418 lines
52 KiB
XML
418 lines
52 KiB
XML
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<mods:title id="B55768D913AFBD286E57A0785C1688CC">Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma)</mods:title>
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<mods:namePart id="EF3D70552F295C7E954DB3CEE2C93C9D">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="2406D48B99D0063D100234B83D8C2C5B">Wood, Perry L.</mods:namePart>
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<mods:namePart id="B370E579CD08A7B3733AE89DCF0A9A77">Zug, George R.</mods:namePart>
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<mods:namePart id="7C949D57316D2BC77D4A38023E43E070">Murdoch, Matthew L.</mods:namePart>
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<mods:namePart id="26215B16F810D1E67218C218E049934A">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="788D2A2AA45403D5D78083A3C1727ABB">Lin, Aung</mods:namePart>
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<treatment id="0390878F67430D2AF1C1F8CEFBF9FABF" ID-DOI="http://doi.org/10.5281/zenodo.5980023" ID-GBIF-Taxon="148403624" ID-Zenodo-Dep="5980023" LSID="urn:lsid:plazi:treatment:0390878F67430D2AF1C1F8CEFBF9FABF" httpUri="http://treatment.plazi.org/id/0390878F67430D2AF1C1F8CEFBF9FABF" lastPageId="17" lastPageNumber="312" pageId="15" pageNumber="310">
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<emphasis id="B94DEA8B67430D34F1C1F8CEFDEFF8A0" bold="true" box="[151,545,1840,1866]" pageId="15" pageNumber="310">
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<taxonomicName id="4C394D1A67430D34F1C1F8CEFE72F8A0" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[151,444,1840,1866]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="310" phylum="Chordata" rank="species" species="uga" status="sp. nov.">
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<emphasis id="B94DEA8B67430D34F1C1F8CEFE72F8A0" bold="true" box="[151,444,1840,1866]" italics="true" pageId="15" pageNumber="310">Hemiphyllodactylus uga</emphasis>
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<taxonomicNameLabel id="A27E57F067430D34F092F8CEFDEFF8A0" box="[452,545,1840,1866]" pageId="15" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B86369967430D34F1C1F8ADFE4BF886" blockId="15.[151,545,1840,1936]" box="[151,389,1875,1900]" pageId="15" pageNumber="310">Uga’s Slender Gecko</paragraph>
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<subSubSection id="C323651267430D2BF1C1F889FD30FF44" lastPageId="16" lastPageNumber="311" pageId="15" pageNumber="310" type="description">
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<paragraph id="8B86369967430D34F1C1F889FEC5F87A" blockId="15.[151,545,1840,1936]" box="[151,267,1911,1936]" pageId="15" pageNumber="310">
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(
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<figureCitation id="13022A1C67430D34F1C9F889FF29F865" box="[159,231,1911,1936]" captionStart="FIGURE 4" captionStartId="12.[151,250,1861,1883]" captionTargetBox="[292,1305,194,1845]" captionTargetId="figure@12.[292,1305,194,1845]" captionTargetPageId="12" captionText="FIGURE 4. A. Adult female paratype (LSUHC 13138) of Hemiphyllodactyus ywanganensis sp. nov. from 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer). B and C: Adult male holotype (USNM 570733) and adult female paratype (USNM 570734) of H. uga sp. nov., respectively, from the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar (Photos by G. R. Zug). D. Adult female (LSUDPC 10666) of H. montawaensis from the new locality of Ruby Cave, 5.1 km northeast of the type locality of Montawa Cave, 3.7 km southwest of Taunggyi, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer)." httpUri="https://zenodo.org/record/1437721/files/figure.png" pageId="15" pageNumber="310">Figs 4</figureCitation>
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,
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<figureCitation id="13022A1C67430D34F1A5F889FECFF87A" box="[243,257,1911,1936]" captionStart="FIGURE 5" captionStartId="13.[151,250,1663,1685]" captionTargetBox="[169,1422,176,1643]" captionTargetId="figure@13.[169,1422,175,1643]" captionTargetPageId="13" captionText="FIGURE 5. Upper row: Type series of Hemiphyllodactyus ywanganensis sp. nov. from 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar. Lower row: Type series of H. uga sp. nov. from the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar. Photos by L. L. Grismer." httpUri="https://zenodo.org/record/1437723/files/figure.png" pageId="15" pageNumber="310">5</figureCitation>
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)
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</paragraph>
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<paragraph id="8B86369967430D2BF1C1F841FD30FF44" blockId="15.[151,1436,1983,2036]" lastBlockId="16.[151,766,152,174]" lastPageId="16" lastPageNumber="311" pageId="15" pageNumber="310">
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<emphasis id="B94DEA8B67430D34F1C1F841FE95F83F" box="[151,347,1983,2005]" italics="true" pageId="15" pageNumber="310">Hemphyllodactylus</emphasis>
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sp.
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<date id="FF87105967430D34F0D5F83EFE08F83F" box="[387,454,1983,2005]" pageId="15" pageNumber="310">nov. 8.</date>
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<bibRefCitation id="EFA84B6867430D34F09AF841FD5CF83F" box="[460,658,1983,2005]" pageId="15" pageNumber="310" refId="ref9307" refString="Grismer, L. L., Wood Junior, P. L., Anuar, S., Muin, M. A., Quah, E. S. H., McGuire, J. A., Brown, R. A., Ngo, V. T. & Pham, H. T. (2013) Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society, 169, 849 - 880. https: // doi. org / 10.1111 / zoj. 12064" type="journal article">
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Grismer
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<emphasis id="B94DEA8B67430D34F373F83EFD99F83F" box="[549,599,1983,2005]" italics="true" pageId="15" pageNumber="310">et al.</emphasis>
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2013
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</bibRefCitation>
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:872,
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<bibRefCitation id="EFA84B6867430D34F39EF841FC53F83F" box="[712,925,1983,2005]" pageId="15" pageNumber="310" refId="ref9428" refString="Grismer, L. L., Riyanto, A., Djoko, I. T. & McGuire, J. A. (2014 a) A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa, 3821 (4), 485 - 495. https: // doi. org / 10.11646 / zootaxa. 3821.4.7" type="journal article">
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Grismer
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<emphasis id="B94DEA8B67430D34F277F83EFC9AF83F" box="[801,852,1983,2005]" italics="true" pageId="15" pageNumber="310">et al.</emphasis>
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2014a
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</bibRefCitation>
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:67,
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<bibRefCitation id="EFA84B6867430D34F292F841FB59F83F" box="[964,1175,1983,2005]" pageId="15" pageNumber="310" refId="ref9502" refString="Grismer, L. L., Wood Junior, P. L. & Cota, M. (2014 b) A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from northwestern Thailand. Zootaxa, 3760 (1), 67 - 68. https: // doi. org / 10.11646 / zootaxa. 3760.1.4" type="journal article">
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Grismer
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<emphasis id="B94DEA8B67430D34F54AF83EFB81F83F" box="[1052,1103,1983,2005]" italics="true" pageId="15" pageNumber="310">et al.</emphasis>
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2014b
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</bibRefCitation>
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:490,
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<bibRefCitation id="EFA84B6867430D34F59AF83EFAA0F83F" box="[1228,1390,1983,2005]" pageId="15" pageNumber="310" refId="ref10727" refString="Ngo, V. T., Grismer, L. L., Thai, P. H. & Wood Junior, P. L. (2014) A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na-Nui Nature reserve, central Vietnam, Zootaxa, 3760 (4), 539 - 552. https: // doi. org / 10.11646 / zootaxa. 3760.4.3" type="journal article">
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Ngo
|
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<emphasis id="B94DEA8B67430D34F5A8F83EFAFFF83F" box="[1278,1329,1983,2005]" italics="true" pageId="15" pageNumber="310">et al.</emphasis>
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2014
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</bibRefCitation>
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:541,
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||
<bibRefCitation id="EFA84B6867430D34F191F823FE41F819" box="[199,399,2013,2036]" pageId="15" pageNumber="310" refId="ref9889" refString="Grismer, L. L., Wood Junior, P. L., Anuar, S., Quah, E. S. H., Muin, M. A., Chan, K. O., Sumarli, A. X. & Loredo, A. I. (2015) Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society, 174, 859 - 876. https: // doi. org / 10.1111 / zoj. 12254" type="journal article">
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Grismer
|
||
<emphasis id="B94DEA8B67430D34F076F821FE9DF819" box="[288,339,2013,2036]" italics="true" pageId="15" pageNumber="310">et al.</emphasis>
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2015
|
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</bibRefCitation>
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:861
|
||
<emphasis id="B94DEA8B675C0D2BF1C1FF66FE98FF44" box="[151,342,152,174]" italics="true" pageId="16" pageNumber="311">Hemiphyllodatylus</emphasis>
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||
<taxonomicName id="4C394D1A675C0D2BF00AFF66FDCCFF44" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[348,514,152,174]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="311" phylum="Chordata" rank="species" species="linnwayensis">
|
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cf.
|
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<emphasis id="B94DEA8B675C0D2BF02AFF66FDCCFF44" box="[380,514,152,174]" italics="true" pageId="16" pageNumber="311">linnwayensis</emphasis>
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</taxonomicName>
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||
<emphasis id="B94DEA8B675C0D2BF354FF66FDC8FF44" box="[514,518,152,174]" italics="true" pageId="16" pageNumber="311">.</emphasis>
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<bibRefCitation id="EFA84B68675C0D2BF35AFF66FD2CFF44" box="[524,738,152,174]" pageId="16" pageNumber="311" refId="ref9686" refString="Grismer, L. L., Wood Junior, P. L., Thura, M. K., Zin, T., Quah, E. S. H., Murdoch, M. L., Grismer, M. S., Lin, A., Kyaw, H. & Ngwe, L. (2017 b) Phylogenetic taxonomy of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) with descriptions of three new species from Myanmar. Journal of Natural History, 52, 881 - 915. https: // doi. org / 10.1080 / 00222933.2017.1367045" type="journal article">
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Grismer
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<emphasis id="B94DEA8B675C0D2BF333FF67FD57FF44" box="[613,665,152,174]" italics="true" pageId="16" pageNumber="311">et al.</emphasis>
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2017b
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</bibRefCitation>
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:31
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</paragraph>
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<paragraph id="8B863699675C0D2BF1C1FF2AFD58FEDF" blockId="16.[151,1437,212,2037]" pageId="16" pageNumber="311">
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<materialsCitation id="3B513CC4675C0D2BF1C1FF2AFD58FEDF" ID-GBIF-Occurrence="1913860789" collectingDate="2003-08-22" collectionCode="USNM" collectorName="George R." country="Myanmar" elevation="1125" latitude="21.9931" location="Mandalay Region" longLatPrecision="7" longitude="96.4711" pageId="16" pageNumber="311" specimenCode="USNM 570733" specimenCount="1" stateProvince="Mandalay Region" typeStatus="holotype">
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<emphasis id="B94DEA8B675C0D2BF1C1FF2AFEC5FF07" bold="true" box="[151,267,212,237]" pageId="16" pageNumber="311">
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<typeStatus id="5482883B675C0D2BF1C1FF2AFEC8FF07" box="[151,262,212,237]" pageId="16" pageNumber="311" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male (
|
||
<specimenCode id="DB9F9EE2675C0D2BF0CBFF2AFD83FF07" box="[413,589,212,237]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="311">USNM 570733</specimenCode>
|
||
) collected on
|
||
<collectingDate id="EFC3E9B1675C0D2BF3BCFF2AFC53FF06" box="[746,925,212,236]" pageId="16" pageNumber="311" value="2003-08-22">
|
||
<date id="FF871059675C0D2BF3BCFF2AFC53FF06" box="[746,925,212,236]" pageId="16" pageNumber="311" value="2003-08-22">22 August 2003</date>
|
||
</collectingDate>
|
||
at 1955 hrs by
|
||
<collectorName id="26CC534F675C0D2BF51CFF2AFB73FF06" box="[1098,1213,212,236]" pageId="16" pageNumber="311">George R.</collectorName>
|
||
Zug, on handrail of elevated timber walkway in Kandawgyi National Gardens, Pyin Oo Lwin,
|
||
<collectingRegion id="49FDF87B675C0D2BF512FF09FAD4FEFA" box="[1092,1306,247,272]" country="Myanmar" name="Mandalay" pageId="16" pageNumber="311">Mandalay Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32E7609675C0D2BF479FF06FA52FEFA" box="[1327,1436,248,272]" name="Myanmar" pageId="16" pageNumber="311">Myanmar</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE0D505E675C0D2BF1C8FEE2FED5FEDE" box="[158,283,284,308]" direction="north" orientation="latitude" pageId="16" pageNumber="311" precision="5" value="21.9931">21.9931°N</geoCoordinate>
|
||
<geoCoordinate id="EE0D505E675C0D2BF074FEE2FE52FEDF" box="[290,412,284,309]" direction="east" orientation="longitude" pageId="16" pageNumber="311" precision="5" value="96.4711">96.4711°E</geoCoordinate>
|
||
;
|
||
<elevation id="0014D1AA675C0D2BF0F0FEE2FE35FEDE" box="[422,507,284,309]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="16" pageNumber="311" unit="m" value="1125.0">
|
||
<quantity id="4CC19B7C675C0D2BF0F0FEE2FE35FEDE" box="[422,507,284,309]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="16" pageNumber="311" unit="m" value="1125.0">1125 m</quantity>
|
||
</elevation>
|
||
in elevation).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B863699675C0D2BF191FEC1FAB2FE2F" blockId="16.[151,1437,212,2037]" pageId="16" pageNumber="311">
|
||
<emphasis id="B94DEA8B675C0D2BF191FEC1FE88FEB2" bold="true" box="[199,326,319,344]" pageId="16" pageNumber="311">
|
||
<typeStatus id="5482883B675C0D2BF191FEC1FE8FFEB2" box="[199,321,319,344]" pageId="16" pageNumber="311" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female (
|
||
<collectionCode id="ED28AE5C675C0D2BF0A4FEBEFD8BFEB2" box="[498,581,320,344]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="311">USNM</collectionCode>
|
||
570732) collected on
|
||
<date id="FF871059675C0D2BF210FEBEFC35FEB2" box="[838,1019,320,344]" pageId="16" pageNumber="311" value="2003-08-14">14 August 2003</date>
|
||
at ~1500 hrs by monastic brother in rockpile,
|
||
<collectingRegion id="49FDF87B675C0D2BF050FE9BFE92FE97" box="[262,348,357,381]" country="Myanmar" name="Yangon" pageId="16" pageNumber="311">Yangon</collectingRegion>
|
||
Monastery,
|
||
<quantity id="4CC19B7C675C0D2BF0A4FE9AFDF0FE97" box="[498,574,356,381]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="311" unit="km" value="11.0">11 km</quantity>
|
||
ENE of Pyin Oo Lwin,
|
||
<collectingRegion id="49FDF87B675C0D2BF239FE9AFB8EFE97" box="[879,1088,356,381]" country="Myanmar" name="Mandalay" pageId="16" pageNumber="311">Mandalay Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32E7609675C0D2BF506FE9BFB73FE97" box="[1104,1213,357,381]" name="Myanmar" pageId="16" pageNumber="311">Myanmar</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE0D505E675C0D2BF584FE9AFAA6FE96" box="[1234,1384,356,381]" direction="north" orientation="latitude" pageId="16" pageNumber="311" precision="1" value="22.0675">22°04.050 N</geoCoordinate>
|
||
<geoCoordinate id="EE0D505E675C0D2BF423FE9AFF39FE4A" direction="east" orientation="longitude" pageId="16" pageNumber="311" precision="1" value="96.563515">96° 33.811E</geoCoordinate>
|
||
, ~
|
||
<quantity id="4CC19B7C675C0D2BF059FE76FE99FE75" box="[271,343,392,416]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="311" unit="m" value="900.0">900 m</quantity>
|
||
elevation); adult females (
|
||
<collectionCode id="ED28AE5C675C0D2BF3D6FE76FD1DFE4A" box="[640,723,392,416]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="311">USNM</collectionCode>
|
||
570734, 570735) same locality as
|
||
<typeStatus id="5482883B675C0D2BF50EFE79FB73FE4A" box="[1112,1213,391,416]" pageId="16" pageNumber="311" type="holotype">holotype</typeStatus>
|
||
,
|
||
<date id="FF871059675C0D2BF591FE76FAB4FE4A" box="[1223,1402,392,416]" pageId="16" pageNumber="311" value="2003-08-23">23 August 2003</date>
|
||
by San Lwin Oo ~1900 hrs and
|
||
<date id="FF871059675C0D2BF08CFE52FD43FE2E" box="[474,653,428,452]" pageId="16" pageNumber="311" value="2003-08-20">20 August 2003</date>
|
||
by Jeremy F. Jacobs from jaws of a spider ~1970 hrs, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236512675C0D2BF191FE31FCFDFC9E" pageId="16" pageNumber="311" type="diagnosis">
|
||
<paragraph id="8B863699675C0D2BF191FE31FCFDFC9E" blockId="16.[151,1437,212,2037]" pageId="16" pageNumber="311">
|
||
<emphasis id="B94DEA8B675C0D2BF191FE31FE8FFE02" bold="true" box="[199,321,463,488]" pageId="16" pageNumber="311">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C394D1A675C0D2BF01EFE31FD97FE02" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[328,601,463,488]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="311" phylum="Chordata" rank="species" species="uga" status="sp. nov.">
|
||
<emphasis id="B94DEA8B675C0D2BF01EFE31FD97FE02" box="[328,601,463,488]" italics="true" pageId="16" pageNumber="311">Hemiphyllodactylus uga</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E57F0675C0D2BF336FE31FD74FE02" box="[608,698,463,488]" pageId="16" pageNumber="311" rank="species">
|
||
<emphasis id="B94DEA8B675C0D2BF336FE31FD74FE02" bold="true" box="[608,698,463,488]" pageId="16" pageNumber="311">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be separated from all other species of
|
||
<taxonomicName id="4C394D1A675C0D2BF5CFFE31FAB7FE02" box="[1177,1401,463,488]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="311" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DEA8B675C0D2BF5CFFE31FAB7FE02" box="[1177,1401,463,488]" italics="true" pageId="16" pageNumber="311">Hemiphyllodactylus</emphasis>
|
||
</taxonomicName>
|
||
by possessing the unique combination of having a maximum SVL of 39.0 mm; 8–11 chin scales; enlarged postmentals; five circumnasal scales; two or three intersupranasals (=postrostrals); nine or 10 supralabials; 8–10 infralabials; 13–15 longitudinally arranged dorsal scales at midbody contained within one eye diameter and 6–8 ventral scales; varied digital formulae on hands and feet (
|
||
<tableCitation id="C6BB0322675C0D2BF264FDA1FC42FD92" box="[818,908,607,632]" captionStart="TABLE 5" captionStartId="9.[151,235,437,459]" captionText="TABLE 5. Descriptive and diagnostic characters among species of the eastern Myanmar clade." pageId="16" pageNumber="311">Table 5</tableCitation>
|
||
); two or three subdigital lamellae on the first finger and toe; 18–22 continuous, pore-bearing, femoroprecloacal scales in males; no plate-like subcaudal scales; adult females not yellow; a dark postorbital stripe extending to base of neck variably present; variable pairs of paravertebral light spots on trunk; no dark, dorsolateral stripe; no dark paravertebral markings on trunk; dorsal body pattern usually with dark zig-zag markings; postsacral marking not bearing light-colored anteriorly projecting arms; and ceacum and gonads unpigmented. These characters are scored across all species of the eastern
|
||
<collectingCountry id="F32E7609675C0D2BF479FCEBFA52FCC7" box="[1327,1436,789,813]" name="Myanmar" pageId="16" pageNumber="311">Myanmar</collectingCountry>
|
||
clade in
|
||
<tableCitation id="C6BB0322675C0D2BF056FCC9FEADFCBA" box="[256,355,823,848]" captionStart="TABLE 5" captionStartId="9.[151,235,437,459]" captionText="TABLE 5. Descriptive and diagnostic characters among species of the eastern Myanmar clade." pageId="16" pageNumber="311">Tables 5</tableCitation>
|
||
and
|
||
<tableCitation id="C6BB0322675C0D2BF0F3FCC9FE7DFCBA" box="[421,435,823,848]" captionStart="TABLE 6" captionStartId="10.[151,239,152,175]" captionText="TABLE 6. Data for the type series of Hemiphyllodactylus ywanganenis sp. nov. and H. uga sp. nov. / = data unavailable." pageId="16" pageNumber="311">6</tableCitation>
|
||
and from all other species of
|
||
<taxonomicName id="4C394D1A675C0D2BF273FCC9FBCBFCBA" box="[805,1029,823,848]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="311" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DEA8B675C0D2BF273FCC9FBCBFCBA" box="[805,1029,823,848]" italics="true" pageId="16" pageNumber="311">Hemiphyllodactylus</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F32E7609675C0D2BF597FCC9FACAFCBA" box="[1217,1284,823,848]" name="China" pageId="16" pageNumber="311">China</collectingCountry>
|
||
and western
|
||
<collectingCountry id="F32E7609675C0D2BF1C1FCA2FF35FC9F" box="[151,251,860,885]" name="Thailand" pageId="16" pageNumber="311">Thailand</collectingCountry>
|
||
(clades 3 and
|
||
<quantity id="4CC19B7C675C0D2BF0CAFCA2FE09FC9F" box="[412,455,860,885]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="16" pageNumber="311" unit="in" value="4.0">4 in</quantity>
|
||
<bibRefCitation id="EFA84B68675C0D2BF098FCA2FD70FC9F" box="[462,702,860,885]" pageId="16" pageNumber="311" refId="ref9686" refString="Grismer, L. L., Wood Junior, P. L., Thura, M. K., Zin, T., Quah, E. S. H., Murdoch, M. L., Grismer, M. S., Lin, A., Kyaw, H. & Ngwe, L. (2017 b) Phylogenetic taxonomy of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) with descriptions of three new species from Myanmar. Journal of Natural History, 52, 881 - 915. https: // doi. org / 10.1080 / 00222933.2017.1367045" type="journal article">
|
||
Grismer
|
||
<emphasis id="B94DEA8B675C0D2BF367FCA3FDA7FC9F" box="[561,617,860,885]" italics="true" pageId="16" pageNumber="311">et al.</emphasis>
|
||
[2017b
|
||
</bibRefCitation>
|
||
:
|
||
<tableCitation id="C6BB0322675C0D2BF397FCA2FCD5FC9E" box="[705,795,860,885]" captionStart="TABLE 3" captionStartId="6.[147,229,508,529]" captionText="TABLE 3. Percent uncorrected pairwise sequence divergence among the individuals of the eastern Myanmar clade." pageId="16" pageNumber="311">Table 3</tableCitation>
|
||
]).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236512675C0D2AF191FC81FF3EFD42" lastPageId="17" lastPageNumber="312" pageId="16" pageNumber="311" type="description">
|
||
<paragraph id="8B863699675C0D2BF191FC81FE41FAA2" blockId="16.[151,1437,212,2037]" pageId="16" pageNumber="311">
|
||
<emphasis id="B94DEA8B675C0D2BF191FC81FE2AFC72" bold="true" box="[199,484,895,920]" pageId="16" pageNumber="311">
|
||
Description of
|
||
<typeStatus id="5482883B675C0D2BF023FC81FE10FC72" box="[373,478,895,920]" pageId="16" pageNumber="311" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male; head triangular in dorsal profile, depressed, distinct from neck; lores and interorbital regions flat; rostrum moderate in length (NarEye/HeadL 0.33); prefrontal region weakly concave; canthus rostralis smoothly rounded, barely discernable; snout moderate, rounded in dorsal profile; eye large; ear opening elliptical, small; eye to ear distance greater than diameter of eye; rostral wider than high, bordered posteriorly by small supranasals; two intersupranasals (=postnasals); external nares bordered anteriorly by rostral, dorsally by supranasal, posteriorly by two postnasals, ventrally by first supralabial (=circumnasals); nine (R,L) rectangular supralabials tapering to below posterior margin of orbit; eight (R,L) rectangular infralabials tapering to below posterior margin of orbit; scales of rostrum, lores, top of head, and occiput small, granular, those of rostrum largest and slightly raised; dorsal superciliaries flat, mostly square, subimbricate, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by two large postmentals; each postmental bordered laterally by a single enlarged, sublabial; eight chin scales; gular scales small, subimbricate, grading posteriorly into slightly larger, subimbricate, throat and even larger pectoral scales which grade into slightly larger, subimbricate ventrals.
|
||
</paragraph>
|
||
<paragraph id="8B863699675C0D2BF191FAAAFCD6F81F" blockId="16.[151,1437,212,2037]" pageId="16" pageNumber="311">
|
||
Body somewhat elongate (Trunk/SVL 0.46), dorsoventrally compressed; ventrolateral folds absent; dorsal scales small, granular, 15 dorsal scales at midbody contained within one eye diameter; ventral scales, flat, subimbricate much larger than dorsal scales, seven ventral scales contained within one eye diameter; precloacal scales slightly larger than abdominal scales; pore-bearing precloacal scales continuous with pore-bearing femoral scales, totaling 18 femoroprecloacal pore-bearing scales; single cloacal spur on anterior margin of hemipenial swelling; forelimbs short, robust in stature, covered with flat, subimbricate scales dorsally and ventrally; palmar scales flat, subimbricate; all digits except digit I well-developed; digit I vestigial, clawless; distal subdigital lamellae of digits II–V undivided, angular and U-shaped, lamellae proximal to these transversely expanded; lamellar formula of digits
|
||
<date id="FF871059675C0D2BF0E9F98BFDEFF966" box="[447,545,1652,1676]" pageId="16" pageNumber="311">II–V 3-3</date>
|
||
-3-3 (R,L); two transversely expanded lamellae on digit I; claws on digits II–V well developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; hind limbs short, more robust than forelimbs, covered with flat, juxtaposed scales dorsally and by larger, flat subimbricate scales ventrally; plantar scales low, flat, subimbricate; all digits except digit I welldeveloped; digit I vestigial, clawless; distal subdigital lamellae of digits II–V undivided, angular and U-shaped, lamellae proximal to these transversely expanded; lamellar formula of digits
|
||
<date id="FF871059675C0D2BF2ABF8D7FB91F8AA" box="[1021,1119,1832,1856]" pageId="16" pageNumber="311">II–V 3-3</date>
|
||
-3-3 (R,L); two transversely expanded lamellae on digit I; claws on digits II–V well developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; dorsal caudal scales of original tail subimbricate, larger than dorsal body scales, grading laterally into larger, slightly flared scales bearing elevated posterior margins; subcaudal scales larger than dorsal caudals, flat, imbricate, not plate-like; posterior
|
||
<quantity id="4CC19B7C675C0D2BF444F846FA9DF83A" box="[1298,1363,1976,2000]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="311" unit="mm" value="3.0">3 mm</quantity>
|
||
of tail regenerated. Morphometric data are presented in
|
||
<tableCitation id="C6BB0322675C0D2BF3E8F822FCDAF81F" box="[702,788,2012,2037]" captionStart="TABLE 6" captionStartId="10.[151,239,152,175]" captionText="TABLE 6. Data for the type series of Hemiphyllodactylus ywanganenis sp. nov. and H. uga sp. nov. / = data unavailable." pageId="16" pageNumber="311">Table 6</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B863699675D0D2AF191FF69FE3FFE3A" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FF69FDCFFF5A" bold="true" box="[199,513,151,176]" pageId="17" pageNumber="312">
|
||
Coloration in life (
|
||
<figureCitation id="13022A1C675D0D2AF0FCFF69FE3DFF5A" box="[426,499,151,176]" captionStart="FIGURE 4" captionStartId="12.[151,250,1861,1883]" captionTargetBox="[292,1305,194,1845]" captionTargetId="figure@12.[292,1305,194,1845]" captionTargetPageId="12" captionText="FIGURE 4. A. Adult female paratype (LSUHC 13138) of Hemiphyllodactyus ywanganensis sp. nov. from 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer). B and C: Adult male holotype (USNM 570733) and adult female paratype (USNM 570734) of H. uga sp. nov., respectively, from the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar (Photos by G. R. Zug). D. Adult female (LSUDPC 10666) of H. montawaensis from the new locality of Ruby Cave, 5.1 km northeast of the type locality of Montawa Cave, 3.7 km southwest of Taunggyi, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer)." httpUri="https://zenodo.org/record/1437721/files/figure.png" pageId="17" pageNumber="312">Fig. 4</figureCitation>
|
||
).
|
||
</emphasis>
|
||
All
|
||
<taxonomicName id="4C394D1A675D0D2AF36EFF69FCD6FF5A" box="[568,792,151,176]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94DEA8B675D0D2AF36EFF69FCD6FF5A" box="[568,792,151,176]" italics="true" pageId="17" pageNumber="312">Hemiphyllodactylus</emphasis>
|
||
</taxonomicName>
|
||
are capable of considerable change in the intensity and boldness of their coloration and pattern. The description below is of that at the time the specimens were photographed. Ground color of top of head, body, limbs, and tail beige; top of head bearing small, diffuse, dark markings; thin, dark, preorbital stripe extends through eye to shoulder region; pairs of irregularly shaped, dark paravertebral markings on dorsum extend from nape to postsacral region; smaller, indistinct dark markings on flanks; limbs bearing irregularly shaped, dark markings; postsacral marking indistinct, not bearing anteriorly projecting arms; dark markings on dorsal surface of tail evenly spaced; gular region generally immaculate, except for darker lateral areas and faint stippling in scales; and pigmentation density increases posteriorly with the abdomen being generally gray.
|
||
</paragraph>
|
||
<paragraph id="8B863699675D0D2AF191FE22FF3EFD42" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FE22FE09FE1F" bold="true" box="[199,455,476,501]" pageId="17" pageNumber="312">
|
||
Variation (
|
||
<figureCitation id="13022A1C675D0D2AF01EFE22FE54FE1F" box="[328,410,476,501]" captionStart="FIGURE 4" captionStartId="12.[151,250,1861,1883]" captionTargetBox="[292,1305,194,1845]" captionTargetId="figure@12.[292,1305,194,1845]" captionTargetPageId="12" captionText="FIGURE 4. A. Adult female paratype (LSUHC 13138) of Hemiphyllodactyus ywanganensis sp. nov. from 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer). B and C: Adult male holotype (USNM 570733) and adult female paratype (USNM 570734) of H. uga sp. nov., respectively, from the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar (Photos by G. R. Zug). D. Adult female (LSUDPC 10666) of H. montawaensis from the new locality of Ruby Cave, 5.1 km northeast of the type locality of Montawa Cave, 3.7 km southwest of Taunggyi, Taunggyi District, Shan State, Myanmar (Photo by L. L. Grismer)." httpUri="https://zenodo.org/record/1437721/files/figure.png" pageId="17" pageNumber="312">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="13022A1C675D0D2AF0FFFE22FE77FE1F" box="[425,441,476,501]" captionStart="FIGURE 5" captionStartId="13.[151,250,1663,1685]" captionTargetBox="[169,1422,176,1643]" captionTargetId="figure@13.[169,1422,175,1643]" captionTargetPageId="13" captionText="FIGURE 5. Upper row: Type series of Hemiphyllodactyus ywanganensis sp. nov. from 2.7 km southwest of Ywangan, Ywangan Township, Taunggyi District, Shan State, Myanmar. Lower row: Type series of H. uga sp. nov. from the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin, Mandalay Region, Myanmar. Photos by L. L. Grismer." httpUri="https://zenodo.org/record/1437723/files/figure.png" pageId="17" pageNumber="312">5</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The color patterns of the
|
||
<typeStatus id="5482883B675D0D2AF3A5FE23FC90FE1F" box="[755,862,477,501]" pageId="17" pageNumber="312" type="paratype">paratypes</typeStatus>
|
||
(USNM 570732, 570735) generally match that of the
|
||
<typeStatus id="5482883B675D0D2AF196FE01FEE8FDF2" box="[192,294,511,536]" pageId="17" pageNumber="312" type="holotype">holotype</typeStatus>
|
||
. USNM 570732 has a slightly bolder zig-zag dorsal pattern and a dark ventrolatral stripe on the flanks and a regenerated, nearly unicolor tail. The color pattern of USNM 570735 is more faint than that of the holoype and the dark dorsal markings both on the dorsum and the tail ae much thinner. USNM 570734 has a much darker, nearly unicolor dorsal pattern and greatly swollen endolymphatic sacs. Differences in scales counts are presented in
|
||
<tableCitation id="C6BB0322675D0D2AF1C1FD71FF23FD42" box="[151,237,655,680]" captionStart="TABLE 6" captionStartId="10.[151,239,152,175]" captionText="TABLE 6. Data for the type series of Hemiphyllodactylus ywanganenis sp. nov. and H. uga sp. nov. / = data unavailable." pageId="17" pageNumber="312">Table 6</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236512675D0D2AF191FD4AFCADFCB7" pageId="17" pageNumber="312" type="distribution">
|
||
<paragraph id="8B863699675D0D2AF191FD4AFB9DFD1A" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FD4AFEAEFD27" bold="true" box="[199,352,692,717]" pageId="17" pageNumber="312">Distribution.</emphasis>
|
||
<taxonomicName id="4C394D1A675D0D2AF03CFD4AFDB1FD26" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[362,639,692,717]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="uga" status="sp. nov.">
|
||
<emphasis id="B94DEA8B675D0D2AF03CFD4AFDB1FD26" box="[362,639,692,717]" italics="true" pageId="17" pageNumber="312">Hemiphyllodactylus uga</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E57F0675D0D2AF3DFFD4BFD2DFD26" box="[649,739,693,716]" pageId="17" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="5482883B675D0D2AF55FFD4BFBF7FD27" box="[1033,1081,693,717]" pageId="17" pageNumber="312">type</typeStatus>
|
||
locality of the Pyin Oo Lwin, Kandawgyi National Gardens, Pyin Oo Lwin,
|
||
<collectingRegion id="49FDF87B675D0D2AF3C8FD29FCC0FD1A" box="[670,782,727,752]" country="Myanmar" name="Mandalay" pageId="17" pageNumber="312">Mandalay</collectingRegion>
|
||
Division,
|
||
<collectingCountry id="F32E7609675D0D2AF2D2FD26FC3FFD1A" box="[900,1009,728,752]" name="Myanmar" pageId="17" pageNumber="312">Myanmar</collectingCountry>
|
||
(
|
||
<figureCitation id="13022A1C675D0D2AF556FD29FB8FFD1A" box="[1024,1089,727,752]" captionStart="FIGURE 1" captionStartId="1.[151,250,1550,1572]" captionTargetBox="[150,1436,555,1526]" captionTargetId="figure@1.[150,1436,555,1526]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the eastern Myanmar clade of Hemiphyllodactylus in Shan State (H. linnwayensis, H. montawaensis, H. tonywitteni, and H. ywnaganensis sp. nov.) and Mandalay Region, Myanmar (H. uga sp. nov.). The green square in the inset delimits the study area." httpUri="https://zenodo.org/record/1437715/files/figure.png" pageId="17" pageNumber="312">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B863699675D0D2AF191FD02FCADFCB7" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FD02FE45FCFF" bold="true" box="[199,395,764,789]" pageId="17" pageNumber="312">Natural History.</emphasis>
|
||
Three individuals were found on the handrails of a wooden canopy walkway built within an evergreen forest of the National Garden. The walkway and forest were visited on five nights but
|
||
<taxonomicName id="4C394D1A675D0D2AF1C1FCBAFE66FCB6" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[151,424,836,861]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="uga">
|
||
<emphasis id="B94DEA8B675D0D2AF1C1FCBAFE66FCB6" box="[151,424,836,861]" italics="true" pageId="17" pageNumber="312">Hemiphyllodactylus uga</emphasis>
|
||
</taxonomicName>
|
||
was found only on two of those nights.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236512675D0D2AF191FC99FBF9FABF" pageId="17" pageNumber="312" type="etymology">
|
||
<paragraph id="8B863699675D0D2AF191FC99FCC6FBDE" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FC99FE80FC6A" bold="true" box="[199,334,871,896]" pageId="17" pageNumber="312">Etymology.</emphasis>
|
||
The specific name recognizes and honors the late U Uga. He was a conservationist and a former director of the Nature and Wildlife Conservation Division (NWCD),
|
||
<collectingCountry id="F32E7609675D0D2AF2A2FC73FBAFFC4F" box="[1012,1121,909,933]" name="Myanmar" pageId="17" pageNumber="312">Myanmar</collectingCountry>
|
||
Forestry Department. He encouraged Joseph B. Slowinski and George R. Zug to do an all-country herpetofaunal survey and established the administrative protocol to establish and support survey teams of NWCD wildlife rangers. These teams working independently and with CAS and USNM collaborators were the essential factor for the high productivity and success of the
|
||
<collectingCountry id="F32E7609675D0D2AF06FFBE3FE68FBDF" box="[313,422,1053,1077]" name="Myanmar" pageId="17" pageNumber="312">Myanmar</collectingCountry>
|
||
Herpetological Survey (MHS).
|
||
</paragraph>
|
||
<paragraph id="8B863699675D0D2AF191FBC1FBF9FABF" blockId="17.[151,1437,151,1365]" pageId="17" pageNumber="312">
|
||
<emphasis id="B94DEA8B675D0D2AF191FBC1FEA2FBB2" bold="true" box="[199,364,1087,1112]" pageId="17" pageNumber="312">Comparisons.</emphasis>
|
||
The molecular analyses indicate that
|
||
<taxonomicName id="4C394D1A675D0D2AF278FBC1FB8DFBB2" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[814,1091,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="uga" status="sp. nov.">
|
||
<emphasis id="B94DEA8B675D0D2AF278FBC1FB8DFBB2" box="[814,1091,1087,1112]" italics="true" pageId="17" pageNumber="312">Hemiphyllodactylus uga</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E57F0675D0D2AF518FBC1FB63FBB2" box="[1102,1197,1087,1112]" pageId="17" pageNumber="312" rank="species">
|
||
<emphasis id="B94DEA8B675D0D2AF518FBC1FB63FBB2" bold="true" box="[1102,1197,1087,1112]" pageId="17" pageNumber="312">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is embedded within eastern
|
||
<collectingCountry id="F32E7609675D0D2AF1B8FB9BFE95FB97" box="[238,347,1125,1149]" name="Myanmar" pageId="17" pageNumber="312">Myanmar</collectingCountry>
|
||
clade and forms a monophyletic group with
|
||
<taxonomicName id="4C394D1A675D0D2AF20DFB9BFBC1FB97" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[859,1039,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="linnwayensis">
|
||
<emphasis id="B94DEA8B675D0D2AF20DFB9BFBC1FB97" box="[859,1039,1124,1149]" italics="true" pageId="17" pageNumber="312">H. linnwayensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D1A675D0D2AF51EFB9BFAC6FB96" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[1096,1288,1125,1148]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="ywnaganensis" status="sp. nov.">
|
||
<emphasis id="B94DEA8B675D0D2AF51EFB9BFAC6FB96" box="[1096,1288,1125,1148]" italics="true" pageId="17" pageNumber="312">H. ywnaganensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E57F0675D0D2AF447FB9AFAA5FB97" box="[1297,1387,1124,1149]" pageId="17" pageNumber="312" rank="species">
|
||
<emphasis id="B94DEA8B675D0D2AF447FB9AFAA5FB97" bold="true" box="[1297,1387,1124,1149]" pageId="17" pageNumber="312">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
See Comparisons section above for comparisons with
|
||
<emphasis id="B94DEA8B675D0D2AF384FB77FB64FB4A" box="[722,1194,1159,1184]" italics="true" pageId="17" pageNumber="312">
|
||
<taxonomicName id="4C394D1A675D0D2AF384FB77FC59FB4A" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[722,919,1161,1184]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="ywanganensis">H. ywanganensis</taxonomicName>
|
||
. Hemiphyllodactlus
|
||
<taxonomicName id="4C394D1A675D0D2AF5D6FB77FB64FB4A" authority="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin, 2018" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[1152,1194,1161,1184]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="uga" status="sp. nov.">uga</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A27E57F0675D0D2AF5E4FB79FAC0FB4A" box="[1202,1294,1159,1184]" pageId="17" pageNumber="312" rank="species">
|
||
<emphasis id="B94DEA8B675D0D2AF5E4FB79FAC0FB4A" bold="true" box="[1202,1294,1159,1184]" pageId="17" pageNumber="312">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is separated from
|
||
<emphasis id="B94DEA8B675D0D2AF183FB53FD94FB2F" box="[213,602,1196,1221]" italics="true" pageId="17" pageNumber="312">
|
||
<taxonomicName id="4C394D1A675D0D2AF183FB53FE54FB2E" box="[213,410,1197,1220]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="montawaensis">H. montawaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D1A675D0D2AF0F3FB53FD98FB2F" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[421,598,1196,1221]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="tonywhitteni">H. tonywhitteni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D1A675D0D2AF3C4FB53FC88FB2F" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[658,838,1196,1221]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="linnwayensis">
|
||
<emphasis id="B94DEA8B675D0D2AF3C4FB53FC88FB2F" box="[658,838,1196,1221]" italics="true" pageId="17" pageNumber="312">H. linnwayensis</emphasis>
|
||
</taxonomicName>
|
||
in have a higher mean number of chin scales (9.3 vs 6.3,
|
||
<emphasis id="B94DEA8B675D0D2AF19BFB2FFF15FB02" box="[205,219,1233,1256]" italics="true" pageId="17" pageNumber="312">p</emphasis>
|
||
= 0.03; 9.3 vs 6.6,
|
||
<emphasis id="B94DEA8B675D0D2AF082FB2FFE2CFB02" box="[468,482,1233,1256]" italics="true" pageId="17" pageNumber="312">p</emphasis>
|
||
= 0.05; 9.3 vs 5.0,
|
||
<emphasis id="B94DEA8B675D0D2AF38CFB2FFD26FB02" box="[730,744,1233,1256]" italics="true" pageId="17" pageNumber="312">p</emphasis>
|
||
= 0.003, respectively;
|
||
<tableCitation id="C6BB0322675D0D2AF2A8FB31FB96FB02" box="[1022,1112,1231,1256]" captionStart="TABLE 7" captionStartId="14.[151,235,153,175]" captionText="TABLE 7. Summary statistics of meristic characters among the species of the eastern Myanmar clade." pageId="17" pageNumber="312">Table 7</tableCitation>
|
||
). It differs further from
|
||
<emphasis id="B94DEA8B675D0D2AF4D7FB2FFE23FAE7" italics="true" pageId="17" pageNumber="312">
|
||
<taxonomicName id="4C394D1A675D0D2AF4D7FB2FFEE5FAE7" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="linnwayensis">H. linnwayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D1A675D0D2AF06EFB0BFE27FAE7" authorityName="Grismer & Wood & Zug & Thura & Grismer & Murdoch & Quah & Lin" authorityYear="2018" box="[312,489,1268,1293]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="tonywhitteni">H. tonywhitteni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D1A675D0D2AF37EFB0BFD25FAE6" box="[552,747,1269,1292]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="montawaensis">
|
||
<emphasis id="B94DEA8B675D0D2AF37EFB0BFD25FAE6" box="[552,747,1269,1292]" italics="true" pageId="17" pageNumber="312">H. montawaensis</emphasis>
|
||
</taxonomicName>
|
||
in having a lower mean number of lamellae on the first toe (2.3 vs 4.5,
|
||
<emphasis id="B94DEA8B675D0D2AF048FAE7FEE2FADA" box="[286,300,1305,1328]" italics="true" pageId="17" pageNumber="312">p</emphasis>
|
||
= 0.03; 2.3 vs 3.3
|
||
<emphasis id="B94DEA8B675D0D2AF352FAE7FDDCFADA" box="[516,530,1305,1328]" italics="true" pageId="17" pageNumber="312">p</emphasis>
|
||
= 0.002, respectively;
|
||
<tableCitation id="C6BB0322675D0D2AF241FAE9FCA3FADA" box="[791,877,1303,1328]" captionStart="TABLE 7" captionStartId="14.[151,235,153,175]" captionText="TABLE 7. Summary statistics of meristic characters among the species of the eastern Myanmar clade." pageId="17" pageNumber="312">Table 7</tableCitation>
|
||
). It differs even further from
|
||
<taxonomicName id="4C394D1A675D0D2AF5EAFAE7FAB0FADA" box="[1212,1406,1305,1328]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="312" phylum="Chordata" rank="species" species="montawaensis">
|
||
<emphasis id="B94DEA8B675D0D2AF5EAFAE7FAB0FADA" box="[1212,1406,1305,1328]" italics="true" pageId="17" pageNumber="312">H. montawaensis</emphasis>
|
||
</taxonomicName>
|
||
in that adult females are not yellow and it lacks a reticulated dorsal pattern (
|
||
<tableCitation id="C6BB0322675D0D2AF29DFAC2FBEFFABF" box="[971,1057,1340,1365]" captionStart="TABLE 5" captionStartId="9.[151,235,437,459]" captionText="TABLE 5. Descriptive and diagnostic characters among species of the eastern Myanmar clade." pageId="17" pageNumber="312">Table 5</tableCitation>
|
||
)..
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |