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<document id="908DA57FF1D34F1ED7374A733FBD0DD1" ID="10.11646/zootaxa.4250.5.2SLASH10743" ID-CLB-Dataset="33278" ID-DOI="10.11646/zootaxa.4250.5.2" ID-GBIF-Dataset="bccefe3e-8531-44a0-aa87-b1544cf3622d" ID-ISSN="1175-5326" ID-Zenodo-Dep="495523" ID-ZooBank="9FEDE52A-7950-449A-B4FB-0859055F1923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491550172294" checkinUser="plazi" docAuthor="Deler-Hernández, Albert, Fikáček, Martin &amp; Delgado, Juan A." docDate="2017" docId="03940D67FFE4FFE3FF36FBE71294F859" docLanguage="en" docName="zootaxa.4250.5.2SLASH10743.pdf" docOrigin="Zootaxa 4250 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydraena (Hydraenopsis) matthiasi Deler-Hernandez, Fikacek &amp; Delgado, new species" docType="treatment" docVersion="7" lastPageNumber="443" masterDocId="FFAD751FFFE1FFEAFFA1FFB9131DFFB7" masterDocTitle="The Hydraenidae of Cuba (Insecta: Coleoptera) III. Description of two new hygropetric species of Hydraena Kugelann" masterLastPageNumber="446" masterPageNumber="434" pageNumber="439" updateTime="1698438768642" updateUser="ExternalLinkService">
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<mods:title id="C69BFFB61560878570B3257836E3C7FB">The Hydraenidae of Cuba (Insecta: Coleoptera) III. Description of two new hygropetric species of Hydraena Kugelann</mods:title>
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<mods:namePart id="CCB9D7E65309CCA9BD4E09300C419C89">Deler-Hernández, Albert</mods:namePart>
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<mods:namePart id="A2977E1F6B34F77C4E4D4331AF512EE8">Fikáček, Martin</mods:namePart>
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<mods:namePart id="BC368E66E9E51A2A88AF6C28363AA92A">Delgado, Juan A.</mods:namePart>
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<mods:title id="185798EC4BD29F3740FC63B150812050">Zootaxa</mods:title>
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<mods:date id="7648FCD1FE2DE62FF693836BA327A432">2017</mods:date>
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<treatment id="03940D67FFE4FFE3FF36FBE71294F859" ID-DOI="http://doi.org/10.5281/zenodo.6000404" ID-GBIF-Taxon="128344202" ID-Zenodo-Dep="6000404" LSID="urn:lsid:plazi:treatment:03940D67FFE4FFE3FF36FBE71294F859" httpUri="http://treatment.plazi.org/id/03940D67FFE4FFE3FF36FBE71294F859" lastPageId="9" lastPageNumber="443" pageId="5" pageNumber="439">
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<paragraph id="8B82BC71FFE4FFEFFF36FBE717D8FBCF" blockId="5.[151,1221,1118,1177]" box="[151,1221,1118,1144]" pageId="5" pageNumber="439">
<heading id="D0CA0B1DFFE4FFEFFF36FBE717D8FBCF" bold="true" box="[151,1221,1118,1144]" fontSize="11" level="1" pageId="5" pageNumber="439" reason="1">
<emphasis id="B9496063FFE4FFEFFF36FBE717D8FBCF" bold="true" box="[151,1221,1118,1144]" pageId="5" pageNumber="439">
<taxonomicName id="4C3DC7F2FFE4FFEFFF36FBE71735FBCF" ID-CoL="3N3DG" authority="Deler-Hernandez, Fikacek &amp; Delgado" authorityName="Deler-Hernandez, Fikacek &amp; Delgado" box="[151,1064,1118,1144]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="matthiasi" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9496063FFE4FFEFFF36FBE7120DFBCF" bold="true" box="[151,272,1118,1144]" italics="true" pageId="5" pageNumber="439">Hydraena</emphasis>
(
<emphasis id="B9496063FFE4FFEFFE81FBE712DBFBCF" bold="true" box="[288,454,1118,1144]" italics="true" pageId="5" pageNumber="439">Hydraenopsis</emphasis>
)
<emphasis id="B9496063FFE4FFEFFE76FBE7115AFBCF" bold="true" box="[471,583,1118,1144]" italics="true" pageId="5" pageNumber="439">matthiasi</emphasis>
Deler-Hernández, Fikáček &amp; Delgado
</taxonomicName>
,
<taxonomicNameLabel id="A27ADD18FFE4FFEFFB94FBE717D8FBCF" box="[1077,1221,1118,1144]" pageId="5" pageNumber="439" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B82BC71FFE4FFEFFF36FB38128AFB2E" blockId="5.[151,1221,1118,1177]" box="[151,407,1153,1177]" pageId="5" pageNumber="439">
(
<figureCitation id="1306A0F4FFE4FFEFFF3EFB3813F4FB2E" box="[159,233,1153,1177]" captionStart="FIGURE 4" captionStartId="6.[151,249,1942,1964]" captionTargetBox="[223,1361,218,1901]" captionTargetId="figure@6.[201,1386,193,1921]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Hydraena matthiasi (holotype). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/495527/files/figure.png" pageId="5" pageNumber="439">Figs 4</figureCitation>
¯6; 7c, d; 8b, e)
</paragraph>
</subSubSection>
<subSubSection id="C327EFFAFFE4FFE2FF36FB7D1782F85C" lastPageId="8" lastPageNumber="442" pageId="5" pageNumber="439" type="materials_examined">
<paragraph id="8B82BC71FFE4FFEFFF36FB7D13A9FB4A" blockId="5.[151,1437,1220,2023]" pageId="5" pageNumber="439">
<materialsCitation id="3B55B62CFFE4FFEFFF36FB7D13A9FB4A" ID-GBIF-Occurrence="1455709080" collectorName="Aguada de Joaquin" country="Cuba" elevation="1359" latitude="20.01447" location="Granma" longLatPrecision="1" longitude="-76.84065" pageId="5" pageNumber="439" specimenCount="1" stateProvince="Granma" typeStatus="holotype">
<emphasis id="B9496063FFE4FFEFFF36FB7D122AFB6A" bold="true" box="[151,311,1220,1245]" pageId="5" pageNumber="439">
<typeStatus id="548602D3FFE4FFEFFF36FB7D13CFFB6A" box="[151,210,1220,1245]" pageId="5" pageNumber="439">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32AFCE1FFE4FFEFFE9EFB7D1260FB6A" box="[319,381,1220,1245]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
,
<collectingRegion id="49F97293FFE4FFEFFE2BFB7D12F9FB6B" box="[394,484,1220,1244]" country="Cuba" name="Granma" pageId="5" pageNumber="439">Granma</collectingRegion>
Prov., Turquino National Park,
<collectorName id="26C8D9A7FFE4FFEFFCF3FB7D1732FB6B" box="[850,1071,1220,1245]" pageId="5" pageNumber="439">Aguada de Joaquín</collectorName>
,
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<geoCoordinate id="EE09DAB6FFE4FFEFFB6AFB7D164AFB6A" box="[1227,1367,1220,1245]" direction="west" orientation="longitude" pageId="5" pageNumber="439" precision="1" value="-76.84065">76.84065°W</geoCoordinate>
,
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<quantity id="4CC51194FFE4FFEFFAC5FB7D13ADFB4A" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="5" pageNumber="439" unit="m" value="1359.0">1359 m</quantity>
</elevation>
.
</materialsCitation>
</paragraph>
<paragraph id="8B82BC71FFE4FFEFFF66FABF10C8FA64" blockId="5.[151,1437,1220,2023]" pageId="5" pageNumber="439">
<materialsCitation id="3B55B62CFFE4FFEFFF66FABF1776FAD0" ID-GBIF-Occurrence="1455709083" collectingDate="2014-09-27" collectionCode="NMPC" collectorName="Aguada de Joaquin &amp; A. Deler-Hernandez &amp; Hygropetric" country="Cuba" elevation="1359" latitude="20.01447" location="Aguada de Joaquin" longLatPrecision="1" longitude="-76.84065" municipality="Turquino National Park" pageId="5" pageNumber="439" specimenCount="3" stateProvince="Granma" typeStatus="holotype">
<emphasis id="B9496063FFE4FFEFFF66FABF1272FAA8" bold="true" box="[199,367,1286,1311]" pageId="5" pageNumber="439">
<typeStatus id="548602D3FFE4FFEFFF66FABF121FFAA8" box="[199,258,1286,1311]" pageId="5" pageNumber="439">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548602D3FFE4FFEFFEDFFABF12F6FAA8" box="[382,491,1286,1311]" pageId="5" pageNumber="439" type="holotype">
<emphasis id="B9496063FFE4FFEFFEDFFABF12F6FAA8" bold="true" box="[382,491,1286,1311]" pageId="5" pageNumber="439">Holotype</emphasis>
</typeStatus>
(male):
<collectingCountry id="F32AFCE1FFE4FFEFFDECFABF1193FAA8" box="[589,654,1286,1311]" name="Cuba" pageId="5" pageNumber="439">
<emphasis id="B9496063FFE4FFEFFDECFABF1193FAA8" bold="true" box="[589,654,1286,1311]" pageId="5" pageNumber="439">Cuba</emphasis>
</collectingCountry>
.
<collectingRegion id="49F97293FFE4FFEFFD3CFABF101FFAA8" box="[669,770,1286,1311]" country="Cuba" name="Granma" pageId="5" pageNumber="439">
<emphasis id="B9496063FFE4FFEFFD3CFABF101FFAA8" bold="true" box="[669,770,1286,1311]" pageId="5" pageNumber="439">Granma</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BE6260BFFE4FFEFFCB3FABF173CFAA8" box="[786,1057,1286,1311]" pageId="5" pageNumber="439">Turquino National Park</collectingMunicipality>
,
<collectorName id="26C8D9A7FFE4FFEFFB8CFABF1614FAA8" box="[1069,1289,1286,1311]" pageId="5" pageNumber="439">
<location id="8EE2EAAAFFE4FFEFFB8CFABF1614FAA8" LSID="urn:lsid:plazi:treatment:03940D67FFE4FFE3FF36FBE71294F859:8EE2EAAAFFE4FFEFFB8CFABF1614FAA8" box="[1069,1289,1286,1311]" country="Cuba" latitude="20.01447" longLatPrecision="1" longitude="-76.84065" municipality="Turquino National Park" name="Aguada de Joaquin" pageId="5" pageNumber="439" stateProvince="Granma">Aguada de Joaquín</location>
</collectorName>
,
<geoCoordinate id="EE09DAB6FFE4FFEFFAB7FABE1681FAA8" box="[1302,1436,1287,1311]" direction="north" orientation="latitude" pageId="5" pageNumber="439" precision="1" value="20.01447">20.01447°N</geoCoordinate>
<geoCoordinate id="EE09DAB6FFE4FFEFFF36FA931239FAF4" box="[151,292,1322,1347]" direction="west" orientation="longitude" pageId="5" pageNumber="439" precision="1" value="-76.84065">76.84065°W</geoCoordinate>
,
<elevation id="00105B42FFE4FFEFFE97FA931288FAF5" box="[310,405,1322,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="5" pageNumber="439" unit="m" value="1359.0">
<quantity id="4CC51194FFE4FFEFFE97FA931288FAF5" box="[310,405,1322,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="5" pageNumber="439" unit="m" value="1359.0">1359 m</quantity>
</elevation>
,
<collectingDate id="EFC76359FFE4FFEFFE09FA93113FFAF4" box="[424,546,1322,1347]" pageId="5" pageNumber="439" value="2014-09-27">
<date id="FF839AB1FFE4FFEFFE09FA93113FFAF4" box="[424,546,1322,1347]" pageId="5" pageNumber="439" value="2014-09-27">27.ix.2014</date>
</collectingDate>
, coll.,
<collectorName id="26C8D9A7FFE4FFEFFDDCFA921077FAF4" box="[637,874,1322,1347]" pageId="5" pageNumber="439">A. Deler-Hernández</collectorName>
,
<collectorName id="26C8D9A7FFE4FFEFFCDCFA931719FAF4" box="[893,1028,1322,1347]" pageId="5" pageNumber="439">Hygropetric</collectorName>
habitat [printed] / “
<typeStatus id="548602D3FFE4FFEFFB5EFA921685FAF4" box="[1279,1432,1323,1347]" pageId="5" pageNumber="439" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C3DC7F2FFE4FFEFFF36FAF612FBFAD1" authority="Det" authorityName="Det" box="[151,486,1358,1383]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="matthiasi">HYDRAENA matthiasi, Det</taxonomicName>
:.
<collectorName id="26C8D9A7FFE4FFEFFE56FAF611C1FAD0" box="[503,732,1358,1383]" pageId="5" pageNumber="439">A. Deler-Hernández</collectorName>
et al. 2016 [red printed] (
<collectionCode id="ED2C24B4FFE4FFEFFBACFAF6177EFAD0" box="[1037,1123,1359,1383]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="439">NMPC</collectionCode>
)
</materialsCitation>
.
<emphasis id="B9496063FFE4FFEFFBD9FAF717EDFAD0" bold="true" box="[1144,1264,1358,1383]" pageId="5" pageNumber="439">Paratypes</emphasis>
.
<materialsCitation id="3B55B62CFFE4FFEFFAA0FAF61139FA3C" ID-GBIF-Occurrence="1455709081" collectingDate="2014-09-27" collectionCode="NMPC" collectorName="Aguada de Joaquin &amp; A. Deler-Hernandez &amp; Hygropetric" country="Cuba" elevation="1359" latitude="20.01447" location="Aguada de Joaquin" longLatPrecision="1" longitude="-76.84065" municipality="Turquino National Park" pageId="5" pageNumber="439" specimenCount="17" stateProvince="Granma" typeStatus="holotype">
<specimenCount id="9D3B77F8FFE4FFEFFAA0FAF61681FAD0" box="[1281,1436,1358,1383]" pageId="5" pageNumber="439" type="generic">17 specimens</specimenCount>
with same data as for the
<typeStatus id="548602D3FFE4FFEFFE1FFACB1139FA3C" box="[446,548,1394,1419]" pageId="5" pageNumber="439" type="holotype">holotype</typeStatus>
</materialsCitation>
: 7 males, 4 females dry-mounted (NMPC, CDUM, NHMW, MCZ); 1 male, 2 females in alcohol (NMPC); 2 females in alcohol (IBE); 1 female, DNA extracted and dry-mounted after extraction (IBE/DNA voucher IBE-AN508). GenBank accession number
<accessionNumber id="946E2192FFE4FFEFFCFBFA0310CCFA64" box="[858,977,1466,1491]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LT627659" pageId="5" pageNumber="439">LT627659</accessionNumber>
.
</paragraph>
<paragraph id="8B82BC71FFE4FFEFFF66FA67129FF930" blockId="5.[151,1437,1220,2023]" pageId="5" pageNumber="439">
<emphasis id="B9496063FFE4FFEFFF66FA671241FA40" bold="true" box="[199,348,1502,1527]" pageId="5" pageNumber="439">Descriptions</emphasis>
. Male. Habitus as in
<figureCitation id="1306A0F4FFE4FFEFFDF3FA671185FA40" box="[594,664,1502,1527]" captionStart="FIGURE 4" captionStartId="6.[151,249,1942,1964]" captionTargetBox="[223,1361,218,1901]" captionTargetId="figure@6.[201,1386,193,1921]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Hydraena matthiasi (holotype). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/495527/files/figure.png" pageId="5" pageNumber="439">Fig. 4</figureCitation>
.
<emphasis id="B9496063FFE4FFEFFD05FA6711C3FA41" box="[676,734,1502,1526]" italics="true" pageId="5" pageNumber="439">Body</emphasis>
length (taken from anterior margin of labrum to elytral apex) about 1.40¯
<quantity id="4CC51194FFE4FFEFFEBCF9BA1295F9AD" box="[285,392,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="5" pageNumber="439" unit="mm" value="1.43">1.43 mm</quantity>
.
<emphasis id="B9496063FFE4FFEFFE34F9BA12F9F9AC" box="[405,484,1539,1563]" italics="true" pageId="5" pageNumber="439">Colour</emphasis>
: Head (dorsal) dark brown to black; pronotal disc dark brown; anterior angles and lateral half of anterior margin of pronotum slightly paler; elytra dark brown, with lateral margin, including the apical region, paler; maxillary palpi, antennae and legs pale brown to testaceous; distal half of terminal maxillary palpomere darkened.
</paragraph>
<paragraph id="8B82BC71FFE4FFEFFF66F92B13ECF959" blockId="5.[151,1437,1220,2023]" pageId="5" pageNumber="439">Head. Labrum moderately excised anteriorly; lobes evenly rounded. Clypeus with fine and sparse punctures. Frons finely sparsely punctuate. Clypeal and frontal punctures denser on lateral areas than medially; interstices shining.</paragraph>
<paragraph id="8B82BC71FFE4FFEDFF66F94F12DAF856" blockId="5.[151,1437,1220,2023]" lastBlockId="7.[151,1436,1776,2017]" lastPageId="7" lastPageNumber="441" pageId="5" pageNumber="439">
Thorax. Pronotum distinctly wider than long, widest near middle; anterior margin straight; anterior angles rounded, sides finely denticulate, slightly curved and convergent to anterior angle, sinuate and convergent to posterior border; anteroexternal and posteroexternal foveolae marked, internal foveolae less evident; punctures moderately impressed, of similar size to those of frons; interstices shining. Elytra elongate oval, lateral explanate margin moderately developed, not reaching apex; apices in dorsal view separately rounded; with 14¯15 (10 between suture and shoulder) rows of impressed punctures, rows quite regular; punctures round, moderately impressed, of approximately same size as on pronotum; interstices not elevated, shining to matt. Mesoventrite with internal and external carinae slightly divergent posteriad; median carina short and thin, not reaching base of intercoxal process, remarkably thinner than external and internal carinae; intercoxal process narrow, sides nearly parallel; apex blunt, width at apex approximately 0.3 distance between internal and median carinae. Base of mesocoxal cavities rounded. Metaventrite with plaques well developed, straight, slightly converging anteriorly; width of each plaque 1.5 width of intercoxal process; plaque separated approximately twice plaque width. Legs moderately short and stout; protibiae broad, with a small spine on inner face near apex (
<figureCitation id="1306A0F4FFE6FFEDFBC5F81D17BAF80B" box="[1124,1191,1956,1981]" captionStart="FIGURE 7" captionStartId="9.[151,249,1445,1467]" captionTargetBox="[177,1418,641,1411]" captionTargetId="figure@9.[151,1435,625,1434]" captionTargetPageId="9" captionText="FIGURE 7. Hydraena blancae (paratype male): a, protibia; b, metatibia. Hydraena matthiasi (paratype male): c, protibia; d, metatibia. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495530/files/figure.png" pageId="7" pageNumber="441">Fig. 7</figureCitation>
c); metatibiae arcuate, enlarged apically (
<figureCitation id="1306A0F4FFE6FFEDFEC7F87112B4F856" box="[358,425,1992,2017]" captionStart="FIGURE 7" captionStartId="9.[151,249,1445,1467]" captionTargetBox="[177,1418,641,1411]" captionTargetId="figure@9.[151,1435,625,1434]" captionTargetPageId="9" captionText="FIGURE 7. Hydraena blancae (paratype male): a, protibia; b, metatibia. Hydraena matthiasi (paratype male): c, protibia; d, metatibia. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495530/files/figure.png" pageId="7" pageNumber="441">Fig. 7</figureCitation>
d).
</paragraph>
<caption id="DF42ECF9FFE7FFECFF36F82F1095F81B" box="[151,904,1942,1964]" httpUri="https://zenodo.org/record/495527/files/figure.png" pageId="6" pageNumber="440" targetBox="[223,1361,218,1901]" targetPageId="6">
<paragraph id="8B82BC71FFE7FFECFF36F82F1095F81B" blockId="6.[151,904,1942,1964]" box="[151,904,1942,1964]" pageId="6" pageNumber="440">
<emphasis id="B9496063FFE7FFECFF36F82F120FF81B" bold="true" box="[151,274,1942,1964]" pageId="6" pageNumber="440">FIGURE 4.</emphasis>
Habitus of
<taxonomicName id="4C3DC7F2FFE7FFECFE28F82F114FF81B" box="[393,594,1942,1964]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="440" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE7FFECFE28F82F114FF81B" box="[393,594,1942,1964]" italics="true" pageId="6" pageNumber="440">Hydraena matthiasi</emphasis>
</taxonomicName>
(holotype). Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="DF42ECF9FFE6FFEDFF36F90A169BF97E" box="[151,1414,1715,1737]" httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="7" pageNumber="441" targetBox="[421,1152,177,1668]" targetPageId="7">
<paragraph id="8B82BC71FFE6FFEDFF36F90A169BF97E" blockId="7.[151,1414,1715,1737]" box="[151,1414,1715,1737]" pageId="7" pageNumber="441">
<emphasis id="B9496063FFE6FFEDFF36F90A120FF97F" bold="true" box="[151,274,1715,1737]" pageId="7" pageNumber="441">FIGURE 5.</emphasis>
<taxonomicName id="4C3DC7F2FFE6FFEDFEB9F90A12FFF97E" box="[280,482,1715,1737]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="441" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE6FFEDFEB9F90A12FFF97E" box="[280,482,1715,1737]" italics="true" pageId="7" pageNumber="441">Hydraena matthiasi</emphasis>
</taxonomicName>
(holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption id="DF42ECF9FFE9FFE2FF36F920104EF979" httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="8" pageNumber="442" targetBox="[228,1364,221,1650]" targetPageId="8">
<paragraph id="8B82BC71FFE9FFE2FF36F920104EF979" blockId="8.[151,1435,1689,1742]" pageId="8" pageNumber="442">
<emphasis id="B9496063FFE9FFE2FF36F920120EF919" bold="true" box="[151,275,1689,1711]" pageId="8" pageNumber="442">FIGURE 6.</emphasis>
<taxonomicName id="4C3DC7F2FFE9FFE2FEBAF92012F8F918" box="[283,485,1689,1711]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="442" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE9FFE2FEBAF92012F8F918" box="[283,485,1689,1711]" italics="true" pageId="8" pageNumber="442">Hydraena matthiasi</emphasis>
</taxonomicName>
male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; de, spermatheca. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B82BC71FFE9FFE2FF66F9431782F85C" blockId="8.[151,1437,1786,2027]" pageId="8" pageNumber="442">
Abdomen. Male terminal sternite and spiculum gastrale as in
<figureCitation id="1306A0F4FFE9FFE2FC30F94310CBF8A4" box="[913,982,1786,1811]" captionStart="FIGURE 6" captionStartId="8.[151,249,1689,1711]" captionTargetBox="[228,1364,221,1650]" captionTargetId="figure@8.[203,1383,193,1668]" captionTargetPageId="8" captionText="FIGURE 6. Hydraena matthiasi male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="8" pageNumber="442">Fig. 6</figureCitation>
a; terminal sternite subtrapezoidal, with anterior margin concave, posterior margin rounded and lateral margins sinuate; spiculum moderately long and curved, not firmly connected with sternite. Male genitalia as in
<figureCitation id="1306A0F4FFE9FFE2FCD3F8FB10A5F8EC" box="[882,952,1858,1883]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="8" pageNumber="442">Figs 5</figureCitation>
ac; main piece long, only slightly dilated distally both in lateral and dorsolateral views (
<figureCitation id="1306A0F4FFE9FFE2FD1CF8DF101BF8C8" box="[701,774,1894,1919]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="8" pageNumber="442">Figs 5</figureCitation>
bc) but not in ventral view (
<figureCitation id="1306A0F4FFE9FFE2FBC9F8DF17B2F8C8" box="[1128,1199,1894,1919]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="8" pageNumber="442">Fig. 5</figureCitation>
a); with two distinct processes at the apex: a slender, elongate gonopore-bearing process (flagellum) and a falcate laminar process; a subdistal lateral crooked projection of main piece (
<figureCitation id="1306A0F4FFE9FFE2FD7DF8171039F870" box="[732,804,1966,1991]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="8" pageNumber="442">Figs 5</figureCitation>
a-c) offers the impression of a third process; parameres inserted near midlenght of main piece, left paramere relatively long and slender, left shorter.
</paragraph>
</subSubSection>
<subSubSection id="C327EFFAFFE8FFE3FF66FF2E110CFED2" pageId="9" pageNumber="443" type="description">
<paragraph id="8B82BC71FFE8FFE3FF66FF2E110CFED2" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF66FF2E1203FF07" bold="true" box="[199,286,151,176]" pageId="9" pageNumber="443">Female</emphasis>
. Similar to male in shape and size; pro- and metatibiae not modified; gonocoxite as in
<figureCitation id="1306A0F4FFE8FFE3FAE3FF2E1697FF07" box="[1346,1418,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,249,1689,1711]" captionTargetBox="[228,1364,221,1650]" captionTargetId="figure@8.[203,1383,193,1668]" captionTargetPageId="8" captionText="FIGURE 6. Hydraena matthiasi male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="9" pageNumber="443">Fig. 6</figureCitation>
c, semicircular, transverse, condyles well developed, outer plate with trichoid setae on disc and lateroposterior areas and two tufts of subapical setae, inner plate distinctly exposed basally, cavea suboval; tergite X as in
<figureCitation id="1306A0F4FFE8FFE3FAE3FF661695FF4F" box="[1346,1416,223,248]" captionStart="FIGURE 6" captionStartId="8.[151,249,1689,1711]" captionTargetBox="[228,1364,221,1650]" captionTargetId="figure@8.[203,1383,193,1668]" captionTargetPageId="8" captionText="FIGURE 6. Hydraena matthiasi male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="9" pageNumber="443">Fig. 6</figureCitation>
b, suboval, with hyaline margin emarginated medially; spermatheca as in
<figureCitation id="1306A0F4FFE8FFE3FC67FEBD1710FEAA" box="[966,1037,260,285]" captionStart="FIGURE 6" captionStartId="8.[151,249,1689,1711]" captionTargetBox="[228,1364,221,1650]" captionTargetId="figure@8.[203,1383,193,1668]" captionTargetPageId="8" captionText="FIGURE 6. Hydraena matthiasi male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="9" pageNumber="443">Figs 6</figureCitation>
de, with proximal portion slightly curved, central portion enlarged and strongly curved and distal portion long; spermathecal gland smooth and rounded; spermathecal duct wide.
</paragraph>
</subSubSection>
<subSubSection id="C327EFFAFFE8FFE3FF66FED61270F88D" pageId="9" pageNumber="443" type="diagnosis">
<paragraph id="8B82BC71FFE8FFE3FF66FED61766FD8A" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF66FED612D6FE3F" bold="true" box="[199,459,367,392]" pageId="9" pageNumber="443">Differential diagnosis</emphasis>
.
<taxonomicName id="4C3DC7F2FFE8FFE3FE7EFEC911DEFE3F" box="[479,707,368,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FE7EFEC911DEFE3F" box="[479,707,368,392]" italics="true" pageId="9" pageNumber="443">Hydraena matthiasi</emphasis>
</taxonomicName>
is externally very close to
<taxonomicName id="4C3DC7F2FFE8FFE3FBB1FEC8178CFE3F" box="[1040,1169,368,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="blancae">
<emphasis id="B9496063FFE8FFE3FBB1FEC8178CFE3F" box="[1040,1169,368,392]" italics="true" pageId="9" pageNumber="443">H. blancae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3DC7F2FFE8FFE3FB70FEC81643FE3F" box="[1233,1374,368,392]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="franklyni">
<emphasis id="B9496063FFE8FFE3FB70FEC81643FE3F" box="[1233,1374,368,392]" italics="true" pageId="9" pageNumber="443">H. franklyni</emphasis>
</taxonomicName>
with similar body coloration and shape. However, it can be externally distinguished from
<taxonomicName id="4C3DC7F2FFE8FFE3FBD5FE2C17EFFE1B" box="[1140,1266,404,428]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="blancae">
<emphasis id="B9496063FFE8FFE3FBD5FE2C17EFFE1B" box="[1140,1266,404,428]" italics="true" pageId="9" pageNumber="443">H. blancae</emphasis>
</taxonomicName>
by its slightly larger size and by modified male metatibiae (
<figureCitation id="1306A0F4FFE8FFE3FD35FE0E11C1FE67" box="[660,732,439,464]" captionStart="FIGURE 7" captionStartId="9.[151,249,1445,1467]" captionTargetBox="[177,1418,641,1411]" captionTargetId="figure@9.[151,1435,625,1434]" captionTargetPageId="9" captionText="FIGURE 7. Hydraena blancae (paratype male): a, protibia; b, metatibia. Hydraena matthiasi (paratype male): c, protibia; d, metatibia. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495530/files/figure.png" pageId="9" pageNumber="443">Figs 7</figureCitation>
b, d). The shape of the metaventral plates can be also useful to distinguish both species, being wider in
<taxonomicName id="4C3DC7F2FFE8FFE3FDC9FE6411FBFE43" box="[616,742,476,500]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="blancae">
<emphasis id="B9496063FFE8FFE3FDC9FE6411FBFE43" box="[616,742,476,500]" italics="true" pageId="9" pageNumber="443">H. blancae</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FD56FE651026FE42" box="[759,827,476,501]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Fig. 8</figureCitation>
d) than in
<taxonomicName id="4C3DC7F2FFE8FFE3FC17FE64175EFE43" box="[950,1091,476,500]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FC17FE64175EFE43" box="[950,1091,476,500]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FBF4FE651784FE43" box="[1109,1177,476,501]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Fig. 8</figureCitation>
e) and, the base of the mesocoxal cavities rounded (
<figureCitation id="1306A0F4FFE8FFE3FE47FE461136FDAF" box="[486,555,511,536]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Fig. 8</figureCitation>
e). Furthermore,
<taxonomicName id="4C3DC7F2FFE8FFE3FD4FFDB81066FDAF" box="[750,891,512,536]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FD4FFDB81066FDAF" box="[750,891,512,536]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="4C3DC7F2FFE8FFE3FB6DFDB81657FDAF" box="[1228,1354,512,536]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="blancae">
<emphasis id="B9496063FFE8FFE3FB6DFDB81657FDAF" box="[1228,1354,512,536]" italics="true" pageId="9" pageNumber="443">H. blancae</emphasis>
</taxonomicName>
by the morphology of their aedeagi (
<figureCitation id="1306A0F4FFE8FFE3FE45FD9D1130FD8B" box="[484,557,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,249,1775,1797]" captionTargetBox="[409,1185,213,1739]" captionTargetId="figure@3.[391,1196,193,1753]" captionTargetPageId="3" captionText="FIGURE 2. Hydraena blancae (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495525/files/figure.png" pageId="9" pageNumber="443">Figs 2</figureCitation>
,
<figureCitation id="1306A0F4FFE8FFE3FD99FD9D115AFD8A" box="[568,583,548,573]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="9" pageNumber="443">5</figureCitation>
), and their male and female terminalia (
<figureCitation id="1306A0F4FFE8FFE3FBA9FD9D174DFD8B" box="[1032,1104,548,573]" captionStart="FIGURE 3" captionStartId="4.[151,249,1982,2004]" captionTargetBox="[220,1357,497,1943]" captionTargetId="figure@4.[203,1383,481,1961]" captionTargetPageId="4" captionText="FIGURE 3. Hydraena blancae male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495526/files/figure.png" pageId="9" pageNumber="443">Figs 3</figureCitation>
,
<figureCitation id="1306A0F4FFE8FFE3FBFCFD9D1770FD8A" box="[1117,1133,548,573]" captionStart="FIGURE 6" captionStartId="8.[151,249,1689,1711]" captionTargetBox="[228,1364,221,1650]" captionTargetId="figure@8.[203,1383,193,1668]" captionTargetPageId="8" captionText="FIGURE 6. Hydraena matthiasi male (holotype) and female (paratype) terminalia: a, male terminal sternite and spiculum; b, female tergite X; c, gonocoxite; d e, spermatheca. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495529/files/figure.png" pageId="9" pageNumber="443">6</figureCitation>
).
</paragraph>
<caption id="DF42ECF9FFE8FFE3FF36FA1C12A7FA6D" httpUri="https://zenodo.org/record/495530/files/figure.png" pageId="9" pageNumber="443" targetBox="[177,1418,641,1411]" targetPageId="9">
<paragraph id="8B82BC71FFE8FFE3FF36FA1C12A7FA6D" blockId="9.[151,1435,1445,1498]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF36FA1C120EFA0C" bold="true" box="[151,275,1445,1467]" pageId="9" pageNumber="443">FIGURE 7.</emphasis>
<taxonomicName id="4C3DC7F2FFE8FFE3FEBDFA1C12C4FA0C" box="[284,473,1445,1467]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="blancae">
<emphasis id="B9496063FFE8FFE3FEBDFA1C12C4FA0C" box="[284,473,1445,1467]" italics="true" pageId="9" pageNumber="443">Hydraena blancae</emphasis>
</taxonomicName>
(paratype male): a, protibia; b, metatibia.
<taxonomicName id="4C3DC7F2FFE8FFE3FC2EFA1C1747FA0C" box="[911,1114,1445,1467]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FC2EFA1C1747FA0C" box="[911,1114,1445,1467]" italics="true" pageId="9" pageNumber="443">Hydraena matthiasi</emphasis>
</taxonomicName>
(paratype male): c, protibia; d, metatibia. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B82BC71FFE8FFE3FF66F9B81270F88D" blockId="9.[151,1436,1537,2030]" pageId="9" pageNumber="443">
The evident similarities in the shape of the male and female genitalia indicate a very close relationship between
<taxonomicName id="4C3DC7F2FFE8FFE3FF36F99F123CF98A" box="[151,289,1573,1597]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FF36F99F123CF98A" box="[151,289,1573,1597]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3DC7F2FFE8FFE3FEF9F99F12C2F98A" box="[344,479,1573,1597]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="franklyni">
<emphasis id="B9496063FFE8FFE3FEF9F99F12C2F98A" box="[344,479,1573,1597]" italics="true" pageId="9" pageNumber="443">H. franklyni</emphasis>
</taxonomicName>
. They can only be reliably separated by a close examination of the male aedeagus. In ventral view, the apical portion of the right process of the distal lobe is clearly more angulate in
<taxonomicName id="4C3DC7F2FFE8FFE3FB7FF9F3167AF9D6" box="[1246,1383,1609,1633]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="franklyni">
<emphasis id="B9496063FFE8FFE3FB7FF9F3167AF9D6" box="[1246,1383,1609,1633]" italics="true" pageId="9" pageNumber="443">H. franklyni</emphasis>
</taxonomicName>
(see fig.
<quantity id="4CC51194FFE8FFE3FF65F9D713E0F931" box="[196,253,1645,1670]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="9" pageNumber="443" unit="in" value="34.0">34 in</quantity>
<bibRefCitation id="EFACC180FFE8FFE3FEA5F9D41197F931" author="Deler-Hernandez" box="[260,650,1645,1670]" pageId="9" pageNumber="443" refString="Deler-Hernandez, A. &amp; Delgado, J. A. (2012) The Hydraenidae of Cuba (Insecta: Coleoptera) I: Contribution to the fauna of eastern Cuba. Zootaxa, 3478, 213 - 238." type="journal article" year="2012">Deler-Hernández &amp; Delgado 2012</bibRefCitation>
), than in
<taxonomicName id="4C3DC7F2FFE8FFE3FD50F9D71066F932" box="[753,891,1645,1669]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FD50F9D71066F932" box="[753,891,1645,1669]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FC2AF9D410D3F931" box="[907,974,1645,1670]" captionStart="FIGURE 5" captionStartId="7.[151,249,1715,1737]" captionTargetBox="[421,1152,177,1668]" captionTargetId="figure@7.[410,1172,169,1683]" captionTargetPageId="7" captionText="FIGURE 5. Hydraena matthiasi (holotype): aedeagus in (a) ventral (b) lateral, and (c) dorsolateral views. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/495528/files/figure.png" pageId="9" pageNumber="443">Fig. 5</figureCitation>
c). In dorsolateral view, this angulation is not so evident but the main piece of
<taxonomicName id="4C3DC7F2FFE8FFE3FD90F92B11A4F91E" box="[561,697,1681,1705]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="franklyni">
<emphasis id="B9496063FFE8FFE3FD90F92B11A4F91E" box="[561,697,1681,1705]" italics="true" pageId="9" pageNumber="443">H. franklyni</emphasis>
</taxonomicName>
is clearly more globose (
<figureCitation id="1306A0F4FFE8FFE3FC74F9281706F91D" box="[981,1051,1681,1706]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Figs 8</figureCitation>
ab). The subdistal lateral process of the main piece is slender and less twisted in
<taxonomicName id="4C3DC7F2FFE8FFE3FD0DF90F102BF97A" box="[684,822,1717,1741]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FD0DF90F102BF97A" box="[684,822,1717,1741]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
and the flagellum is also slender in
<taxonomicName id="4C3DC7F2FFE8FFE3FB71F90F1646F97A" box="[1232,1371,1717,1741]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FB71F90F1646F97A" box="[1232,1371,1717,1741]" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FACBF90C13B9F945" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Figs 8</figureCitation>
ab). A less evident but useful character is the length of the mesoventral median carina, which reaches the base of the intercoxal process in
<taxonomicName id="4C3DC7F2FFE8FFE3FE12F9471126F8A2" box="[435,571,1789,1813]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="franklyni">
<emphasis id="B9496063FFE8FFE3FE12F9471126F8A2" box="[435,571,1789,1813]" italics="true" pageId="9" pageNumber="443">H. franklyni</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FDEAF9441190F8A1" box="[587,653,1789,1814]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Fig. 8</figureCitation>
c) being shorter, not reaching the base of the intercoxal process in
<taxonomicName id="4C3DC7F2FFE8FFE3FA20F94713E2F88E" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FA20F94713E2F88E" italics="true" pageId="9" pageNumber="443">H. matthiasi</emphasis>
</taxonomicName>
(
<figureCitation id="1306A0F4FFE8FFE3FEAFF898124CF88D" box="[270,337,1825,1850]" captionStart="FIGURE 8" captionStartId="10.[151,249,1228,1250]" captionTargetBox="[160,1424,204,1193]" captionTargetId="figure@10.[150,1436,195,1206]" captionTargetPageId="10" captionText="FIGURE 8. Aedeagi in dorsolateral views: a, Hydraena franklyni; b, Hydraena matthiasi. Thoracic venter (SEM images): c, Hydraena franklyni; d, Hydraena blancae; e, Hydraena matthiasi." httpUri="https://zenodo.org/record/495531/files/figure.png" pageId="9" pageNumber="443">Fig. 8</figureCitation>
e).
</paragraph>
</subSubSection>
<subSubSection id="C327EFFAFFE8FFE3FF66F8FC10E2F835" pageId="9" pageNumber="443" type="etymology">
<paragraph id="8B82BC71FFE8FFE3FF66F8FC10E2F835" blockId="9.[151,1436,1537,2030]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF66F8FC125BF8E9" bold="true" box="[199,326,1861,1886]" pageId="9" pageNumber="443">Etymology</emphasis>
. The new species is dedicated to Matthias Seidel, fellow co-worker of the first author at Charles University and a very enthusiastic entomologist specializing on
<taxonomicName id="4C3DC7F2FFE8FFE3FCC5F8D010E6F835" box="[868,1019,1897,1922]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C327EFFAFFE8FFE3FF66F8341294F859" pageId="9" pageNumber="443" type="distribution">
<paragraph id="8B82BC71FFE8FFE3FF66F834168CF811" blockId="9.[151,1436,1537,2030]" box="[199,1425,1933,1958]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF66F8341244F811" bold="true" box="[199,345,1933,1958]" pageId="9" pageNumber="443">Distribution</emphasis>
. Currently only known from the
<typeStatus id="548602D3FFE8FFE3FD6DF83711E1F811" box="[716,764,1934,1958]" pageId="9" pageNumber="443">type</typeStatus>
locality (Turquino National Park) in eastern
<collectingCountry id="F32AFCE1FFE8FFE3FB55F8341632F811" box="[1268,1327,1933,1958]" name="Cuba" pageId="9" pageNumber="443">Cuba</collectingCountry>
(
<figureCitation id="1306A0F4FFE8FFE3FA9FF834169EF811" box="[1342,1411,1933,1958]" captionStart="FIGURE 9" captionStartId="10.[151,249,1966,1988]" captionTargetBox="[320,1240,1343,1933]" captionTargetId="figure@10.[308,1278,1331,1945]" captionTargetPageId="10" captionText="FIGURE 9. Geographical distribution: Hydraena matthiasi (1); Hydraena franklyni (2); Hydraena blancae (3)." httpUri="https://zenodo.org/record/495532/files/figure.png" pageId="9" pageNumber="443">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B82BC71FFE8FFE3FF66F8081294F859" blockId="9.[151,1436,1537,2030]" pageId="9" pageNumber="443">
<emphasis id="B9496063FFE8FFE3FF66F808123EF87D" bold="true" box="[199,291,1969,1994]" pageId="9" pageNumber="443">Habitat</emphasis>
. Hygropetric.
<taxonomicName id="4C3DC7F2FFE8FFE3FE6CF80811ADF87E" box="[461,688,1969,1993]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="443" phylum="Arthropoda" rank="species" species="matthiasi">
<emphasis id="B9496063FFE8FFE3FE6CF80811ADF87E" box="[461,688,1969,1993]" italics="true" pageId="9" pageNumber="443">Hydraena matthiasi</emphasis>
</taxonomicName>
has been collected on wet rock faces in a well preserved cloud forest at
<quantity id="4CC51194FFE8FFE3FF58F86C1253F85A" box="[249,334,2005,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.359" pageId="9" pageNumber="443" unit="m" value="1359.0">1359 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>