223 lines
34 KiB
XML
223 lines
34 KiB
XML
<document id="08A71DAE4F2B91EB90FC20E76D5D06CE" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2023" docId="03877623626DFFCF2DEFB08BFB985B45" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Crossoglossa) sciakyi Shi & Liang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="44" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="A5E032222F43C57761B7E0ABF7A297BA">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="DBCB282532638704AF0039CF94AABA40">Shi, Hongliang</mods:namePart>
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<mods:namePart id="51A651C22FC43D609322E38699A97E2F">Liang, Hongbin</mods:namePart>
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<mods:title id="AA10A3B7211B55C7F31646468431D4D1">Zootaxa</mods:title>
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<mods:date id="2CED8D4370E0337603F6EA9C56472E87">2023</mods:date>
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<treatment id="03877623626DFFCF2DEFB08BFB985B45" ID-DOI="http://doi.org/10.5281/zenodo.7963733" ID-GBIF-Taxon="209628960" ID-Zenodo-Dep="7963733" LSID="urn:lsid:plazi:treatment:03877623626DFFCF2DEFB08BFB985B45" httpUri="http://treatment.plazi.org/id/03877623626DFFCF2DEFB08BFB985B45" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
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<subSubSection id="C33494BE626DFFC82DEFB08BFD765E50" box="[151,657,751,778]" pageId="43" pageNumber="44" type="nomenclature">
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<paragraph id="8B91C735626DFFC82DEFB08BFD765E50" blockId="43.[151,751,751,815]" box="[151,657,751,778]" pageId="43" pageNumber="44">
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<emphasis id="B95A1B27626DFFC82DEFB08BFD765E50" bold="true" box="[151,657,751,778]" pageId="43" pageNumber="44">
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[9]
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<taxonomicName id="4C2EBCB6626DFFC82DB9B08BFDCB5E50" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[193,556,751,778]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov." subGenus="Crossoglossa">
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<emphasis id="B95A1B27626DFFC82DB9B08BFEFE5E50" bold="true" box="[193,281,752,778]" italics="true" pageId="43" pageNumber="44">Parena</emphasis>
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(
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<emphasis id="B95A1B27626DFFC82C52B094FE2F5E50" bold="true" box="[298,456,751,778]" italics="true" pageId="43" pageNumber="44">Crossoglossa</emphasis>
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)
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<emphasis id="B95A1B27626DFFC82CA1B094FDCB5E50" bold="true" box="[473,556,751,778]" italics="true" pageId="43" pageNumber="44">sciakyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A269A65C626DFFC82F4BB08BFD765E50" box="[563,657,752,778]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33494BE626DFFC82DEFB16FFD085E74" box="[151,751,788,815]" pageId="43" pageNumber="44" type="description">
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<paragraph id="8B91C735626DFFC82DEFB16FFD085E74" blockId="43.[151,751,751,815]" box="[151,751,788,815]" pageId="43" pageNumber="44">
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Habitus:
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<figureCitation id="1315DBB0626DFFC82D85B16FFEA85E75" box="[253,335,788,815]" captionStart="FIGURE 25" captionStartId="44.[151,250,1404,1428]" captionTargetBox="[157,1431,533,1372]" captionTargetId="figure-202@44.[151,1436,525,1381]" captionTargetPageId="44" captionText="FIGURE 25. Habitus of Parena (Crossoglossa) sciakyi sp. n., Holotype. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958989" httpUri="https://zenodo.org/record/7958989/files/figure.png" pageId="43" pageNumber="44">Fig. 25</figureCitation>
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. Gonocoxites of ovipositor:
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<figureCitation id="1315DBB0626DFFC82FF6B16FFD3B5E74" box="[654,732,788,814]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="43" pageNumber="44">Fig. 11</figureCitation>
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γ.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33494BE626DFFC82DEFB127FEB5595C" pageId="43" pageNumber="44" type="materials_examined">
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<paragraph id="8B91C735626DFFC82DEFB127FD3D5E2C" blockId="43.[151,1437,860,2039]" box="[151,730,860,886]" pageId="43" pageNumber="44">
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<materialsCitation id="3B46CD68626DFFC82DEFB127FD315E2C" ID-GBIF-Occurrence="4115871387" box="[151,726,860,886]" country="Solomon Islands" location="Guadalcanal" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Guadalcanal" typeStatus="holotype">
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<emphasis id="B95A1B27626DFFC82DEFB127FED15E2C" bold="true" box="[151,310,860,886]" pageId="43" pageNumber="44">
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<typeStatus id="54957997626DFFC82DEFB127FF355E2C" box="[151,210,860,886]" pageId="43" pageNumber="44">Type</typeStatus>
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locality.
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</emphasis>
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<collectingRegion id="49EA09D7626DFFC82C44B127FE2F5E2C" box="[316,456,860,886]" country="Solomon Islands" name="Guadalcanal" pageId="43" pageNumber="44">Guadalcanal</collectingRegion>
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of the
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<collectingCountry id="F33987A5626DFFC82F6FB127FD315E2C" box="[535,726,860,886]" name="Solomon Islands" pageId="43" pageNumber="44">Solomon Islands</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B91C735626DFFC82DBFB1FBFEB5595C" blockId="43.[151,1437,860,2039]" pageId="43" pageNumber="44">
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<emphasis id="B95A1B27626DFFC82DBFB1FBFE965EC0" bold="true" box="[199,369,896,922]" pageId="43" pageNumber="44">Type material</emphasis>
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.
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<emphasis id="B95A1B27626DFFC82CFAB1FBFE085EC0" bold="true" box="[386,495,896,922]" pageId="43" pageNumber="44">
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<typeStatus id="54957997626DFFC82CFAB1FBFE085EC0" box="[386,495,896,922]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
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</emphasis>
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(CRS,
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<figureCitation id="1315DBB0626DFFC82F30B1FBFD785EC1" box="[584,671,896,923]" captionStart="FIGURE 25" captionStartId="44.[151,250,1404,1428]" captionTargetBox="[157,1431,533,1372]" captionTargetId="figure-202@44.[151,1436,525,1381]" captionTargetPageId="44" captionText="FIGURE 25. Habitus of Parena (Crossoglossa) sciakyi sp. n., Holotype. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958989" httpUri="https://zenodo.org/record/7958989/files/figure.png" pageId="43" pageNumber="44">Fig. 25</figureCitation>
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): female, body length =
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<quantity id="4CD66AD0626DFFC82EC7B1FBFBC05EC1" box="[959,1063,896,923]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="43" pageNumber="44" unit="mm" value="11.3">11.3 mm</quantity>
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, board mounted, "South Pasific, Solomon Is. /
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<collectingRegion id="49EA09D7626DFFC82C41B1DFFDF75EE4" box="[313,528,932,958]" country="Solomon Islands" name="Guadalcanal" pageId="43" pageNumber="44">GUADALCANAL</collectingRegion>
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I.,
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<quantity id="4CD66AD0626DFFC82F40B1DFFD5F5EE5" box="[568,696,932,959]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="43" pageNumber="44" unit="m" value="825.0" valueMax="900.0" valueMin="750.0">750–900 m</quantity>
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/ Karukiki env.,
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<quantity id="4CD66AD0626DFFC82E03B1DFFC0E5EE4" box="[891,1001,932,959]" metricMagnitude="4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="43" pageNumber="44" unit="km" value="22.5" valueMax="25.0" valueMin="20.0">20-25 km</quantity>
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SSE / of
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<collectingRegion id="49EA09D7626DFFC82923B1DFFB5F5EE4" box="[1115,1208,932,958]" country="Solomon Islands" name="Honiara" pageId="43" pageNumber="44">Honiara</collectingRegion>
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,
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<date id="FF90E1F5626DFFC829BDB1DFFA825EE4" box="[1221,1381,932,958]" pageId="43" pageNumber="44" value="2016-12-01" valueMax="2016-12-18" valueMin="2016-12-01">1-18.XII.2016</date>
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/ St. Jakl leg."; "
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<typeStatus id="54957997626DFFC82C6FB1B3FE485EB8" box="[279,431,968,994]" pageId="43" pageNumber="44" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="4C2EBCB6626DFFC82CBDB1B2FD275EB8" box="[453,704,968,995]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subGenus" subGenus="Crossoglossa">Parena (Crossoglossa)</taxonomicName>
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/ sciakyi
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<taxonomicNameLabel id="A269A65C626DFFC82E54B1B2FC645EB9" box="[812,899,969,995]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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/ des. Shi H.L. 2022" [red label] <
|
||
<figureCitation id="1315DBB0626DFFC82872B1B3FA805EB8" box="[1290,1383,968,994]" captionStart="FIGURE 6" captionStartId="13.[151,250,724,748]" captionTargetBox="[209,1380,181,700]" captionTargetId="figure-19@13.[199,1388,181,700]" captionTargetPageId="13" captionText="FIGURE 6. A Hind wing, P. (Crossoglossa) cavipennis (Bates), Beijing, scale bar = 1mm. Abbreviations: oc oblongum cell; wc wedge cell. B–G Tarsals claws of right hindlegs for Metallicina spp., scale bar = 0.2mm: B Pachycallida rufoplagiata Jeannel, Madagascar, Sept Lacs, female. C Metallica viridipennis Chaudoir, Cameroon, Bois des Singes, male. D P. (Crossoglossa) sciakyi sp. n., holotype. E P. (Bothynoptera) kurosai Habu, Nepal. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Parena) latecincta (Bates), Vietnam. H Protibia of P. (Parena) nigrolineata (Chaudoir), scale bar = 0.5 mm. I Mesotarsomeres of male of P. (Parena) nigrolineata (Chaudoir), adhesive hairs present on mesotarsomere 1, scale bar = 0.2mm. J Mesotarsomeres of male for P. (Parena) latecincta (Bates), adhesive hairs absent from mesotarsomere 1, scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7958949" httpUri="https://zenodo.org/record/7958949/files/figure.png" pageId="43" pageNumber="44">Figs 6D</figureCitation>
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,
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<figureCitation id="1315DBB0626DFFC8280BB1B3FA705EB8" box="[1395,1431,968,994]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. A–D females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. E–J males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="43" pageNumber="44">8A</figureCitation>
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,
|
||
<figureCitation id="1315DBB0626DFFC82DEFB197FF55595C" box="[151,178,1004,1030]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="43" pageNumber="44">11</figureCitation>
|
||
γ, 14H, 14K>.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE626DFFC82DBFB66BFF175834" pageId="43" pageNumber="44" type="diagnosis">
|
||
<paragraph id="8B91C735626DFFC82DBFB66BFE3559E0" blockId="43.[151,1437,860,2039]" pageId="43" pageNumber="44">
|
||
<emphasis id="B95A1B27626DFFC82DBFB66BFE2D5970" bold="true" box="[199,458,1040,1066]" pageId="43" pageNumber="44">Diagnostic characters</emphasis>
|
||
. Dorsum pale yellow, elytra with large median black patch; antennomeres 1 to base of 4 yellow, apical antennomeres black; tibiae yellowish brown except apex dark brown; pronotum strongly transverse; elytra striae not incised, replaced by rows of fine punctures; disc shallowly depressed near middle of intervals 3 to 6; sutural angles simple, without pointed denticles (
|
||
<figureCitation id="1315DBB0626DFFC82F9FB607FCB659CC" box="[743,849,1148,1174]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="43" pageNumber="44">Fig. 14H</figureCitation>
|
||
); female abdominal sternite VII with apical margin straight at middle (
|
||
<figureCitation id="1315DBB0626DFFC82C12B6DBFE2359E0" box="[362,452,1184,1210]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. A–D females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. E–J males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="43" pageNumber="44">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B91C735626DFFC82DBFB6BFFF175834" blockId="43.[151,1437,860,2039]" pageId="43" pageNumber="44">
|
||
<emphasis id="B95A1B27626DFFC82DBFB6BFFE815984" bold="true" box="[199,358,1220,1246]" pageId="43" pageNumber="44">Comparisons</emphasis>
|
||
.
|
||
<taxonomicName id="4C2EBCB6626DFFC82C0FB6BEFE045984" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[375,483,1220,1246]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
|
||
<emphasis id="B95A1B27626DFFC82C0FB6BEFE045984" box="[375,483,1220,1246]" italics="true" pageId="43" pageNumber="44">P. sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B27626DFFC82C95B6BFFDC85984" bold="true" box="[493,559,1220,1246]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="A269A65C626DFFC82C95B6BFFDC85984" box="[493,559,1220,1246]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the only known species in the subgenus
|
||
<taxonomicName id="4C2EBCB6626DFFC82954B6BFFB255984" authority="Chaudoir, 1873" box="[1068,1218,1220,1246]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27626DFFC82954B6BFFB255984" box="[1068,1218,1220,1246]" italics="true" pageId="43" pageNumber="44">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
which has a black patch on elytra. Besides the conspicuous large patch, the new species is also different from all other species of the
|
||
<taxonomicName id="4C2EBCB6626DFFC82DEFB776FEE8587C" box="[151,271,1293,1318]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27626DFFC82DEFB776FEE8587C" box="[151,271,1293,1318]" italics="true" pageId="43" pageNumber="44">P. testacea</emphasis>
|
||
</taxonomicName>
|
||
species group by: (1) elytral sutural angles not at all pointed, without denticles or spines; (2) pronotum much wider (PW/PL 1.82) than other species (PW/PL 1.60–1.71); (3) elytra microsculpture stronger than in other species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE626DFFCF2DBFB703FE245CAF" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
|
||
<paragraph id="8B91C735626DFFCF2DBFB703FE245CAF" blockId="43.[151,1437,860,2039]" lastBlockId="44.[151,1436,151,501]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
|
||
<emphasis id="B95A1B27626DFFC82DBFB703FEB658C8" bold="true" box="[199,337,1400,1426]" pageId="43" pageNumber="44">Description</emphasis>
|
||
. Body length
|
||
<quantity id="4CD66AD0626DFFC82C80B703FD8658C9" box="[504,609,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="43" pageNumber="44" unit="mm" value="11.3">11.3 mm</quantity>
|
||
. Body wide and stout, elytra subconvex.
|
||
<emphasis id="B95A1B27626DFFC82938B703FB6E58C8" bold="true" box="[1088,1161,1400,1426]" pageId="43" pageNumber="44">Color.</emphasis>
|
||
Head yellowish brown, slightly darker than pronotum, pronotum pale yellow; antennomeres 1 to 3 yellowish brown, apex of 2 and 3 slightly darker, antennomere 4 gradually darkened from brown base to black apex, remaining antennomeres black, apex of antennomere 11 yellow; apex and inner margins of mandibles black; apex of laciniae and galea black; terminal labial palpomere black except apex, terminal and penultimate maxillary palpomeres black except apex. Elytra with margins pale yellow, disc with a conspicuous trapezoidal black patch, extended from basal pore basally to apical pore on interval 3 apically, extended to interval 6 near humerus and to interval 8 on outer apical angle. Scutellum yellow. Tibiae yellowish brown, apex dark brown; tarsomeres black; claws yellow. Venter of head reddish brown; metasternum and metaepisterna black; remaining parts of venter pale yellow.
|
||
<emphasis id="B95A1B27626DFFC82E85B4E3FBDA5BE8" bold="true" box="[1021,1085,1688,1714]" pageId="43" pageNumber="44">Head</emphasis>
|
||
without microsculpture, vertex and frons with fine punctures; eyes large and strongly prominent; tempora very short, abruptly narrowed behind eyes, length of tempora plus neck-constriction slightly less than one-third of eyes' diameter; postgenae with a pair of long suborbital setae. Antennae barely reaching pronotum base.
|
||
<collectingRegion id="49EA09D7626DFFC82EC4B57EFBFC5A45" box="[956,1051,1797,1823]" country="Myanmar" name="Chin" pageId="43" pageNumber="44">Mentum</collectingRegion>
|
||
without median seta, lateral lobes extremely large, inner margins nearly straight, outer and apical margins widely rounded, epilobes very narrow; submentum with two setae on each side.
|
||
<emphasis id="B95A1B27626DFFC82F1AB537FD3C5A3C" bold="true" box="[610,731,1868,1894]" pageId="43" pageNumber="44">Pronotum</emphasis>
|
||
strongly transverse, PW/PL = 1.82, distinctly wider than head, PW/HW = 1.26; widest at anterior third; lateral margin largely rounded, very weakly sinuate before posterior angles; posterior angles rounded, obtuse; anterior margin slightly curved inward at middle; anterior angles widely rounded. Disc slightly convex, without microsculpture, with very fine and sparse punctures; lateral explanations very wide.
|
||
<emphasis id="B95A1B27626DFFC82DEFB5A7FF045AAC" bold="true" box="[151,227,2012,2038]" pageId="43" pageNumber="44">Elytra</emphasis>
|
||
with isodiametric microsculpture, very faint near base, fine but distinct in other regions. Striae well defined, not incised, replaced by rows of very fine punctures; intervals completely flat, with punctures much sparser than in striae; disc shallowly depressed near middle of intervals 3 to 6, depressions subtriangular; lateral sides slightly depressed near anterior third. Elytral basal pore present on base of stria 1; interval 3 with three discal pores: first one on level of scutellar apex, adjacent to stria 3, second one on basal two-fifth, adjacent to stria 3, third one on apical twelfth, adjacent to stria 2; interval 9 with 27 umbilicular pores.Apical truncation distinct, nearly straight; outer apical angles completely rounded; sutural angles not pointed, without denticles.
|
||
<emphasis id="B95A1B27626AFFCF2E8DB330FBA25C3F" bold="true" box="[1013,1093,331,357]" pageId="44" pageNumber="45">Venter</emphasis>
|
||
. Metasternum with short setae behind mesocoxae; abdominal sternite III to VI with long setae medially; in female, apex of abdominal sternite VII straight, with two long setae on each side (
|
||
<figureCitation id="1315DBB0626AFFCF2F07B3E8FD3D5CF7" box="[639,730,403,429]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. A–D females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. E–J males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="44" pageNumber="45">Fig. 8A</figureCitation>
|
||
).
|
||
<emphasis id="B95A1B27626AFFCF2F88B3E8FC515CF6" bold="true" box="[752,950,402,429]" pageId="44" pageNumber="45">Female genitalia</emphasis>
|
||
. Gonocoxite II of ovipositor dichotomous, outer branch slightly wider than inner branch, similar in length, each branch with two long ensiform setae on apex (
|
||
<figureCitation id="1315DBB0626AFFCF2DE7B3A0FF085CAF" box="[159,239,475,501]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="44" pageNumber="45">Fig. 11</figureCitation>
|
||
γ). Male unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5197BD626AFFCF2DEFB707FBDA58CE" ID-DOI="http://doi.org/10.5281/zenodo.7958989" ID-Zenodo-Dep="7958989" box="[151,1085,1404,1428]" httpUri="https://zenodo.org/record/7958989/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1404,1428]" targetBox="[157,1431,533,1372]" targetPageId="44" targetType="figure">
|
||
<paragraph id="8B91C735626AFFCF2DEFB707FBDA58CE" blockId="44.[151,1085,1404,1428]" box="[151,1085,1404,1428]" pageId="44" pageNumber="45">
|
||
<emphasis id="B95A1B27626AFFCF2DEFB707FEFE58CE" bold="true" box="[151,281,1404,1428]" pageId="44" pageNumber="45">FIGURE 25</emphasis>
|
||
. Habitus of
|
||
<taxonomicName id="4C2EBCB6626AFFCF2CEFB706FD2C58CE" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[407,715,1404,1428]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov." subGenus="Crossoglossa">
|
||
<emphasis id="B95A1B27626AFFCF2CEFB706FE0658CE" box="[407,481,1405,1428]" italics="true" pageId="44" pageNumber="45">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B27626AFFCF2C97B707FD9058CE" box="[495,631,1404,1428]" italics="true" pageId="44" pageNumber="45">Crossoglossa</emphasis>
|
||
)
|
||
<emphasis id="B95A1B27626AFFCF2FFDB707FD2C58CE" box="[645,715,1404,1428]" italics="true" pageId="44" pageNumber="45">sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B27626AFFCF2FAAB707FCEC58CE" bold="true" box="[722,779,1404,1428]" pageId="44" pageNumber="45">
|
||
<taxonomicNameLabel id="A269A65C626AFFCF2FAAB707FCEC58CE" box="[722,779,1404,1428]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype. Scale bar = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33494BE626AFFCF2DBFB7C6FB72588D" box="[199,1173,1469,1495]" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="8B91C735626AFFCF2DBFB7C6FB72588D" blockId="44.[151,1437,1469,1568]" box="[199,1173,1469,1495]" pageId="44" pageNumber="45">
|
||
<emphasis id="B95A1B27626AFFCF2DBFB7C6FEBE588D" bold="true" box="[199,345,1469,1495]" pageId="44" pageNumber="45">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1315DBB0626AFFCF2C10B7C6FE53588D" box="[360,436,1469,1495]" captionStart="MAP 4" captionStartId="36.[152,211,1895,1919]" captionTargetBox="[158,1430,1002,1871]" captionTargetId="figure-71@36.[158,1430,1002,1871]" captionTargetPageId="36" captionText="MAP 4. Distributions for species in the Parena (Crossoglossa) testacea group." figureDoi="http://doi.org/10.5281/zenodo.7963663" httpUri="https://zenodo.org/record/7963663/files/figure.png" pageId="44" pageNumber="45">Map 4</figureCitation>
|
||
, orange). Only known by the
|
||
<typeStatus id="54957997626AFFCF2E7AB7C6FC83588D" box="[770,868,1469,1495]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
from the
|
||
<collectingCountry id="F33987A5626AFFCF2EAAB7C6FB76588D" box="[978,1169,1469,1495]" name="Solomon Islands" pageId="44" pageNumber="45">Solomon Islands</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE626AFFCF2DBFB79AFB985B45" pageId="44" pageNumber="45" type="etymology">
|
||
<paragraph id="8B91C735626AFFCF2DBFB79AFB985B45" blockId="44.[151,1437,1469,1568]" pageId="44" pageNumber="45">
|
||
<emphasis id="B95A1B27626AFFCF2DBFB79AFEAF58A1" bold="true" box="[199,328,1505,1531]" pageId="44" pageNumber="45">Etymology</emphasis>
|
||
. The new species is named for Dr. Riccardo Sciaky (Milano), who kindly provided several important specimens of
|
||
<taxonomicName id="4C2EBCB6626AFFCF2C1AB47DFE545B45" box="[354,435,1542,1567]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27626AFFCF2C1AB47DFE545B45" box="[354,435,1542,1567]" italics="true" pageId="44" pageNumber="45">Parena</emphasis>
|
||
</taxonomicName>
|
||
under his care for the present study, including this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |