186 lines
21 KiB
XML
186 lines
21 KiB
XML
<document id="A8395FBC03B3642700412F553F3FB856" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni & Giachino, Pier Mauro" docDate="2015" docId="039087C7FFBFFF96E28C483934656AF2" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) tardus Tschitscherine 1898" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="17" updateTime="1698599231025" updateUser="plazi">
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<mods:title id="ACD5274FF09BEAC5DFA3B726072B0EEC">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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<mods:namePart id="1CC70EADDA3A79083BFB57974AC539E5">Allegro, Gianni</mods:namePart>
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<mods:namePart id="0F216384BD22CF6475A35A349BD826A0">Giachino, Pier Mauro</mods:namePart>
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<mods:title id="1804731F9E59511CF888059C744A0516">Zootaxa</mods:title>
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<mods:date id="DBC6D24B6B7CBA3E2CB21A12407DC56B">2015</mods:date>
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<treatment id="039087C7FFBFFF96E28C483934656AF2" ID-DOI="http://doi.org/10.5281/zenodo.6096079" ID-GBIF-Taxon="127692725" ID-Zenodo-Dep="6096079" LSID="urn:lsid:plazi:treatment:039087C7FFBFFF96E28C483934656AF2" httpUri="http://treatment.plazi.org/id/039087C7FFBFFF96E28C483934656AF2" lastPageNumber="17" pageId="16" pageNumber="17">
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<subSubSection id="C323655AFFBFFF96E28C483934046CFD" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8B8636D1FFBFFF96E28C4839366C6C1C" blockId="16.[151,837,844,903]" box="[151,837,844,870]" pageId="16" pageNumber="17">
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<heading id="D0CE81BDFFBFFF96E28C4839366C6C1C" bold="true" box="[151,837,844,870]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="4C394D52FFBFFF96E28C4839366C6C1C" ID-CoL="M577" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[151,837,844,870]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tardus" subGenus="Agraphoderus">
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<emphasis id="B94DEAC3FFBFFF96E28C4839366C6C1C" bold="true" box="[151,837,844,870]" pageId="16" pageNumber="17">
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<emphasis id="B94DEAC3FFBFFF96E28C4839343A6C1C" bold="true" box="[151,275,844,870]" italics="true" pageId="16" pageNumber="17">Blennidus</emphasis>
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(
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<emphasis id="B94DEAC3FFBFFF96E33F483934F96C1C" bold="true" box="[292,464,844,870]" italics="true" pageId="16" pageNumber="17">Agraphoderus</emphasis>
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)
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<emphasis id="B94DEAC3FFBFFF96E3FA483937046C1C" bold="true" box="[481,557,844,870]" italics="true" pageId="16" pageNumber="17">tardus</emphasis>
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(
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<bibRefCitation id="EFA84B20FFBFFF96E026483936146C1C" author="Tschitscherine" box="[573,829,844,870]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8636D1FFBFFF96E28C481B34046CFD" blockId="16.[151,837,844,903]" box="[151,301,878,903]" pageId="16" pageNumber="17">
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(
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<figureCitation id="13022A54FFBFFF96E284481B35D56CFD" box="[159,252,878,903]" captionStart="FIGURES 44 – 47" captionStartId="35.[151,264,1953,1976]" captionTargetBox="[228,1350,210,1908]" captionTargetId="figure@35.[210,1377,193,1932]" captionTargetPageId="35" captionText="FIGURES 44 – 47. Habitus of Blennidus: sublaevis female PT (44); tardus female HT (45); tenenbaumi male HT (46); ticlianus male HT (47)." httpUri="https://zenodo.org/record/289231/files/figure.png" pageId="16" pageNumber="17">Figs. 45</figureCitation>
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,
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<figureCitation id="13022A54FFBFFF96E313481A340C6CFD" box="[264,293,879,903]" captionStart="FIGURES 82 – 87" captionStartId="41.[151,264,1498,1521]" captionTargetBox="[165,1418,199,1467]" captionTargetId="figure@41.[151,1436,193,1477]" captionTargetPageId="41" captionText="FIGURES 82 – 87. Pronotum of Blennidus: sublaevis female PT (82); tardus female HT (83); tenenbaumi male HT (84); ticlianus male HT (85); unistria male HT (86); vereshaginae male HT (87)." httpUri="https://zenodo.org/record/289236/files/figure.png" pageId="16" pageNumber="17">83</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C323655AFFBFFF96E28C48CD37D66B3D" pageId="16" pageNumber="17" type="reference_group">
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<paragraph id="8B8636D1FFBFFF96E28C48CD37076CB6" blockId="16.[151,1092,950,1095]" box="[151,558,950,973]" pageId="16" pageNumber="17">
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<treatmentCitationGroup id="AB2911FFFFBFFF96E28C48CD37076CB6" box="[151,558,950,973]" pageId="16" pageNumber="17">
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<taxonomicName id="4C394D52FFBFFF96E28C48CD37076CB6" authority="Tschitscherine, 1898: 161" authorityName="Tschitscherine" authorityPageNumber="161" authorityYear="1898" box="[151,558,950,973]" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tarda">
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<emphasis id="B94DEAC3FFBFFF96E28C48CD340D6CB6" box="[151,292,950,973]" italics="true" pageId="16" pageNumber="17">Feronia tarda</emphasis>
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<treatmentCitation id="0A9810C0FFBFFF96E33148C337076CB6" author="Tschitscherine" box="[298,558,950,972]" page="161" pageId="16" pageNumber="17" year="1898">
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<bibRefCitation id="EFA84B20FFBFFF96E33148C334D66CB6" author="Tschitscherine" box="[298,511,950,972]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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: 161
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</treatmentCitation>
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||
</taxonomicName>
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8636D1FFBFFF96E28C48A037D46B52" blockId="16.[151,1092,950,1095]" pageId="16" pageNumber="17">
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<treatmentCitationGroup id="AB2911FFFFBFFF96E28C48A037D46B52" pageId="16" pageNumber="17">
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<taxonomicName id="4C394D52FFBFFF96E28C48A037166C91" ID-CoL="48Z7Q" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[151,575,981,1003]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tarda">
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<emphasis id="B94DEAC3FFBFFF96E28C48A0347A6C91" box="[151,339,981,1003]" italics="true" pageId="16" pageNumber="17">Ogmopleura tarda</emphasis>
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(
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<bibRefCitation id="EFA84B20FFBFFF96E37B48A0371E6C91" author="Tschitscherine" box="[352,567,981,1003]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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)
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</taxonomicName>
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:
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<treatmentCitation id="0A9810C0FFBFFF96E05248A0368D6C91" author="Straneo" box="[585,932,981,1003]" page="199" pageId="16" pageNumber="17" year="1991">
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<bibRefCitation id="EFA84B20FFBFFF96E05248A0365C6C91" author="Straneo" box="[585,885,981,1003]" pageId="16" pageNumber="17" refString="Straneo, S. L. & Vereshagina, T. (1991) Sui tipi del genere Ogmopleura Tschitscherine descritti da Tschitscherine nel 1896 e 1898. Bollettino della Societa entomologica italiana, Genova, 122 (3), 195 - 204." type="journal article" year="1991">Straneo & Vereshagina, 1991</bibRefCitation>
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: 199
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||
</treatmentCitation>
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||
<taxonomicName id="4C394D52FFBFFF96E28C488137166B70" ID-CoL="48Z7Q" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[151,575,1012,1034]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tarda">
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<emphasis id="B94DEAC3FFBFFF96E28C4881347A6B70" box="[151,339,1012,1034]" italics="true" pageId="16" pageNumber="17">Ogmopleura tarda</emphasis>
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(
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<bibRefCitation id="EFA84B20FFBFFF96E37B4881371E6B70" author="Tschitscherine" box="[352,567,1012,1034]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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)
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</taxonomicName>
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:
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<treatmentCitation id="0A9810C0FFBFFF96E052488136216B70" author="Straneo" box="[585,776,1012,1034]" page="378" pageId="16" pageNumber="17" year="1993">
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<bibRefCitation id="EFA84B20FFBFFF96E052488137F16B70" author="Straneo" box="[585,728,1012,1034]" pageId="16" pageNumber="17" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale " G. Doria ", 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
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||
: 378
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||
</treatmentCitation>
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||
[=
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<taxonomicName id="4C394D52FFBFFF96E137488031176B70" ID-CoL="48Z66" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[812,1086,1012,1034]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anxia">
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<emphasis id="B94DEAC3FFBFFF96E1374880364D6B70" box="[812,868,1013,1034]" italics="true" pageId="16" pageNumber="17">anxia</emphasis>
|
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<bibRefCitation id="EFA84B20FFBFFF96E171488131176B70" author="Tschitscherine" box="[874,1086,1012,1034]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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</taxonomicName>
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]
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<taxonomicName id="4C394D52FFBFFF96E28C4F6737066B52" ID-CoL="M577" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[151,559,1042,1064]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tardus">
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<emphasis id="B94DEAC3FFBFFF96E28C4F67346B6B52" box="[151,322,1042,1064]" italics="true" pageId="16" pageNumber="17">Blennidus tardus</emphasis>
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(
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<bibRefCitation id="EFA84B20FFBFFF96E3544F67370F6B52" author="Tschitscherine" box="[335,550,1042,1064]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
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)
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||
</taxonomicName>
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||
:
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||
<treatmentCitation id="0A9810C0FFBFFF96E0234F6637D46B52" author="Lorenz" box="[568,765,1042,1064]" page="263" pageId="16" pageNumber="17" year="2005">
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<bibRefCitation id="EFA84B20FFBFFF96E0234F6637E76B52" author="Lorenz" box="[568,718,1042,1064]" pageId="16" pageNumber="17" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005" yearSuffix="a">Lorenz, 2005a</bibRefCitation>
|
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: 263
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</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8B8636D1FFBFFF96E28C4F4437D66B3D" blockId="16.[151,1092,950,1095]" box="[151,767,1073,1095]" pageId="16" pageNumber="17">
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<treatmentCitationGroup id="AB2911FFFFBFFF96E28C4F4437D66B3D" box="[151,767,1073,1095]" pageId="16" pageNumber="17">
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<taxonomicName id="4C394D52FFBFFF96E28C4F4437066B3D" ID-CoL="M577" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[151,559,1073,1095]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tardus">
|
||
<emphasis id="B94DEAC3FFBFFF96E28C4F44346B6B3D" box="[151,322,1073,1095]" italics="true" pageId="16" pageNumber="17">Blennidus tardus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA84B20FFBFFF96E3544F44370F6B3D" author="Tschitscherine" box="[335,550,1073,1095]" pageId="16" pageNumber="17" refString="Tschitscherine, T. (1898) Materiaux pour servir a l'etude des Feroniens. IV. Horae Societatis Entomologicae Rossicae, 32, 1 - 224." type="journal article" year="1898">Tschitschérine, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A9810C0FFBFFF96E0234F4737D66B3D" author="Lorenz" box="[568,767,1073,1095]" page="907" pageId="16" pageNumber="17" year="2005">
|
||
<bibRefCitation id="EFA84B20FFBFFF96E0234F4737E66B3D" author="Lorenz" box="[568,719,1073,1095]" pageId="16" pageNumber="17" refString="Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp." type="book" year="2005" yearSuffix="b">Lorenz, 2005b</bibRefCitation>
|
||
: 907
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||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323655AFFBFFF96E28C4F0134656AF2" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="8B8636D1FFBFFF96E28C4F01345E6BF6" blockId="16.[151,1437,1140,1416]" box="[151,375,1140,1165]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94DEAC3FFBFFF96E28C4F01341C6BF7" bold="true" box="[151,309,1140,1165]" pageId="16" pageNumber="17">
|
||
<typeStatus id="54828873FFBFFF96E28C4F0135FB6BF7" box="[151,210,1140,1165]" pageId="16" pageNumber="17">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<collectingCountry id="F32E7641FFBFFF96E3274F00345A6BF6" box="[316,371,1141,1164]" name="Peru" pageId="16" pageNumber="17">Peru</collectingCountry>
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||
.
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</paragraph>
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||
<paragraph id="8B8636D1FFBFFF96E2DC4FE237846BCA" blockId="16.[151,1437,1140,1416]" box="[199,685,1175,1200]" pageId="16" pageNumber="17">
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<emphasis id="B94DEAC3FFBFFF96E2DC4FE2349A6BCA" bold="true" box="[199,435,1175,1200]" pageId="16" pageNumber="17">Examined material.</emphasis>
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HT ♀ (CTs at
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<collectionCode id="ED28AE14FFBFFF96E0404FED37886BCA" box="[603,673,1176,1200]" pageId="16" pageNumber="17">ZIRA</collectionCode>
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||
).
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||
</paragraph>
|
||
<paragraph id="8B8636D1FFBFFF96E2DC4FC934656AF2" blockId="16.[151,1437,1140,1416]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94DEAC3FFBFFF96E2DC4FC9342F6BAF" bold="true" box="[199,262,1212,1237]" pageId="16" pageNumber="17">Note.</emphasis>
|
||
This species was described from a single ♀ specimen deposited in the Tschitschérine Collection (
|
||
<collectionCode id="ED28AE14FFBFFF96E7524FC930B96BAE" box="[1353,1424,1212,1236]" pageId="16" pageNumber="17">ZIRA</collectionCode>
|
||
).
|
||
<bibRefCitation id="EFA84B20FFBFFF96E28C4F9534C06B82" author="Straneo" box="[151,489,1247,1272]" pageId="16" pageNumber="17" refString="Straneo, S. L. & Vereshagina, T. (1991) Sui tipi del genere Ogmopleura Tschitscherine descritti da Tschitscherine nel 1896 e 1898. Bollettino della Societa entomologica italiana, Genova, 122 (3), 195 - 204." type="journal article" year="1991">Straneo & Vereshagina (1991)</bibRefCitation>
|
||
compared the HTs of
|
||
<taxonomicName id="4C394D52FFBFFF96E0F94F9536B66B82" box="[738,927,1248,1272]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tardus">
|
||
<emphasis id="B94DEAC3FFBFFF96E0F94F9536B66B82" box="[738,927,1248,1272]" italics="true" pageId="16" pageNumber="17">Blennidus tardus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D52FFBFFF96E1CE4F9531BC6B8D" box="[981,1173,1248,1272]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anxius">
|
||
<emphasis id="B94DEAC3FFBFFF96E1CE4F9531BC6B8D" box="[981,1173,1248,1272]" italics="true" pageId="16" pageNumber="17">Blennidus anxius</emphasis>
|
||
</taxonomicName>
|
||
noting some consistent differences; later
|
||
<bibRefCitation id="EFA84B20FFBFFF96E3474E71372A6A66" author="Straneo" box="[348,515,1284,1308]" pageId="16" pageNumber="17" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale " G. Doria ", 89, 351 - 399." type="journal article" year="1993">Straneo (1993)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="4C394D52FFBFFF96E0974E7037DA6A66" box="[652,755,1284,1308]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tardus">
|
||
<emphasis id="B94DEAC3FFBFFF96E0974E7037DA6A66" box="[652,755,1284,1308]" italics="true" pageId="16" pageNumber="17">B. tardus</emphasis>
|
||
</taxonomicName>
|
||
a synonym of
|
||
<taxonomicName id="4C394D52FFBFFF96E1874E70312C6A66" box="[924,1029,1285,1308]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anxius">
|
||
<emphasis id="B94DEAC3FFBFFF96E1874E70312C6A66" box="[924,1029,1285,1308]" italics="true" pageId="16" pageNumber="17">B. anxius</emphasis>
|
||
</taxonomicName>
|
||
, evaluating the differences between the 2 HT as intraspecific variability. Our study of the HTs of both species emphasized some evident differences, mainly in the morphology of pronotum; therefore we prefer to keep these species separate, in agreement with
|
||
<bibRefCitation id="EFA84B20FFBFFF96E28C4E05346E6AF2" author="Lorenz" box="[151,327,1391,1416]" pageId="16" pageNumber="17" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005" yearSuffix="a">Lorenz (2005a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |