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<document id="B10C202A2F42BF5C9A57CB758FD389D9" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623620FFFA82DEFB2ECFB7B5A13" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Bothynoptera) malaisei" docType="treatment" docVersion="8" lastPageNumber="76" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="74" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="9B3E5AFD18077BB8EBD8D90C01F9BA56">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
</mods:titleInfo>
<mods:name id="63737A310D9416F41F4E5516FA4C78EC" type="personal">
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<mods:namePart id="186D75C8BE862388F619B98D328F2A1F">Shi, Hongliang</mods:namePart>
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<mods:nameIdentifier id="B1FAF4844F79320B41EF9A109DF8B2C0" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="B70CC70075626CC66101387A8A6514A8">Liang, Hongbin</mods:namePart>
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<mods:title id="AA6F56466AB577839BEF155B48166DB1">Zootaxa</mods:title>
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<mods:date id="C0B90C5CDC6DDAC850F372C1113A426F">2023</mods:date>
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<mods:number id="74F68735CA7F339B3FA54B69E9B66DFD">2023-05-18</mods:number>
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<treatment id="03877623620FFFA82DEFB2ECFB7B5A13" ID-DOI="http://doi.org/10.5281/zenodo.7963989" ID-GBIF-Taxon="209628958" ID-Zenodo-Dep="7963989" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13" httpUri="http://treatment.plazi.org/id/03877623620FFFA82DEFB2ECFB7B5A13" lastPageId="75" lastPageNumber="76" pageId="73" pageNumber="74">
<subSubSection id="C33494BE620FFFAA2DEFB2ECFCD35DEB" box="[151,820,151,178]" pageId="73" pageNumber="74" type="nomenclature">
<paragraph id="8B91C735620FFFAA2DEFB2ECFCD35DEB" blockId="73.[151,1174,151,214]" box="[151,820,151,178]" pageId="73" pageNumber="74">
<emphasis id="B95A1B27620FFFAA2DEFB2ECFCD35DEB" bold="true" box="[151,820,151,178]" pageId="73" pageNumber="74">
[19]
<taxonomicName id="4C2EBCB6620FFFAA2DB7B2ECFCD35DEB" ID-CoL="4DKPS" authority="(Andrewes, 1947)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[207,820,151,178]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei" subGenus="Bothynoptera">
<emphasis id="B95A1B27620FFFAA2DB7B2ECFEC05DEB" bold="true" box="[207,295,151,177]" italics="true" pageId="73" pageNumber="74">Parena</emphasis>
(
<emphasis id="B95A1B27620FFFAA2C40B2ECFE3A5DE8" bold="true" box="[312,477,151,178]" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27620FFFAA2C96B2ECFDB45DE8" bold="true" box="[494,595,151,178]" italics="true" pageId="73" pageNumber="74">malaisei</emphasis>
(
<bibRefCitation id="EFBFBAC4620FFFAA2F1BB2ECFCCB5DEB" author="Andrewes, H. E." box="[611,812,151,177]" pageId="73" pageNumber="74" pagination="1 - 49" refId="ref109781" refString="Andrewes, H. E. (1947) Entomological results from the Swedish expedition 1934 to Burma and British India. Coleoptera: Carabidae. Collected by Rene Malaise. Arkiv f ˆ r Zoologi, 38 A nr. 20, 1 - 49." type="journal article" year="1947">Andrewes, 1947</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620FFFAA2DEFB2C0FB715D8C" box="[151,1174,187,214]" pageId="73" pageNumber="74" type="description">
<paragraph id="8B91C735620FFFAA2DEFB2C0FB715D8C" blockId="73.[151,1174,151,214]" box="[151,1174,187,214]" pageId="73" pageNumber="74">
Habitus:
<figureCitation id="1315DBB0620FFFAA2D85B2C0FE065D8F" box="[253,481,187,213]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="73" pageNumber="74">Figs 43A, 43B, 43C</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB0620FFFAA2FE4B2C0FD095D8F" box="[668,750,187,213]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="73" pageNumber="74">Fig. 44</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB0620FFFAA2954B2C0FB695D8F" box="[1068,1166,187,213]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="73" pageNumber="74">Fig. 11G</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620FFFAA2DEFB37BFCC45C2E" pageId="73" pageNumber="74" type="reference_group">
<paragraph id="8B91C735620FFFAA2DEFB37BFAE25C6C" blockId="73.[151,1437,255,372]" pageId="73" pageNumber="74">
<bibRefCitation id="EFBFBAC4620FFFAA2DEFB37BFE835C4D" author="Andrewes, H. E." box="[151,356,255,280]" pageId="73" pageNumber="74" pagination="1 - 49" refId="ref109781" refString="Andrewes, H. E. (1947) Entomological results from the Swedish expedition 1934 to Burma and British India. Coleoptera: Carabidae. Collected by Rene Malaise. Arkiv f ˆ r Zoologi, 38 A nr. 20, 1 - 49." type="journal article" year="1947">Andrewes, 1947: 46</bibRefCitation>
(original:
<taxonomicName id="4C2EBCB6620FFFAA2CA8B37BFDBD5C42" ID-CoL="8LR5S" authority="Schaum, 1863" box="[464,602,256,280]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27620FFFAA2CA8B37BFDBD5C42" box="[464,602,256,280]" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
</taxonomicName>
; type locality:
<collectingCountry id="F33987A5620FFFAA2F8BB37BFCDB5C4D" box="[755,828,256,279]" name="Myanmar" pageId="73" pageNumber="74">Burma</collectingCountry>
: Kambaiti;
<typeStatus id="54957997620FFFAA2EC9B37BFBED5C42" box="[945,1034,256,280]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
in NHRS);
<bibRefCitation id="EFBFBAC4620FFFAA29FCB37BFADA5C4D" author="Jedlicka, A." box="[1156,1341,255,280]" pageId="73" pageNumber="74" pagination="59 - 60" refId="ref111384" refString="Jedlicka, A. (1951) Stukie O. Carabidech Palaearcticke Fauny. Etudes sur les Carabides de la faune palaearctique. Acta Musei Silesiae, (A) 1, 59 - 60." type="journal article" year="1951">Jedlička, 1951: 60</bibRefCitation>
(
<taxonomicName id="4C2EBCB6620FFFAA2834B37BFEEE5C6C" ID-CoL="8LR5S" authority="Schaum, 1863" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27620FFFAA2834B37BFEEE5C6C" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC4620FFFAA2C67B365FE025C6C" author="Jedlicka, A." box="[287,485,286,310]" pageId="73" pageNumber="74" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 447</bibRefCitation>
(
<taxonomicName id="4C2EBCB6620FFFAA2C8CB365FD995C6C" ID-CoL="8LR5S" authority="Schaum, 1863" box="[500,638,286,310]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27620FFFAA2C8CB365FD995C6C" box="[500,638,286,310]" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
</taxonomicName>
); Kirschenhofer, 1996: 781 (Yunnan);
<bibRefCitation id="EFBFBAC4620FFFAA2970B365FAE75C6C" author="Kirschenhofer, E." box="[1032,1280,286,310]" pageId="73" pageNumber="74" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
.
</paragraph>
<paragraph id="8B91C735620FFFAA2DEFB346FCC45C2E" blockId="73.[151,1437,255,372]" pageId="73" pageNumber="74">
<taxonomicName id="4C2EBCB6620FFFAA2DEFB346FDC45C0F" ID-CoL="4DKNZ" authority="Habu, 1979: 55" authorityName="Habu" authorityPageNumber="55" authorityYear="1979" box="[151,547,316,341]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="B95A1B27620FFFAA2DEFB346FE9D5C0E" bold="true" box="[151,378,316,340]" italics="true" pageId="73" pageNumber="74">Parena albomaculata</emphasis>
<materialsCitation id="3B46CD68620FFFAA2CFAB346FDC45C0F" ID-GBIF-Occurrence="4115871427" box="[386,547,317,341]" country="Taiwan" location="Science" municipality="National Museum of Nature" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Tokyo" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4620FFFAA2CFAB346FDC45C0F" author="Habu, A." box="[386,547,317,341]" pageId="73" pageNumber="74" pagination="51 - 59" refId="ref110845" refString="Habu, A. (1979) Formosan Lebiini collected by Professor T. Shirozu in 1965 (Coleoptera, Carabidae). The Entomological Review of Japan, 33, 51 - 59." type="journal article" year="1979">Habu, 1979: 55</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality:
<collectingCountry id="F33987A5620FFFAA2FBBB346FCF65C0F" box="[707,785,317,341]" name="Taiwan" pageId="73" pageNumber="74">Taiwan</collectingCountry>
: Tottaka;
<typeStatus id="54957997620FFFAA2E0FB346FC375C0F" box="[887,976,317,341]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
in NMNS:
<collectingMunicipality id="6BF55D4F620FFFAA2934B346FA885C0E" box="[1100,1391,317,341]" pageId="73" pageNumber="74">National Museum of Nature</collectingMunicipality>
and
<location id="8EF191EE620FFFAA2DBFB327FEFF5C2E" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620FFFAA2DBFB327FEFF5C2E" box="[199,280,348,372]" country="Taiwan" municipality="National Museum of Nature" name="Science" pageId="73" pageNumber="74" stateProvince="Tokyo">Science</location>
,
<collectingRegion id="49EA09D7620FFFAA2C5AB327FE825C2E" box="[290,357,348,372]" country="Japan" name="Tokyo" pageId="73" pageNumber="74">Tokyo</collectingRegion>
,
<collectingCountry id="F33987A5620FFFAA2C17B327FE4A5C2E" box="[367,429,348,372]" name="Japan" pageId="73" pageNumber="74">Japan</collectingCountry>
);
<bibRefCitation id="EFBFBAC4620FFFAA2CC7B327FD505C2E" author="Kirschenhofer, E." box="[447,695,347,372]" pageId="73" pageNumber="74" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 88</bibRefCitation>
.
<emphasis id="B95A1B27620FFFAA2FB9B320FCC45C2E" bold="true" box="[705,803,347,372]" pageId="73" pageNumber="74">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<caption id="DF5197BD620FFFAA2DEFB4AEFC265A03" ID-DOI="http://doi.org/10.5281/zenodo.7959025" ID-Zenodo-Dep="7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,250,1749,1773]" targetBox="[157,1434,397,1717]" targetPageId="73" targetType="figure">
<paragraph id="8B91C735620FFFAA2DEFB4AEFC265A03" blockId="73.[151,1437,1749,1882]" pageId="73" pageNumber="74">
<emphasis id="B95A1B27620FFFAA2DEFB4AEFEFB5BB7" bold="true" box="[151,284,1749,1773]" pageId="73" pageNumber="74">FIGURE 43</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB6620FFFAA2CDCB4ADFC955BB7" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[420,882,1749,1774]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei" subGenus="Bothynoptera">
<emphasis id="B95A1B27620FFFAA2CDCB4ADFE095BB7" box="[420,494,1750,1773]" italics="true" pageId="73" pageNumber="74">Parena</emphasis>
(
<emphasis id="B95A1B27620FFFAA2C87B4AEFD6E5BB7" box="[511,649,1749,1773]" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27620FFFAA2FE3B4AEFD165BB7" box="[667,753,1749,1773]" italics="true" pageId="73" pageNumber="74">malaisei</emphasis>
(Andrewes)
</taxonomicName>
and
<taxonomicName id="4C2EBCB6620FFFAA2ED2B4ADFA7B5BB7" authority="Mateu. A" authorityName="Mateu. A" authorityYear="1977" box="[938,1436,1749,1774]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="quadrisignata" subGenus="Bothynoptera">
<emphasis id="B95A1B27620FFFAA2ED2B4ADFC135BB7" box="[938,1012,1750,1773]" italics="true" pageId="73" pageNumber="74">Parena</emphasis>
(
<emphasis id="B95A1B27620FFFAA297EB4AEFB775BB7" box="[1030,1168,1749,1773]" italics="true" pageId="73" pageNumber="74">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27620FFFAA29DAB4AEFAD75BB7" box="[1186,1328,1749,1773]" italics="true" pageId="73" pageNumber="74">quadrisignata</emphasis>
Mateu.
<emphasis id="B95A1B27620FFFAA28F2B4AEFA7B5BB7" bold="true" box="[1418,1436,1749,1773]" pageId="73" pageNumber="74">A</emphasis>
</taxonomicName>
Holotype of
<taxonomicName id="4C2EBCB6620FFFAA2C6FB481FDDA5A4B" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[279,573,1785,1809]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620FFFAA2C6FB481FE5B5A4B" box="[279,444,1785,1809]" italics="true" pageId="73" pageNumber="74">Parena malaisei</emphasis>
(Andrewes)
</taxonomicName>
.
<emphasis id="B95A1B27620FFFAA2F3EB481FDB05A48" bold="true" box="[582,599,1786,1810]" pageId="73" pageNumber="74">B</emphasis>
<taxonomicName id="4C2EBCB6620FFFAA2F25B481FCA85A4B" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[605,847,1785,1809]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620FFFAA2F25B481FD8B5A4B" box="[605,620,1786,1809]" italics="true" pageId="73" pageNumber="74">P</emphasis>
.
<emphasis id="B95A1B27620FFFAA2F01B482FD285A4B" box="[633,719,1785,1809]" italics="true" pageId="73" pageNumber="74">malaisei</emphasis>
(Andrewes)
</taxonomicName>
, male from Yunnan.
<emphasis id="B95A1B27620FFFAA2958B482FBD55A4B" bold="true" box="[1056,1074,1785,1809]" pageId="73" pageNumber="74">C</emphasis>
<taxonomicName id="4C2EBCB6620FFFAA2940B481FACD5A4B" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[1080,1322,1785,1809]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620FFFAA2940B481FBA05A4B" box="[1080,1095,1786,1809]" italics="true" pageId="73" pageNumber="74">P</emphasis>
.
<emphasis id="B95A1B27620FFFAA292CB482FB4D5A4B" box="[1108,1194,1785,1809]" italics="true" pageId="73" pageNumber="74">malaisei</emphasis>
(Andrewes)
</taxonomicName>
, male from Nepal.
<emphasis id="B95A1B27620FFFAA2D98B565FF155A6C" bold="true" box="[224,242,1822,1846]" pageId="73" pageNumber="74">D</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6620FFFAA2C00B565FD5D5A6C" authority="Mateu. E" authorityName="Mateu. E" box="[376,698,1821,1846]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B95A1B27620FFFAA2C00B565FDB25A6F" box="[376,597,1821,1845]" italics="true" pageId="73" pageNumber="74">Parena quadrisignata</emphasis>
Mateu.
<emphasis id="B95A1B27620FFFAA2FD1B565FD5D5A6C" bold="true" box="[681,698,1822,1846]" pageId="73" pageNumber="74">E</emphasis>
</taxonomicName>
Holotype of
<taxonomicName id="4C2EBCB6620FFFAA2E47B565FB275A6F" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2011" box="[831,1216,1821,1845]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="phongsalyensis">
<emphasis id="B95A1B27620FFFAA2E47B565FBCD5A6F" box="[831,1066,1821,1845]" italics="true" pageId="73" pageNumber="74">Parena phongsalyensis</emphasis>
Kirschenhofer
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6620FFFAA299BB565FEF65A03" authority="Mateu" authorityName="Mateu. A" authorityYear="1977" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B95A1B27620FFFAA299BB565FF235A03" italics="true" pageId="73" pageNumber="74">Parena quadrisignata</emphasis>
Mateu
</taxonomicName>
].
<emphasis id="B95A1B27620FFFAA2C58B539FEC85A00" bold="true" box="[288,303,1858,1882]" pageId="73" pageNumber="74">F</emphasis>
<taxonomicName id="4C2EBCB6620FFFAA2C4EB539FDCD5A03" authority="Mateu" authorityName="Mateu. A" authorityYear="1977" box="[310,554,1857,1881]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B95A1B27620FFFAA2C4EB539FEA25A03" box="[310,325,1858,1881]" italics="true" pageId="73" pageNumber="74">P</emphasis>
.
<emphasis id="B95A1B27620FFFAA2C2AB53AFE075A03" box="[338,480,1857,1881]" italics="true" pageId="73" pageNumber="74">quadrisignata</emphasis>
Mateu
</taxonomicName>
, male from Yunnan. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE620FFFA92DEFB5F3FC455C47" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="8B91C735620FFFA92DEFB5F3FC455C47" blockId="73.[151,1436,1927,2026]" lastBlockId="74.[151,1437,150,2013]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<emphasis id="B95A1B27620FFFAA2DEFB5F3FE585AFB" bold="true" box="[151,447,1927,1954]" pageId="73" pageNumber="74">Type material examined</emphasis>
.
<taxonomicName id="4C2EBCB6620FFFAA2CA8B5FCFCBD5AFB" authority="Andrewes" authorityName="Andrewes" authorityYear="1947" box="[464,858,1927,1954]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620FFFAA2CA8B5FCFCBD5AFB" bold="true" box="[464,858,1927,1954]" pageId="73" pageNumber="74">
<emphasis id="B95A1B27620FFFAA2CA8B5FCFD3F5AF8" bold="true" box="[464,728,1927,1954]" italics="true" pageId="73" pageNumber="74">Bothynoptera malaisei</emphasis>
Andrewes
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27620FFFAA2E14B5F3FC3E5AF8" bold="true" box="[876,985,1928,1954]" pageId="73" pageNumber="74">
<typeStatus id="54957997620FFFAA2E14B5F3FC3E5AF8" box="[876,985,1928,1954]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
(NHRS,
<figureCitation id="1315DBB0620FFFAA2931B5F3FB525AF8" box="[1097,1205,1928,1954]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="73" pageNumber="74">Fig. 43A</figureCitation>
): male, body length =
<quantity id="4CD66AD0620FFFAA2DD7B5D7FEEF5A9C" box="[175,264,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="73" pageNumber="74" unit="mm" value="7.1">7.1 mm</quantity>
, board mounted, &quot;N.E.
<collectingCountry id="F33987A5620FFFAA2F6DB5D7FD9D5A9C" box="[533,634,1964,1990]" name="Myanmar" pageId="73" pageNumber="74">BURMA</collectingCountry>
/ Kambaiti,
<quantity id="4CD66AD0620FFFAA2E72B5D7FC855A9C" box="[778,866,1964,1990]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="73" pageNumber="74" unit="ft" value="7000.0">7000 ft.</quantity>
/
<emphasis id="B95A1B27620FFFAA2E02B5D7FC0B5A9C" box="[890,1004,1964,1990]" italics="true" pageId="73" pageNumber="74">
<date id="FF90E1F5620FFFAA2E02B5D7FC0B5A9C" box="[890,1004,1964,1990]" pageId="73" pageNumber="74" value="1934-05-23">23/5 1934</date>
</emphasis>
/ R. MALAISE&quot;, &quot;Typus&quot; [red label], &quot;7882 / E91+&quot; [blue label], &quot;Figured / Specimen&quot;, &quot;
<emphasis id="B95A1B27620FFFAA2F96B5ABFB465AB0" box="[750,1185,2000,2026]" italics="true" pageId="73" pageNumber="74">
<taxonomicName id="4C2EBCB6620FFFAA2F96B5ABFC615AB0" authority="Schaum, 1863" box="[750,902,2000,2026]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">Bothynoptera</taxonomicName>
/
<taxonomicName id="4C2EBCB6620FFFAA2EDAB5ABFBE65AB0" authorityName="Andrewes" authorityYear="1947" box="[930,1025,2000,2026]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="malaisei">malaisei</taxonomicName>
/ type Andr. /
</emphasis>
H.E. Andrewes det.&quot;, &quot;NHRS-JLKB / 000020251&quot;.
<emphasis id="B95A1B27620CFFA92C9EB2ECFDB95DEB" bold="true" box="[486,606,151,177]" pageId="74" pageNumber="75">
<typeStatus id="54957997620CFFA92C9EB2ECFDB95DEB" box="[486,606,151,177]" pageId="74" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
(NHML): 1 ex, &quot;N.E.
<collectingCountry id="F33987A5620CFFA92E24B2ECFC4F5DEB" box="[860,936,151,177]" name="Myanmar" pageId="74" pageNumber="75">Burma</collectingCountry>
/ Kambaiti,
<quantity id="4CD66AD0620CFFA9294BB2ECFB635DEB" box="[1075,1156,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="74" pageNumber="75" unit="ft" value="7000.0">7000ft.</quantity>
/
<emphasis id="B95A1B27620CFFA929E1B2ECFAED5DEB" box="[1177,1290,151,177]" italics="true" pageId="74" pageNumber="75">
<date id="FF90E1F5620CFFA929E1B2ECFAED5DEB" box="[1177,1290,151,177]" pageId="74" pageNumber="75" value="1934-04-18">18/4 1934</date>
</emphasis>
R. Malaise&quot;, &quot;Para-type&quot; [round label with yellow circle], &quot;Brit. Mus. / 1947-14&quot;, &quot;
<emphasis id="B95A1B27620CFFA92ED3B2C0FA855D8F" box="[939,1378,187,213]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="4C2EBCB6620CFFA92ED3B2C0FBD95D8F" authority="Schaum, 1863" box="[939,1086,187,213]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">Bothynoptera</taxonomicName>
/
<taxonomicName id="4C2EBCB6620CFFA9292AB2C0FB4B5D8F" authorityName="Andrewes" authorityYear="1947" box="[1106,1196,187,213]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="malaisei">malaisei</taxonomicName>
/ Cotype Andr. /
</emphasis>
H.E. Andrewes det.&quot;. 1 ex, &quot;N.E.
<collectingCountry id="F33987A5620CFFA92C98B2A4FDCB5DA3" box="[480,556,223,249]" name="Myanmar" pageId="74" pageNumber="75">Burma</collectingCountry>
/ Kambaiti,
<quantity id="4CD66AD0620CFFA92FC7B2A4FCF15DA3" box="[703,790,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="74" pageNumber="75" unit="m" value="2000.0">2000 m</quantity>
/ 23/5.1934 Malaise&quot;, &quot;Co-type&quot;[round label with green circle]; &quot;
<emphasis id="B95A1B27620CFFA92D81B378FD495C47" box="[249,686,259,285]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="4C2EBCB6620CFFA92D81B378FE6B5C47" authority="Schaum, 1863" box="[249,396,259,285]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">Bothynoptera</taxonomicName>
/
<taxonomicName id="4C2EBCB6620CFFA92CD8B378FE1D5C47" authorityName="Andrewes" authorityYear="1947" box="[416,506,259,285]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="malaisei">malaisei</taxonomicName>
/ Cotype Andr. /
</emphasis>
H.E. Andrewes det.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620CFFA92DBFB35CFB085CF7" pageId="74" pageNumber="75" type="discussion">
<paragraph id="8B91C735620CFFA92DBFB35CFB085CF7" blockId="74.[151,1437,150,2013]" pageId="74" pageNumber="75">
<emphasis id="B95A1B27620CFFA92DBFB35CFDC35C1B" bold="true" box="[199,548,295,321]" pageId="74" pageNumber="75">
Notes on
<typeStatus id="54957997620CFFA92C4DB35CFE945C1B" box="[309,371,295,321]" pageId="74" pageNumber="75">types</typeStatus>
and synonyms
</emphasis>
.
<taxonomicName id="4C2EBCB6620CFFA92F4AB35CFC945C1B" authority="Habu" authorityName="Habu" authorityYear="1979" box="[562,883,294,321]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="B95A1B27620CFFA92F4AB35CFC945C1B" bold="true" box="[562,883,294,321]" pageId="74" pageNumber="75">
<emphasis id="B95A1B27620CFFA92F4AB35CFCCE5C1B" bold="true" box="[562,809,294,321]" italics="true" pageId="74" pageNumber="75">Parena albomaculata</emphasis>
Habu
</emphasis>
</taxonomicName>
: Habu described this species without comparison with
<taxonomicName id="4C2EBCB6620CFFA92DB7B330FEAE5C3F" authorityName="Andrewes" authorityYear="1947" box="[207,329,331,357]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620CFFA92DB7B330FEAE5C3F" box="[207,329,331,357]" italics="true" pageId="74" pageNumber="75">P. malaisei</emphasis>
</taxonomicName>
, which is very similar in external features. We studied male genitalia of
<taxonomicName id="4C2EBCB6620CFFA92913B330FB025C3F" authorityName="Andrewes" authorityYear="1947" box="[1131,1253,331,357]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620CFFA92913B330FB025C3F" box="[1131,1253,331,357]" italics="true" pageId="74" pageNumber="75">P. malaisei</emphasis>
</taxonomicName>
from several localities including
<collectingCountry id="F33987A5620CFFA92C25B314FE485CD3" box="[349,431,367,393]" name="Taiwan" pageId="74" pageNumber="75">Taiwan</collectingCountry>
(
<figureCitation id="1315DBB0620CFFA92CC5B314FDC45CD3" box="[445,547,367,393]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="74" pageNumber="75">Fig. 44B</figureCitation>
), the
<typeStatus id="54957997620CFFA92F27B314FD685CD3" box="[607,655,367,393]" pageId="74" pageNumber="75">type</typeStatus>
locality of
<taxonomicName id="4C2EBCB6620CFFA92E76B314FC245CD3" authorityName="Habu" authorityYear="1979" box="[782,963,367,393]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="B95A1B27620CFFA92E76B314FC245CD3" box="[782,963,367,393]" italics="true" pageId="74" pageNumber="75">P. albomaculata</emphasis>
</taxonomicName>
, and found that they have only very subtle differences in the apical lamella. Thus we treat
<taxonomicName id="4C2EBCB6620CFFA92FD1B3E8FCB85CF7" authorityName="Habu" authorityYear="1979" box="[681,863,403,429]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="albomaculata">
<emphasis id="B95A1B27620CFFA92FD1B3E8FCB85CF7" box="[681,863,403,429]" italics="true" pageId="74" pageNumber="75">P. albomaculata</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C2EBCB6620CFFA92915B3E8FB0F5CF7" authorityName="Andrewes" authorityYear="1947" box="[1133,1256,403,429]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B95A1B27620CFFA92915B3E8FB0F5CF7" box="[1133,1256,403,429]" italics="true" pageId="74" pageNumber="75">P. malaisei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620CFFA92DBFB3CCFA9758BF" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="8B91C735620CFFA92DBFB3CCFA9758BF" blockId="74.[151,1437,150,2013]" pageId="74" pageNumber="75">
<emphasis id="B95A1B27620CFFA92DBFB3CCFDF85C8A" bold="true" box="[199,543,438,465]" pageId="74" pageNumber="75">Non-type material examined</emphasis>
.
<materialsCitation id="3B46CD68620CFFA92F57B3CDFCCA5CAF" ID-GBIF-Occurrence="4115871318" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<emphasis id="B95A1B27620CFFA92F57B3CDFD665C8A" bold="true" box="[559,641,438,464]" pageId="74" pageNumber="75">China:</emphasis>
1 ex (IZAS), &quot;Hubei,
<collectingCounty id="62F0BFB9620CFFA92EFCB3CCFC005C8B" box="[900,999,439,465]" pageId="74" pageNumber="75">Xuan-en</collectingCounty>
,
<collectingMunicipality id="6BF55D4F620CFFA92E8DB3CCFB935C8B" box="[1013,1140,439,465]" pageId="74" pageNumber="75">Shadaogou</collectingMunicipality>
,
<location id="8EF191EE620CFFA929F9B3CCFAD35C8B" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA929F9B3CCFAD35C8B" box="[1153,1332,439,465]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Longtan station" pageId="74" pageNumber="75" stateProvince="Hubei">Longtan station</location>
,
<geoCoordinate id="EE1AA1F2620CFFA92839B3CCFA7B5C8B" box="[1345,1436,439,465]" degrees="29.6998" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="5" value="29.6998">29.6998</geoCoordinate>
<geoCoordinate id="EE1AA1F2620CFFA92DEFB3A1FEE55CAF" box="[151,258,474,501]" degrees="109.6511" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="5" value="109.6511">109.6511</geoCoordinate>
,
<quantity id="4CD66AD0620CFFA92C76B3A0FEBF5CAF" box="[270,344,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="74" pageNumber="75" unit="m" value="631.0">
<elevation id="00032006620CFFA92C76B3A0FEBF5CAF" box="[270,344,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.31" pageId="74" pageNumber="75" unit="m" value="631.0">631 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92C1CB3A0FE335CAF" box="[356,468,475,501]" pageId="74" pageNumber="75" value="2016-04-30">
<collectingDate id="EFD4181D620CFFA92C1CB3A0FE335CAF" box="[356,468,475,501]" pageId="74" pageNumber="75" value="2016-04-30">2016.4.30</collectingDate>
</date>
D Liang HB., Zhao KD. lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92E4DB3A0FD575F43" ID-GBIF-Occurrence="4115871320" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Hunan, Sangzhi, Tianpingshan,
<quantity id="4CD66AD0620CFFA92838B3A0FA705CAF" box="[1344,1431,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="74" pageNumber="75" unit="m" value="1330.0">
<elevation id="00032006620CFFA92838B3A0FA705CAF" box="[1344,1431,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="74" pageNumber="75" unit="m" value="1330.0">1330 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92DEFB384FEE85F43" box="[151,271,511,537]" pageId="74" pageNumber="75" value="2014-10-24">
<collectingDate id="EFD4181D620CFFA92DEFB384FEE85F43" box="[151,271,511,537]" pageId="74" pageNumber="75" value="2014-10-24">2014.X.24</collectingDate>
</date>
D,
<geoCoordinate id="EE1AA1F2620CFFA92C4AB384FE6A5F43" box="[306,397,511,537]" degrees="29.7712" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="5" value="29.7712">29.7712</geoCoordinate>
<geoCoordinate id="EE1AA1F2620CFFA92CEEB384FDE65F43" box="[406,513,511,537]" degrees="110.0680" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="5" value="110.068">110.0680</geoCoordinate>
, Yao Jian lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92FC2B384FE1B5F67" ID-GBIF-Occurrence="4115871434" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (HBUM), &quot;Sichuan, Baoxing, Fengtongzhai,
<date id="FF90E1F5620CFFA92982B384FA7F5F43" box="[1274,1432,511,537]" pageId="74" pageNumber="75" value="2004-08-01" valueMax="2004-08-03" valueMin="2004-08-01">
<collectingDate id="EFD4181D620CFFA92982B384FA7F5F43" box="[1274,1432,511,537]" pageId="74" pageNumber="75" value="2004-08-01" valueMax="2004-08-03" valueMin="2004-08-01">2004.VIII.1-3</collectingDate>
</date>
, Yang Xiujuan, Hua Huiran lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92F7AB058FEB25F3B" ID-GBIF-Occurrence="4115871527" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<specimenCount id="9D280CBC620CFFA92F7AB058FDAC5F67" box="[514,587,547,573]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(HBUM), &quot; Sichuan, Baoxing, Fengtongzhai,,
<date id="FF90E1F5620CFFA9292CB058FB155F67" box="[1108,1266,547,573]" pageId="74" pageNumber="75" value="2004-08-01" valueMax="2004-08-03" valueMin="2004-08-01">
<collectingDate id="EFD4181D620CFFA9292CB058FB155F67" box="[1108,1266,547,573]" pageId="74" pageNumber="75" value="2004-08-01" valueMax="2004-08-03" valueMin="2004-08-01">2004.VIII.1-3</collectingDate>
</date>
, Yang Xiujuan, Hua Huiran lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92C23B03CFA7B5F3B" ID-GBIF-Occurrence="4115871505" box="[347,1436,583,609]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (ZWWC), &quot;Chongqing, Fuling, Damu forestry station,
<quantity id="4CD66AD0620CFFA92E96B03CFBA45F3B" box="[1006,1091,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="m" value="1600.0">
<elevation id="00032006620CFFA92E96B03CFBA45F3B" box="[1006,1091,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92935B03CFB555F3B" box="[1101,1202,583,609]" pageId="74" pageNumber="75" value="2006-05-01">
<collectingDate id="EFD4181D620CFFA92935B03CFB555F3B" box="[1101,1202,583,609]" pageId="74" pageNumber="75" value="2006-05-01">2006.V.1</collectingDate>
</date>
, Zhang Weiwei lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92DEFB010FB5D5FDF" ID-GBIF-Occurrence="4115871575" box="[151,1210,618,645]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
2 ex (HBUM), &quot;Guizhou, Leishan, Fangxiang,
<date id="FF90E1F5620CFFA92FC9B011FCBF5FDF" box="[689,856,618,645]" pageId="74" pageNumber="75" value="2005-09-15" valueMax="2005-09-16" valueMin="2005-09-15">
<collectingDate id="EFD4181D620CFFA92FC9B011FCBF5FDF" box="[689,856,618,645]" pageId="74" pageNumber="75" value="2005-09-15" valueMax="2005-09-16" valueMin="2005-09-15">2005.IX.15-16</collectingDate>
</date>
, Wang Jiliang, Gao Chao lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA929BBB010FC565FF3" ID-GBIF-Occurrence="4115871523" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Guizhou, Daozhen, Xiannyudong,
<quantity id="4CD66AD0620CFFA92C89B0F5FDDD5FF3" box="[497,570,654,681]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="74" pageNumber="75" unit="m" value="650.0">
<elevation id="00032006620CFFA92C89B0F5FDDD5FF3" box="[497,570,654,681]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="74" pageNumber="75" unit="m" value="650.0">650 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92F3DB0F4FD5E5FF3" box="[581,697,655,681]" pageId="74" pageNumber="75" value="2006-05-17">
<collectingDate id="EFD4181D620CFFA92F3DB0F4FD5E5FF3" box="[581,697,655,681]" pageId="74" pageNumber="75" value="2006-05-17">2006.V.17</collectingDate>
</date>
, Zhou Huizhong lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92EC0B0F4FD6A5F97" ID-GBIF-Occurrence="4115871325" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Guangxi, Tianlin, Linaoshan,
<quantity id="4CD66AD0620CFFA92DEFB0C8FED65F97" box="[151,305,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="74" pageNumber="75" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
<elevation id="00032006620CFFA92DEFB0C8FED65F97" box="[151,305,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="74" pageNumber="75" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300-1400 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92C47B0C8FE535F97" box="[319,436,691,717]" pageId="74" pageNumber="75" value="2002-05-28">
<collectingDate id="EFD4181D620CFFA92C47B0C8FE535F97" box="[319,436,691,717]" pageId="74" pageNumber="75" value="2002-05-28">2002.V.28</collectingDate>
</date>
, Liu Jianwen lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92FEEB0C8FC905FAB" ID-GBIF-Occurrence="4115871522" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Guangxi, Xing'an, Maoershan,
<collectingMethod id="526FBF22620CFFA929D8B0C8FAEC5F97" box="[1184,1291,691,717]" pageId="74" pageNumber="75">light trap</collectingMethod>
,
<geoCoordinate id="EE1AA1F2620CFFA92860B0C9FA7F5F97" box="[1304,1432,690,717]" degrees="25.87001" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="25.87001">N25.87001</geoCoordinate>
,
<geoCoordinate id="EE1AA1F2620CFFA92DEFB0ACFEC65FAB" box="[151,289,727,753]" degrees="110.41660" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="110.4166">E110.41660</geoCoordinate>
,
<quantity id="4CD66AD0620CFFA92C57B0ADFE6E5FAB" box="[303,393,726,753]" metricMagnitude="3" metricUnit="m" metricValue="2.005" pageId="74" pageNumber="75" unit="m" value="2005.0">
<elevation id="00032006620CFFA92C57B0ADFE6E5FAB" box="[303,393,726,753]" metricMagnitude="3" metricUnit="m" metricValue="2.005" pageId="74" pageNumber="75" unit="m" value="2005.0">2005 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92CEFB0ACFDFE5FAB" box="[407,537,726,753]" pageId="74" pageNumber="75" value="2006-06-25">
<collectingDate id="EFD4181D620CFFA92CEFB0ACFDFE5FAB" box="[407,537,726,753]" pageId="74" pageNumber="75" value="2006-06-25">2006.VI.25</collectingDate>
</date>
, Liang Hongbin,
<collectorName id="26DBA2E3620CFFA92F9DB0ACFCD05FAB" box="[741,823,727,753]" pageId="74" pageNumber="75">T. Sota</collectorName>
lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92EF9B0ACFC895E4F" ID-GBIF-Occurrence="4115871426" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<specimenCount id="9D280CBC620CFFA92EF9B0ACFC285FAB" box="[897,975,727,753]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(CCCC), &quot;Guangxi, Jinxiu, Dayaoshan, Tiantoutun,
<quantity id="4CD66AD0620CFFA92C5AB080FE9B5E4F" box="[290,380,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="74" pageNumber="75" unit="m" value="1100.0">
<elevation id="00032006620CFFA92C5AB080FE9B5E4F" box="[290,380,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="74" pageNumber="75" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92CF3B080FDC55E4F" box="[395,546,763,789]" pageId="74" pageNumber="75" value="2016-08-06">
<collectingDate id="EFD4181D620CFFA92CF3B080FDC55E4F" box="[395,546,763,789]" pageId="74" pageNumber="75" value="2016-08-06">2016.VIII-06</collectingDate>
</date>
, Zhao J.T. lgt.&quot; &lt;
<figureCitation id="1315DBB0620CFFA92F89B080FCBC5E4F" box="[753,859,763,789]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="74" pageNumber="75">Fig. 44A</figureCitation>
&gt;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92E00B080FDAF5E63" ID-GBIF-Occurrence="4115871595" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CCCC), &quot;Yunnan, Gongshan, Xiongdang,
<quantity id="4CD66AD0620CFFA92DEFB164FF095E63" box="[151,238,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.919" pageId="74" pageNumber="75" unit="m" value="1919.0">
<elevation id="00032006620CFFA92DEFB164FF095E63" box="[151,238,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.919" pageId="74" pageNumber="75" unit="m" value="1919.0">1919 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92D81B165FE6E5E63" box="[249,393,798,825]" pageId="74" pageNumber="75" value="2015-07-13">
<collectingDate id="EFD4181D620CFFA92D81B165FE6E5E63" box="[249,393,798,825]" pageId="74" pageNumber="75" value="2015-07-13">2015-VII-13</collectingDate>
</date>
, Yang X.D. lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92F37B164FEEA5E07" ID-GBIF-Occurrence="4115871362" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CCCC), &quot;Yunnan, Gongshan, Menggaguo,
<quantity id="4CD66AD0620CFFA92909B164FB205E63" box="[1137,1223,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.8339999999999996" pageId="74" pageNumber="75" unit="m" value="2834.0">
<elevation id="00032006620CFFA92909B164FB205E63" box="[1137,1223,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.8339999999999996" pageId="74" pageNumber="75" unit="m" value="2834.0">2834 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA929ABB164FABE5E63" box="[1235,1369,799,825]" pageId="74" pageNumber="75" value="2016-06-11">
<collectingDate id="EFD4181D620CFFA929ABB164FABE5E63" box="[1235,1369,799,825]" pageId="74" pageNumber="75" value="2016-06-11">2016-VI-11</collectingDate>
</date>
, Yang X.D. lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92C6DB138FB7C5E07" ID-GBIF-Occurrence="4115871398" box="[277,1179,835,861]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CCCC), &quot;Yunnan, Lushui, Pianma,
<quantity id="4CD66AD0620CFFA92F93B138FCA45E07" box="[747,835,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.643" pageId="74" pageNumber="75" unit="m" value="2643.0">
<elevation id="00032006620CFFA92F93B138FCA45E07" box="[747,835,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.643" pageId="74" pageNumber="75" unit="m" value="2643.0">2643 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92E29B138FC305E07" box="[849,983,835,861]" pageId="74" pageNumber="75" value="2014-06-17">
<collectingDate id="EFD4181D620CFFA92E29B138FC305E07" box="[849,983,835,861]" pageId="74" pageNumber="75" value="2014-06-17">2014-VI-17</collectingDate>
</date>
, Yang X.D. lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA929DBB138FF0F5EFF" ID-GBIF-Occurrence="4115871561" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CAS), &quot;Yunnan, Baoshan, Longyang, Baihualing, beating on vegetation,
<geoCoordinate id="EE1AA1F2620CFFA92E6CB11CFC735ED8" box="[788,916,871,898]" degrees="25.30764" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="25.30764">N25.30764</geoCoordinate>
,
<geoCoordinate id="EE1AA1F2620CFFA92ED8B11CFBFA5EDB" box="[928,1053,871,897]" degrees="98.79367" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="98.79367">E98.79367</geoCoordinate>
,
<quantity id="4CD66AD0620CFFA92951B11CFB665EDB" box="[1065,1153,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="74" pageNumber="75" unit="m" value="1625.0">
<elevation id="00032006620CFFA92951B11CFB665EDB" box="[1065,1153,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="74" pageNumber="75" unit="m" value="1625.0">1625 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA929F6B11CFAE65EDB" box="[1166,1281,871,898]" pageId="74" pageNumber="75" value="2005-06-02">
<collectingDate id="EFD4181D620CFFA929F6B11CFAE65EDB" box="[1166,1281,871,898]" pageId="74" pageNumber="75" value="2005-06-02">2005.VI.2</collectingDate>
</date>
d, Kavanaugh D lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92D97B1F0FB445EFF" ID-GBIF-Occurrence="4115871392" box="[239,1187,907,933]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Yunnan, Tengchong, Dahaoping,
<quantity id="4CD66AD0620CFFA92E79B1F0FCB05EFF" box="[769,855,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="74" pageNumber="75" unit="m" value="1980.0">
<elevation id="00032006620CFFA92E79B1F0FCB05EFF" box="[769,855,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="74" pageNumber="75" unit="m" value="1980.0">1980 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92E1AB1F0FC315EFF" box="[866,982,907,933]" pageId="74" pageNumber="75" value="2002-05-04">
<collectingDate id="EFD4181D620CFFA92E1AB1F0FC315EFF" box="[866,982,907,933]" pageId="74" pageNumber="75" value="2002-05-04">2002.V.04</collectingDate>
</date>
, Song Jinxin lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA929D1B1F0FC6B5E93" ID-GBIF-Occurrence="4115871413" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CAS), &quot;Yunnan, Houqiao, Guyong, on vegetation,
<date id="FF90E1F5620CFFA92F66B1D5FD765E93" box="[542,657,942,969]" pageId="74" pageNumber="75" value="2006-05-27">
<collectingDate id="EFD4181D620CFFA92F66B1D5FD765E93" box="[542,657,942,969]" pageId="74" pageNumber="75" value="2006-05-27">2006.5.27</collectingDate>
</date>
, Liang Hongbin lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92EEEB1D4FD835EB7" ID-GBIF-Occurrence="4115871400" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Yunnan, Houqiao, Yangjiatian, beating,
<date id="FF90E1F5620CFFA92D81B1A9FE8C5EB7" box="[249,363,978,1005]" pageId="74" pageNumber="75" value="2006-05-28">
<collectingDate id="EFD4181D620CFFA92D81B1A9FE8C5EB7" box="[249,363,978,1005]" pageId="74" pageNumber="75" value="2006-05-28">2006.5.28</collectingDate>
</date>
, Liang Hongbin lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92F14B1A8FA615EB7" ID-GBIF-Occurrence="4115871404" box="[620,1414,979,1005]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Yunnan, Tengchong, Qushi,
<date id="FF90E1F5620CFFA9292CB1A8FB325EB7" box="[1108,1237,979,1005]" pageId="74" pageNumber="75" value="2006-08-14">
<collectingDate id="EFD4181D620CFFA9292CB1A8FB325EB7" box="[1108,1237,979,1005]" pageId="74" pageNumber="75" value="2006-08-14">2006.08.14</collectingDate>
</date>
, Liu Biao lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA928F6B1A8FB31594B" ID-GBIF-Occurrence="4115871448" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<specimenCount id="9D280CBC620CFFA928F6B1A8FF2A594B" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(IZAS), &quot;Yunnan, Jinping, Fenshuiling,
<quantity id="4CD66AD0620CFFA92FE2B18CFD15594B" box="[666,754,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="74" pageNumber="75" unit="m" value="1870.0">
<elevation id="00032006620CFFA92FE2B18CFD15594B" box="[666,754,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="74" pageNumber="75" unit="m" value="1870.0">1870 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92F87B18CFC93594B" box="[767,884,1015,1041]" pageId="74" pageNumber="75" value="2009-05-27">
<collectingDate id="EFD4181D620CFFA92F87B18CFC93594B" box="[767,884,1015,1041]" pageId="74" pageNumber="75" value="2009-05-27">2009.V.27</collectingDate>
</date>
, Bi Wenxuan lgt.&quot; &lt;
<figureCitation id="1315DBB0620CFFA92924B18CFB26594B" box="[1116,1217,1015,1041]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="74" pageNumber="75">Fig. 43B</figureCitation>
&gt;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA929A5B18CFCE1596F" ID-GBIF-Occurrence="4115871334" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<specimenCount id="9D280CBC620CFFA929A5B18CFAD8594B" box="[1245,1343,1015,1041]" pageId="74" pageNumber="75" type="female">1 female</specimenCount>
(CAU), &quot;Xizang, Zayu,
<quantity id="4CD66AD0620CFFA92C37B660FE4E596F" box="[335,425,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="74" pageNumber="75" unit="m" value="1700.0">
<elevation id="00032006620CFFA92C37B660FE4E596F" box="[335,425,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="74" pageNumber="75" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92CC0B660FDD1596F" box="[440,566,1051,1077]" pageId="74" pageNumber="75" value="1978-06-29">
<collectingDate id="EFD4181D620CFFA92CC0B660FDD1596F" box="[440,566,1051,1077]" pageId="74" pageNumber="75" value="1978-06-29">1978-vi-29</collectingDate>
</date>
, Li Fasheng lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92E68B660FEBE5903" ID-GBIF-Occurrence="4115871411" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
<specimenCount id="9D280CBC620CFFA92E68B660FC90596F" box="[784,887,1051,1077]" pageId="74" pageNumber="75" type="female">1 female</specimenCount>
, (CAU), &quot;Xizang, Yigong,
<quantity id="4CD66AD0620CFFA929CBB660FAEA596F" box="[1203,1293,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="74" pageNumber="75" unit="m" value="2300.0">
<elevation id="00032006620CFFA929CBB660FAEA596F" box="[1203,1293,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="74" pageNumber="75" unit="m" value="2300.0">2300 m</elevation>
</quantity>
, Li Fasheng lgt.,
<date id="FF90E1F5620CFFA92DB2B644FEAA5903" box="[202,333,1087,1113]" pageId="74" pageNumber="75" value="1978-06-16">
<collectingDate id="EFD4181D620CFFA92DB2B644FEAA5903" box="[202,333,1087,1113]" pageId="74" pageNumber="75" value="1978-06-16">1978-vi-16</collectingDate>
</date>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92C19B644FDEC5927" ID-GBIF-Occurrence="4115871432" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Xizang, Mêdog, Gedang Xiang, Xingkai vill,
<quantity id="4CD66AD0620CFFA9297EB645FBB95903" box="[1030,1118,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.025" pageId="74" pageNumber="75" unit="m" value="2025.0">
<elevation id="00032006620CFFA9297EB645FBB95903" box="[1030,1118,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.025" pageId="74" pageNumber="75" unit="m" value="2025.0">2025 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92912B645FB1A5903" box="[1130,1277,1086,1113]" pageId="74" pageNumber="75" value="2019-08-15">
<collectingDate id="EFD4181D620CFFA92912B645FB1A5903" box="[1130,1277,1086,1113]" pageId="74" pageNumber="75" value="2019-08-15">2019.VIII.15</collectingDate>
</date>
N,
<geoCoordinate id="EE1AA1F2620CFFA92867B644FA7B5903" box="[1311,1436,1087,1113]" degrees="29.46414" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="29.46414">N29.46414</geoCoordinate>
<geoCoordinate id="EE1AA1F2620CFFA92DEFB619FEF35927" box="[151,276,1122,1149]" degrees="95.73563" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="95.73563">E95.73563</geoCoordinate>
, Ma Z, Zhou R lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92F6DB618FD4E59FB" ID-GBIF-Occurrence="4115871483" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (IZAS), &quot;Xizang,
<location id="8EF191EE620CFFA92E67B618FC535927" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92E67B618FC535927" box="[799,948,1123,1149]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Bomi county" pageId="74" pageNumber="75" stateProvince="Hubei">Bomi county</location>
, Tongmai bridge,
<quantity id="4CD66AD0620CFFA929FFB619FB075927" box="[1159,1248,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.025" pageId="74" pageNumber="75" unit="m" value="2025.0">
<elevation id="00032006620CFFA929FFB619FB075927" box="[1159,1248,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.025" pageId="74" pageNumber="75" unit="m" value="2025.0">2025 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92997B618FA665927" box="[1263,1409,1123,1149]" pageId="74" pageNumber="75" value="2019-08-19">
<collectingDate id="EFD4181D620CFFA92997B618FA665927" box="[1263,1409,1123,1149]" pageId="74" pageNumber="75" value="2019-08-19">2019.VIII.19</collectingDate>
</date>
N,
<geoCoordinate id="EE1AA1F2620CFFA92DEFB6FCFEF359FB" box="[151,276,1159,1185]" degrees="30.09632" direction="north" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="30.09632">N30.09632</geoCoordinate>
<geoCoordinate id="EE1AA1F2620CFFA92C65B6FDFE7D59FB" box="[285,410,1158,1185]" degrees="95.06610" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="95.0661">E95.06610</geoCoordinate>
, Liang HB, Xu Y. lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92FCAB6FCFEF459B3" ID-GBIF-Occurrence="4115871459" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (MTMB), &quot;Taiwan,
<location id="8EF191EE620CFFA92EA9B6FCFB9E59FB" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92EA9B6FCFB9E59FB" box="[977,1145,1159,1185]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Nantou county" pageId="74" pageNumber="75" stateProvince="Hubei">Nantou county</location>
, Kao-Leng Dyi,
<locationDeviation id="9A23DE17620CFFA92842B6FCFEE2599F" location="Wushe" pageId="74" pageNumber="75">
<quantity id="4CD66AD0620CFFA92842B6FCFA9C59FB" box="[1338,1403,1159,1185]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="74" pageNumber="75" unit="km" value="18.0">18km</quantity>
W of Wushe
</locationDeviation>
,
<geoCoordinate id="EE1AA1F2620CFFA92C68B6D1FE76599F" box="[272,401,1194,1221]" degrees="24" direction="north" minutes="4.561" orientation="latitude" pageId="74" pageNumber="75" precision="1" value="24.076017">N24°4.561'</geoCoordinate>
<geoCoordinate id="EE1AA1F2620CFFA92CEFB6D0FDC2599F" box="[407,549,1195,1221]" degrees="121" direction="east" minutes="8.046" orientation="longitude" pageId="74" pageNumber="75" precision="1" value="121.1341">E121°8.046'</geoCoordinate>
,
<quantity id="4CD66AD0620CFFA92F48B6D1FD60599F" box="[560,647,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="74" pageNumber="75" unit="m" value="1945.0">
<elevation id="00032006620CFFA92F48B6D1FD60599F" box="[560,647,1194,1221]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="74" pageNumber="75" unit="m" value="1945.0">1945 m</elevation>
</quantity>
, swept from vegetation,
<date id="FF90E1F5620CFFA92EE5B6D0FBA6599F" box="[925,1089,1195,1221]" pageId="74" pageNumber="75" value="2002-04-18" valueMax="2002-04-19" valueMin="2002-04-18">
<collectingDate id="EFD4181D620CFFA92EE5B6D0FBA6599F" box="[925,1089,1195,1221]" pageId="74" pageNumber="75" value="2002-04-18" valueMax="2002-04-19" valueMin="2002-04-18">18-19.IV.2002</collectingDate>
</date>
, leg.
<collectorName id="26DBA2E3620CFFA92905B6D0FB1E599F" box="[1149,1273,1195,1221]" pageId="74" pageNumber="75">D. Anstine</collectorName>
,
<collectorName id="26DBA2E3620CFFA9287CB6D0FA98599F" box="[1284,1407,1195,1221]" pageId="74" pageNumber="75">Gy. Fabian</collectorName>
&amp;
<collectorName id="26DBA2E3620CFFA92DEFB6B4FEE059B3" box="[151,263,1231,1257]" pageId="74" pageNumber="75">O. Merkl</collectorName>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92C64B6B5FB5459B3" ID-GBIF-Occurrence="4115871470" box="[284,1203,1230,1257]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
5 ex (CCCC), &quot;Taiwan, Hualien, Bilyu scarce tree,
<date id="FF90E1F5620CFFA92E13B6B5FC3659B3" box="[875,977,1230,1257]" pageId="74" pageNumber="75" value="1995-05-02">
<collectingDate id="EFD4181D620CFFA92E13B6B5FC3659B3" box="[875,977,1230,1257]" pageId="74" pageNumber="75" value="1995-05-02">1995.V.2</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB0620CFFA92E8CB6B4FBBA59B3" box="[1012,1117,1231,1257]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="74" pageNumber="75">Figs 11G</figureCitation>
,
<figureCitation id="1315DBB0620CFFA92915B6B4FB7859B3" box="[1133,1183,1231,1257]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="74" pageNumber="75">44B</figureCitation>
&gt;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA929C4B6B4FD275857" ID-GBIF-Occurrence="4115871322" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CCCC), &quot;Taiwan, Hualien, Xiulin, Guanyuan,
<date id="FF90E1F5620CFFA92F63B688FD535857" box="[539,692,1267,1293]" pageId="74" pageNumber="75" value="2006-08-29">
<collectingDate id="EFD4181D620CFFA92F63B688FD535857" box="[539,692,1267,1293]" pageId="74" pageNumber="75" value="2006-08-29">2006.VIII.29</collectingDate>
</date>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92FB1B688FEE9586B" ID-GBIF-Occurrence="4115871504" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
1 ex (CCCC), &quot;Taiwan,
<location id="8EF191EE620CFFA92E98B688FB7B5857" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92E98B688FB7B5857" box="[992,1180,1267,1293]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Pingtung county" pageId="74" pageNumber="75" stateProvince="Hubei">Pingtung county</location>
, Tai24 Road,
<quantity id="4CD66AD0620CFFA92846B689FA705857" box="[1342,1431,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="74" pageNumber="75" unit="m" value="1500.0">
<elevation id="00032006620CFFA92846B689FA705857" box="[1342,1431,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="74" pageNumber="75" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92DEFB76CFF18586B" box="[151,255,1303,1329]" pageId="74" pageNumber="75" value="1988-05-02">
<collectingDate id="EFD4181D620CFFA92DEFB76CFF18586B" box="[151,255,1303,1329]" pageId="74" pageNumber="75" value="1988-05-02">1988.V.2</collectingDate>
</date>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92C6FB76CFA7B586B" ID-GBIF-Occurrence="4115871385" box="[279,1436,1303,1329]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
2 ex (CCCC), &quot;Taiwan,
<location id="8EF191EE620CFFA92F55B76CFD06586B" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92F55B76CFD06586B" box="[557,737,1303,1329]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Hsinchu county" pageId="74" pageNumber="75" stateProvince="Hubei">Hsinchu county</location>
, Wufeng,
<location id="8EF191EE620CFFA92E2EB76CFC36586B" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92E2EB76CFC36586B" box="[854,977,1303,1329]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Talu forest" pageId="74" pageNumber="75" stateProvince="Hubei">Talu forest</location>
road,
<date id="FF90E1F5620CFFA92964B76CFB77586B" box="[1052,1168,1303,1329]" pageId="74" pageNumber="75" value="1997-05-31">
<collectingDate id="EFD4181D620CFFA92964B76CFB77586B" box="[1052,1168,1303,1329]" pageId="74" pageNumber="75" value="1997-05-31">1997.V.31</collectingDate>
</date>
, Chen Changchin lgt.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA92DEFB740FAF4580F" ID-GBIF-Occurrence="4115871428" box="[151,1299,1339,1365]" collectingDate="1978-06-16" collectingDateMax="2020-06-10" collectingDateMin="1978-06-16" country="China" county="Xuan-en" elevation="631" latitude="29.6998" location="Longtan station" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" pageId="74" pageNumber="75" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei">
3 ex (IZAS), &quot; Taiwan,
<location id="8EF191EE620CFFA92CDEB740FDC0580F" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92CDEB740FDC0580F" box="[422,551,1339,1365]" country="China" county="Xuan-en" latitude="29.6998" longLatPrecision="6" longitude="109.6511" municipality="Shadaogou" name="Ilan county" pageId="74" pageNumber="75" stateProvince="Hubei">Ilan county</location>
, Taiping Shan, Beech Path 38 k,
<date id="FF90E1F5620CFFA92EDCB740FBC1580F" box="[932,1062,1339,1365]" pageId="74" pageNumber="75" value="2020-06-10">
<collectingDate id="EFD4181D620CFFA92EDCB740FBC1580F" box="[932,1062,1339,1365]" pageId="74" pageNumber="75" value="2020-06-10">2020.VI.10</collectingDate>
</date>
, Huang Jialong lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68620CFFA92858B740FF13589B" ID-GBIF-Occurrence="4115871421" collectingDate="2009-06-01" collectingDateMax="2009-06-16" collectingDateMin="2009-06-01" country="Laos" elevation="1425" location="Phou Pane Mt." municipality="Houa Phan prov." pageId="74" pageNumber="75" specimenCount="1">
<emphasis id="B95A1B27620CFFA92858B740FA85580F" bold="true" box="[1312,1378,1339,1365]" pageId="74" pageNumber="75">Laos:</emphasis>
1 ex (NHMB), &quot;Laos-NE,
<collectingMunicipality id="6BF55D4F620CFFA92CF4B724FDAE5823" box="[396,585,1375,1401]" pageId="74" pageNumber="75">Houa Phan prov.</collectingMunicipality>
~20°13N 104°00'E,
<location id="8EF191EE620CFFA92E4DB724FC395823" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92E4DB724FC395823" box="[821,990,1375,1401]" country="Laos" latitude="104.0" municipality="Houa Phan prov." name="Phou Pane Mt." pageId="74" pageNumber="75">Phou Pane Mt.</location>
,
<date id="FF90E1F5620CFFA92E92B724FB655823" box="[1002,1154,1375,1401]" pageId="74" pageNumber="75" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">
<collectingDate id="EFD4181D620CFFA92E92B724FB655823" box="[1002,1154,1375,1401]" pageId="74" pageNumber="75" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">1.-16.vi.2009</collectingDate>
</date>
,
<quantity id="4CD66AD0620CFFA929F7B725FACF5823" box="[1167,1320,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.5" metricValueMin="1.35" pageId="74" pageNumber="75" unit="m" value="1425.0" valueMax="1500.0" valueMin="1350.0">
<elevation id="00032006620CFFA929F7B725FACF5823" box="[1167,1320,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.5" metricValueMin="1.35" pageId="74" pageNumber="75" unit="m" value="1425.0" valueMax="1500.0" valueMin="1350.0">1350-1500 m</elevation>
</quantity>
,
<collectorName id="26DBA2E3620CFFA9284DB724FF3558C7" pageId="74" pageNumber="75">M. Brancucci</collectorName>
leg.&quot;, &quot;NHMB Basel, NMPC Prague, Laos 2009 Expediton,
<collectorName id="26DBA2E3620CFFA92E03B7F8FBF158C7" box="[891,1046,1411,1437]" pageId="74" pageNumber="75">M. Brancucci</collectorName>
,
<collectorName id="26DBA2E3620CFFA92959B7F8FB7458C7" box="[1057,1171,1411,1437]" pageId="74" pageNumber="75">M. Geiser</collectorName>
,
<collectorName id="26DBA2E3620CFFA929E5B7F8FAE558C7" box="[1181,1282,1411,1437]" pageId="74" pageNumber="75">Z. Kraus</collectorName>
,
<collectorName id="26DBA2E3620CFFA92874B7F8FA9D58C7" box="[1292,1402,1411,1437]" pageId="74" pageNumber="75">D. Hauck</collectorName>
,
<collectorName id="26DBA2E3620CFFA928FDB7F8FF0B589B" pageId="74" pageNumber="75">V. Kubari</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68620CFFA92C78B7DCFA61589B" ID-GBIF-Occurrence="4115871340" box="[256,1414,1446,1473]" collectingDate="1983-06-04" collectingDateMax="1985-06-06" collectingDateMin="1983-06-04" collectorName="M. Brancucci" country="Nepal" elevation="700" location="Gorza Dobhan" pageId="74" pageNumber="75" specimenCount="2" specimenCount-male="2" stateProvince="Mechi">
<emphasis id="B95A1B27620CFFA92C78B7DCFEA8589B" bold="true" box="[256,335,1447,1473]" pageId="74" pageNumber="75">Nepal:</emphasis>
<specimenCount id="9D280CBC620CFFA92C20B7DCFE43589B" box="[344,420,1447,1473]" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(NHMB), &quot;
<location id="8EF191EE620CFFA92F57B7DCFD3F589B" LSID="urn:lsid:plazi:treatment:03877623620FFFA82DEFB2ECFB7B5A13:8EF191EE620CFFA92F57B7DCFD3F589B" box="[559,728,1447,1473]" country="Nepal" name="Gorza Dobhan" pageId="74" pageNumber="75" stateProvince="Mechi">Gorza Dobhan</location>
, 2100-
<quantity id="4CD66AD0620CFFA92E5DB7DCFC97589B" box="[805,880,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="700.0">
<elevation id="00032006620CFFA92E5DB7DCFC97589B" box="[805,880,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92E05B7DCFC13589B" box="[893,1012,1446,1473]" pageId="74" pageNumber="75" value="1985-06-06">
<collectingDate id="EFD4181D620CFFA92E05B7DCFC13589B" box="[893,1012,1446,1473]" pageId="74" pageNumber="75" value="1985-06-06">6.VI.1985</collectingDate>
</date>
&quot;, &quot;E-Nepal, Mechi,
<collectorName id="26DBA2E3620CFFA929A1B7DCFA9C589B" box="[1241,1403,1447,1473]" pageId="74" pageNumber="75">M. Brancucci</collectorName>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68620CFFA928F6B7DCFABB58BF" ID-GBIF-Occurrence="4115871542" collectingDate="1983-06-04" collectingDateMax="1985-06-06" collectingDateMin="1983-06-04" collectorName="M. Brancucci" country="Nepal" elevation="700" location="Gorza Dobhan" pageId="74" pageNumber="75" specimenCount="2" specimenCount-male="2" stateProvince="Mechi">
<specimenCount id="9D280CBC620CFFA928F6B7DCFF2A58BF" pageId="74" pageNumber="75" type="male">1 male</specimenCount>
(NHMB), &quot;Mure Num, 2000-
<quantity id="4CD66AD0620CFFA92F58B7B1FD9058BF" box="[544,631,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="74" pageNumber="75" unit="m" value="1550.0">
<elevation id="00032006620CFFA92F58B7B1FD9058BF" box="[544,631,1482,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="74" pageNumber="75" unit="m" value="1550.0">1550 m</elevation>
</quantity>
,
<date id="FF90E1F5620CFFA92FFAB7B0FCF658BF" box="[642,785,1483,1509]" pageId="74" pageNumber="75" value="1983-06-04" valueMax="1983-06-07" valueMin="1983-06-04">
<collectingDate id="EFD4181D620CFFA92FFAB7B0FCF658BF" box="[642,785,1483,1509]" pageId="74" pageNumber="75" value="1983-06-04" valueMax="1983-06-07" valueMin="1983-06-04">4-7.VI.1983</collectingDate>
</date>
&quot;, &quot;E-Nepal, Arun v.,
<collectorName id="26DBA2E3620CFFA92E84B7B0FB7F58BF" box="[1020,1176,1483,1509]" pageId="74" pageNumber="75">M. Brancucci</collectorName>
&quot; &lt;
<figureCitation id="1315DBB0620CFFA929CEB7B0FAF858BF" box="[1206,1311,1483,1509]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="74" pageNumber="75">Figs 43C</figureCitation>
,
<figureCitation id="1315DBB0620CFFA92852B7B0FABB58BF" box="[1322,1372,1483,1509]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="74" pageNumber="75">44C</figureCitation>
</materialsCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620CFFA92DBFB795FD445B2F" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="8B91C735620CFFA92DBFB795FD445B2F" blockId="74.[151,1437,150,2013]" pageId="74" pageNumber="75">
<emphasis id="B95A1B27620CFFA92DBFB795FE815B52" bold="true" box="[199,358,1518,1544]" pageId="74" pageNumber="75">Comparisons</emphasis>
. This species is easily distinguished from all other species of subgenus
<taxonomicName id="4C2EBCB6620CFFA929C0B794FAB75B53" authority="Schaum, 1863" box="[1208,1360,1519,1545]" class="Insecta" family="Carabidae" genus="Bothynoptera" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27620CFFA929C0B794FAB75B53" box="[1208,1360,1519,1545]" italics="true" pageId="74" pageNumber="75">Bothynoptera</emphasis>
</taxonomicName>
by its exceptional elytra pattern (with serrate pale band). The male genitalia of this species are very similar to those of
<taxonomicName id="4C2EBCB6620CFFA928F0B468FEC75B0A" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B27620CFFA928F0B468FEC75B0A" italics="true" pageId="74" pageNumber="75">P. monostigma</emphasis>
</taxonomicName>
with the apex of median lobe only weakly bent to the right side, but they have small differences in the shape of the apical lamella and ventral margin.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE620CFFA82DBFB405FA8F5C47" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="description">
<paragraph id="8B91C735620CFFA82DBFB405FA8F5C47" blockId="74.[151,1437,150,2013]" lastBlockId="75.[151,1437,151,285]" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
<emphasis id="B95A1B27620CFFA92DBFB405FEB65BC2" bold="true" box="[199,337,1662,1688]" pageId="74" pageNumber="75">Description</emphasis>
. Body length
<quantity id="4CD66AD0620CFFA92C82B404FD605BC3" box="[506,647,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="7.449999999999999" metricValueMax="8.2" metricValueMin="6.7" pageId="74" pageNumber="75" unit="mm" value="7.449999999999999" valueMax="8.2" valueMin="6.7">6.7–8.2 mm</quantity>
. Head reddish brown to piceous, frons generally with red triangular patch; pronotum with disc dark brown, lateral margins light brown; elytra background reddish brown, lateral intervals darker in some specimens, with a splayed, serrate, pale band behind middle, narrowest on interval 4, a pair of small elongate spots often present on basal fifth of interval 5, elytral pale patches with borders much darker than background in most specimens; legs, antennae, and mouthparts yellowish brown; venter yellowish brown. Labrum weakly dilated to apex, apex very weakly emarginate. Pronotum nearly quadrate, PW/PL = 1.20–1.32, slightly narrower than head, PW/HW = 0.89–0.98; lateral explanations narrow; widest at anterior third, weakly expanded at anterior half, and then slightly sinuate before posterior angles; posterior angles nearly rectangular with rounded apex. Elytra slightly dilated to apex, surface without microsculpture; striae shallowly incised, with fine puncture rows; intervals with very sparse punctures. Males with biseriate adhesive setae almost on full length of mesotarsomeres 2 and 3. Median lobe of aedeagus very stout, in dorsal view apex only slightly bent to right; in lateral view ventral margin well expanded near middle; apical lamella thick and short, LL usually much smaller than LW, apex rounded-truncate. Endophallus with squamate sac not divided. Gonocoxite II of ovipositor nearly quadrate, slightly shorter than basal width, apex slightly concave near middle, with six or seven ensiform setae, nearly evenly arranged.
</paragraph>
</subSubSection>
<caption id="DF5197BD620DFFA82DEFB7B3FC185B5E" ID-DOI="http://doi.org/10.5281/zenodo.7959027" ID-Zenodo-Dep="7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,250,1480,1504]" targetBox="[156,1423,320,1448]" targetPageId="75" targetType="figure">
<paragraph id="8B91C735620DFFA82DEFB7B3FC185B5E" blockId="75.[151,1436,1480,1541]" pageId="75" pageNumber="76">
<emphasis id="B95A1B27620DFFA82DEFB7B3FEFE58BA" bold="true" box="[151,281,1480,1504]" pageId="75" pageNumber="76">FIGURE 44</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB6620DFFA82FFBB7B2FBAC58BB" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[643,1099,1481,1505]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="malaisei" subGenus="Bothynoptera">
<emphasis id="B95A1B27620DFFA82FFBB7B2FD2A58BA" box="[643,717,1481,1504]" italics="true" pageId="75" pageNumber="76">Parena</emphasis>
(
<emphasis id="B95A1B27620DFFA82FA4B7B2FC8158BB" box="[732,870,1481,1505]" italics="true" pageId="75" pageNumber="76">Bothynoptera</emphasis>
)
<emphasis id="B95A1B27620DFFA82E0DB7B2FC2C58BB" box="[885,971,1481,1505]" italics="true" pageId="75" pageNumber="76">malaisei</emphasis>
(Andrewes)
</taxonomicName>
, right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views.
<emphasis id="B95A1B27620DFFA82CA4B797FE095B5E" bold="true" box="[476,494,1516,1540]" pageId="75" pageNumber="76">A</emphasis>
Guangxi.
<emphasis id="B95A1B27620DFFA82F21B796FD8D5B5F" bold="true" box="[601,618,1517,1541]" pageId="75" pageNumber="76">B</emphasis>
Taiwan.
<emphasis id="B95A1B27620DFFA82FBEB797FD3F5B5E" bold="true" box="[710,728,1516,1540]" pageId="75" pageNumber="76">C</emphasis>
Nepal. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE620DFFA82DBFB448FB7B5A13" pageId="75" pageNumber="76" type="distribution">
<paragraph id="8B91C735620DFFA82DBFB448FDEF5B2B" blockId="75.[151,1437,1587,1865]" pageId="75" pageNumber="76">
<emphasis id="B95A1B27620DFFA82DBFB448FEBE5B17" bold="true" box="[199,345,1587,1613]" pageId="75" pageNumber="76">Distribution</emphasis>
(
<figureCitation id="1315DBB0620DFFA82C11B448FE525B17" box="[361,437,1587,1613]" captionStart="MAP 8" captionStartId="65.[152,211,1703,1727]" captionTargetBox="[180,1408,802,1680]" captionTargetId="figure-242@65.[180,1408,802,1680]" captionText="MAP 8. Distributions for species in the Parena (Bothynoptera) tripunctata group." figureDoi="http://doi.org/10.5281/zenodo.7963675" httpUri="https://zenodo.org/record/7963675/files/figure.png" pageId="75" pageNumber="76">Map 8</figureCitation>
, blue).
<collectingCountry id="F33987A5620DFFA82F74B448FDA85B17" box="[524,591,1587,1613]" name="China" pageId="75" pageNumber="76">China</collectingCountry>
(
<collectingRegion id="49EA09D7620DFFA82F26B448FD405B17" box="[606,679,1587,1613]" country="China" name="Hubei" pageId="75" pageNumber="76">Hubei</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA82FCBB448FCE75B17" box="[691,768,1587,1613]" country="China" name="Hunan" pageId="75" pageNumber="76">Hunan</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA82E75B448FC6B5B17" box="[781,908,1587,1613]" country="China" name="Chongqing" pageId="75" pageNumber="76">Chongqing</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA82EE0B448FC125B17" box="[920,1013,1587,1613]" country="China" name="Sichuan" pageId="75" pageNumber="76">Sichuan</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA82979B448FB835B17" box="[1025,1124,1587,1613]" country="China" name="Guangxi" pageId="75" pageNumber="76">Guangxi</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA82909B448FB335B17" box="[1137,1236,1587,1613]" country="China" name="Guizhou" pageId="75" pageNumber="76">Guizhou</collectingRegion>
,
<collectingRegion id="49EA09D7620DFFA829A7B448FADF5B17" box="[1247,1336,1587,1613]" country="China" name="Yunnan" pageId="75" pageNumber="76">Yunnan</collectingRegion>
,
<collectingCountry id="F33987A5620DFFA8283BB448FA7F5B17" box="[1347,1432,1587,1613]" name="Taiwan" pageId="75" pageNumber="76">Taiwan</collectingCountry>
, Xizang),
<collectingCountry id="F33987A5620DFFA82D87B42CFE895B2B" box="[255,366,1623,1649]" name="Myanmar" pageId="75" pageNumber="76">Myanmar</collectingCountry>
,
<collectingCountry id="F33987A5620DFFA82C01B42CFE555B2B" box="[377,434,1623,1649]" name="Laos" pageId="75" pageNumber="76">Laos</collectingCountry>
,
<collectingCountry id="F33987A5620DFFA82CC6B42CFDE25B2B" box="[446,517,1623,1649]" name="Nepal" pageId="75" pageNumber="76">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8B91C735620DFFA82DBFB400FB7B5A13" blockId="75.[151,1437,1587,1865]" pageId="75" pageNumber="76">
<emphasis id="B95A1B27620DFFA82DBFB400FE395BCF" bold="true" box="[199,478,1659,1685]" pageId="75" pageNumber="76">Geographical variation</emphasis>
. We examined
<specimenCount id="9D280CBC620DFFA82FF5B400FD185BCF" box="[653,767,1659,1685]" pageId="75" pageNumber="76" type="male">two males</specimenCount>
from
<collectingCountry id="F33987A5620DFFA82E3EB400FC6E5BCF" box="[838,905,1659,1685]" name="Nepal" pageId="75" pageNumber="76">Nepal</collectingCountry>
representing the westernmost limit of this species (
<figureCitation id="1315DBB0620DFFA82DACB4E4FEDA5BE3" box="[212,317,1695,1721]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="75" pageNumber="76">Fig. 43C</figureCitation>
). They slightly differ from specimens from eastern localities (
<figureCitation id="1315DBB0620DFFA8297DB4E4FB8C5BE3" box="[1029,1131,1695,1721]" captionStart="FIGURE 43" captionStartId="73.[151,250,1749,1773]" captionTargetBox="[157,1434,397,1717]" captionTargetId="figure-103@73.[151,1436,396,1726]" captionTargetPageId="73" captionText="FIGURE 43. Habitus of Parena (Bothynoptera) malaisei (Andrewes) and Parena (Bothynoptera) quadrisignata Mateu. A Holotype of Parena malaisei (Andrewes). B P. malaisei (Andrewes), male from Yunnan. C P. malaisei (Andrewes), male from Nepal. D Holotype of Parena quadrisignata Mateu. E Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. F P. quadrisignata Mateu, male from Yunnan. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959025" httpUri="https://zenodo.org/record/7959025/files/figure.png" pageId="75" pageNumber="76">Fig. 43B</figureCitation>
) in having the elytral pale band strongly extended to elytra base on interval 1 and the pale spots on the interval 5 rounded, not elongate. The male genitalia of this species also vary slightly among specimens from different localities, mainly in the shape of the apical lamella. Among specimens from eastern localities (e.g. Taiwan) to western localities (e.g.
<collectingCountry id="F33987A5620DFFA829ADB570FAFB5A7F" box="[1237,1308,1803,1829]" name="Nepal" pageId="75" pageNumber="76">Nepal</collectingCountry>
), the apical lamella varies from shorter and wider (
<figureCitation id="1315DBB0620DFFA82F33B554FD575A13" box="[587,688,1839,1865]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="75" pageNumber="76">Fig. 44B</figureCitation>
) to slightly longer and narrower (
<figureCitation id="1315DBB0620DFFA82951B554FB775A13" box="[1065,1168,1839,1865]" captionStart="FIGURE 44" captionStartId="75.[151,250,1480,1504]" captionTargetBox="[156,1423,320,1448]" captionTargetId="figure-93@75.[151,1436,309,1457]" captionTargetPageId="75" captionText="FIGURE 44. Median lobe of male genitalia for Parena (Bothynoptera) malaisei (Andrewes), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Guangxi. B Taiwan. C Nepal. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959027" httpUri="https://zenodo.org/record/7959027/files/figure.png" pageId="75" pageNumber="76">Fig. 44C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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