id="76DD0F43F67800368754FAFAFD7C1F5F" box="[584,683,1382,1406]" gridcol="1" gridrow="18" pageId="5" pageNumber="168">6.1</td>
<td id="76DD0F43F678003687E1FAFAFBDA1F5F" box="[765,1037,1382,1406]" gridcol="2" gridrow="18" pageId="5" pageNumber="168">6.1 ± 0.7</td>
<td id="76DD0F43F67800368197FAFAFA8E1F5F" box="[1163,1369,1382,1406]" gridcol="3" gridrow="18" pageId="5" pageNumber="168">6.6 ± 0.8</td>
</tr>
<tr id="350C663FF678003685BFFA13FA8E1F86" box="[163,1369,1423,1447]" gridrow="19" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFA13FE531F86" box="[163,388,1423,1447]" gridcol="0" gridrow="19" pageId="5" pageNumber="168">Toe 3</th>
<td id="76DD0F43F67800368754FA13FD7C1F86" box="[584,683,1423,1447]" gridcol="1" gridrow="19" pageId="5" pageNumber="168">11.2</td>
<td id="76DD0F43F678003687E1FA13FBDA1F86" box="[765,1037,1423,1447]" gridcol="2" gridrow="19" pageId="5" pageNumber="168">10.8 ± 0.8</td>
<td id="76DD0F43F67800368197FA13FA8E1F86" box="[1163,1369,1423,1447]" gridcol="3" gridrow="19" pageId="5" pageNumber="168">11.7 ± 0.9</td>
</tr>
<tr id="350C663FF678003685BFFA2BFA8E1FEE" box="[163,1369,1463,1487]" gridrow="20" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFA2BFE531FEE" box="[163,388,1463,1487]" gridcol="0" gridrow="20" pageId="5" pageNumber="168">Toe 4</th>
<td id="76DD0F43F67800368754FA2BFD7C1FEE" box="[584,683,1463,1487]" gridcol="1" gridrow="20" pageId="5" pageNumber="168">6</td>
<td id="76DD0F43F678003687E1FA2BFBDA1FEE" box="[765,1037,1463,1487]" gridcol="2" gridrow="20" pageId="5" pageNumber="168">5.9 ± 0.4</td>
<td id="76DD0F43F67800368197FA2BFA8E1FEE" box="[1163,1369,1463,1487]" gridcol="3" gridrow="20" pageId="5" pageNumber="168">6.0 ± 0.6</td>
</tr>
<tr id="350C663FF678003685BFFA43FA8E1FD6" box="[163,1369,1503,1527]" gridrow="21" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFFA43FE531FD6" box="[163,388,1503,1527]" gridcol="0" gridrow="21" pageId="5" pageNumber="168">Toe 5</th>
<td id="76DD0F43F67800368754FA43FD7C1FD6" box="[584,683,1503,1527]" gridcol="1" gridrow="21" pageId="5" pageNumber="168">17.1</td>
<td id="76DD0F43F678003687E1FA43FBDA1FD6" box="[765,1037,1503,1527]" gridcol="2" gridrow="21" pageId="5" pageNumber="168">15.5 ± 1.0</td>
<td id="76DD0F43F67800368197FA43FA8E1FD6" box="[1163,1369,1503,1527]" gridcol="3" gridrow="21" pageId="5" pageNumber="168">16.1 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFF99BFA8E1C3E" box="[163,1369,1543,1567]" gridrow="22" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF99BFE531C3E" box="[163,388,1543,1567]" gridcol="0" gridrow="22" pageId="5" pageNumber="168">Toe 6</th>
<td id="76DD0F43F67800368754F99BFD7C1C3E" box="[584,683,1543,1567]" gridcol="1" gridrow="22" pageId="5" pageNumber="168">11.8</td>
<td id="76DD0F43F678003687E1F99BFBDA1C3E" box="[765,1037,1543,1567]" gridcol="2" gridrow="22" pageId="5" pageNumber="168">10.1 ± 0.9</td>
<td id="76DD0F43F67800368197F99BFA8E1C3E" box="[1163,1369,1543,1567]" gridcol="3" gridrow="22" pageId="5" pageNumber="168">9.4 ± 1</td>
</tr>
<tr id="350C663FF678003685BFF9B3FA8E1C69" box="[163,1369,1583,1608]" gridrow="23" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF9B3FE531C69" box="[163,388,1583,1608]" gridcol="0" gridrow="23" pageId="5" pageNumber="168">Leg IV: Toe 1</th>
<td id="76DD0F43F67800368754F9B3FD7C1C69" box="[584,683,1583,1608]" gridcol="1" gridrow="23" pageId="5" pageNumber="168">7.4</td>
<td id="76DD0F43F678003687E1F9B3FBDA1C69" box="[765,1037,1583,1608]" gridcol="2" gridrow="23" pageId="5" pageNumber="168">7.6 ± 1.0</td>
<td id="76DD0F43F67800368197F9B3FA8E1C69" box="[1163,1369,1583,1608]" gridcol="3" gridrow="23" pageId="5" pageNumber="168">7.7 ± 1.1</td>
</tr>
<tr id="350C663FF678003685BFF9C4FA8E1C51" box="[163,1369,1624,1648]" gridrow="24" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF9C4FE531C51" box="[163,388,1624,1648]" gridcol="0" gridrow="24" pageId="5" pageNumber="168">Toe 2</th>
<td id="76DD0F43F67800368754F9C4FD7C1C51" box="[584,683,1624,1648]" gridcol="1" gridrow="24" pageId="5" pageNumber="168">11.9</td>
<td id="76DD0F43F678003687E1F9C4FBDA1C51" box="[765,1037,1624,1648]" gridcol="2" gridrow="24" pageId="5" pageNumber="168">11.7 ± 1.1</td>
<td id="76DD0F43F67800368197F9C4FA8E1C51" box="[1163,1369,1624,1648]" gridcol="3" gridrow="24" pageId="5" pageNumber="168">12.1 ± 0.7</td>
</tr>
<tr id="350C663FF678003685BFF91CFA8E1CB9" box="[163,1369,1664,1688]" gridrow="25" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF91CFE531CB9" box="[163,388,1664,1688]" gridcol="0" gridrow="25" pageId="5" pageNumber="168">Toe 3</th>
<td id="76DD0F43F67800368754F91CFD7C1CB9" box="[584,683,1664,1688]" gridcol="1" gridrow="25" pageId="5" pageNumber="168">5.6</td>
<td id="76DD0F43F678003687E1F91CFBDA1CB9" box="[765,1037,1664,1688]" gridcol="2" gridrow="25" pageId="5" pageNumber="168">5.9 ± 0.9</td>
<td id="76DD0F43F67800368197F91CFA8E1CB9" box="[1163,1369,1664,1688]" gridcol="3" gridrow="25" pageId="5" pageNumber="168">6.1 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFF934FA8E1CE1" box="[163,1369,1704,1728]" gridrow="26" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF934FE531CE1" box="[163,388,1704,1728]" gridcol="0" gridrow="26" pageId="5" pageNumber="168">Toe 4</th>
<td id="76DD0F43F67800368754F934FD7C1CE1" box="[584,683,1704,1728]" gridcol="1" gridrow="26" pageId="5" pageNumber="168">5.5</td>
<td id="76DD0F43F678003687E1F934FBDA1CE1" box="[765,1037,1704,1728]" gridcol="2" gridrow="26" pageId="5" pageNumber="168">5.6 ± 1.0</td>
<td id="76DD0F43F67800368197F934FA8E1CE1" box="[1163,1369,1704,1728]" gridcol="3" gridrow="26" pageId="5" pageNumber="168">5.4 ± 0.8</td>
</tr>
<tr id="350C663FF678003685BFF94CFA8E1CC9" box="[163,1369,1744,1768]" gridrow="27" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF94CFE531CC9" box="[163,388,1744,1768]" gridcol="0" gridrow="27" pageId="5" pageNumber="168">Toe 5</th>
<td id="76DD0F43F67800368754F94CFD7C1CC9" box="[584,683,1744,1768]" gridcol="1" gridrow="27" pageId="5" pageNumber="168">12.9</td>
<td id="76DD0F43F678003687E1F94CFBDA1CC9" box="[765,1037,1744,1768]" gridcol="2" gridrow="27" pageId="5" pageNumber="168">13.0 ± 1.2</td>
<td id="76DD0F43F67800368197F94CFA8E1CC9" box="[1163,1369,1744,1768]" gridcol="3" gridrow="27" pageId="5" pageNumber="168">12.2 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFF965FA8E1D30" box="[163,1369,1785,1809]" gridrow="28" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF965FE531D30" box="[163,388,1785,1809]" gridcol="0" gridrow="28" pageId="5" pageNumber="168">Toe 6</th>
<td id="76DD0F43F67800368754F965FD7C1D30" box="[584,683,1785,1809]" gridcol="1" gridrow="28" pageId="5" pageNumber="168">7.3</td>
<td id="76DD0F43F678003687E1F965FBDA1D30" box="[765,1037,1785,1809]" gridcol="2" gridrow="28" pageId="5" pageNumber="168">7.3 ± 0.8</td>
<td id="76DD0F43F67800368197F965FA8E1D30" box="[1163,1369,1785,1809]" gridcol="3" gridrow="28" pageId="5" pageNumber="168">7.4 ± 0.9</td>
</tr>
<tr id="350C663FF678003685BFF8BDFA8E1D18" box="[163,1369,1825,1849]" gridrow="29" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF8BDFE531D18" box="[163,388,1825,1849]" gridcol="0" gridrow="29" pageId="5" pageNumber="168">Leg I spine</th>
<td id="76DD0F43F67800368754F8BDFD7C1D18" box="[584,683,1825,1849]" gridcol="1" gridrow="29" pageId="5" pageNumber="168">9.5</td>
<td id="76DD0F43F678003687E1F8BDFBDA1D18" box="[765,1037,1825,1849]" gridcol="2" gridrow="29" pageId="5" pageNumber="168">10.0 ± 1.6</td>
<td id="76DD0F43F67800368197F8BDFA8E1D18" box="[1163,1369,1825,1849]" gridcol="3" gridrow="29" pageId="5" pageNumber="168">9.9 ± 1.0</td>
</tr>
<tr id="350C663FF678003685BFF8D5FA8E1D43" box="[163,1369,1865,1890]" gridrow="30" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF8D5FE531D43" box="[163,388,1865,1890]" gridcol="0" gridrow="30" pageId="5" pageNumber="168">Leg II spine</th>
<td id="76DD0F43F67800368754F8D5FD7C1D43" box="[584,683,1865,1890]" gridcol="1" gridrow="30" pageId="5" pageNumber="168">12.8</td>
<td id="76DD0F43F678003687E1F8D5FBDA1D43" box="[765,1037,1865,1890]" gridcol="2" gridrow="30" pageId="5" pageNumber="168">12.7 ± 2.1</td>
<td id="76DD0F43F67800368197F8D5FA8E1D43" box="[1163,1369,1865,1890]" gridcol="3" gridrow="30" pageId="5" pageNumber="168">12.5 ± 1.8</td>
</tr>
<tr id="350C663FF678003685BFF8EDFA8E1DAB" box="[163,1369,1905,1930]" gridrow="31" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF8EDFE531DAB" box="[163,388,1905,1930]" gridcol="0" gridrow="31" pageId="5" pageNumber="168">Leg III spine</th>
<td id="76DD0F43F67800368754F8EDFD7C1DAB" box="[584,683,1905,1930]" gridcol="1" gridrow="31" pageId="5" pageNumber="168">10.1</td>
<td id="76DD0F43F678003687E1F8EDFBDA1DAB" box="[765,1037,1905,1930]" gridcol="2" gridrow="31" pageId="5" pageNumber="168">11.2 ± 1.4</td>
<td id="76DD0F43F67800368197F8EDFA8E1DAB" box="[1163,1369,1905,1930]" gridcol="3" gridrow="31" pageId="5" pageNumber="168">11.9 ± 1.3</td>
</tr>
<tr id="350C663FF678003685BFF806FA8E1D93" box="[163,1369,1946,1970]" gridrow="32" pageId="5" pageNumber="168">
<th id="76DD0F43F678003685BFF806FE531D93" box="[163,388,1946,1970]" gridcol="0" gridrow="32" pageId="5" pageNumber="168">Leg IV sense organ</th>
<td id="76DD0F43F67800368754F806FD7C1D93" box="[584,683,1946,1970]" gridcol="1" gridrow="32" pageId="5" pageNumber="168">11.7</td>
<td id="76DD0F43F678003687E1F806FBDA1D93" box="[765,1037,1946,1970]" gridcol="2" gridrow="32" pageId="5" pageNumber="168">11.3 ± 1.8</td>
<td id="76DD0F43F67800368197F806FA8E1D93" box="[1163,1369,1946,1970]" gridcol="3" gridrow="32" pageId="5" pageNumber="168">12.1 ± 1.9</td>
</tr>
</table>
</paragraph>
<caption id="DF4334F5F67BFFCF858BFC41FE781E40" ID-DOI="http://doi.org/10.5281/zenodo.8354622" ID-Zenodo-Dep="8354622" httpUri="https://zenodo.org/record/8354622/files/figure.png" pageId="6" pageNumber="169" startId="6.[151,250,989,1013]" targetBox="[151,1436,179,964]" targetPageId="6" targetType="figure">
<paragraph id="8B83647DF67BFFCF858BFC41FE781E40" blockId="6.[151,1437,989,1122]" pageId="6" pageNumber="169">
<emphasis id="B948B86FF67BFFCF858BFC41FEC219D4" bold="true" box="[151,277,989,1013]" pageId="6" pageNumber="169">FIGURE 4.</emphasis>
<taxonomicName id="4C3C1FFEF67BFFCF8402FC42FE1A19D7" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[286,461,990,1014]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF8402FC42FE1A19D7" box="[286,461,990,1014]" italics="true" pageId="6" pageNumber="169">Batillipes kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF84C9FC42FDFF19D7" bold="true" box="[469,552,990,1014]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF84C9FC42FDFF19D7" box="[469,552,990,1014]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (A) DIC image of cephalic appendages; (B) LM image of the buccal apparatus and fine punctations on ventral cuticle; (C) PCM image of leg IV sensory organ; (D) DIC image of caudal region showing cirrus E; mc, median cirrus; ec, external cirrus; pc, primary clava; st, stylet; bt, buccal tube; pl, placoids; p, punctations; L–IV s, leg IV sensory organ; cE, cirrus E.
</paragraph>
</caption>
<subSubSection id="C32637F6F67BFFCF85DBFB0CFD641DBF" pageId="6" pageNumber="169" type="diagnosis">
<paragraph id="8B83647DF67BFFCF85DBFB0CFA8E1F37" blockId="6.[151,1437,1168,1950]" pageId="6" pageNumber="169">
<emphasis id="B948B86FF67BFFCF85DBFB0CFE061E8B" bold="true" box="[199,465,1168,1194]" pageId="6" pageNumber="169">Differential diagnosis.</emphasis>
<taxonomicName id="4C3C1FFEF67BFFCF84C7FB0CFD4B1E8B" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[475,668,1168,1194]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF84C7FB0CFD4B1E8B" box="[475,668,1168,1194]" italics="true" pageId="6" pageNumber="169">Batillipes kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF87BBFB0CFCD41E8B" bold="true" box="[679,771,1168,1194]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF87BBFB0CFCD41E8B" box="[679,771,1168,1194]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the group A species proposed by 
<bibRefCitation id="EFAD198CF67BFFCF800FFB0CFF331EEF" author="Santos, E. &amp; Rubal, M. &amp; Veiga, P. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." pageId="6" pageNumber="169" pagination="1 - 32" refId="ref4574" refString="Santos, E., Rubal, M., Veiga, P., Da Rocha, C. M. C. &amp; Fontoura, P. (2018) Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species. European Journal of Taxonomy, 425, 1 - 32. https: // doi. org / 10.5852 / ejt. 2018.425" type="journal article" year="2018">
Santos 
<emphasis id="B948B86FF67BFFCF8074FB0DFA4B1E8B" box="[1384,1436,1168,1194]" italics="true" pageId="6" pageNumber="169">et al</emphasis>
(2018)
</bibRefCitation>
, based on the toe pattern on the feet of leg IV. This major group contains 26 described species (
<bibRefCitation id="EFAD198CF67BFFCF800DFB28FF031ED3" author="Santos, E. &amp; Rubal, M. &amp; Veiga, P. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." pageId="6" pageNumber="169" pagination="1 - 32" refId="ref4574" refString="Santos, E., Rubal, M., Veiga, P., Da Rocha, C. M. C. &amp; Fontoura, P. (2018) Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species. European Journal of Taxonomy, 425, 1 - 32. https: // doi. org / 10.5852 / ejt. 2018.425" type="journal article" year="2018">
Santos 
<emphasis id="B948B86FF67BFFCF8078FB29FA4A1EEF" box="[1380,1437,1204,1230]" italics="true" pageId="6" pageNumber="169">et al.</emphasis>
2018
</bibRefCitation>
). However, only five species within this group have toes 3 and 4 sessile like 
<taxonomicName id="4C3C1FFEF67BFFCF810FFB45FBAA1ED3" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[1043,1149,1240,1266]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF810FFB45FBAA1ED3" box="[1043,1149,1240,1266]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF819EFB44FB0D1ED3" bold="true" box="[1154,1242,1240,1266]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF819EFB44FB0D1ED3" box="[1154,1242,1240,1266]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(i.e. 
<taxonomicName id="4C3C1FFEF67BFFCF800CFB45FA421ED3" authorityName="Tchesunov &amp; Mokievsky" authorityYear="1995" box="[1296,1429,1241,1266]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="crassipes">
<emphasis id="B948B86FF67BFFCF800CFB45FA421ED3" box="[1296,1429,1241,1266]" italics="true" pageId="6" pageNumber="169">B. crassipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C1FFEF67BFFCF858BFB61FECC1F37" authorityName="McGinty" authorityYear="1969" box="[151,283,1276,1302]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="gilmatini">
<emphasis id="B948B86FF67BFFCF858BFB61FECC1F37" box="[151,283,1276,1302]" italics="true" pageId="6" pageNumber="169">B. gilmatini</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C1FFEF67BFFCF8435FB61FE751F37" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[297,418,1276,1302]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="B948B86FF67BFFCF8435FB61FE751F37" box="[297,418,1276,1302]" italics="true" pageId="6" pageNumber="169">B. pennaki</emphasis>
</taxonomicName>
and two undescribed similar species to 
<taxonomicName id="4C3C1FFEF67BFFCF867CFB61FC0F1F37" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[864,984,1276,1302]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="B948B86FF67BFFCF867CFB61FC0F1F37" box="[864,984,1276,1302]" italics="true" pageId="6" pageNumber="169">B. pennaki</emphasis>
</taxonomicName>
reported by 
<bibRefCitation id="EFAD198CF67BFFCF817AFB60FA821F37" author="Santos, E. &amp; Veiga, P. &amp; Rubal, M. &amp; Bartels, P. J. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." box="[1126,1365,1276,1302]" pageId="6" pageNumber="169" pagination="549 - 567" refId="ref4655" refString="Santos, E., Veiga, P., Rubal, M., Bartels, P. J., Da Rocha, C. M. C. &amp; Fontoura, P. (2019 b) Batillipes pennaki Marcus, 1946 (Arthrotardigrada: Batillipedidae): Deciphering a species complex. Zootaxa, 4648, 549 - 567. https: // doi. org / 10.11646 / zootaxa. 4648.3.9" type="journal article" year="2019">
Santos 
<emphasis id="B948B86FF67BFFCF81A4FB61FB3E1F37" box="[1208,1257,1276,1302]" italics="true" pageId="6" pageNumber="169">et al</emphasis>
(2019b))
</bibRefCitation>
.
</paragraph>
<paragraph id="8B83647DF67BFFCF858BFABCFD5D1FEB" blockId="6.[151,1437,1168,1950]" pageId="6" pageNumber="169">
<taxonomicName id="4C3C1FFEF67BFFCF858BFABCFEA71F1B" authorityName="Tchesunov &amp; Mokievsky" authorityYear="1995" box="[151,368,1312,1338]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="crassipes">
<emphasis id="B948B86FF67BFFCF858BFABCFEA71F1B" box="[151,368,1312,1338]" italics="true" pageId="6" pageNumber="169">Batillipes crassipes</emphasis>
</taxonomicName>
can be easily distinguished from 
<taxonomicName id="4C3C1FFEF67BFFCF87FCFABDFC9D1F1B" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[736,842,1312,1338]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF87FCFABDFC9D1F1B" box="[736,842,1312,1338]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF8653FABCFC7F1F1B" bold="true" box="[847,936,1312,1338]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF8653FABCFC7F1F1B" box="[847,936,1312,1338]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the absence of lateral projections between legs III and IV, different shape and size of sensory organ on leg IV along with very short caudal spike (
<bibRefCitation id="EFAD198CF67BFFCF803DFAD8FEAD1FA2" author="Tchesunov, A. V. &amp; Mokievsky, V. O." pageId="6" pageNumber="169" pagination="153 - 158" refId="ref4731" refString="Tchesunov, A. V. &amp; Mokievsky, V. O. (1995) A new marine tardigrade, Batillipes crassipes sp. nov., from the Japan Sea (Tardigrada Arthrotardigrada, Batillipedidae). Cahiers de biologie Marine, 36 (2), 153 - 158." type="journal article" year="1995">Tchesunov &amp; Mokievsky, 1995</bibRefCitation>
) while 
<taxonomicName id="4C3C1FFEF67BFFCF84CDFAF5FDEA1FA3" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[465,573,1384,1410]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF84CDFAF5FDEA1FA3" box="[465,573,1384,1410]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF875AFAF4FD761FA3" bold="true" box="[582,673,1384,1410]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF875AFAF4FD761FA3" box="[582,673,1384,1410]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have distinct lateral projections between leg III and leg IV and the leg IV sensory organ is a finger like structure terminated by slightly swollen tip and the caudal spike is consistently conspicuous in all the examined individuals.
</paragraph>
<paragraph id="8B83647DF67BFFCF85DBFA48FDE51C5F" blockId="6.[151,1437,1168,1950]" pageId="6" pageNumber="169">
<taxonomicName id="4C3C1FFEF67BFFCF85DBFA48FE7A1FCF" authorityName="McGinty" authorityYear="1969" box="[199,429,1492,1518]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="gilmartini">
<emphasis id="B948B86FF67BFFCF85DBFA48FE7A1FCF" box="[199,429,1492,1518]" italics="true" pageId="6" pageNumber="169">Batillipes gilmartini</emphasis>
</taxonomicName>
can be differentiated from 
<taxonomicName id="4C3C1FFEF67BFFCF87ECFA49FC881FCF" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[752,863,1492,1518]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF87ECFA49FC881FCF" box="[752,863,1492,1518]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF8676FA48FC101FCF" bold="true" box="[874,967,1492,1518]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF8676FA48FC101FCF" box="[874,967,1492,1518]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the presence of a constriction in the primary clavae and incomplete dorsal plates that are absent in 
<taxonomicName id="4C3C1FFEF67BFFCF867AFA65FB4A1C33" authority="(McGinty, 1969)" baseAuthorityName="McGinty" baseAuthorityYear="1969" box="[870,1181,1528,1554]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami">
<emphasis id="B948B86FF67BFFCF867AFA65FC041C33" box="[870,979,1528,1554]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
(
<bibRefCitation id="EFAD198CF67BFFCF86F8FA64FB421C33" author="McGinty, M." box="[996,1173,1528,1554]" pageId="6" pageNumber="169" pagination="394 - 396" refId="ref4205" refString="McGinty, M. (1969) Notes. Batillipes gilmartini, a new marine tardigrade from a California Beach. Pacific Science, 23 (3), 394 - 396. https: // doi. org / 10.1016 / 0025 - 5416 (69) 90045 - 7" type="journal article" year="1969">McGinty, 1969</bibRefCitation>
)
</taxonomicName>
. The description of 
<taxonomicName id="4C3C1FFEF67BFFCF8098FA65FED01C17" authorityName="McGinty" authorityYear="1969" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="gilmartini">
<emphasis id="B948B86FF67BFFCF8098FA65FED01C17" italics="true" pageId="6" pageNumber="169">B. gilmartini</emphasis>
</taxonomicName>
is very incomplete and does not provide details about some characters, but the drawing of 
<taxonomicName id="4C3C1FFEF67BFFCF8010F981FA4B1C17" authorityName="McGinty" authorityYear="1969" box="[1292,1436,1564,1590]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="gilmartini">
<emphasis id="B948B86FF67BFFCF8010F981FA4B1C17" box="[1292,1436,1564,1590]" italics="true" pageId="6" pageNumber="169">B. gilmartini</emphasis>
</taxonomicName>
shows a single small lateral projection between legs III and IV while 
<taxonomicName id="4C3C1FFEF67BFFCF86BFF9DDFBD91C7B" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[931,1038,1600,1626]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF86BFF9DDFBD91C7B" box="[931,1038,1600,1626]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF810AF9DCFBA71C7B" bold="true" box="[1046,1136,1600,1626]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF810AF9DCFBA71C7B" box="[1046,1136,1600,1626]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows conspicuous lateral projections between all pairs of legs.
</paragraph>
<paragraph id="8B83647DF67BFFCF85DBF914FD641DBF" blockId="6.[151,1437,1168,1950]" pageId="6" pageNumber="169">
Finally, 
<taxonomicName id="4C3C1FFEF67BFFCF8438F915FE471C83" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[292,400,1672,1698]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF8438F915FE471C83" box="[292,400,1672,1698]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF8484F914FE241C83" bold="true" box="[408,499,1672,1698]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF8484F914FE241C83" box="[408,499,1672,1698]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="4C3C1FFEF67BFFCF87E0F915FBFC1C83" authority="(Marcus, 1946)" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[764,1067,1672,1698]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="B948B86FF67BFFCF87E0F915FCA21C83" box="[764,885,1672,1698]" italics="true" pageId="6" pageNumber="169">B. pennaki</emphasis>
(
<bibRefCitation id="EFAD198CF67BFFCF8699F914FBF51C83" author="Marcus, E." box="[901,1058,1672,1698]" pageId="6" pageNumber="169" pagination="1 - 3" refId="ref4166" refString="Marcus, E. (1946) Batillipes pennaki, a new marine tardigrade from the North and South American Atlantic coast. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 2, 1 - 3." type="journal article" year="1946">Marcus, 1946</bibRefCitation>
)
</taxonomicName>
and the two similar undescribed species by 
<bibRefCitation id="EFAD198CF67BFFCF8409F930FE2B1CE7" author="Santos, E. &amp; Veiga, P. &amp; Rubal, M. &amp; Bartels, P. J. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." box="[277,508,1708,1734]" pageId="6" pageNumber="169" pagination="549 - 567" refId="ref4655" refString="Santos, E., Veiga, P., Rubal, M., Bartels, P. J., Da Rocha, C. M. C. &amp; Fontoura, P. (2019 b) Batillipes pennaki Marcus, 1946 (Arthrotardigrada: Batillipedidae): Deciphering a species complex. Zootaxa, 4648, 549 - 567. https: // doi. org / 10.11646 / zootaxa. 4648.3.9" type="journal article" year="2019">
Santos 
<emphasis id="B948B86FF67BFFCF8474F931FE4C1CE7" box="[360,411,1708,1734]" italics="true" pageId="6" pageNumber="169">et al</emphasis>
(2019b)
</bibRefCitation>
by the lack of any constriction in the primary clavae in 
<taxonomicName id="4C3C1FFEF67BFFCF819DF931FB3A1CE7" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[1153,1261,1708,1734]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami">
<emphasis id="B948B86FF67BFFCF819DF931FB3A1CE7" box="[1153,1261,1708,1734]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
. Moreover, the shape of the sensory organ on leg IV in 
<taxonomicName id="4C3C1FFEF67BFFCF8777F94DFD0E1CCB" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[619,729,1744,1770]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF8777F94DFD0E1CCB" box="[619,729,1744,1770]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF87FEF94CFCE91CCB" bold="true" box="[738,830,1744,1770]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF87FEF94CFCE91CCB" box="[738,830,1744,1770]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to the undescribed species from European Atlantic shores but, it is different from 
<taxonomicName id="4C3C1FFEF67BFFCF874FF969FD1C1D2F" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[595,715,1780,1806]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="B948B86FF67BFFCF874FF969FD1C1D2F" box="[595,715,1780,1806]" italics="true" pageId="6" pageNumber="169">B. pennaki</emphasis>
</taxonomicName>
and the similar Mediterranean species that both have a conicalshaped distal portion terminated by a lance-like tip (
<bibRefCitation id="EFAD198CF67BFFCF87CDF884FC7D1D13" author="Santos, E. &amp; Veiga, P. &amp; Rubal, M. &amp; Bartels, P. J. &amp; Da Rocha, C. M. C. &amp; Fontoura, P." box="[721,938,1816,1842]" pageId="6" pageNumber="169" pagination="549 - 567" refId="ref4655" refString="Santos, E., Veiga, P., Rubal, M., Bartels, P. J., Da Rocha, C. M. C. &amp; Fontoura, P. (2019 b) Batillipes pennaki Marcus, 1946 (Arthrotardigrada: Batillipedidae): Deciphering a species complex. Zootaxa, 4648, 549 - 567. https: // doi. org / 10.11646 / zootaxa. 4648.3.9" type="journal article" year="2019">
Santos 
<emphasis id="B948B86FF67BFFCF863FF885FC8D1D13" box="[803,858,1816,1842]" italics="true" pageId="6" pageNumber="169">et al.</emphasis>
2019b
</bibRefCitation>
). Finally, 
<taxonomicName id="4C3C1FFEF67BFFCF8109F885FB5C1D13" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[1045,1163,1816,1842]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">
<emphasis id="B948B86FF67BFFCF8109F885FB5C1D13" box="[1045,1163,1816,1842]" italics="true" pageId="6" pageNumber="169">B. pennaki</emphasis>
</taxonomicName>
and the two undescribed similar species have a conspicuous lateral projection between legs III and IV. The lateral projection between legs III and IV in 
<taxonomicName id="4C3C1FFEF67BFFCF841AF8FDFEA61D5B" authority="Vishnudattan &amp; Rubal &amp; Nandan, 2023" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[262,369,1888,1914]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami" status="sp. nov.">
<emphasis id="B948B86FF67BFFCF841AF8FDFEA61D5B" box="[262,369,1888,1914]" italics="true" pageId="6" pageNumber="169">B. kalami</emphasis>
</taxonomicName>
<emphasis id="B948B86FF67BFFCF8464F8FCFE061D5B" bold="true" box="[376,465,1888,1914]" pageId="6" pageNumber="169">
<taxonomicNameLabel id="A27B0514F67BFFCF8464F8FCFE061D5B" box="[376,465,1888,1914]" pageId="6" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is smaller and more pointed than the one of 
<emphasis id="B948B86FF67BFFCF86DDF8FDFB971D5B" box="[961,1088,1888,1914]" italics="true" pageId="6" pageNumber="169">
<taxonomicName id="4C3C1FFEF67BFFCF86DDF8FDFBEB1D5B" baseAuthorityName="Marcus" baseAuthorityYear="1946" box="[961,1084,1888,1914]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="pennaki">B. pennaki</taxonomicName>
.
</emphasis>
Moreover, in 
<emphasis id="B948B86FF67BFFCF81C2F8FDFA981D5B" box="[1246,1359,1888,1914]" italics="true" pageId="6" pageNumber="169">
<taxonomicName id="4C3C1FFEF67BFFCF81C2F8FDFA9C1D5B" authorityName="Vishnudattan &amp; Rubal &amp; Nandan" authorityYear="2023" box="[1246,1355,1888,1914]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="169" phylum="Tardigrada" rank="species" species="kalami">B. kalami</taxonomicName>
,
</emphasis>
lateral projections are present between all pairs of legs.
</paragraph>
</subSubSection>
</treatment>
</document>