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<mods:title id="DCE93E0912186ADD66091E1AED6E9EBA">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
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<taxonomicName id="4C2AED86FFA28D7BA3DAF94457229607" ID-CoL="3T8VK" authority="(Macleay, 1888)" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[151,587,1778,1805]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pygmaeus">
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<emphasis id="B95E4A17FFA28D7BA3DAF94454179606" bold="true" box="[151,382,1778,1804]" italics="true" pageId="9" pageNumber="50">Lepanus pygmaeus</emphasis>
(
<bibRefCitation id="EFBBEBF4FFA28D7BA2C3F945572A9607" author="Macleay, W." box="[398,579,1779,1805]" pageId="9" pageNumber="50" pagination="897 - 904" refId="ref29625" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. II. The lamellicornes, Proceedings of the Linnean Society of New South Wales, 3, 897 - 904." type="journal article" year="1888">Macleay, 1888</bibRefCitation>
)
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<paragraph id="8B959605FFA28D7BA3DAF8A3549A9624" blockId="9.[151,587,1778,1838]" box="[151,499,1813,1838]" pageId="9" pageNumber="50">
(
<figureCitation id="13118A80FFA28D7BA3D2F8A355859624" box="[159,236,1813,1838]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="9" pageNumber="50">Figs. 4</figureCitation>
GH, 7A, 9D, 12C, 13)
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<taxonomicName id="4C2AED86FFA28D7BA3DAF8EB571B9679" ID-CoL="559X6" authority="Macleay, 1888: 898" authorityName="Macleay" authorityPageNumber="898" authorityYear="1888" box="[151,626,1885,1907]" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pygmaeum">
<emphasis id="B95E4A17FFA28D7BA3DAF8EB54C99679" box="[151,416,1885,1907]" italics="true" pageId="9" pageNumber="50">Temnoplectron pygmaeum</emphasis>
<treatmentCitation id="0A8BB014FFA28D7BA2EAF8EB571B9679" author="Macleay, W." box="[423,626,1885,1907]" page="898" pageId="9" pageNumber="50" year="1888">
<bibRefCitation id="EFBBEBF4FFA28D7BA2EAF8EB57569679" author="Macleay, W." box="[423,575,1885,1907]" pageId="9" pageNumber="50" pagination="897 - 904" refId="ref29625" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. II. The lamellicornes, Proceedings of the Linnean Society of New South Wales, 3, 897 - 904." type="journal article" year="1888">Macleay, 1888</bibRefCitation>
: 898
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;
<treatmentCitation id="0A8BB014FFA28D7BA131F8EB563C9679" author="Blackburn, T." box="[636,853,1885,1907]" page="212" pageId="9" pageNumber="50" year="1900">
<bibRefCitation id="EFBBEBF4FFA28D7BA131F8EB564A9679" author="Blackburn, T." box="[636,803,1885,1907]" pageId="9" pageNumber="50" pagination="206 - 233" refId="ref28969" refString="Blackburn, T. (1900) On some new genera and species of Australian Coleoptera. Proceedings of the Royal Society of Uictoria, 12, 206 - 233." type="journal article" year="1900">Blackburn 1900</bibRefCitation>
: 212
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;
<treatmentCitation id="0A8BB014FFA28D7BA02DF8EB56929679" author="Gillet, J. J. E." box="[864,1019,1885,1907]" page="40" pageId="9" pageNumber="50" year="1911">
<bibRefCitation id="EFBBEBF4FFA28D7BA02DF8EB56BC9679" author="Gillet, J. J. E." box="[864,981,1885,1907]" pageId="9" pageNumber="50" pagination="1 - 100" refId="ref29144" refString="Gillet, J. J. E. (1911) Scarabaeidae. Coprinae I. Coleopterorum Catalogus, 19, 1 - 100." type="journal article" year="1911">Gillet 1911</bibRefCitation>
: 40
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.
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</paragraph>
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<taxonomicName id="4C2AED86FFA28D7BA3DAF8CA549E9698" authority=": Lea 1904: 89" authorityName="Lea" authorityPageNumber="89" authorityYear="1904" box="[151,503,1916,1938]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA28D7BA3DAF8CA543E9698" box="[151,343,1916,1938]" italics="true" pageId="9" pageNumber="50">Panelus pygmaeus</emphasis>
:
<treatmentCitation id="0A8BB014FFA28D7BA225F8CB549E9698" author="Lea, A. M." box="[360,503,1916,1938]" page="89" pageId="9" pageNumber="50" year="1904">
<bibRefCitation id="EFBBEBF4FFA28D7BA225F8CB54A79698" author="Lea, A. M." box="[360,462,1916,1938]" pageId="9" pageNumber="50" pagination="60 - 107" refId="ref29459" refString="Lea, A. M. (1904) Descriptions of new species of Australian Coleoptera Part VIII. Proceedings of the Linnean Society of New South Wales, 29, 60 - 107. https: // doi. org / 10.5962 / bhl. part. 20155" type="journal article" year="1904">Lea 1904</bibRefCitation>
: 89
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</taxonomicName>
;
<treatmentCitation id="0A8BB014FFA28D7BA148F8CA57DE9698" author="Wilson, F. E." box="[517,695,1916,1938]" page="39" pageId="9" pageNumber="50" year="1922">
<bibRefCitation id="EFBBEBF4FFA28D7BA148F8CA57E49698" author="Wilson, F. E." box="[517,653,1916,1938]" pageId="9" pageNumber="50" pagination="33 - 41" refId="ref30164" refString="Wilson, F. E. (1922) New Australian Coleoptera with notes on some previously described species. Part I, Proceedings of the Royal Society of Uictoria, 34, 33 - 41." type="journal article" year="1922">Wilson 1922</bibRefCitation>
: 39
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;
<treatmentCitation id="0A8BB014FFA28D7BA188F8CB56089698" author="Lea, A. M." box="[709,865,1916,1938]" page="355" pageId="9" pageNumber="50" year="1923">
<bibRefCitation id="EFBBEBF4FFA28D7BA188F8CB56429698" author="Lea, A. M." box="[709,811,1916,1938]" pageId="9" pageNumber="50" pagination="353 - 396" refId="ref29548" refString="Lea, A. M. (1923) Australian dung beetles of the subfamily Coprides. Records of the South Australian Museum, 2, 353 - 396." type="journal article" year="1923">Lea 1923</bibRefCitation>
: 355
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;
<treatmentCitation id="0A8BB014FFA28D7BA022F8CA516C9698" author="Gillet, J. J. E." box="[879,1029,1916,1938]" page="5" pageId="9" pageNumber="50" year="1925">
<bibRefCitation id="EFBBEBF4FFA28D7BA022F8CA56809698" author="Gillet, J. J. E." box="[879,1001,1916,1938]" pageId="9" pageNumber="50" pagination="1 - 16" refId="ref29169" refString="Gillet, J. J. E. (1925) Results of Dr E. Mjoberg Swedish scientific expeditions to Australia, 1910 - 1913, 40. Scarabaeidae: Geotrupinae et Coprinae, Arkiv for Zoologi, 17, 1 - 16." type="journal article" year="1925">Gillet 1925</bibRefCitation>
: 5
</treatmentCitation>
;
<treatmentCitation id="0A8BB014FFA28D7BA75EF8CA51BD9698" author="Paulian, R." box="[1043,1236,1916,1938]" page="119" pageId="9" pageNumber="50" year="1935">
<bibRefCitation id="EFBBEBF4FFA28D7BA75EF8CA51C99698" author="Paulian, R." box="[1043,1184,1916,1938]" pageId="9" pageNumber="50" pagination="114 - 125" refId="ref29941" refString="Paulian, R. (1935) Essai sur les Canthonides de la region australien II, Bulletin de la Societe entomologique de France, 40, 114 - 125." type="journal article" year="1935">Paulian 1935</bibRefCitation>
: 119
</treatmentCitation>
;
<treatmentCitation id="0A8BB014FFA28D7BA7AFF8CA50F19698" author="Carter, H. J." box="[1250,1432,1916,1938]" page="101" pageId="9" pageNumber="50" year="1936">
<bibRefCitation id="EFBBEBF4FFA28D7BA7AFF8CA50089698" author="Carter, H. J." box="[1250,1377,1916,1938]" pageId="9" pageNumber="50" pagination="99 - 102" refId="ref29053" refString="Carter, H. J. (1936) Australian Coleoptera. Notes and new species. X. Proceedings of the Linnean Society of New South Wales, 61, 99 - 102." type="journal article" year="1936">Carter 1936</bibRefCitation>
: 101
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;
<treatmentCitation id="0A8BB014FFA28D7BA38AF82C541C96BA" author="Paulian, R." box="[199,373,1946,1968]" page="70" pageId="9" pageNumber="50" year="1939">
<bibRefCitation id="EFBBEBF4FFA28D7BA38AF82C542696BA" author="Paulian, R." box="[199,335,1946,1968]" pageId="9" pageNumber="50" pagination="68 - 74" refId="ref29972" refString="Paulian, R. (1939) Quelques novelles especes de lamellicornes coprophages, Bulletin de la Societe entomologique de France, 44, 68 - 74." type="journal article" year="1939">Paulian 1939</bibRefCitation>
: 70
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.
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</paragraph>
<paragraph id="8B959605FFA28D7BA3DAF80F561C96C5" blockId="9.[151,1435,1885,2030]" box="[151,885,1977,1999]" pageId="9" pageNumber="50">
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<taxonomicName id="4C2AED86FFA28D7BA3DAF80F577296C5" ID-CoL="4CC79" authority="Blackburn, 1900: 211" authorityName="Blackburn" authorityPageNumber="211" authorityYear="1900" box="[151,539,1977,1999]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA28D7BA3DAF80F545E96C5" box="[151,311,1977,1999]" italics="true" pageId="9" pageNumber="50">Panelus arthuri</emphasis>
<treatmentCitation id="0A8BB014FFA28D7BA270F80F577296C5" author="Blackburn, T." box="[317,539,1977,1999]" page="211" pageId="9" pageNumber="50" year="1900">
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: 211
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;
<treatmentCitation id="0A8BB014FFA28D7BA168F80C57D196C5" author="Lea, A. M." box="[549,696,1977,1999]" page="389" pageId="9" pageNumber="50" year="1904">
<bibRefCitation id="EFBBEBF4FFA28D7BA168F80C57E196C5" author="Lea, A. M." box="[549,648,1977,1999]" pageId="9" pageNumber="50" pagination="60 - 107" refId="ref29459" refString="Lea, A. M. (1904) Descriptions of new species of Australian Coleoptera Part VIII. Proceedings of the Linnean Society of New South Wales, 29, 60 - 107. https: // doi. org / 10.5962 / bhl. part. 20155" type="journal article" year="1904">Lea 1904</bibRefCitation>
: 389
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(junior synonym).
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<taxonomicName id="4C2AED86FFA28D7BA3DAF86F575196E4" ID-CoL="3T8VK" authority=": Matthews 1974: 112" authorityName=": Matthews" authorityPageNumber="112" authorityYear="1974" box="[151,568,2008,2030]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA28D7BA3DAF86F543196E4" box="[151,344,2009,2030]" italics="true" pageId="9" pageNumber="50">Lepanus pygmaeus</emphasis>
:
<treatmentCitation id="0A8BB014FFA28D7BA228F86E575196E4" author="Matthews, E. G." box="[357,568,2008,2030]" page="112" pageId="9" pageNumber="50" year="1974">
<bibRefCitation id="EFBBEBF4FFA28D7BA228F86E576F96E4" author="Matthews, E. G." box="[357,518,2008,2030]" pageId="9" pageNumber="50" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews 1974</bibRefCitation>
: 112
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<emphasis id="B95E4A17FFA18D78A3DAFF21510491BA" bold="true" box="[151,1133,151,176]" pageId="10" pageNumber="51">
Type material examined.
<taxonomicName id="4C2AED86FFA18D78A29BFF21566491A5" authorityName="Macleay" authorityYear="1888" box="[470,781,151,176]" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeum">
<emphasis id="B95E4A17FFA18D78A29BFF21566491A5" bold="true" box="[470,781,151,176]" italics="true" pageId="10" pageNumber="51">Temnoplectron pygmaeum</emphasis>
</taxonomicName>
<materialsCitation id="3B429C58FFA18D78A057FF21510491BA" ID-GBIF-Occurrence="2012745342" box="[794,1133,151,176]" collectionCode="ANIC" collectorName="N. W. Austr" location="On" municipality="University of Sydney" pageId="10" pageNumber="51" specimenCount="2" specimenCount-female="2" typeStatus="lectotype">
<typeStatus id="549128A7FFA18D78A057FF2156EC91BA" box="[794,901,151,176]" pageId="10" pageNumber="51" type="lectotype">lectotype</typeStatus>
(here designated):
</materialsCitation>
</emphasis>
<collectorName id="26DFF3D3FFA18D78A7C8FF2E506791BA" box="[1157,1294,152,176]" pageId="10" pageNumber="51">N.W. Austr</collectorName>
” / “
<taxonomicName id="4C2AED86FFA18D78A60FFF21564091DE" authority=", Macl. Kings Sound. N. W. A" authorityName="Macl. Kings Sound. N. W. A" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeus" subGenus="Temnoplectron">Panelus (Temnoplectron) pygmaeus, Macl. Kings Sound. N.W.A</taxonomicName>
” / “
<location id="8EF5C0DEFFA18D78A010FF0A56E891DE" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA18D78A010FF0A56E891DE" box="[861,897,188,212]" municipality="University of Sydney" name="On" pageId="10" pageNumber="51">On</location>
permanent loan from MACLEAY MUSEUM
<collectingMunicipality id="6BF10C7FFFA18D78A3DAFF6954F991F2" box="[151,400,223,248]" pageId="10" pageNumber="51">University of Sydney</collectingMunicipality>
” / “
<typeStatus id="549128A7FFA18D78A28FFF56574891F2" box="[450,545,224,248]" pageId="10" pageNumber="51" type="syntype">Syntype</typeStatus>
”/ “ANIC Database No. 25-057970” / “
<typeStatus id="549128A7FFA18D78A0A3FF56510891F2" box="[1006,1121,224,248]" pageId="10" pageNumber="51" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C2AED86FFA18D78A721FF69559E9017" authority="Macleay" authorityName="Macleay" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeum">Temnoplectron pygmaeum Macleay</taxonomicName>
<specimenCount id="9D2C5D8CFFA18D78A24FFEB3547E9016" box="[258,279,261,284]" pageId="10" pageNumber="51" type="female"></specimenCount>
top designated by Gunter &amp; Weir 2019” (25-057970 in ANIC);
<specimenCount id="9D2C5D8CFFA18D78A741FEB351489016" box="[1036,1057,261,284]" pageId="10" pageNumber="51" type="female"></specimenCount>
<materialsCitation id="3B429C58FFA18D78A761FEB2558190A6" ID-GBIF-Occurrence="2012745347" collectionCode="ANIC" collectorName="N. W. Austr" location="On" municipality="University of Sydney" pageId="10" pageNumber="51" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="B95E4A17FFA18D78A761FEB251B19017" bold="true" box="[1068,1240,260,285]" pageId="10" pageNumber="51">
<typeStatus id="549128A7FFA18D78A761FEB251BD9017" box="[1068,1236,260,285]" pageId="10" pageNumber="51" type="paralectotype">paralectotype</typeStatus>
:
</emphasis>
(on same pin as
<typeStatus id="549128A7FFA18D78A3DAFE915469904A" box="[151,256,295,320]" pageId="10" pageNumber="51" type="lectotype">lectotype</typeStatus>
) “
<collectorName id="26DFF3D3FFA18D78A255FE9E54F2904A" box="[280,411,296,320]" pageId="10" pageNumber="51">N.W. Austr</collectorName>
” / “
<taxonomicName id="4C2AED86FFA18D78A289FE9151F2904A" authority=", Macl. Kings Sound. N. W. A" authorityName="Macl. Kings Sound. N. W. A" box="[452,1179,295,320]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeus" subGenus="Temnoplectron">Panelus (Temnoplectron) pygmaeus, Macl. Kings Sound. N.W.A</taxonomicName>
” / “
<location id="8EF5C0DEFFA18D78A788FE9E5180904A" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA18D78A788FE9E5180904A" box="[1221,1257,296,320]" municipality="University of Sydney" name="On" pageId="10" pageNumber="51">On</location>
permanent loan from MACLEAY MUSEUM
<collectingMunicipality id="6BF10C7FFFA18D78A140FEFA5670906F" box="[525,793,332,357]" pageId="10" pageNumber="51">University of Sydney</collectingMunicipality>
” / “
<typeStatus id="549128A7FFA18D78A013FEFA56D7906E" box="[862,958,332,356]" pageId="10" pageNumber="51" type="syntype">Syntype</typeStatus>
” / “
<collectionCode id="ED3B0EC0FFA18D78A74EFEFA5123906E" box="[1027,1098,332,356]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="51">ANIC</collectionCode>
Database No. 25-057970” “
<typeStatus id="549128A7FFA18D78A3ECFED954559082" box="[161,316,367,392]" pageId="10" pageNumber="51" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C2AED86FFA18D78A208FED957B59082" authority="Macleay" authorityName="Macleay" authorityYear="1888" box="[325,732,367,392]" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeum">Temnoplectron pygmaeum Macleay</taxonomicName>
<specimenCount id="9D2C5D8CFFA18D78A1ABFEC65792908D" box="[742,763,368,391]" pageId="10" pageNumber="51" type="female"></specimenCount>
bottom designated by Gunter &amp;Weir 2019” (25-057970 in
<collectionCode id="ED3B0EC0FFA18D78A3DAFE22558890A6" box="[151,225,404,428]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="51">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF55C68DFFA18D78A3DAF95B560F96E4" httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="10" pageNumber="51" startId="10.[151,250,1773,1795]" targetBox="[153,1436,509,1747]" targetPageId="10">
<paragraph id="8B959605FFA18D78A3DAF95B560F96E4" blockId="10.[151,1436,1773,2030]" pageId="10" pageNumber="51">
<emphasis id="B95E4A17FFA18D78A3DAF95B54799608" bold="true" box="[151,272,1773,1795]" pageId="10" pageNumber="51">FIGURE 4</emphasis>
. Some diagnostic characters of the
<taxonomicName id="4C2AED86FFA18D78A1C6F95856399609" authorityName=": Matthews" authorityYear="1974" box="[651,848,1774,1795]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA18D78A1C6F95856399609" box="[651,848,1774,1795]" italics="true" pageId="10" pageNumber="51">Lepanus pygmaeus</emphasis>
</taxonomicName>
species group.
<emphasis id="B95E4A17FFA18D78A0BAF95B51609608" bold="true" box="[1015,1033,1773,1794]" pageId="10" pageNumber="51">A</emphasis>
, Head of
<taxonomicName id="4C2AED86FFA18D78A73AF958505E9609" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1143,1335,1773,1795]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFA18D78A73AF958505E9609" box="[1143,1335,1773,1795]" italics="true" pageId="10" pageNumber="51">Lepanus guthrieae</emphasis>
</taxonomicName>
featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow);
<emphasis id="B95E4A17FFA18D78A7D1F8B951C4962E" bold="true" box="[1180,1197,1807,1828]" pageId="10" pageNumber="51">B</emphasis>
, head of
<taxonomicName id="4C2AED86FFA18D78A641F8B950F5962E" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1292,1436,1806,1828]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="lentil">
<emphasis id="B95E4A17FFA18D78A641F8B950F5962E" box="[1292,1436,1806,1828]" italics="true" pageId="10" pageNumber="51">Lepanus lentil</emphasis>
</taxonomicName>
featuring angulate and complete basal margin (white arrow);
<emphasis id="B95E4A17FFA18D78A054F8995642964F" bold="true" box="[793,811,1839,1861]" pageId="10" pageNumber="51">C</emphasis>
, pterothorax of
<taxonomicName id="4C2AED86FFA18D78A09AF88751A3964F" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[983,1226,1839,1862]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="andersonorum">
<emphasis id="B95E4A17FFA18D78A09AF88751A3964F" box="[983,1226,1839,1862]" italics="true" pageId="10" pageNumber="51">Lepanus andersonorum</emphasis>
</taxonomicName>
showing fine, short carina extending backwards from the meso-metaventral suture (white arrow);
<emphasis id="B95E4A17FFA18D78A0EAF8E756D0966C" bold="true" box="[935,953,1873,1894]" pageId="10" pageNumber="51">D</emphasis>
, posterior aspect of metatibia of
<taxonomicName id="4C2AED86FFA18D78A649F8E450FD966D" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1284,1428,1873,1895]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FFA18D78A649F8E450FD966D" box="[1284,1428,1873,1895]" italics="true" pageId="10" pageNumber="51">Lepanus pecki</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A3DAF8C555C19682" bold="true" box="[151,168,1907,1928]" pageId="10" pageNumber="51">E</emphasis>
, anterior aspect of metatibia of
<taxonomicName id="4C2AED86FFA18D78A2BBF8C557E09682" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[502,649,1906,1928]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FFA18D78A2BBF8C557E09682" box="[502,649,1906,1928]" italics="true" pageId="10" pageNumber="51">Lepanus pecki</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A1D4F8C557CF9682" bold="true" box="[665,678,1907,1928]" pageId="10" pageNumber="51">F</emphasis>
, anterior aspect of metatibia of
<taxonomicName id="4C2AED86FFA18D78A0B8F8C551DD9682" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1013,1204,1906,1928]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFA18D78A0B8F8C551DD9682" box="[1013,1204,1906,1928]" italics="true" pageId="10" pageNumber="51">Lepanus guthrieae</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A78EF8C451BD9682" bold="true" box="[1219,1236,1906,1928]" pageId="10" pageNumber="51">G</emphasis>
, female protibia of
<taxonomicName id="4C2AED86FFA18D78A3DAF823543096A0" authorityName=": Matthews" authorityYear="1974" box="[151,345,1941,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA18D78A3DAF823543096A0" box="[151,345,1941,1962]" italics="true" pageId="10" pageNumber="51">Lepanus pygmaeus</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A225F822541596A3" bold="true" box="[360,380,1940,1961]" pageId="10" pageNumber="51">H</emphasis>
, male protibia of
<taxonomicName id="4C2AED86FFA18D78A178F823579196A0" authorityName=": Matthews" authorityYear="1974" box="[565,760,1941,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA18D78A178F823579196A0" box="[565,760,1941,1962]" italics="true" pageId="10" pageNumber="51">Lepanus pygmaeus</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A04AF822567896A3" bold="true" box="[775,785,1940,1961]" pageId="10" pageNumber="51">I</emphasis>
, female protibia of
<taxonomicName id="4C2AED86FFA18D78A093F823511896A0" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[990,1137,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FFA18D78A093F823511896A0" box="[990,1137,1940,1962]" italics="true" pageId="10" pageNumber="51">Lepanus pecki</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A7CDF82251E596A3" bold="true" box="[1152,1164,1940,1961]" pageId="10" pageNumber="51">J</emphasis>
, male protibia of
<taxonomicName id="4C2AED86FFA18D78A60BF82355A596C1" authorityName="Gunter &amp; Weir" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B95E4A17FFA18D78A60BF82355A596C1" italics="true" pageId="10" pageNumber="51">Lepanus pecki</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A393F803559B96C0" bold="true" box="[222,242,1973,1994]" pageId="10" pageNumber="51">K</emphasis>
, male protibia of
<taxonomicName id="4C2AED86FFA18D78A2FBF800571196C1" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[438,632,1973,1995]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="crenidens">
<emphasis id="B95E4A17FFA18D78A2FBF800571196C1" box="[438,632,1973,1995]" italics="true" pageId="10" pageNumber="51">Lepanus crenidens</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A1C7F80357F296C0" bold="true" box="[650,667,1973,1994]" pageId="10" pageNumber="51">L</emphasis>
, male protibia of
<taxonomicName id="4C2AED86FFA18D78A02DF800514E96C1" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[864,1063,1973,1995]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pungalina">
<emphasis id="B95E4A17FFA18D78A02DF800514E96C1" box="[864,1063,1973,1995]" italics="true" pageId="10" pageNumber="51">Lepanus pungalina</emphasis>
</taxonomicName>
;
<emphasis id="B95E4A17FFA18D78A775F803513996C0" bold="true" box="[1080,1104,1973,1994]" pageId="10" pageNumber="51">M</emphasis>
, inner laterodorsal aspect male protibia of
<taxonomicName id="4C2AED86FFA18D78A24AF86F54A396E4" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[263,458,2008,2030]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="pungalina">
<emphasis id="B95E4A17FFA18D78A24AF86F54A396E4" box="[263,458,2008,2030]" italics="true" pageId="10" pageNumber="51">Lepanus pungalina</emphasis>
</taxonomicName>
showing bent apical digit (black arrow).
</paragraph>
</caption>
<paragraph id="8B959605FFA08D79A38AFF2151649034" blockId="11.[151,1437,151,2021]" pageId="11" pageNumber="52">
<emphasis id="B95E4A17FFA08D79A38AFF2157A791BA" bold="true" box="[199,718,151,176]" pageId="11" pageNumber="52">
<taxonomicName id="4C2AED86FFA08D79A38AFF21541791BA" authorityName="Blackburn" authorityYear="1900" box="[199,382,151,176]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA08D79A38AFF21541791BA" bold="true" box="[199,382,151,176]" italics="true" pageId="11" pageNumber="52">Panelus arthuri</emphasis>
</taxonomicName>
<typeStatus id="549128A7FFA08D79A2C8FF21549991BA" box="[389,496,151,176]" pageId="11" pageNumber="52" type="lectotype">lectotype</typeStatus>
(here designated):
</emphasis>
“6612 T., N.W.A.” (6612 and N.W.A handwritten in red, and T. handwritten in black) / “Type. H. T.” (round label with red border, typed) / “Blackburn coll. 1910-236” (typed) / “
<taxonomicName id="4C2AED86FFA08D79A3ECFF68543F91FD" authorityName="Blackburn" authorityYear="1900" box="[161,342,222,247]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="arthuri">Panelus arthuri</taxonomicName>
. Blackb.” (handwritten) / “=
<taxonomicName id="4C2AED86FFA08D79A1E5FF6851CD91FD" authority="Macleay, Det Matthews" authorityName="Macleay, Det Matthews" box="[680,1188,222,248]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="pygmaeus">Panelus pygmaeus Macleay, Det Matthews</taxonomicName>
<date id="FF94B0C5FFA08D79A7FCFF68519C91FD" box="[1201,1269,222,247]" pageId="11" pageNumber="52" value="1968-06">VI-68</date>
” (handwritten possibly
<date id="FF94B0C5FFA08D79A24DFEB454259011" box="[256,332,258,283]" pageId="11" pageNumber="52" value="1968-07">VII-68</date>
but the second “I” appears crossed out) / “NHMUK011525636” / “
<typeStatus id="549128A7FFA08D79A73CFEB5518A9011" box="[1137,1251,259,283]" pageId="11" pageNumber="52" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C2AED86FFA08D79A7A2FEB454659034" authority="Blackburn" authorityName="Blackburn" authorityYear="1900" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="arthuri">Panelus arthuri Blackburn</taxonomicName>
designated by Gunter &amp; Weir 2019” (in BMNH), sex undetermined.
</paragraph>
<paragraph id="8B959605FFA08D79A38AFEFF5166908F" blockId="11.[151,1437,151,2021]" pageId="11" pageNumber="52">
<emphasis id="B95E4A17FFA08D79A38AFEFF576F9068" bold="true" box="[199,518,329,354]" pageId="11" pageNumber="52">Other material examined.</emphasis>
An additional
<specimenCount id="9D2C5D8CFFA08D79A1F0FEFF56019068" box="[701,872,329,354]" pageId="11" pageNumber="52" type="generic">141 specimens</specimenCount>
from 35 localities from
<collectingRegion id="49EE58E7FFA08D79A7DEFEFF500E9068" box="[1171,1383,329,354]" country="Australia" name="Northern Territory" pageId="11" pageNumber="52">Northern Territory</collectingRegion>
and
<collectingRegion id="49EE58E7FFA08D79A3DAFEDA5472908F" box="[151,283,364,389]" country="Australia" name="Queensland" pageId="11" pageNumber="52">Queensland</collectingRegion>
including Groote Eylandt and Torres Strait Islands were examined.
</paragraph>
<paragraph id="8B959605FFA08D7EA38AFE2651B6908F" blockId="11.[151,1437,151,2021]" lastBlockId="12.[151,1437,151,2009]" lastPageId="12" lastPageNumber="53" pageId="11" pageNumber="52">
<materialsCitation id="3B429C58FFA08D79A38AFE2655B990C6" ID-GBIF-Occurrence="2012745337" collectingDate="1975-07-17" collectionCode="QM" collectorName="H. Heatwole" latitude="-9.395278" location="Torres Strait" longLatPrecision="21" longitude="142.69084" municipality="Saibai Island" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Queensland">
<emphasis id="B95E4A17FFA08D79A38AFE26543790A3" bold="true" box="[199,350,400,425]" pageId="11" pageNumber="52">
<collectingRegion id="49EE58E7FFA08D79A38AFE26543090A3" box="[199,345,400,425]" country="Australia" name="Queensland" pageId="11" pageNumber="52">Queensland</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BF10C7FFFA08D79A225FE26549690A3" box="[360,511,400,425]" pageId="11" pageNumber="52">Saibai Island</collectingMunicipality>
,
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<date id="FF94B0C5FFA08D79A0AFFE26510A90A3" box="[994,1123,400,425]" pageId="11" pageNumber="52" value="1975-07-17">
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<collectorName id="26DFF3D3FFA08D79A73EFE27506A90A3" box="[1139,1283,400,425]" pageId="11" pageNumber="52">H. Heatwole</collectorName>
(T
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<collectionCode id="ED3B0EC0FFA08D79A3DAFE0255A090C6" box="[151,201,436,460]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A3A9FE0257B690E5" ID-GBIF-Occurrence="2012745340" collectingDate="1977-07-09" collectingDateMax="1977-07-13" collectingDateMin="1977-07-09" collectionCode="QM" collectorName="G. B. Monteith &amp; D. J. Cook" county="Eet Hill" latitude="-10.16" location="Banks Island" longLatPrecision="21" longitude="142.29973" municipality="Moa" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Queensland">
<collectingCounty id="62F4EE89FFA08D79A3A9FE02542F90C6" box="[228,326,435,460]" pageId="11" pageNumber="52">Eet Hill</collectingCounty>
, vicinity,
<collectingMunicipality id="6BF10C7FFFA08D79A28FFE02549C90C1" box="[450,501,436,459]" pageId="11" pageNumber="52">Moa</collectingMunicipality>
(
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),
<location id="8EF5C0DEFFA08D79A18BFE02563290C6" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A18BFE02563290C6" box="[710,859,435,460]" county="Eet Hill" latitude="-10.16" longLatPrecision="21" longitude="142.29973" municipality="Moa" name="Torres Strait" pageId="11" pageNumber="52" stateProvince="Queensland">Torres Strait</location>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A741FE0551F690C6" box="[1036,1183,435,460]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="142.29973">142°17'59&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A7F4FE05503E90C6" box="[1209,1367,435,460]" pageId="11" pageNumber="52" value="1977-07-09" valueMax="1977-07-13" valueMin="1977-07-09">
<collectingDate id="EFD0492DFFA08D79A7F4FE05503E90C6" box="[1209,1367,435,460]" pageId="11" pageNumber="52" value="1977-07-09" valueMax="1977-07-13" valueMin="1977-07-09">913.vii.1977</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A627FE02559790FA" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A260FE6E54CD90FA" box="[301,420,471,496]" pageId="11" pageNumber="52">D.J. Cook</collectorName>
(T84016T
<quantity id="4CD23BE0FFA08D79A162FE61571C90FA" box="[559,629,471,496]" metricMagnitude="3" metricUnit="m" metricValue="2.1340572" pageId="11" pageNumber="52" unit="in" value="84018.0">84018</quantity>
in
<collectionCode id="ED3B0EC0FFA08D79A1E8FE6157BF90E5" box="[677,726,471,495]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A1BDFE61512D9319" ID-GBIF-Occurrence="2012745325" collectingDate="1977-07-14" collectingDateMax="1977-07-17" collectingDateMin="1977-07-14" collectionCode="QM" collectorName="G. B. Monteith &amp; D. J. Cook" county="St Pauls" latitude="-10.188056" location="Banks Island" longLatPrecision="21" longitude="142.3336" municipality="Moa" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Queensland">
<collectingCounty id="62F4EE89FFA08D79A1BDFE61563A90FA" box="[752,851,471,496]" pageId="11" pageNumber="52">St Pauls</collectingCounty>
,
<collectingMunicipality id="6BF10C7FFFA08D79A028FE6E56F290E5" box="[869,923,472,495]" pageId="11" pageNumber="52">Moa</collectingMunicipality>
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),
<location id="8EF5C0DEFFA08D79A724FE6E519590FA" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A724FE6E519590FA" box="[1129,1276,471,496]" county="St Pauls" latitude="-10.188056" longLatPrecision="21" longitude="142.3336" municipality="Moa" name="Torres Strait" pageId="11" pageNumber="52" stateProvince="Queensland">Torres Strait</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A658FE6150FE90E5" box="[1301,1431,471,495]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-10.188056">10°11'17&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A3DAFE4D54759319" box="[151,284,507,531]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="142.3336">142°20'1&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A262FE4C54B29319" box="[303,475,506,531]" pageId="11" pageNumber="52" value="1977-07-14" valueMax="1977-07-17" valueMin="1977-07-14">
<collectingDate id="EFD0492DFFA08D79A262FE4C54B29319" box="[303,475,506,531]" pageId="11" pageNumber="52" value="1977-07-14" valueMax="1977-07-17" valueMin="1977-07-14">1417.vii.1977</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A2ABFE4D57EF9319" box="[486,646,506,531]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A1E7FE4D56739319" box="[682,794,506,531]" pageId="11" pageNumber="52">D.J. Cook</collectorName>
(T84007T
<quantity id="4CD23BE0FFA08D79A0EDFE4D568E9319" box="[928,999,507,531]" metricMagnitude="3" metricUnit="m" metricValue="2.1339556" pageId="11" pageNumber="52" unit="in" value="84014.0">84014</quantity>
in
<collectionCode id="ED3B0EC0FFA08D79A746FE4D51549319" box="[1035,1085,507,531]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A71DFE4C50F19319" ID-GBIF-Occurrence="2012745353" box="[1104,1432,506,531]" collectorName="Wincheura" location="Lake Boronto" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Queensland">
<location id="8EF5C0DEFFA08D79A71DFE4C51829318" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A71DFE4C51829318" box="[1104,1259,506,531]" name="Lake Boronto" pageId="11" pageNumber="52" stateProvince="Queensland">Lake Boronto</location>
(=
<collectorName id="26DFF3D3FFA08D79A65FFE4C50F99319" box="[1298,1424,506,531]" pageId="11" pageNumber="52">Wincheura</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B429C58FFA08D79A3DAFDAB54E2933C" ID-GBIF-Occurrence="2012745334" box="[151,395,541,567]" collectorName="Cape" location="Bay" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Newcastle upon Tyne">
<collectingRegion id="49EE58E7FFA08D79A3DAFDAB5465933C" box="[151,268,541,566]" country="United Kingdom" name="Newcastle upon Tyne" pageId="11" pageNumber="52">Newcastle</collectingRegion>
<location id="8EF5C0DEFFA08D79A258FDA9542A933D" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A258FDA9542A933D" box="[277,323,543,567]" name="Bay" pageId="11" pageNumber="52" stateProvince="Newcastle upon Tyne">Bay</location>
,
<collectorName id="26DFF3D3FFA08D79A21CFDA854E2933C" box="[337,395,542,566]" pageId="11" pageNumber="52">Cape</collectorName>
</materialsCitation>
<materialsCitation id="3B429C58FFA08D79A2D9FDAB56D89350" ID-GBIF-Occurrence="2012745341" collectingDate="1927-10" collectingDateMax="1974-09-26" collectingDateMin="1927-10" collectionCode="QM" collectorName="G. B. Monteith &amp; Wincheura" elevation="50" latitude="-10.766666" location="Lake Boronto" longLatPrecision="21" longitude="142.55" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<collectingRegion id="49EE58E7FFA08D79A2D9FDAB54A7933C" box="[404,462,541,566]" country="United Kingdom" name="York" pageId="11" pageNumber="52">York</collectingRegion>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A139FDA8566E933C" box="[628,775,542,566]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="142.55">142°33'00&quot;E</geoCoordinate>
],
<quantity id="4CD23BE0FFA08D79A050FDAB5632933C" box="[797,859,541,566]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="52" unit="m" value="50.0">
<elevation id="00077136FFA08D79A050FDAB5632933C" box="[797,859,541,566]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="52" unit="m" value="50.0">50 m</elevation>
</quantity>
,
<date id="FF94B0C5FFA08D79A024FDAB5167933C" box="[873,1038,541,566]" pageId="11" pageNumber="52" value="1974-09-21" valueMax="1974-09-26" valueMin="1974-09-21">
<collectingDate id="EFD0492DFFA08D79A024FDAB5167933C" box="[873,1038,541,566]" pageId="11" pageNumber="52" value="1974-09-21" valueMax="1974-09-26" valueMin="1974-09-21">2126.ix.1974</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A756FDA851D4933C" box="[1051,1213,541,566]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
(25-015584
<date id="FF94B0C5FFA08D79A61EFDA85014933C" box="[1363,1405,542,566]" pageId="11" pageNumber="52" value="1927-10">
<collectingDate id="EFD0492DFFA08D79A61EFDA85014933C" box="[1363,1405,542,566]" pageId="11" pageNumber="52" value="1927-10">
x
<quantity id="4CD23BE0FFA08D79A62CFDA85014933C" box="[1377,1405,542,566]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="11" pageNumber="52" unit="in" value="27.0">27</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A3DAFDF455AD9350" box="[151,196,578,602]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
also as T50139T
<quantity id="4CD23BE0FFA08D79A2ECFDF7548E9350" box="[417,487,577,602]" metricMagnitude="3" metricUnit="m" metricValue="1.274191" pageId="11" pageNumber="52" unit="in" value="50165.0">50165</quantity>
in
<collectionCode id="ED3B0EC0FFA08D79A15EFDF4572C9350" box="[531,581,578,602]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
);
<location id="8EF5C0DEFFA08D79A110FDF757959353" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A110FDF757959353" box="[605,764,577,602]" latitude="-10.766666" longLatPrecision="21" longitude="142.55" name="Lake Boronto" pageId="11" pageNumber="52" stateProvince="York">Lake Boronto</location>
(=
<collectorName id="26DFF3D3FFA08D79A061FDF756C39350" box="[812,938,577,602]" pageId="11" pageNumber="52">Wincheura</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B429C58FFA08D79A08CFDF751D09350" ID-GBIF-Occurrence="2012745329" box="[961,1209,577,602]" collectorName="Cape" location="Bay" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Newcastle upon Tyne">
<collectingRegion id="49EE58E7FFA08D79A08CFDF7515F9350" box="[961,1078,577,602]" country="United Kingdom" name="Newcastle upon Tyne" pageId="11" pageNumber="52">Newcastle</collectingRegion>
<location id="8EF5C0DEFFA08D79A70CFDF451069350" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A70CFDF451069350" box="[1089,1135,578,602]" name="Bay" pageId="11" pageNumber="52" stateProvince="Newcastle upon Tyne">Bay</location>
,
<collectorName id="26DFF3D3FFA08D79A732FDF451D09350" box="[1151,1209,578,602]" pageId="11" pageNumber="52">Cape</collectorName>
</materialsCitation>
<materialsCitation id="3B429C58FFA08D79A78EFDF7545A93CE" ID-GBIF-Occurrence="2012745352" collectingDate="1925-10-02" collectingDateMax="1986-07-08" collectingDateMin="1925-10-02" collectionCode="QM, ANIC" collectorName="G. B. Monteith &amp; T. A. Weir" latitude="-12.75" location="8 km E of Mount Tozer" longLatPrecision="21" longitude="143.28334" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<collectingRegion id="49EE58E7FFA08D79A78EFDF751949350" box="[1219,1277,577,602]" country="United Kingdom" name="York" pageId="11" pageNumber="52">York</collectingRegion>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A3DAFDD354439377" box="[151,298,613,637]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="142.55">142°33'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A272FDD354BF9374" box="[319,470,613,638]" pageId="11" pageNumber="52" value="1974-09-02" valueMax="1974-09-26" valueMin="1974-09-02">
<collectingDate id="EFD0492DFFA08D79A272FDD354BF9374" box="[319,470,613,638]" pageId="11" pageNumber="52" value="1974-09-02" valueMax="1974-09-26" valueMin="1974-09-02">226.ix.1974</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A2A9FDD357EC9374" box="[484,645,613,638]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
(25-015585
<date id="FF94B0C5FFA08D79A051FDD3560A9374" box="[796,867,613,638]" pageId="11" pageNumber="52" value="1925-10-03">
<collectingDate id="EFD0492DFFA08D79A051FDD3560A9374" box="[796,867,613,638]" pageId="11" pageNumber="52" value="1925-10-03">x3, 25</collectingDate>
</date>
-015586
<date id="FF94B0C5FFA08D79A086FDD3517A9374" box="[971,1043,613,638]" pageId="11" pageNumber="52" value="1925-10-02">
<collectingDate id="EFD0492DFFA08D79A086FDD3517A9374" box="[971,1043,613,638]" pageId="11" pageNumber="52" value="1925-10-02">x2, 25</collectingDate>
</date>
-015587
<date id="FF94B0C5FFA08D79A736FDD351AB9374" box="[1147,1218,613,638]" pageId="11" pageNumber="52" value="1925-10-08">
<collectingDate id="EFD0492DFFA08D79A736FDD351AB9374" box="[1147,1218,613,638]" pageId="11" pageNumber="52" value="1925-10-08">x8, 25</collectingDate>
</date>
-015588
<date id="FF94B0C5FFA08D79A667FDD3502E9377" box="[1322,1351,613,637]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A667FDD3502E9377" box="[1322,1351,613,637]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A671FDD3502E9377" box="[1340,1351,613,637]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A622FDD350F59377" box="[1391,1436,613,637]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
also as T50166T50179);
<location id="8EF5C0DEFFA08D79A28FFD3F57BA93AA" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A28FFD3F57BA93AA" box="[450,723,648,673]" latitude="-12.75" longLatPrecision="21" longitude="143.28334" name="8 km E of Mount Tozer" pageId="11" pageNumber="52" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A28FFD3F549493AB" box="[450,509,648,673]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="km" value="8.0">8 km</quantity>
E of Mount Tozer
</location>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A035FD3F516593AB" box="[888,1036,649,673]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="143.28334">143°17'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A76DFD3E51FD93AB" box="[1056,1172,648,673]" pageId="11" pageNumber="52" value="1986-07-08">
<collectingDate id="EFD0492DFFA08D79A76DFD3E51FD93AB" box="[1056,1172,648,673]" pageId="11" pageNumber="52" value="1986-07-08">8.vii.1986</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A7ECFD3F507893AB" box="[1185,1297,648,673]" pageId="11" pageNumber="52">T.A. Weir</collectorName>
(25-057972
<date id="FF94B0C5FFA08D79A3DAFD1A55DA93CE" box="[151,179,684,708]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A3DAFD1A55DA93CE" box="[151,179,684,708]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A3E5FD1A55DA93CE" box="[168,179,684,708]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A3AFFD1A544293CE" box="[226,299,684,708]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A209FD1A563393CE" ID-GBIF-Occurrence="2012745314" box="[324,858,684,709]" location="Cape" municipality="Archer River" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<collectingMunicipality id="6BF10C7FFFA08D79A209FD1A54B593CF" box="[324,476,684,709]" pageId="11" pageNumber="52">Archer River</collectingMunicipality>
Xing,
<locationDeviation id="9A278F27FFA08D79A179FD1A567993CE" box="[564,784,684,709]" location="Coen" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A179FD1A57E893CF" box="[564,641,684,709]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="52" unit="km" value="70.0">70 km</quantity>
N of Coen
</locationDeviation>
,
<location id="8EF5C0DEFFA08D79A06DFD1A563393CE" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A06DFD1A563393CE" box="[800,858,684,708]" municipality="Archer River" name="Cape" pageId="11" pageNumber="52" stateProvince="York">Cape</location>
</materialsCitation>
<materialsCitation id="3B429C58FFA08D79A02AFD1A547E9293" ID-GBIF-Occurrence="2012745321" collectingDate="1925-10-03" collectingDateMax="1984-02-23" collectingDateMin="1925-10-03" collectionCode="QM, CMNC" collectorName="G. B. Monteith &amp; G. B. &amp; S. R. Monteith &amp; L. Masner" county="Peninsula" elevation="2" latitude="-16.482779" location="McIvor River" longLatPrecision="21" longitude="145.46472" municipality="McIvor River Crossing" pageId="11" pageNumber="52" specimenCode="CMNEN-00030239-00030240" specimenCount="1" stateProvince="York">
<collectingRegion id="49EE58E7FFA08D79A02AFD1A56F793CF" box="[871,926,684,709]" country="United Kingdom" name="York" pageId="11" pageNumber="52">York</collectingRegion>
<collectingCounty id="62F4EE89FFA08D79A0E7FD1A517393CF" box="[938,1050,684,709]" pageId="11" pageNumber="52">Peninsula</collectingCounty>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A77FFD1A51DC93CE" box="[1074,1205,684,708]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-13.45">13°27'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A78AFD1A503393CF" box="[1223,1370,684,709]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="142.93333">142°56'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A63FFD1A547093E2" pageId="11" pageNumber="52" value="1975-07-17" valueMax="1975-07-18" valueMin="1975-07-17">
<collectingDate id="EFD0492DFFA08D79A63FFD1A547093E2" pageId="11" pageNumber="52" value="1975-07-17" valueMax="1975-07-18" valueMin="1975-07-17">17 18.vii.1975</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A267FD6654A693E2" box="[298,463,719,744]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
(25-015578
<date id="FF94B0C5FFA08D79A126FD6657E193E2" box="[619,648,720,744]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A126FD6657E193E2" box="[619,648,720,744]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A130FD6657E193E2" box="[637,648,720,744]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A1FAFD66578D93E2" box="[695,740,720,744]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
also as T50109);
<locationDeviation id="9A278F27FFA08D79A089FD66519593E2" box="[964,1276,719,744]" location="Port Stewart" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A089FD66517993E2" box="[964,1040,719,744]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="52" unit="km" value="18.0">18 km</quantity>
W of Port Stewart
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A659FD6650FE93E2" box="[1300,1431,720,744]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-14.116667">14°07'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A3DAFD4554439201" box="[151,298,755,779]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="143.51666">143°31'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A270FD45548B9206" box="[317,482,755,780]" pageId="11" pageNumber="52" value="1976-06-25" valueMax="1976-06-28" valueMin="1976-06-25">
<collectingDate id="EFD0492DFFA08D79A270FD45548B9206" box="[317,482,755,780]" pageId="11" pageNumber="52" value="1976-06-25" valueMax="1976-06-28" valueMin="1976-06-25">2528.vi.1976</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A2A3FD45578D9206" box="[494,740,755,780]" pageId="11" pageNumber="52">G.B. &amp; S.R. Monteith</collectorName>
(25-015576
<date id="FF94B0C5FFA08D79A03BFD4556D29206" box="[886,955,755,780]" pageId="11" pageNumber="52" value="1925-10-05">
<collectingDate id="EFD0492DFFA08D79A03BFD4556D29206" box="[886,955,755,780]" pageId="11" pageNumber="52" value="1925-10-05">x5, 25</collectingDate>
</date>
-015581
<date id="FF94B0C5FFA08D79A76CFD45510E9206" box="[1057,1127,755,780]" pageId="11" pageNumber="52" value="1925-10-03">
<collectingDate id="EFD0492DFFA08D79A76CFD45510E9206" box="[1057,1127,755,780]" pageId="11" pageNumber="52" value="1925-10-03">x3, 25</collectingDate>
</date>
-015582
<date id="FF94B0C5FFA08D79A780FD45507A9206" box="[1229,1299,755,780]" pageId="11" pageNumber="52" value="1925-10-08">
<collectingDate id="EFD0492DFFA08D79A780FD45507A9206" box="[1229,1299,755,780]" pageId="11" pageNumber="52" value="1925-10-08">x8, 25</collectingDate>
</date>
-017239
<date id="FF94B0C5FFA08D79A634FD4555DB9225" pageId="11" pageNumber="52" value="1925-10-03">
<collectingDate id="EFD0492DFFA08D79A634FD4555DB9225" pageId="11" pageNumber="52" value="1925-10-03">x3, 25</collectingDate>
</date>
-017240
<date id="FF94B0C5FFA08D79A25AFCA1545A9225" box="[279,307,791,815]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A25AFCA1545A9225" box="[279,307,791,815]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A268FCA1545A9225" box="[293,307,791,815]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="52" unit="in" value="2.0">2</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A21AFCA154ED9225" box="[343,388,791,815]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
also as T50102T50106, T50112T50115, T50117T50124);
<collectingMunicipality id="6BF10C7FFFA08D79A70EFCA150239225" box="[1091,1354,790,815]" pageId="11" pageNumber="52">
<location id="8EF5C0DEFFA08D79A70EFCA151B49225" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A70EFCA151B49225" box="[1091,1245,790,815]" county="Peninsula" latitude="-16.482779" longLatPrecision="21" longitude="145.46472" municipality="McIvor River Crossing" name="McIvor River" pageId="11" pageNumber="52" stateProvince="York">
McIvor
<location id="8EF5C0DEFFA08D79A7D2FCA051B49225" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A7D2FCA051B49225" box="[1183,1245,790,815]" county="Peninsula" latitude="-16.482779" longLatPrecision="21" longitude="145.46472" municipality="McIvor River Crossing" name="River Crossing" pageId="11" pageNumber="52" stateProvince="York">River</location>
</location>
Crossing
</collectingMunicipality>
,
<locationDeviation id="9A278F27FFA08D79A618FCA154229258" location="Cooktown" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A618FCA150F59225" box="[1365,1436,790,815]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="52" unit="km" value="40.0">40 km</quantity>
N of Cooktown
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A213FC8F54889258" box="[350,481,825,850]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-15.15">15°09'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A2A3FC8F57EB9258" box="[494,642,825,850]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.1">145°06'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A1D8FC8F56289258" box="[661,833,825,850]" pageId="11" pageNumber="52" value="1976-07-15" valueMax="1976-07-18" valueMin="1976-07-15">
<collectingDate id="EFD0492DFFA08D79A1D8FC8F56289258" box="[661,833,825,850]" pageId="11" pageNumber="52" value="1976-07-15" valueMax="1976-07-18" valueMin="1976-07-15">1518.vii.1976</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A003FC8C512D9258" box="[846,1092,825,850]" pageId="11" pageNumber="52">G.B. &amp; S.R. Monteith</collectorName>
(25-015580
<date id="FF94B0C5FFA08D79A795FC8C519D9258" box="[1240,1268,826,850]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A795FC8C519D9258" box="[1240,1268,826,850]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A7A4FC8C519D9258" box="[1257,1268,826,850]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A657FC8C502E9258" box="[1306,1351,826,850]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
also as T50111);
<location id="8EF5C0DEFFA08D79A249FCE854F0927C" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A249FCE854F0927C" box="[260,409,861,886]" county="Peninsula" latitude="-16.482779" longLatPrecision="21" longitude="145.46472" municipality="McIvor River Crossing" name="Port Douglas" pageId="11" pageNumber="52" stateProvince="York">Port Douglas</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A2E6FCEB5746927C" box="[427,559,861,886]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.482779">16°28'58&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A177FCEB57A7927C" box="[570,718,861,886]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.46472">145°27'53&quot;E</geoCoordinate>
],
<quantity id="4CD23BE0FFA08D79A1ADFCE85664927F" box="[736,781,862,886]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="52" unit="m" value="2.0">
<elevation id="00077136FFA08D79A1ADFCE85664927F" box="[736,781,862,886]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="52" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<date id="FF94B0C5FFA08D79A055FCEB56DF927C" box="[792,950,861,886]" pageId="11" pageNumber="52" value="1984-02-21" valueMax="1984-02-23" valueMin="1984-02-21">
<collectingDate id="EFD0492DFFA08D79A055FCEB56DF927C" box="[792,950,861,886]" pageId="11" pageNumber="52" value="1984-02-21" valueMax="1984-02-23" valueMin="1984-02-21">2123.ii.1984</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A08CFCE8515D927F" box="[961,1076,862,885]" pageId="11" pageNumber="52">L. Masner</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A70FFCE850F5927C" box="[1090,1436,862,886]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030239-00030240</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A3FAFC3754669293" box="[183,271,897,921]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A26BFC36559F92B7" ID-GBIF-Occurrence="2012745331" collectingDate="1978-10-15" collectionCode="CMNC" collectorName="A. &amp; M. Walford-Huggins" latitude="-16.567223" location="Julatten" longLatPrecision="21" longitude="145.36057" pageId="11" pageNumber="52" specimenCode="CMNEN-00030224" specimenCount="1" stateProvince="York">
<location id="8EF5C0DEFFA08D79A26BFC3654E99293" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A26BFC3654E99293" box="[294,384,896,921]" latitude="-16.567223" longLatPrecision="21" longitude="145.36057" name="Julatten" pageId="11" pageNumber="52" stateProvince="York">Julatten</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A2DBFC3657709293" box="[406,537,896,921]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.567223">16°34'02&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A164FC3657D59293" box="[553,700,896,921]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.36057">145°21'38&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A19EFC36562C9293" box="[723,837,896,921]" pageId="11" pageNumber="52" value="1978-10-15">
<collectingDate id="EFD0492DFFA08D79A19EFC36562C9293" box="[723,837,896,921]" pageId="11" pageNumber="52" value="1978-10-15">15.x.1978</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A018FC3751E39293" box="[853,1162,896,921]" pageId="11" pageNumber="52">A. &amp; M. Walford-Huggins</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A7D6FC3750129293" box="[1179,1403,897,921]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030224</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A3DAFC13558692B7" box="[151,239,933,957]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A24EFC13547D92EA" ID-GBIF-Occurrence="2012745320" collectingDate="1987-02-11" collectingDateMax="1987-02-13" collectingDateMin="1987-02-11" collectionCode="CMNC" collectorName="H. F. &amp; A. T. Howden" latitude="-16.924444" location="Mareeba" longLatPrecision="21" longitude="145.42139" pageId="11" pageNumber="52" specimenCode="CMNEN-00030246" specimenCount="1" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A24EFC13545492B7" box="[259,317,932,957]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="km" value="8.0">8 km</quantity>
N
<location id="8EF5C0DEFFA08D79A22FFC1254AE92B7" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A22FFC1254AE92B7" box="[354,455,932,957]" latitude="-16.924444" longLatPrecision="21" longitude="145.42139" name="Mareeba" pageId="11" pageNumber="52" stateProvince="York">Mareeba</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A296FC12573692B7" box="[475,607,932,957]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.924444">16°55'28&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A121FC12579692B7" box="[620,767,932,957]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.42139">145°25'17&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A05EFC1256D992B7" box="[787,944,932,957]" pageId="11" pageNumber="52" value="1987-02-11" valueMax="1987-02-13" valueMin="1987-02-11">
<collectingDate id="EFD0492DFFA08D79A05EFC1256D992B7" box="[787,944,932,957]" pageId="11" pageNumber="52" value="1987-02-11" valueMax="1987-02-13" valueMin="1987-02-11">1113.ii.1987</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A0F0FC1351C592B7" box="[957,1196,932,957]" pageId="11" pageNumber="52">H.F. &amp; A.T. Howden</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A7F1FC1250F592B7" box="[1212,1436,932,957]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030246</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A3F9FC7E546592EA" box="[180,268,968,992]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A26DFC7E559F9509" ID-GBIF-Occurrence="2012745327" collectingDate="1997-02-06" collectionCode="CMNC" collectorName="H. F. &amp; A. T. Howden" latitude="-16.924444" location="Mareeba" longLatPrecision="21" longitude="145.42139" pageId="11" pageNumber="52" specimenCode="CMNEN-00030247" specimenCount="1" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A26DFC7E543392EA" box="[288,346,967,992]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="km" value="8.0">8 km</quantity>
N
<location id="8EF5C0DEFFA08D79A233FC71548A92EA" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A233FC71548A92EA" box="[382,483,967,992]" latitude="-16.924444" longLatPrecision="21" longitude="145.42139" name="Mareeba" pageId="11" pageNumber="52" stateProvince="York">Mareeba</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A2BAFC71571292EA" box="[503,635,967,992]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.924444">16°55'28&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A1CAFC71567392EA" box="[647,794,967,992]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.42139">145°25'17&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A063FC7156FD92EA" box="[814,916,967,992]" pageId="11" pageNumber="52" value="1997-02-06">
<collectingDate id="EFD0492DFFA08D79A063FC7156FD92EA" box="[814,916,967,992]" pageId="11" pageNumber="52" value="1997-02-06">6.ii.1997</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A0ECFC7F51E692EA" box="[929,1167,967,992]" pageId="11" pageNumber="52">H.F. &amp; A.T. Howden</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A7D3FC7E501792EA" box="[1182,1406,968,992]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030247</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A3DAFC5D55869509" box="[151,239,1003,1027]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A248FC5D559F952D" ID-GBIF-Occurrence="2012745322" collectingDate="1997-02-08" collectionCode="CMNC" collectorName="H. F. &amp; A. T. Howden" latitude="-16.924444" location="Mareeba" longLatPrecision="21" longitude="145.42139" pageId="11" pageNumber="52" specimenCode="CMNEN-00030248" specimenCount="1" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A248FC5D5428950E" box="[261,321,1003,1028]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="km" value="8.0">8 km</quantity>
N
<location id="8EF5C0DEFFA08D79A227FC5D54A6950E" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A227FC5D54A6950E" box="[362,463,1003,1028]" latitude="-16.924444" longLatPrecision="21" longitude="145.42139" name="Mareeba" pageId="11" pageNumber="52" stateProvince="York">Mareeba</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A2A8FC5D5701950E" box="[485,616,1003,1028]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.924444">16°55'28&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A13AFC5D5663950E" box="[631,778,1003,1028]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.42139">145°25'17&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A06DFC5D56EF950E" box="[800,902,1003,1028]" pageId="11" pageNumber="52" value="1997-02-08">
<collectingDate id="EFD0492DFFA08D79A06DFC5D56EF950E" box="[800,902,1003,1028]" pageId="11" pageNumber="52" value="1997-02-08">8.ii.1997</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A0D8FC5A51E3950E" box="[917,1162,1003,1028]" pageId="11" pageNumber="52">H.F. &amp; A.T. Howden</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A7D1FC5D50159509" box="[1180,1404,1003,1027]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030248</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A3DAFBB95586952D" box="[151,239,1039,1063]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A24CFBB9562F9540" ID-GBIF-Occurrence="2012745328" collectingDate="1986-12-28" collectionCode="CMNC" collectorName="H. F. &amp; A. T. Howden" latitude="-16.978611" location="Southedge Tobacco Research Station." longLatPrecision="21" longitude="145.3514" pageId="11" pageNumber="52" specimenCode="CMNEN-00030241-00030242" specimenCount="1" stateProvince="York">
<locationDeviation id="9A278F27FFA08D79A24CFBB9548E952D" box="[257,487,1038,1063]" location="Mareeba" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A24CFBB9542F952D" box="[257,326,1038,1063]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="52" unit="km" value="11.0">11 km</quantity>
NW Mareeba
</locationDeviation>
,
<location id="8EF5C0DEFFA08D79A2BCFBB856F3952D" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A2BCFBB856F3952D" box="[497,922,1038,1063]" latitude="-16.978611" longLatPrecision="21" longitude="145.3514" name="Southedge Tobacco Research Station." pageId="11" pageNumber="52" stateProvince="York">Southedge Tobacco Research Station.</location>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A777FBB851A4952D" box="[1082,1229,1038,1063]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.3514">145°21'05&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A7ADFBB85008952D" box="[1248,1377,1038,1063]" pageId="11" pageNumber="52" value="1986-12-28">
<collectingDate id="EFD0492DFFA08D79A7ADFBB85008952D" box="[1248,1377,1038,1063]" pageId="11" pageNumber="52" value="1986-12-28">28.xii.1986</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A620FBB954389541" pageId="11" pageNumber="52">H.F. &amp; A.T. Howden</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A22EFB8457D59540" box="[355,700,1074,1098]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030241-00030242</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A1AAFB8456569540" box="[743,831,1074,1098]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A018FB84544695D2" ID-GBIF-Occurrence="2012745313" collectingDate="1987-03" collectingDateMax="1989-02-03" collectingDateMin="1987-03" collectionCode="CMNC, CMNEN-, QDPC" collectorName="Howden &amp; H. F. &amp; A. T. Howden &amp; R. I. Storey &amp; De Faveri" county="Southedge Research Station" latitude="-17.15" location="Mareeba" longLatPrecision="21" longitude="145.51666" municipality="Southedge Research Station" pageId="11" pageNumber="52" specimenCode="CMNEN-00030243" specimenCount="1" stateProvince="York">
<collectingCounty id="62F4EE89FFA08D79A018FB8451F19541" box="[853,1176,1074,1099]" pageId="11" pageNumber="52">Southedge Research Station</collectingCounty>
,
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<quantity id="4CD23BE0FFA08D79A7EBFB8451999541" box="[1190,1264,1074,1099]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="52" unit="km" value="12.0">12 km</quantity>
NW Mareeba
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A3D3FBE3544B9564" box="[158,290,1109,1134]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.978611">16°58'43&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A263FBE354AB9564" box="[302,450,1109,1134]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.3514">145°21'05&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A298FBE357529564" box="[469,571,1109,1134]" pageId="11" pageNumber="52" value="1989-02-03">
<collectingDate id="EFD0492DFFA08D79A298FBE357529564" box="[469,571,1109,1134]" pageId="11" pageNumber="52" value="1989-02-03">3.ii.1989</collectingDate>
</date>
, H.F. &amp; A.T.
<collectorName id="26DFF3D3FFA08D79A19AFBE3565F9564" box="[727,822,1109,1134]" pageId="11" pageNumber="52">Howden</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A008FBE0514F9564" box="[837,1062,1110,1134]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030243</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A701FBE051CD9564" box="[1100,1188,1110,1134]" pageId="11" pageNumber="52">CMNC</collectionCode>
);
<collectingMunicipality id="6BF10C7FFFA08D79A7FAFBE355809598" pageId="11" pageNumber="52">Southedge Research Station</collectingMunicipality>
,
<quantity id="4CD23BE0FFA08D79A3B6FBCF54209598" box="[251,329,1145,1170]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="52" unit="km" value="12.0">12 km</quantity>
NW
<location id="8EF5C0DEFFA08D79A2D9FBCF54909598" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A2D9FBCF54909598" box="[404,505,1145,1170]" county="Southedge Research Station" latitude="-17.15" longLatPrecision="21" longitude="145.51666" municipality="Southedge Research Station" name="Mareeba" pageId="11" pageNumber="52" stateProvince="York">Mareeba</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A15EFBCF57FE9598" box="[531,663,1145,1170]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-16.978611">16°58'43&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A1E4FBCF56549598" box="[681,829,1145,1170]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.3514">145°21'05&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A01AFBCF516A9598" box="[855,1027,1145,1170]" pageId="11" pageNumber="52" value="1988-12-26" valueMax="1988-12-29" valueMin="1988-12-26">
<collectingDate id="EFD0492DFFA08D79A01AFBCF516A9598" box="[855,1027,1145,1170]" pageId="11" pageNumber="52" value="1988-12-26" valueMax="1988-12-29" valueMin="1988-12-26">2629.xii.1988</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A75BFBCC507F9598" box="[1046,1302,1145,1170]" pageId="11" pageNumber="52">H.F. &amp; A.T. Howden</collectorName>
(
<collectionCode id="ED3B0EC0FFA08D79A661FBCF50F5959B" box="[1324,1436,1145,1169]" pageId="11" pageNumber="52">CMNEN-</collectionCode>
0 0 0
<quantity id="4CD23BE0FFA08D79A3F2FB2B544A95BF" box="[191,291,1180,1205]" metricMagnitude="2" metricUnit="m" metricValue="7.681976" pageId="11" pageNumber="52" unit="in" value="30244.0">30244 in</quantity>
<collectionCode id="ED3B0EC0FFA08D79A267FB2B54EB95BF" box="[298,386,1181,1205]" pageId="11" pageNumber="52">CMNC</collectionCode>
);
<locationDeviation id="9A278F27FFA08D79A2D9FB2B573795BF" box="[404,606,1180,1205]" location="Tolga" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A2D9FB2B54A595BF" box="[404,460,1180,1205]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="52" unit="km" value="7.0">7 km</quantity>
NE of Tolga
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A13DFB2B579A95BF" box="[624,755,1181,1205]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-17.15">17°09'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A1B2FB2A56FB95BF" box="[767,914,1180,1205]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.51666">145°31'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A0E8FB2A569495BF" box="[933,1021,1180,1205]" pageId="11" pageNumber="52" value="1987-03">
<collectingDate id="EFD0492DFFA08D79A0E8FB2A569495BF" box="[933,1021,1180,1205]" pageId="11" pageNumber="52" value="1987-03">iii.1987</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A745FB2B51EB95BF" box="[1032,1154,1181,1205]" pageId="11" pageNumber="52">R.I. Storey</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A7E9FB2B507A95BF" box="[1188,1299,1180,1205]" pageId="11" pageNumber="52">De Faveri</collectorName>
(25-
<collectingDate id="EFD0492DFFA08D79A609FB2A55DA95D2" isEnumeration="true" pageId="11" pageNumber="52">
<date id="FF94B0C5FFA08D79A609FB2A50F595BF" box="[1348,1436,1180,1205]" pageId="11" pageNumber="52" value="35-03-26">032635</date>
<date id="FF94B0C5FFA08D79A3DAFB7655DA95D2" box="[151,179,1216,1240]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A3E5FB7655DA95D2" box="[168,179,1216,1240]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</date>
</collectingDate>
in
<collectionCode id="ED3B0EC0FFA08D79A39AFB76544E95D2" box="[215,295,1216,1240]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34157" lsid="urn:lsid:biocol.org:col:34157" name="Queensland Primary Industries Insect Collection" pageId="11" pageNumber="52" type="Other type of research institution/biorepository">QDPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A276FB7754D39415" ID-GBIF-Occurrence="2012745309" collectingDate="1987-01-19" collectingDateMax="1987-02-02" collectingDateMin="1987-01-19" collectionCode="QDPC, ANIC" collectorName="R. I. Storey &amp; De Faveri" latitude="-17.15" location="Tolga" longLatPrecision="21" longitude="145.51666" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A276FB77541D95D2" box="[315,372,1215,1240]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="52" unit="km" value="7.0">7 km</quantity>
NE of
<location id="8EF5C0DEFFA08D79A28BFB09576195D2" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A28BFB09576195D2" box="[454,520,1215,1240]" latitude="-17.15" longLatPrecision="21" longitude="145.51666" name="Tolga" pageId="11" pageNumber="52" stateProvince="York">Tolga</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A156FB7657F795D2" box="[539,670,1216,1240]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-17.15">17°09'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A1E6FB09565695D2" box="[683,831,1215,1240]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.51666">145°31'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A01CFB0956DE95D2" box="[849,951,1215,1240]" pageId="11" pageNumber="52" value="1987-02-02">
<collectingDate id="EFD0492DFFA08D79A01CFB0956DE95D2" box="[849,951,1215,1240]" pageId="11" pageNumber="52" value="1987-02-02">2.ii.1987</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A08EFB77515795D2" box="[963,1086,1216,1240]" pageId="11" pageNumber="52">R.I. Storey</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A72FFB7751B895D2" box="[1122,1233,1215,1240]" pageId="11" pageNumber="52">De Faveri</collectorName>
(25-
<collectingDate id="EFD0492DFFA08D79A64EFB09501695D2" box="[1283,1407,1215,1240]" isEnumeration="true" pageId="11" pageNumber="52">
<date id="FF94B0C5FFA08D79A64EFB09503295D2" box="[1283,1371,1215,1240]" pageId="11" pageNumber="52" value="37-03-26">032637</date>
<date id="FF94B0C5FFA08D79A62EFB76501695D2" box="[1379,1407,1216,1240]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A639FB76501695D2" box="[1396,1407,1216,1240]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</date>
</collectingDate>
in
<collectionCode id="ED3B0EC0FFA08D79A3DAFB52558C95F6" box="[151,229,1252,1276]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34157" lsid="urn:lsid:biocol.org:col:34157" name="Queensland Primary Industries Insect Collection" pageId="11" pageNumber="52" type="Other type of research institution/biorepository">QDPC</collectionCode>
, 25-058982
<date id="FF94B0C5FFA08D79A222FB5254E295F6" box="[367,395,1252,1276]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A222FB5254E295F6" box="[367,395,1252,1276]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A2CDFB5254E295F6" box="[384,395,1252,1276]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A2E3FB52549E95F6" box="[430,503,1252,1276]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
);
<locationDeviation id="9A278F27FFA08D79A147FB5257BD95F6" box="[522,724,1251,1276]" location="Tolga" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A147FB52572B95F6" box="[522,578,1251,1276]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="52" unit="km" value="7.0">7 km</quantity>
NE of Tolga
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A1ABFB52560095F6" box="[742,873,1252,1276]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-17.15">17°09'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A038FB55516195F6" box="[885,1032,1251,1276]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.51666">145°31'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A757FB5551EF95F6" box="[1050,1158,1251,1276]" pageId="11" pageNumber="52" value="1987-01-19">
<collectingDate id="EFD0492DFFA08D79A757FB5551EF95F6" box="[1050,1158,1251,1276]" pageId="11" pageNumber="52" value="1987-01-19">19.i.1987</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A7DCFB52506295F6" box="[1169,1291,1252,1276]" pageId="11" pageNumber="52">R.I. Storey</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A660FB5250F595F6" box="[1325,1436,1251,1276]" pageId="11" pageNumber="52">De Faveri</collectorName>
(25-
<collectingDate id="EFD0492DFFA08D79A38CFAB054549415" box="[193,317,1286,1311]" isEnumeration="true" pageId="11" pageNumber="52">
<date id="FF94B0C5FFA08D79A38CFAB054709415" box="[193,281,1286,1311]" pageId="11" pageNumber="52" value="36-03-26">032636</date>
<date id="FF94B0C5FFA08D79A26CFAB154549415" box="[289,317,1287,1311]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A27FFAB154549415" box="[306,317,1287,1311]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</date>
</collectingDate>
in
<collectionCode id="ED3B0EC0FFA08D79A22FFAB154DB9415" box="[354,434,1287,1311]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34157" lsid="urn:lsid:biocol.org:col:34157" name="Queensland Primary Industries Insect Collection" pageId="11" pageNumber="52" type="Other type of research institution/biorepository">QDPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A288FABE54EA9449" ID-GBIF-Occurrence="2012745312" collectingDate="1987-01-27" collectionCode="ANIC" collectorName="R. I. Storey &amp; De Faveri" latitude="-17.15" location="Tolga" longLatPrecision="21" longitude="145.51666" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<quantity id="4CD23BE0FFA08D79A288FABE54969415" box="[453,511,1286,1311]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="52" unit="km" value="7.0">7 km</quantity>
NE of
<location id="8EF5C0DEFFA08D79A11FFAB057FD9415" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A11FFAB057FD9415" box="[594,660,1286,1311]" latitude="-17.15" longLatPrecision="21" longitude="145.51666" name="Tolga" pageId="11" pageNumber="52" stateProvince="York">Tolga</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A1E5FAB156429415" box="[680,811,1287,1311]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-17.15">17°09'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A07AFAB056A29415" box="[823,971,1286,1311]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="145.51666">145°31'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A093FAB051229415" box="[990,1099,1286,1311]" pageId="11" pageNumber="52" value="1987-01-27">
<collectingDate id="EFD0492DFFA08D79A093FAB051229415" box="[990,1099,1286,1311]" pageId="11" pageNumber="52" value="1987-01-27">27.i.1987</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A71AFABE51B99415" box="[1111,1232,1287,1311]" pageId="11" pageNumber="52">R.I. Storey</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A7BBFABE500F9415" box="[1270,1382,1286,1311]" pageId="11" pageNumber="52">De Faveri</collectorName>
(25- 0 58984
<date id="FF94B0C5FFA08D79A3BFFA9D54679449" box="[242,270,1323,1347]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A3BFFA9D54679449" box="[242,270,1323,1347]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A24EFA9D54679449" box="[259,270,1323,1347]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A27CFA9D54129449" box="[305,379,1323,1347]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
)
</materialsCitation>
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<date id="FF94B0C5FFA08D79A195F994561F9731" box="[728,886,1570,1595]" pageId="11" pageNumber="52" value="2001-02-20" valueMax="2001-02-21" valueMin="2001-02-20">
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,
<collectorName id="26DFF3D3FFA08D79A0C5F99551449731" box="[904,1069,1570,1595]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
(T
<date id="FF94B0C5FFA08D79A71FF99451CF9731" box="[1106,1190,1570,1595]" pageId="11" pageNumber="52" value="59-11-19">
<collectingDate id="EFD0492DFFA08D79A71FF99451CF9731" box="[1106,1190,1570,1595]" pageId="11" pageNumber="52" value="59-11-19">111959</collectingDate>
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T
<date id="FF94B0C5FFA08D79A78BF99450739731" box="[1222,1306,1570,1595]" pageId="11" pageNumber="52" value="60-11-19">
<collectingDate id="EFD0492DFFA08D79A78BF99450739731" box="[1222,1306,1570,1595]" pageId="11" pageNumber="52" value="60-11-19">111960</collectingDate>
</date>
, T
<date id="FF94B0C5FFA08D79A676F99450E49731" box="[1339,1421,1570,1595]" pageId="11" pageNumber="52" value="62-11-19">
<collectingDate id="EFD0492DFFA08D79A676F99450E49731" box="[1339,1421,1570,1595]" pageId="11" pageNumber="52" value="62-11-19">111962</collectingDate>
</date>
T
<date id="FF94B0C5FFA08D79A3E5F9F055949755" box="[168,253,1606,1631]" pageId="11" pageNumber="52" value="64-11-19">
<collectingDate id="EFD0492DFFA08D79A3E5F9F055949755" box="[168,253,1606,1631]" pageId="11" pageNumber="52" value="64-11-19">111964</collectingDate>
</date>
, T
<date id="FF94B0C5FFA08D79A26CF9F0541D9755" box="[289,372,1606,1631]" pageId="11" pageNumber="52" value="66-11-19">
<collectingDate id="EFD0492DFFA08D79A26CF9F0541D9755" box="[289,372,1606,1631]" pageId="11" pageNumber="52" value="66-11-19">111966</collectingDate>
</date>
T
<quantity id="4CD23BE0FFA08D79A2DEF9F057639755" box="[403,522,1606,1631]" metricMagnitude="3" metricUnit="m" metricValue="2.8439618" pageId="11" pageNumber="52" unit="in" value="111967.0">
<date id="FF94B0C5FFA08D79A2DEF9F0548C9755" box="[403,485,1606,1631]" pageId="11" pageNumber="52" value="67-11-19">
<collectingDate id="EFD0492DFFA08D79A2DEF9F0548C9755" box="[403,485,1606,1631]" pageId="11" pageNumber="52" value="67-11-19">111967</collectingDate>
</date>
in
</quantity>
<collectionCode id="ED3B0EC0FFA08D79A157F9F157259755" box="[538,588,1607,1631]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A12AF9F0577D97AF" ID-GBIF-Occurrence="2012745310" collectingDate="1974-05-06" collectingDateMax="2012-03-16" collectingDateMin="1974-05-06" collectionCode="ANIC" collectorName="G. B. Monteith &amp; R. I. Storey" elevation="60" latitude="-19.016666" location="Balgal Beach" longLatPrecision="21" longitude="146.4" municipality="Paluma Road" pageId="11" pageNumber="52" specimenCount="1" stateProvince="York">
<collectingMunicipality id="6BF10C7FFFA08D79A12AF9F056609755" box="[615,777,1606,1631]" pageId="11" pageNumber="52">Paluma Road</collectingMunicipality>
,
<geoCoordinate id="EE1EF0C2FFA08D79A050F9F056F79755" box="[797,926,1606,1631]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-18.99">18°59'24&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A0FEF9F0512D9755" box="[947,1092,1606,1631]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="146.29501">146°17'42&quot;E</geoCoordinate>
,
<quantity id="4CD23BE0FFA08D79A715F9F051F59754" box="[1112,1180,1606,1631]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="52" unit="m" value="60.0">
<elevation id="00077136FFA08D79A715F9F051F59754" box="[1112,1180,1606,1631]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="52" unit="m" value="60.0">60 m</elevation>
</quantity>
,
<date id="FF94B0C5FFA08D79A7FCF9F0503F9755" box="[1201,1366,1606,1631]" pageId="11" pageNumber="52" value="2012-03-14" valueMax="2012-03-16" valueMin="2012-03-14">
<collectingDate id="EFD0492DFFA08D79A7FCF9F0503F9755" box="[1201,1366,1606,1631]" pageId="11" pageNumber="52" value="2012-03-14" valueMax="2012-03-16" valueMin="2012-03-14">1416.iii.2012</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A626F9F155979788" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
(25-053918
<date id="FF94B0C5FFA08D79A2D8F9DC54DB9788" box="[405,434,1642,1666]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A2D8F9DC54DB9788" box="[405,434,1642,1666]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A2EAF9DC54DB9788" box="[423,434,1642,1666]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A290F9DC574B9788" box="[477,546,1642,1666]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
also as COL2404);
<location id="8EF5C0DEFFA08D79A05CF9DF56C79788" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A05CF9DF56C79788" box="[785,942,1641,1666]" latitude="-19.016666" longLatPrecision="21" longitude="146.4" municipality="Paluma Road" name="Balgal Beach" pageId="11" pageNumber="52" stateProvince="York">Balgal Beach</location>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A71AF9DF51839788" box="[1111,1258,1641,1666]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="146.4">146°24'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A64DF9DF500A9788" box="[1280,1379,1641,1666]" pageId="11" pageNumber="52" value="1974-05-06">
<collectingDate id="EFD0492DFFA08D79A64DF9DF500A9788" box="[1280,1379,1641,1666]" pageId="11" pageNumber="52" value="1974-05-06">6.v.1974</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A63FF9DC558997AF" pageId="11" pageNumber="52">R.I. Storey</collectorName>
(25-058519
<date id="FF94B0C5FFA08D79A234F93B54FF97AF" box="[377,406,1677,1701]" pageId="11" pageNumber="52">
<collectingDate id="EFD0492DFFA08D79A234F93B54FF97AF" box="[377,406,1677,1701]" pageId="11" pageNumber="52">
x
<quantity id="4CD23BE0FFA08D79A2C6F93B54FF97AF" box="[395,406,1677,1701]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="52" unit="in" value="1.0">1</quantity>
</collectingDate>
</date>
in
<collectionCode id="ED3B0EC0FFA08D79A28EF93B576597AF" box="[451,524,1677,1701]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A169F93B558197E6" ID-GBIF-Occurrence="2012745324" collectingDate="2014-08-14" collectionCode="ANIC" collectorName="M. Braby" latitude="-11.267777" location="Site" longLatPrecision="21" longitude="132.26611" municipality="Black Point" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Northern Territory">
<emphasis id="B95E4A17FFA08D79A169F93B567E97AC" bold="true" box="[548,791,1677,1702]" pageId="11" pageNumber="52">
<collectingRegion id="49EE58E7FFA08D79A169F93B567B97AC" box="[548,786,1677,1702]" country="Australia" name="Northern Territory" pageId="11" pageNumber="52">Northern Territory</collectingRegion>
:
</emphasis>
<geoCoordinate id="EE1EF0C2FFA08D79A06EF93B56CE97AC" box="[803,935,1677,1702]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="1" value="-11.26774">11.26774°S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FFA08D79A0FFF93B512197AC" box="[946,1096,1677,1702]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="1" value="132.26614">132.26615°E</geoCoordinate>
, near
<collectingMunicipality id="6BF10C7FFFA08D79A7D9F93B507797AC" box="[1172,1310,1677,1702]" pageId="11" pageNumber="52">Black Point</collectingMunicipality>
,
<location id="8EF5C0DEFFA08D79A663F93B503397AC" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A663F93B503397AC" box="[1326,1370,1677,1702]" latitude="-11.267777" longLatPrecision="21" longitude="132.26611" municipality="Black Point" name="Site" pageId="11" pageNumber="52" stateProvince="Northern Territory">Site</location>
c002 mushroom paperbark with rainforest understorey,
<geoCoordinate id="EE1EF0C2FFA08D79A187F906562297C3" box="[714,843,1712,1737]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-11.267777">11°16'04&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A01AF906568197C3" box="[855,1000,1712,1737]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="132.26611">132°15'58&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFA08D79A0B9F906511497C3" box="[1012,1149,1712,1737]" pageId="11" pageNumber="52" value="2014-08-14">
<collectingDate id="EFD0492DFFA08D79A0B9F906511497C3" box="[1012,1149,1712,1737]" pageId="11" pageNumber="52" value="2014-08-14">14.viii.2014</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A7C4F907519D97C3" box="[1161,1268,1712,1737]" pageId="11" pageNumber="52">M. Braby</collectorName>
(25-065548 in
<collectionCode id="ED3B0EC0FFA08D79A3DAF962558997E6" box="[151,224,1748,1772]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="52">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A3BAF963543E961A" ID-GBIF-Occurrence="2012745332" collectingDate="1979-07-22" collectingDateMax="46-11-19" collectingDateMin="1979-07-22" collectionCode="QM" collectorName="G. B. Monteith &amp; D. J. Cook" latitude="-12.233334" location="West Alligator Mouth" longLatPrecision="21" longitude="132.26666" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Northern Territory">
<location id="8EF5C0DEFFA08D79A3BAF963549E97E7" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A3BAF963549E97E7" box="[247,503,1748,1773]" latitude="-12.233334" longLatPrecision="21" longitude="132.26666" name="West Alligator Mouth" pageId="11" pageNumber="52" stateProvince="Northern Territory">West Alligator Mouth</location>
,
<geoCoordinate id="EE1EF0C2FFA08D79A14BF962571E97E6" box="[518,631,1748,1772]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-12.233334">12°14'0&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A1CAF962566E97E7" box="[647,775,1748,1773]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="132.26666">132°16'0&quot;E</geoCoordinate>
,
<date id="FF94B0C5FFA08D79A05AF96256AA97E7" box="[791,963,1748,1773]" pageId="11" pageNumber="52" value="1979-07-22" valueMax="1979-07-24" valueMin="1979-07-22">
<collectingDate id="EFD0492DFFA08D79A05AF96256AA97E7" box="[791,963,1748,1773]" pageId="11" pageNumber="52" value="1979-07-22" valueMax="1979-07-24" valueMin="1979-07-22">2224.vii.1979</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A09EF962511F97E7" box="[979,1142,1748,1773]" pageId="11" pageNumber="52">G.B. Monteith</collectorName>
&amp;
<collectorName id="26DFF3D3FFA08D79A7EFF963507E97E7" box="[1186,1303,1748,1773]" pageId="11" pageNumber="52">D.J. Cook</collectorName>
(T
<date id="FF94B0C5FFA08D79A677F96250E497E7" box="[1338,1421,1748,1773]" pageId="11" pageNumber="52" value="45-11-19">
<collectingDate id="EFD0492DFFA08D79A677F96250E497E7" box="[1338,1421,1748,1773]" pageId="11" pageNumber="52" value="45-11-19">111945</collectingDate>
</date>
T
<quantity id="4CD23BE0FFA08D79A3EAF941547E961A" box="[167,279,1783,1808]" metricMagnitude="3" metricUnit="m" metricValue="2.8434284" pageId="11" pageNumber="52" unit="in" value="111946.0">
<date id="FF94B0C5FFA08D79A3EAF9415593961A" box="[167,250,1783,1808]" pageId="11" pageNumber="52" value="46-11-19">
<collectingDate id="EFD0492DFFA08D79A3EAF9415593961A" box="[167,250,1783,1808]" pageId="11" pageNumber="52" value="46-11-19">111946</collectingDate>
</date>
in
</quantity>
<collectionCode id="ED3B0EC0FFA08D79A250F94E5427961A" box="[285,334,1784,1808]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="52" type="Museum">QM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A22FF941560A9639" ID-GBIF-Occurrence="2012745315" collectingDate="1993-12-12" collectingDateMax="1993-12-25" collectingDateMin="1993-12-12" collectionCode="CMNC" collectorName="S. B. &amp; J. Peck" county="Kakadu National Park" latitude="-12.672778" location="Gungaree Rainforest Trail" longLatPrecision="21" longitude="132.48138" municipality="Alligator River" pageId="11" pageNumber="52" specimenCode="CMNEN-00030231" specimenCount="1" stateProvince="Northern Territory">
<collectingCounty id="62F4EE89FFA08D79A22FF9415734961A" box="[354,605,1783,1808]" pageId="11" pageNumber="52">Kakadu National Park</collectingCounty>
, S
<collectingMunicipality id="6BF10C7FFFA08D79A133F9415643961A" box="[638,810,1783,1808]" pageId="11" pageNumber="52">Alligator River</collectingMunicipality>
,
<location id="8EF5C0DEFFA08D79A078F94E5135961A" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A078F94E5135961A" box="[821,1116,1783,1808]" county="Kakadu National Park" latitude="-12.672778" longLatPrecision="21" longitude="132.48138" municipality="Alligator River" name="Gungaree Rainforest Trail" pageId="11" pageNumber="52" stateProvince="Northern Territory">Gungaree Rainforest Trail</location>
, [
<geoCoordinate id="EE1EF0C2FFA08D79A723F94E5198961A" box="[1134,1265,1784,1808]" direction="south" orientation="latitude" pageId="11" pageNumber="52" precision="15" value="-12.672778">12°40'22&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA08D79A7B0F94150F9961A" box="[1277,1424,1783,1808]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="132.48138">132°28'53&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A3DAF8AD542A963E" box="[151,323,1819,1844]" pageId="11" pageNumber="52" value="1993-12-12" valueMax="1993-12-25" valueMin="1993-12-12">
<collectingDate id="EFD0492DFFA08D79A3DAF8AD542A963E" box="[151,323,1819,1844]" pageId="11" pageNumber="52" value="1993-12-12" valueMax="1993-12-25" valueMin="1993-12-12">1225.xii.1993</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A203F8AD5498963E" box="[334,497,1819,1844]" pageId="11" pageNumber="52">S.B. &amp; J. Peck</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A14DF8AD57899639" box="[512,736,1819,1843]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030231</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A04EF8AD56329639" box="[771,859,1819,1843]" pageId="11" pageNumber="52">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B429C58FFA08D79A023F8AD507F9670" ID-GBIF-Occurrence="2012745316" collectingDate="1973-05-21" collectingDateMax="1993-12-27" collectingDateMin="1973-05-21" collectionCode="CMNC, ANIC" collectorName="S. B. &amp; J. Peck &amp; E. G. Matthews" county="Darwin" latitude="-12.8" location="Nourlangie Creek" longLatPrecision="21" longitude="132.7" municipality="Berry Spring Park" pageId="11" pageNumber="52" specimenCode="CMNEN-00030236-00030238" specimenCount="1" stateProvince="Northern Territory">
<quantity id="4CD23BE0FFA08D79A023F8AD56DD963E" box="[878,948,1819,1844]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="52" unit="km" value="50.0">50 km</quantity>
S
<collectingCounty id="62F4EE89FFA08D79A09CF8AD5141963E" box="[977,1064,1819,1844]" pageId="11" pageNumber="52">Darwin</collectingCounty>
,
<collectingMunicipality id="6BF10C7FFFA08D79A77EF8AA5068963E" box="[1075,1281,1819,1844]" pageId="11" pageNumber="52">Berry Spring Park</collectingMunicipality>
, [
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,
<geoCoordinate id="EE1EF0C2FFA08D79A3DAF8895443965D" box="[151,298,1855,1879]" direction="east" orientation="longitude" pageId="11" pageNumber="52" precision="15" value="131.0">131°00'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA08D79A273F88854B5965D" box="[318,476,1854,1879]" pageId="11" pageNumber="52" value="1993-12-04" valueMax="1993-12-27" valueMin="1993-12-04">
<collectingDate id="EFD0492DFFA08D79A273F88854B5965D" box="[318,476,1854,1879]" pageId="11" pageNumber="52" value="1993-12-04" valueMax="1993-12-27" valueMin="1993-12-04">427.xii.1993</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA08D79A2A4F88957E6965D" box="[489,655,1854,1879]" pageId="11" pageNumber="52">S.B. &amp; J. Peck</collectorName>
(
<specimenCode id="DB8C3E7EFFA08D79A1D2F8885691965D" box="[671,1016,1854,1879]" collectionCode="CMNEN-" pageId="11" pageNumber="52">CMNEN-00030236-00030238</specimenCode>
in
<collectionCode id="ED3B0EC0FFA08D79A752F889511E965D" box="[1055,1143,1855,1879]" pageId="11" pageNumber="52">CMNC</collectionCode>
);
<location id="8EF5C0DEFFA08D79A7C7F888503F965D" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA08D79A7C7F888503F965D" box="[1162,1366,1854,1879]" county="Darwin" latitude="-12.8" longLatPrecision="21" longitude="132.7" municipality="Berry Spring Park" name="Nourlangie Creek" pageId="11" pageNumber="52" stateProvince="Northern Territory">Nourlangie Creek</location>
,
<locationDeviation id="9A278F27FFA08D79A62FF88954189670" location="Mount Cahill" pageId="11" pageNumber="52">
<quantity id="4CD23BE0FFA08D79A62FF88950F5965D" box="[1378,1436,1854,1879]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="km" value="8.0">8 km</quantity>
N of Mount Cahill
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</date>
,
<collectorName id="26DFF3D3FFA78D7EA25AFEFC54C8906B" box="[279,417,329,353]" pageId="12" pageNumber="53">N.L. Gunter</collectorName>
(25-057633
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x
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in
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)
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;
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<collectingMunicipality id="6BF10C7FFFA78D7EA199FEFF562E9068" box="[724,839,329,354]" pageId="12" pageNumber="53">Pungalina</collectingMunicipality>
,
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SSE of Pungalina Homestead
</locationDeviation>
,
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,
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,
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,
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</collectingDate>
,
<collectorName id="26DFF3D3FFA78D7EA1CAFEDB5666908F" box="[647,783,365,389]" pageId="12" pageNumber="53">N.L. Gunter</collectorName>
(25-057634
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<collectingDate id="EFD0492DFFA78D7EA0EDFEDB568C908F" box="[928,997,364,389]" pageId="12" pageNumber="53" value="1925-10-01">x1, 25</collectingDate>
</date>
-057635
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<collectingDate id="EFD0492DFFA78D7EA707FEDB510F908F" box="[1098,1126,365,389]" pageId="12" pageNumber="53">
x
<quantity id="4CD23BE0FFA78D7EA715FEDB510F908F" box="[1112,1126,365,389]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="53" unit="in" value="2.0">2</quantity>
</collectingDate>
</date>
in
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA78D7EA38AFE26541E9403" pageId="12" pageNumber="53" type="description">
<paragraph id="8B959605FFA78D7EA38AFE26511390A3" blockId="12.[151,1437,151,2009]" box="[199,1146,400,425]" pageId="12" pageNumber="53">
<emphasis id="B95E4A17FFA78D7EA38AFE26543090A3" bold="true" box="[199,345,400,425]" pageId="12" pageNumber="53">Description.</emphasis>
Dark brown in colour, sometimes with red humeri. Antennal club grey.
</paragraph>
<paragraph id="8B959605FFA78D7EA38AFE0254EB931F" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">
Total length:
<quantity id="4CD23BE0FFA78D7EA213FE02548090C6" box="[350,489,436,461]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="12" pageNumber="53" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.22.6 mm</quantity>
.
<typeStatus id="549128A7FFA78D7EA2B8FE03570C90C7" box="[501,613,437,461]" pageId="12" pageNumber="53" type="lectotype">Lectotype</typeStatus>
measurements (of
<taxonomicName id="4C2AED86FFA78D7EA073FE02510C90C6" authorityName="Macleay" authorityYear="1888" box="[830,1125,436,461]" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="pygmaeum">
<emphasis id="B95E4A17FFA78D7EA073FE02510C90C6" box="[830,1125,436,461]" italics="true" pageId="12" pageNumber="53">Temnoplectron pygmaeum</emphasis>
</taxonomicName>
): (
<specimenCount id="9D2C5D8CFFA78D7EA7C4FE0351F290C7" box="[1161,1179,437,461]" pageId="12" pageNumber="53" type="female"></specimenCount>
) Total length
<quantity id="4CD23BE0FFA78D7EA673FE0350FE90C6" box="[1342,1431,437,461]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="53" unit="mm" value="2.2">2.2 mm</quantity>
, elytral width
<quantity id="4CD23BE0FFA78D7EA278FE6E54FB90FA" box="[309,402,472,497]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="53" unit="mm" value="1.5">1.5 mm</quantity>
.
<typeStatus id="549128A7FFA78D7EA2ECFE6F577890FB" box="[417,529,473,497]" pageId="12" pageNumber="53" type="lectotype">Lectotype</typeStatus>
measurements (of
<taxonomicName id="4C2AED86FFA78D7EA1B9FE6E56C090FB" authorityName="Blackburn" authorityYear="1900" box="[756,937,472,497]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA78D7EA1B9FE6E56C090FB" box="[756,937,472,497]" italics="true" pageId="12" pageNumber="53">Panelus arthuri</emphasis>
</taxonomicName>
): (sex not recorded) Total length
<quantity id="4CD23BE0FFA78D7EA676FE6E50FE90FA" box="[1339,1431,472,496]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="53" unit="mm" value="2.4">2.4 mm</quantity>
, elytral width
<quantity id="4CD23BE0FFA78D7EA261FE4A5417931F" box="[300,382,508,533]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="53" unit="mm" value="1.5">1.5mm</quantity>
.
</paragraph>
<paragraph id="8B959605FFA78D7EA38AFD96578793AF" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Margin slightly angulate at clypeogenal suture. Genal angle prominent. Basal carina present and interrupted medially. Dorsal part of eyes moderate in size, separated by an interocular space approximately 1617 times eye width (3234: 2); eye canthus not dividing eye. Head 1.4 × wider than long (44:32).</paragraph>
<paragraph id="8B959605FFA78D7EA38AFD065481921A" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Prothorax: Hypomeral surface finely reticulate with very light punctation, punctures along basal margin slightly elongate and different from the disc. Hypomeral striae about half the length of the hypomeron. Pronotum 1.6 × wider than long (70:44).</paragraph>
<paragraph id="8B959605FFA78D7EA38AFCAA50369277" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Elytra: Surface smooth. Striae 16 or 15 appearing double with very finely punctate edges. All striae on disc not equal width, striae 57 not as wide, striae 6 and/or 7 appearing single (sometimes stria 5 also appears single). Apically, with a short groove extending from end of epipleuron to apex. Elytra 1.2 × wider than long (85:70).</paragraph>
<paragraph id="8B959605FFA78D7EA38AFC3F559A92E3" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Legs: Front edge of protibiae truncate; apical digit somewhat elongate and sharp; two teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws small and toothed.</paragraph>
<paragraph id="8B959605FFA78D7EA38AFC4256F5955E" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Abdomen: Pygidium with wide pear-shape depression containing a somewhat triangular tubercle in upper third. The depression takes up over half of surface and extends under the slightly concave upper edge. Abdominal surface very finely reticulate right across. Segment 6 finely punctate.</paragraph>
<paragraph id="8B959605FFA78D7EA38AFBD651BE95CB" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Mesometaventral suture very slightly arched. Lateral lobe of metaventrite smooth and punctate with setae. Mesoventrite with slight notch anteriorly. Mesoventrite and mesepimeron smooth. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FFA78D7EA38AFB7A541E9403" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">Female: Apical digit absent; apical spur extending straight from front edge. Abdominal ventrite 6 slightly longer than in male.</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA78D7EA38AFAA25011979F" pageId="12" pageNumber="53" type="distribution">
<paragraph id="8B959605FFA78D7EA38AFAA25011979F" blockId="12.[151,1437,151,2009]" pageId="12" pageNumber="53">
<emphasis id="B95E4A17FFA78D7EA38AFAA254099427" bold="true" box="[199,352,1300,1325]" pageId="12" pageNumber="53">Distribution.</emphasis>
A widespread species known from coast to coast in equatorial latitudes north of about 19°S, the vast majority of specimens have been collected in open forest (
<figureCitation id="13118A80FFA78D7EA025FA8E56D4945A" box="[872,957,1336,1361]" captionStart="FIGURE 13" captionStartId="29.[151,250,1893,1915]" captionTargetBox="[151,1436,1204,1872]" captionTargetId="figure@29.[151,1436,1204,1872]" captionTargetPageId="29" captionText="FIGURE 13. Map of known records of Lepanus pygmaeus including predicted distribution overlaid with predicted distribution boundary of L. guthrieae (dashed lines) taken from Fig. 16 map." httpUri="https://zenodo.org/record/2588647/files/figure.png" pageId="12" pageNumber="53">Fig. 13</figureCitation>
). Specimens have been recorded from the IBRA regions Dampierland (DAL), Darwin Coastal (DAC), Pine Creek (PCK), Daly Basin (DAB), Arnhem Coast (ARC), Gulf Coastal (GUC), Cape York Peninsula (CYP), and Wet Tropics (WET) on mainland
<collectingCountry id="F33DD695FFA78D7EA796FA36502C9493" box="[1243,1349,1408,1433]" name="Australia" pageId="12" pageNumber="53">Australia</collectingCountry>
, and on Groote Eylandt (NT), Saibai and Moa Islands (Torres Strait,
<collectingRegion id="49EE58E7FFA78D7EA0C0FA12517E94B7" box="[909,1047,1444,1469]" country="Australia" name="Queensland" pageId="12" pageNumber="53">Queensland</collectingRegion>
). BIOCLIM modelling indicates appropriate climate for
<taxonomicName id="4C2AED86FFA78D7EA2E7FA7F575594EA" box="[426,572,1481,1504]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA78D7EA2E7FA7F575594EA" box="[426,572,1481,1504]" italics="true" pageId="12" pageNumber="53">L. pygmaeus</emphasis>
</taxonomicName>
exists in Central Kimberley (CEK), Northern Kimberley (NOK),
<collectingRegion id="49EE58E7FFA78D7EA60FFA7E50F594EB" box="[1346,1436,1480,1505]" country="Australia" name="Victoria" pageId="12" pageNumber="53">Victoria</collectingRegion>
Bonaparte (VIB), Ord
<collectingRegion id="49EE58E7FFA78D7EA2DBFA5A5499970F" box="[406,496,1516,1541]" country="Australia" name="Victoria" pageId="12" pageNumber="53">Victoria</collectingRegion>
Plain (OVP), Sturt Plateau (STU), Mount Isa Inlier (MII), Central Arnhem (CEA), Arnhem Plateau (ARP), Gulf Fall and Uplands (GFU), Gulf Plains (GUP), Einasleigh Uplands (EIU), Desert Uplands (DEU), Brigalow Belt North (BBN), Central Mackay Coast (CMC), and South Eastern
<collectingRegion id="49EE58E7FFA78D7EA787F98250279747" box="[1226,1358,1588,1613]" country="Australia" name="Queensland" pageId="12" pageNumber="53">Queensland</collectingRegion>
(SEQ) IBRA regions. While much of northern
<collectingCountry id="F33DD695FFA78D7EA125F9EE57A6977B" box="[616,719,1624,1649]" name="Australia" pageId="12" pageNumber="53">Australia</collectingCountry>
remains under sampled, numerous habitat
<typeStatus id="549128A7FFA78D7EA78FF9EF5194977B" box="[1218,1277,1625,1649]" pageId="12" pageNumber="53">types</typeStatus>
across CMC, BBN, and SEQ have been extensively surveyed and
<taxonomicName id="4C2AED86FFA78D7EA1AEF9CB5618979E" box="[739,881,1661,1684]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA78D7EA1AEF9CB5618979E" box="[739,881,1661,1684]" italics="true" pageId="12" pageNumber="53">L. pygmaeus</emphasis>
</taxonomicName>
has not been collected in these IBRA regions.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA78D7FA38AF91651EF90A7" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53" type="materials_examined">
<paragraph id="8B959605FFA78D7FA38AF91651EF90A7" blockId="12.[151,1437,151,2009]" lastBlockId="13.[151,1437,151,2014]" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53">
<emphasis id="B95E4A17FFA78D7EA38AF91656A797B3" bold="true" box="[199,974,1696,1721]" pageId="12" pageNumber="53">
Status of type specimens.
<taxonomicName id="4C2AED86FFA78D7EA2BEF91656A797B3" authority="Macleay, 1888" authorityName="Macleay" authorityYear="1888" box="[499,974,1696,1721]" class="Insecta" family="Scarabaeidae" genus="Temnoplectron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="pygmaeum">
<emphasis id="B95E4A17FFA78D7EA2BEF916564997B2" bold="true" box="[499,800,1696,1721]" italics="true" pageId="12" pageNumber="53">Temnoplectron pygmaeum</emphasis>
<bibRefCitation id="EFBBEBF4FFA78D7EA06AF91656A797B3" author="Macleay, W." box="[807,974,1696,1721]" pageId="12" pageNumber="53" pagination="897 - 904" refId="ref29625" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. II. The lamellicornes, Proceedings of the Linnean Society of New South Wales, 3, 897 - 904." type="journal article" year="1888">Macleay, 1888</bibRefCitation>
</taxonomicName>
</emphasis>
. Macleay described this species from one or more specimens collected by Froggatt in the vicinity of Kings Sound,
<collectingRegion id="49EE58E7FFA78D7EA0B5F97351A897D7" box="[1016,1217,1732,1757]" country="Australia" name="Western Australia" pageId="12" pageNumber="53">Western Australia</collectingRegion>
and housed in the Macleay Museum;
<bibRefCitation id="EFBBEBF4FFA78D7EA220F95E57F3960A" author="Monteith, G. B. &amp; Rossini, M." box="[365,666,1768,1793]" pageId="12" pageNumber="53" pagination="161 - 171" refId="ref29777" refString="Monteith, G. B. &amp; Rossini, M. (2017) A name for the Australian dung beetle Onthophagus bicornis Macleay, 1888 (Coleoptera: Scarabaeidae), with notes on type locality, distribution and biology, Australian Entomologist, 44, 161 - 171." type="journal article" year="2017">Monteith &amp; Rossini (2017)</bibRefCitation>
detailed the Froggatt expedition and history of Macleays specimens. In his revision of the genus
<taxonomicName id="4C2AED86FFA78D7EA291F8BB5663962F" authority=", Matthews (1974)" authorityName=", Matthews" authorityYear="1974" box="[476,778,1804,1829]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA78D7EA291F8BB5729962E" box="[476,576,1805,1828]" italics="true" pageId="12" pageNumber="53">Lepanus,</emphasis>
<bibRefCitation id="EFBBEBF4FFA78D7EA106F8BA5663962F" author="Matthews, E. G." box="[587,778,1804,1829]" pageId="12" pageNumber="53" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
</taxonomicName>
stated that he saw the
<typeStatus id="549128A7FFA78D7EA755F8BA5111962F" box="[1048,1144,1804,1829]" pageId="12" pageNumber="53" type="holotype">holotype</typeStatus>
(sex not recorded) in the Macleay Museum, but when the coleopteran types from that museum were transferred to ANIC, only two female specimens on the one card from that locality and labelled as
<taxonomicName id="4C2AED86FFA78D7EA061F8E250F19667" authority="Macl., Kings Sound" authorityName="Macl., Kings Sound" box="[812,1432,1876,1901]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="pygmaeus" subGenus="Temnoplectron">
<emphasis id="B95E4A17FFA78D7EA061F8E251DA9666" box="[812,1203,1876,1901]" italics="true" pageId="12" pageNumber="53">Panelus (Temnoplectron) pygmaeus</emphasis>
Macl., Kings Sound
</taxonomicName>
,
<collectingRegion id="49EE58E7FFA78D7EA3DAF8CF5437969B" box="[151,350,1912,1937]" country="Australia" name="Western Australia" pageId="12" pageNumber="53">Western Australia</collectingRegion>
were located. This label may have been added by Macleay Museums curator, George Masters, following the transfer of the species to the genus
<taxonomicName id="4C2AED86FFA78D7EA181F82A56C596BF" authority="(Lea 1904)" baseAuthorityName="Lea" baseAuthorityYear="1904" box="[716,940,1948,1973]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA78D7EA181F82A564D96BF" box="[716,804,1948,1973]" italics="true" pageId="12" pageNumber="53">Panelus</emphasis>
(
<bibRefCitation id="EFBBEBF4FFA78D7EA078F82B56CD96BF" author="Lea, A. M." box="[821,932,1949,1973]" pageId="12" pageNumber="53" pagination="60 - 107" refId="ref29459" refString="Lea, A. M. (1904) Descriptions of new species of Australian Coleoptera Part VIII. Proceedings of the Linnean Society of New South Wales, 29, 60 - 107. https: // doi. org / 10.5962 / bhl. part. 20155" type="journal article" year="1904">Lea 1904</bibRefCitation>
)
</taxonomicName>
. Interestingly, this label appears with added locality details compared to the smaller “N.W. Austr” label and may either be taken from Macleays description listing material from the vicinity of Kings Sound or represent a replaced label. These two specimens were presumed to be
<typeStatus id="549128A7FFA68D7FA3DAFF0B559E91DF" box="[151,247,189,213]" pageId="13" pageNumber="54" type="syntype">syntypes</typeStatus>
by
<bibRefCitation id="EFBBEBF4FFA68D7FA267FF0A571991DE" author="Britton, E. B. &amp; Stanbury, P. J." box="[298,624,188,213]" pageId="13" pageNumber="54" pagination="241 - 293" refId="ref29000" refString="Britton, E. B. &amp; Stanbury, P. J. (1981) Type specimens in the Macleay Museum, University of Sydney VII. Insects: Beetles (Insecta: Coleoptera). Proceedings of the Linnaean Society of New South Wales, 105, 241 - 293." type="journal article" year="1981">Britton and Stanbury (1981)</bibRefCitation>
, but there was no label to indicate that they were seen by Matthews. Subsequently,
<bibRefCitation id="EFBBEBF4FFA68D7FA27BFF69575F91F2" author="Cassis, G. &amp; Weir, T. A." box="[310,566,223,248]" pageId="13" pageNumber="54" pagination="106 - 173" refId="ref29088" refString="Cassis, G. &amp; Weir, T. A. (1992) Scarabaeinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Uol. 9. Australian Government Publishing Service, Canberra, pp. 106 - 173" type="book chapter" year="1992">Cassis and Weir (1992)</bibRefCitation>
interpreted the incorrect reference to a
<typeStatus id="549128A7FFA68D7FA0A8FF69512C91F2" box="[997,1093,223,248]" pageId="13" pageNumber="54" type="holotype">holotype</typeStatus>
as a
<typeStatus id="549128A7FFA68D7FA731FF69518991F2" box="[1148,1248,223,248]" pageId="13" pageNumber="54" type="lectotype">lectotype</typeStatus>
designation, but, as Matthews did not explicitly state which specimen or give characteristics of the specimen he called
<typeStatus id="549128A7FFA68D7FA640FEB250069017" box="[1293,1391,260,285]" pageId="13" pageNumber="54" type="holotype">holotype</typeStatus>
, we consider this not to be a valid
<typeStatus id="549128A7FFA68D7FA2B7FE915737904A" box="[506,606,295,320]" pageId="13" pageNumber="54" type="lectotype">lectotype</typeStatus>
designation under Article 74.5 (International Commission on
<bibRefCitation id="EFBBEBF4FFA68D7FA66BFE91541D906E" author="International Commission on Zoological Nomenclature &amp; ICZN" pageId="13" pageNumber="54" refId="ref29404" refString="International Commission on Zoological Nomenclature [ICZN] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">Zoological Nomenclature 1999</bibRefCitation>
). The two female specimens have subsequently been remounted on to separate points on the one pin and we hereby designate the female on the upper point with all tarsi intact as the
<typeStatus id="549128A7FFA68D7FA771FED951C99082" box="[1084,1184,367,392]" pageId="13" pageNumber="54" type="lectotype">lectotype</typeStatus>
and have labelled it as such. The female on the lower point with the protarsi missing thus becomes a
<typeStatus id="549128A7FFA68D7FA0A6FE2251EB90A7" box="[1003,1154,404,429]" pageId="13" pageNumber="54" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA68D7FA38AFE0154B79552" pageId="13" pageNumber="54" type="reference_group">
<paragraph id="8B959605FFA68D7FA38AFE0154B79552" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="54">
<taxonomicName id="4C2AED86FFA68D7FA38AFE01573990DA" ID-CoL="4CC79" authority="Blackburn, 1900" authorityName="Blackburn" authorityYear="1900" box="[199,592,439,464]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA68D7FA38AFE01573990DA" bold="true" box="[199,592,439,464]" pageId="13" pageNumber="54">
<emphasis id="B95E4A17FFA68D7FA38AFE0154E990DA" bold="true" box="[199,384,439,464]" italics="true" pageId="13" pageNumber="54">Panelus arthuri</emphasis>
<bibRefCitation id="EFBBEBF4FFA68D7FA2C5FE01573990DA" author="Blackburn, T." box="[392,592,439,464]" pageId="13" pageNumber="54" pagination="206 - 233" refId="ref28969" refString="Blackburn, T. (1900) On some new genera and species of Australian Coleoptera. Proceedings of the Royal Society of Uictoria, 12, 206 - 233." type="journal article" year="1900">Blackburn, 1900</bibRefCitation>
</emphasis>
</taxonomicName>
. Blackburn described this species from one or more specimens from NW
<collectingCountry id="F33DD695FFA68D7FA3DAFE6A559790FF" box="[151,254,476,501]" name="Australia" pageId="13" pageNumber="54">Australia</collectingCountry>
given to him by AM Lea and listed as from the Macleay Museum. In his revision of the genus
<taxonomicName id="4C2AED86FFA68D7FA67BFE6B543F9312" ID-CoL="5CD4" authority=", Matthews (1974)" authorityName=", Matthews" authorityYear="1974" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA68D7FA67BFE6B50F590FE" box="[1334,1436,477,500]" italics="true" pageId="13" pageNumber="54">Lepanus,</emphasis>
<bibRefCitation id="EFBBEBF4FFA68D7FA3DAFE49543F9312" author="Matthews, E. G." box="[151,342,511,536]" pageId="13" pageNumber="54" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
</taxonomicName>
stated that he saw the
<typeStatus id="549128A7FFA68D7FA11DFE4957DB9312" box="[592,690,511,536]" pageId="13" pageNumber="54" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2AED86FFA68D7FA19BFE4956EF9312" ID-CoL="4CC79" authorityName="Blackburn" authorityYear="1900" box="[726,902,511,536]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA68D7FA19BFE4956EF9312" box="[726,902,511,536]" italics="true" pageId="13" pageNumber="54">Panelus arthuri</emphasis>
</taxonomicName>
(sex not recorded) in the Macleay Museum, but when the coleopteran types from that museum were transferred to ANIC, no such specimen was found nor was it listed in Britton and Stanburys (1982) review of
<taxonomicName id="4C2AED86FFA68D7FA196FDF1563F936A" ID-CoL="C2L" authorityName="" authorityYear="1758" box="[731,854,583,608]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
types from Macleay Museum. A literature search revealed that the type was sent to the BMNH (NHM) along with many other Blackburn types in 1910 and after his death in 1912 (
<bibRefCitation id="EFBBEBF4FFA68D7FA207FD2654D593A2" author="Lea, A. M." box="[330,444,656,680]" pageId="13" pageNumber="54" refId="ref29508" refString="Lea, A. M. (1912) The late Rev. Canon Thomas Blackburn, B. A., and his entomological work. Transactions of the Royal Society of South Australia, 36, v-xl." type="book" year="1912">Lea 1912</bibRefCitation>
). Subsequent contact with that institution resulted in the specimen being located and borrowed by us.
<bibRefCitation id="EFBBEBF4FFA68D7FA21BFD02570993C6" author="Cassis, G. &amp; Weir, T. A." box="[342,608,692,717]" pageId="13" pageNumber="54" pagination="106 - 173" refId="ref29088" refString="Cassis, G. &amp; Weir, T. A. (1992) Scarabaeinae. In: Houston, W. W. K. (Ed.), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Uol. 9. Australian Government Publishing Service, Canberra, pp. 106 - 173" type="book chapter" year="1992">Cassis and Weir (1992)</bibRefCitation>
, following
<bibRefCitation id="EFBBEBF4FFA68D7FA1AFFD0256CC93C6" author="Matthews, E. G." box="[738,933,692,717]" pageId="13" pageNumber="54" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
, listed a
<typeStatus id="549128A7FFA68D7FA746FD02510493C7" box="[1035,1133,692,717]" pageId="13" pageNumber="54" type="holotype">holotype</typeStatus>
(probable). This specimen located in BMNH is certainly the type of
<taxonomicName id="4C2AED86FFA68D7FA1C5FD61565593FA" ID-CoL="4CC79" authorityName="Blackburn" authorityYear="1900" box="[648,828,727,752]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA68D7FA1C5FD61565593FA" box="[648,828,727,752]" italics="true" pageId="13" pageNumber="54">Panelus arthuri</emphasis>
</taxonomicName>
as evidenced by the characteristic Blackburn labels detailed above (Blackburn marked his types with a “T.” (see
<bibRefCitation id="EFBBEBF4FFA68D7FA027FD4B56B3921E" author="Lea, A. M." box="[874,986,764,788]" pageId="13" pageNumber="54" refId="ref29508" refString="Lea, A. M. (1912) The late Rev. Canon Thomas Blackburn, B. A., and his entomological work. Transactions of the Royal Society of South Australia, 36, v-xl." type="book" year="1912">Lea 1912</bibRefCitation>
)), and by the attached Matthews label indicating it was synonymous with
<emphasis id="B95E4A17FFA68D7FA162FCA956639232" box="[559,778,799,824]" italics="true" pageId="13" pageNumber="54">
<taxonomicName id="4C2AED86FFA68D7FA162FCA9566F9232" box="[559,774,799,824]" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">Panelus pygmaeus</taxonomicName>
.
</emphasis>
Blackburn did not specifically say in his description that there was only a single specimen examined so this specimen is hereby designated as the
<typeStatus id="549128A7FFA68D7FA7E2FCF2507C9257" box="[1199,1301,836,861]" pageId="13" pageNumber="54" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C2AED86FFA68D7FA60FFCF254C5928A" ID-CoL="4CC79" authority="Blackburn, 1900" authorityName="Blackburn" authorityYear="1900" class="Insecta" family="Scarabaeidae" genus="Panelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="arthuri">
<emphasis id="B95E4A17FFA68D7FA60FFCF2558F928A" italics="true" pageId="13" pageNumber="54">Panelus arthuri</emphasis>
<bibRefCitation id="EFBBEBF4FFA68D7FA3A0FCD154C5928A" author="Blackburn, T." box="[237,428,871,896]" pageId="13" pageNumber="54" pagination="206 - 233" refId="ref28969" refString="Blackburn, T. (1900) On some new genera and species of Australian Coleoptera. Proceedings of the Royal Society of Uictoria, 12, 206 - 233." type="journal article" year="1900">Blackburn, 1900</bibRefCitation>
</taxonomicName>
. The specimen has been remounted and lacks both protibiae and protarsi, which precludes the sex being determined without dissection, but exhibits enough other characters to show that it conforms to the species concept of
<emphasis id="B95E4A17FFA68D7FA226FC07572192C2" box="[363,584,945,968]" italics="true" pageId="13" pageNumber="54">
<taxonomicName id="4C2AED86FFA68D7FA226FC07572A92C2" ID-CoL="3T8VK" authorityName=": Matthews" authorityYear="1974" box="[363,579,945,968]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">Lepanus pygmaeus</taxonomicName>
.
</emphasis>
Further confirmation of the synonymy of
<bibRefCitation id="EFBBEBF4FFA68D7FA76DFC0651F792C2" author="Lea, A. M." box="[1056,1182,944,968]" pageId="13" pageNumber="54" pagination="60 - 107" refId="ref29459" refString="Lea, A. M. (1904) Descriptions of new species of Australian Coleoptera Part VIII. Proceedings of the Linnean Society of New South Wales, 29, 60 - 107. https: // doi. org / 10.5962 / bhl. part. 20155" type="journal article" year="1904">Lea (1904)</bibRefCitation>
, can be taken from the original justification that “both descriptions were drawn up from specimens taken at Kings Sound by W.W. Froggatt”. There is no reason to question this line of evidence given Leas involvement in the transfer of material to Blackburn, Blackburns reference that the material was from Macleay Museum, and that Froggatts King Sound material was deposited there.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA68D7FA38AFBD2575896B3" pageId="13" pageNumber="54" type="materials_examined">
<paragraph id="8B959605FFA68D7FA38AFBD257E494CA" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="54">
<bibRefCitation id="EFBBEBF4FFA68D7FA38AFBD254E09576" author="Matthews, E. G." box="[199,393,1124,1149]" pageId="13" pageNumber="54" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
examined almost
<specimenCount id="9D2C5D8CFFA68D7FA110FBD2566C9577" box="[605,773,1124,1149]" pageId="13" pageNumber="54" type="generic">500 specimens</specimenCount>
of
<taxonomicName id="4C2AED86FFA68D7FA063FBD356E49576" authorityName=", Matthews" authorityYear="1974" box="[814,909,1125,1148]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA68D7FA063FBD356E49576" box="[814,909,1125,1148]" italics="true" pageId="13" pageNumber="54">Lepanus</emphasis>
</taxonomicName>
during his revision but only had access to
<specimenCount id="9D2C5D8CFFA68D7FA6CDFBD2546595AA" pageId="13" pageNumber="54" type="generic">14 specimens</specimenCount>
of
<taxonomicName id="4C2AED86FFA68D7FA27FFB3F54A995AA" authorityName=": Matthews" authorityYear="1974" box="[306,448,1161,1184]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA27FFB3F54A995AA" box="[306,448,1161,1184]" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
from eight localities. Fortunately, hundreds of specimens belonging to the
<taxonomicName id="4C2AED86FFA68D7FA643FB3F50F595AA" authorityName=": Matthews" authorityYear="1974" box="[1294,1436,1161,1184]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA643FB3F50F595AA" box="[1294,1436,1161,1184]" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
species group have since been collected allowing for rigorous assessment of morphological variation within and between species. Of the material examined by Matthews, the specimen listed from Charters Towers, collected
<date id="FF94B0C5FFA68D7FA62CFB7955A69407" pageId="13" pageNumber="54" value="1965-04">April 1965</date>
by Bornemissza &amp; Yapp (25-
<date id="FF94B0C5FFA68D7FA15BFB4257079407" box="[534,622,1268,1293]" pageId="13" pageNumber="54" value="85-06-03">060385</date>
in ANIC) is
<taxonomicName id="4C2AED86FFA68D7FA1B2FB4356CC9407" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[767,933,1268,1293]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="menendezae">
<emphasis id="B95E4A17FFA68D7FA1B2FB4356CC9407" box="[767,933,1268,1293]" italics="true" pageId="13" pageNumber="54">L. menendezae</emphasis>
</taxonomicName>
, while the specimen from Beames Brook, 15 Mi SW Burketown collected
<date id="FF94B0C5FFA68D7FA2AEFAAF573F943A" box="[483,598,1304,1329]" pageId="13" pageNumber="54" value="1972-05">May 1972</date>
by Monteith &amp; Monteith (25-058521 in ANIC) is
<taxonomicName id="4C2AED86FFA68D7FA7D8FAAF5077943A" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1173,1310,1303,1328]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FFA68D7FA7D8FAAF5077943A" box="[1173,1310,1303,1328]" italics="true" pageId="13" pageNumber="54">L. guthrieae</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFBBEBF4FFA68D7FA660FAA15588945E" author="Matthews, E. G." pageId="13" pageNumber="54" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews (1974)</bibRefCitation>
noted the morphological variation of
<collectingRegion id="49EE58E7FFA68D7FA1CAFA8A5662945F" box="[647,779,1340,1365]" country="Australia" name="Queensland" pageId="13" pageNumber="54">Queensland</collectingRegion>
specimens of
<taxonomicName id="4C2AED86FFA68D7FA0E6FA8B5151945E" authorityName=": Matthews" authorityYear="1974" box="[939,1080,1341,1364]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA0E6FA8B5151945E" box="[939,1080,1341,1364]" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
and listed diagnostic characters of the species group within the remarks section. Based on these remarks, it is likely he was referring to the variation observed in these specimens from Charters Towers and Beames Brook, or the specimen from Mornington Island, which we did not examine (
<bibRefCitation id="EFBBEBF4FFA68D7FA280FA1157E894CA" author="Matthews, E. G." box="[461,641,1447,1472]" pageId="13" pageNumber="54" pagination="1 - 211" refId="ref29693" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211." type="journal article" year="1974">Matthews 1974</bibRefCitation>
).
</paragraph>
<paragraph id="8B959605FFA68D7FA38AFA7A564D9792" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="54">
Specimens from the
<collectingRegion id="49EE58E7FFA68D7FA2F2FA7A57FA94EF" box="[447,659,1484,1509]" country="Australia" name="Northern Territory" pageId="13" pageNumber="54">Northern Territory</collectingRegion>
(Fish River and Pungalina) and
<collectingRegion id="49EE58E7FFA68D7FA76CFA7A51CC94EF" box="[1057,1189,1484,1509]" country="Australia" name="Queensland" pageId="13" pageNumber="54">Queensland</collectingRegion>
(Paluma) have been sequenced and were included in the phylogenies of
<bibRefCitation id="EFBBEBF4FFA68D7FA1B4FA4656949702" author="Gunter, N. L. &amp; Weir, T. A." box="[761,1021,1519,1544]" pageId="13" pageNumber="54" pagination="201 - 243" refId="ref29214" refString="Gunter, N. L. &amp; Weir, T. A. (2017) Two new genera of Australian dung beetles (Scarabaeidae: Scarabaeinae), with the description of six new species and transfer of six described species, Zootaxa, 4290 (2), 201 - 243. https: // doi. org / 10.11646 / zootaxa. 4290.2.1" type="journal article" year="2017">Gunter &amp; Weir (2017)</bibRefCitation>
and
<bibRefCitation id="EFBBEBF4FFA68D7FA771FA4650409702" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[1084,1321,1519,1544]" pageId="13" pageNumber="54" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">
Gunter
<emphasis id="B95E4A17FFA68D7FA7D9FA4751B99702" box="[1172,1232,1519,1544]" italics="true" pageId="13" pageNumber="54">et al.</emphasis>
(2018)
</bibRefCitation>
. These
<taxonomicName id="4C2AED86FFA68D7FA6C8FA47546E9726" authorityName=": Matthews" authorityYear="1974" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA6C8FA47546E9726" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
specimens formed a clade within a larger clade containing other members of the
<taxonomicName id="4C2AED86FFA68D7FA795F9A350219726" authorityName=": Matthews" authorityYear="1974" box="[1240,1352,1557,1580]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA795F9A350219726" box="[1240,1352,1557,1580]" italics="true" pageId="13" pageNumber="54">pygmaeus</emphasis>
</taxonomicName>
group. Intraspecific variation is slightly higher than observed for other species of
<taxonomicName id="4C2AED86FFA68D7FA09EF98F515B975A" authorityName=", Matthews" authorityYear="1974" box="[979,1074,1593,1616]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA68D7FA09EF98F515B975A" box="[979,1074,1593,1616]" italics="true" pageId="13" pageNumber="54">Lepanus</emphasis>
</taxonomicName>
, however most other species are more narrowly distributed and other broadly distributed species (
<emphasis id="B95E4A17FFA68D7FA026F9EB56FA977E" box="[875,915,1629,1652]" italics="true" pageId="13" pageNumber="54">e.g.</emphasis>
,
<taxonomicName id="4C2AED86FFA68D7FA0ECF9EB5101977F" authorityName="Matthews" authorityYear="1974" box="[929,1128,1628,1653]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B95E4A17FFA68D7FA0ECF9EB5101977F" box="[929,1128,1628,1653]" italics="true" pageId="13" pageNumber="54">Lepanus australis</emphasis>
</taxonomicName>
) also exhibit similarly high levels of intraspecific variation (see
<bibRefCitation id="EFBBEBF4FFA68D7FA163F93656709792" author="Gunter, N. L. &amp; Weir, T. A." box="[558,793,1663,1688]" pageId="13" pageNumber="54" pagination="201 - 243" refId="ref29214" refString="Gunter, N. L. &amp; Weir, T. A. (2017) Two new genera of Australian dung beetles (Scarabaeidae: Scarabaeinae), with the description of six new species and transfer of six described species, Zootaxa, 4290 (2), 201 - 243. https: // doi. org / 10.11646 / zootaxa. 4290.2.1" type="journal article" year="2017">Gunter &amp; Weir 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8B959605FFA68D7FA38AF913575896B3" blockId="13.[151,1437,151,2014]" pageId="13" pageNumber="54">
<taxonomicName id="4C2AED86FFA68D7FA38AF91354CA97B6" authorityName=": Matthews" authorityYear="1974" box="[199,419,1701,1724]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA38AF91354CA97B6" box="[199,419,1701,1724]" italics="true" pageId="13" pageNumber="54">Lepanus pygmaeus</emphasis>
</taxonomicName>
has been collected in a variety of habitats including rainforest, heath, open and dry sclerophyll forests, and pasture. Most specimens have been collected in dung baited pitfall traps or at light but a handful of specimens have also been collected in mushroom baited pitfall traps suggesting they may be generalist feeders. Recently collected specimens of
<taxonomicName id="4C2AED86FFA68D7FA13FF8A756689622" authorityName=": Matthews" authorityYear="1974" box="[626,769,1809,1832]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA13FF8A756689622" box="[626,769,1809,1832]" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
from Tinaroo Waters on the northern end of the Atherton Tableland were attracted to traps baited with mealworm/cricket carrion. These specimens were collected during a survey that explored bait preferences and interestingly,
<taxonomicName id="4C2AED86FFA68D7FA045F8EF56FF967A" authorityName=": Matthews" authorityYear="1974" box="[776,918,1881,1904]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA68D7FA045F8EF56FF967A" box="[776,918,1881,1904]" italics="true" pageId="13" pageNumber="54">L. pygmaeus</emphasis>
</taxonomicName>
was only collected in the invertebrate carrion traps and not recorded from the traps baited with dung, fruit, mushroom, or vertebrate carrion (K. Ebert &amp; G.B. Monteith, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA68D7CA3DAF87254B891BA" lastPageId="14" lastPageNumber="55" pageId="13" pageNumber="54" type="description">
<paragraph id="8B959605FFA68D7FA3DAF872567296D4" blockId="13.[151,1437,151,2014]" box="[151,795,1988,2014]" pageId="13" pageNumber="54">
<emphasis id="B95E4A17FFA68D7FA3DAF872567296D4" bold="true" box="[151,795,1988,2014]" pageId="13" pageNumber="54">
<taxonomicName id="4C2AED86FFA68D7FA3DAF872571796D4" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,638,1988,2014]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="andersonorum" status="sp. nov.">
<emphasis id="B95E4A17FFA68D7FA3DAF87254D196D4" bold="true" box="[151,440,1988,2014]" italics="true" pageId="13" pageNumber="54">Lepanus andersonorum</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFFA68D7FA1C6F872567296D4" box="[651,795,1988,2014]" pageId="13" pageNumber="54" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B959605FFA58D7CA3DAFF2154B891BA" blockId="14.[151,465,151,176]" box="[151,465,151,176]" pageId="14" pageNumber="55">
(
<figureCitation id="13118A80FFA58D7CA3D2FF21546B91BA" box="[159,258,151,176]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="14" pageNumber="55">Figs. 4C</figureCitation>
,
<figureCitation id="13118A80FFA58D7CA240FF21545B91BA" box="[269,306,151,176]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1967]" captionTargetBox="[245,1336,223,1911]" captionTargetId="figure@15.[233,1355,193,1926]" captionTargetPageId="15" captionText="FIGURE 5. Habitus of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil. All specimens in comparative scale. Scale bar 1 mm." httpUri="https://zenodo.org/record/2588631/files/figure.png" pageId="14" pageNumber="55">5A</figureCitation>
,
<figureCitation id="13118A80FFA58D7CA270FF2E540891BA" box="[317,353,152,176]" captionStart="FIGURE 8" captionStartId="18.[151,250,1896,1918]" captionTargetBox="[190,1384,223,1858]" captionTargetId="figure@18.[173,1427,195,1873]" captionTargetPageId="18" captionText="FIGURE 8. Pygidial configuration of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae; F, Lepanus lentil; G, Lepanus lingziae; H, Lepanus mckenziei; I, Lepanus menendezae. All specimens in comparative scale. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/2588637/files/figure.png" pageId="14" pageNumber="55">8A</figureCitation>
,
<figureCitation id="13118A80FFA58D7CA220FF2E54C991BA" box="[365,416,152,176]" captionStart="FIGURE 10" captionStartId="23.[151,250,1909,1931]" captionTargetBox="[167,1333,208,1895]" captionTargetId="figure@23.[164,1428,193,1895]" captionTargetPageId="23" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus pygmaeus species group. A, Lepanus andersonorum; B, Lepanus crenidens; C, Lepanus feehani; D, Lepanus gubara; E, Lepanus guthrieae. Not in comparative scale, given size difference aedegi are roughly grouped in three size classes as per lower left schematic and scaled accordingly. Size class indicated by shaded silhouette and 0.2 mm scale bar." httpUri="https://zenodo.org/record/2588641/files/figure.png" pageId="14" pageNumber="55">10A</figureCitation>
,
<figureCitation id="13118A80FFA58D7CA2E1FF2E54A091BA" box="[428,457,152,176]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="14" pageNumber="55">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA58D7CA3DAFF69506093A2" pageId="14" pageNumber="55" type="materials_examined">
<paragraph id="8B959605FFA58D7CA3DAFF69506093A2" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">
<emphasis id="B95E4A17FFA58D7CA3DAFF6954C791F2" bold="true" box="[151,430,223,248]" pageId="14" pageNumber="55">
Type series.
<materialsCitation id="3B429C58FFA58D7CA277FF6954C791F2" ID-GBIF-Occurrence="2012745355" box="[314,430,223,248]" collectingDate="1975-07-04" collectingDateMax="2003-05-23" collectingDateMin="1975-07-04" collectionCode="ANIC, ABRS, QM" collectorName="T. A. Weir &amp; I. D. Naumann &amp; T. McLeod &amp; G. B. Monteith" county="Area" determinerName="T. A. Weir" latitude="-11.85" location="Cape" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" pageId="14" pageNumber="55" specimenCode="CYP7" specimenCount="1" specimenCount-male="1" stateProvince="York" typeStatus="holotype">
<typeStatus id="549128A7FFA58D7CA277FF6954C091F2" box="[314,425,223,248]" pageId="14" pageNumber="55" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFFA58D7CA2F2FF5654BD91FD" box="[447,468,224,247]" pageId="14" pageNumber="55" type="male"></specimenCount>
<locationDeviation id="9A278F27FFA58D7CA2A2FF69566F91F2" box="[495,774,223,248]" location="Heathlands" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA2A2FF69575891F2" box="[495,561,223,248]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="55" unit="km" value="12.0">12km</quantity>
SSE Heathlands
</locationDeviation>
,
<date id="FF94B0C5FFA58D7CA056FF6956DE91F2" box="[795,951,223,248]" pageId="14" pageNumber="55" value="1992-01-15">
<collectingDate id="EFD0492DFFA58D7CA056FF6956DE91F2" box="[795,951,223,248]" pageId="14" pageNumber="55" value="1992-01-15">15 Jan 1992</collectingDate>
</date>
<date id="FF94B0C5FFA58D7CA08AFF69510C91F2" box="[967,1125,223,248]" pageId="14" pageNumber="55" value="1992-01-26">
<collectingDate id="EFD0492DFFA58D7CA08AFF69510C91F2" box="[967,1125,223,248]" pageId="14" pageNumber="55" value="1992-01-26">26 Jan 1992</collectingDate>
</date>
, Weir,T.A.” [
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,
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] / “
<collectingMethod id="526BEE12FFA58D7CA21FFEB2577F9016" box="[338,534,260,285]" pageId="14" pageNumber="55">human dung trap</collectingMethod>
closed forest” / “ANIC Database No. 25-023701” / “ANIC Image” / “
<taxonomicName id="4C2AED86FFA58D7CA676FEB250F59016" authorityName=", Matthews" authorityYear="1974" box="[1339,1436,260,284]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
<specimenCode id="DB8C3E7EFFA58D7CA3DAFE9E55B2904A" box="[151,219,296,320]" collectionCode="CYP" country="Cyprus" lsid="urn:lsid:biocol.org:col:13586" name="Ministry of Agriculture, Natural Resources and Environment" pageId="14" pageNumber="55" type="Herbarium">CYP7</specimenCode>
det.
<determinerName id="7E497AC1FFA58D7CA25BFE9E54E2904A" box="[278,395,295,320]" pageId="14" pageNumber="55">T.A. Weir</determinerName>
2013” in ANIC” in ANIC.
<emphasis id="B95E4A17FFA58D7CA19EFE915638904A" bold="true" box="[723,849,295,320]" pageId="14" pageNumber="55">
<typeStatus id="549128A7FFA58D7CA19EFE915625904A" box="[723,844,295,320]" pageId="14" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<locationDeviation id="9A278F27FFA58D7CA017FE9E5109904A" box="[858,1120,295,320]" location="Heathlands" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA017FE9E56C8904A" box="[858,929,295,320]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="55" unit="km" value="12.0">12 km</quantity>
SSE Heathlands
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA739FE91519B904A" box="[1140,1266,295,320]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.85">11°51'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA64DFE9E50F8904A" box="[1280,1425,296,320]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.63333">142°38'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA3DAFEFA5443906F" box="[151,298,332,357]" pageId="14" pageNumber="55" value="1992-01-15" valueMax="1992-01-26" valueMin="1992-01-15">
<collectingDate id="EFD0492DFFA58D7CA3DAFEFA5443906F" box="[151,298,332,357]" pageId="14" pageNumber="55" value="1992-01-15" valueMax="1992-01-26" valueMin="1992-01-15">1526.i.1992</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA275FEFA54CE906F" box="[312,423,332,357]" pageId="14" pageNumber="55">T.A. Weir</collectorName>
(25-023701
<date id="FF94B0C5FFA58D7CA176FEFA573E906E" box="[571,599,332,356]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA176FEFA573E906E" box="[571,599,332,356]" pageId="14" pageNumber="55">x2</collectingDate>
</date>
♂,
<date id="FF94B0C5FFA58D7CA1CBFEFA57C8906E" box="[646,673,332,356]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA1CBFEFA57C8906E" box="[646,673,332,356]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
sex undetermined in ANIC);
<locationDeviation id="9A278F27FFA58D7CA0B5FEFA506A906F" box="[1016,1283,332,357]" location="Heathlands" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA0B5FEFA5128906F" box="[1016,1089,332,357]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="55" unit="km" value="12.0">12 km</quantity>
SSE Heathlands
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA655FEFA50FE906F" box="[1304,1431,332,357]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.85">11°51'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA3DAFEC6544E9082" box="[151,295,368,392]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.63333">142°38'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA271FED954CF9082" box="[316,422,367,392]" pageId="14" pageNumber="55" value="1992-01-20">
<collectingDate id="EFD0492DFFA58D7CA271FED954CF9082" box="[316,422,367,392]" pageId="14" pageNumber="55" value="1992-01-20">20.i.1992</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA2F9FEC6574A9082" box="[436,547,367,392]" pageId="14" pageNumber="55">T.A. Weir</collectorName>
&amp;
<collectorName id="26DFF3D3FFA58D7CA100FEC65784908D" box="[589,749,368,391]" pageId="14" pageNumber="55">I.D. Naumann</collectorName>
(25-023700
<date id="FF94B0C5FFA58D7CA0CCFEC656F59082" box="[897,924,368,392]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA0CCFEC656F59082" box="[897,924,368,392]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂,
<date id="FF94B0C5FFA58D7CA081FEC6568E9082" box="[972,999,368,392]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA081FEC6568E9082" box="[972,999,368,392]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♀ in ANIC);
<locationDeviation id="9A278F27FFA58D7CA7C3FEC650F19082" box="[1166,1432,367,392]" location="Heathlands" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA7C3FEC651BF9082" box="[1166,1238,367,392]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="55" unit="km" value="12.0">12 km</quantity>
SSE Heathlands
</locationDeviation>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA3D3FE22547490A7" box="[158,285,404,429]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.85">11°51'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA262FE2254D690A6" box="[303,447,404,428]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.63333">142°38'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA29AFE2257FA90A7" box="[471,659,404,429]" pageId="14" pageNumber="55" value="1992-03-22" valueMax="1992-04-25" valueMin="1992-03-22">
<collectingDate id="EFD0492DFFA58D7CA29AFE2257FA90A7" box="[471,659,404,429]" pageId="14" pageNumber="55" value="1992-03-22" valueMax="1992-04-25" valueMin="1992-03-22">22.iii25.iv.1992</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA1E9FE23564A90A7" box="[676,803,404,429]" pageId="14" pageNumber="55">T. McLeod</collectorName>
(25-023703
<date id="FF94B0C5FFA58D7CA0F1FE2256BE90A6" box="[956,983,404,428]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA0F1FE2256BE90A6" box="[956,983,404,428]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂,
<date id="FF94B0C5FFA58D7CA740FE22514190A6" box="[1037,1064,404,428]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA740FE22514190A6" box="[1037,1064,404,428]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♀ in ANIC); “ABRS
<collectingCounty id="62F4EE89FFA58D7CA674FE22500790A6" box="[1337,1390,404,428]" pageId="14" pageNumber="55">Area</collectingCounty>
2”,
<collectingMunicipality id="6BF10C7FFFA58D7CA3DAFE01542390DA" box="[151,330,439,464]" pageId="14" pageNumber="55">Dividing Range</collectingMunicipality>
,
<locationDeviation id="9A278F27FFA58D7CA21AFE0157A590DA" box="[343,716,439,464]" location="Captain Billy Creek" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA21AFE0154F790DA" box="[343,414,439,464]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="55" unit="km" value="15.0">15 km</quantity>
W of Captain Billy Creek
</locationDeviation>
,
<location id="8EF5C0DEFFA58D7CA194FE0E567890DA" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA194FE0E567890DA" box="[729,785,440,464]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Cape" pageId="14" pageNumber="55" stateProvince="York">Cape</location>
<collectingRegion id="49EE58E7FFA58D7CA057FE01563990DA" box="[794,848,439,464]" country="United Kingdom" name="York" pageId="14" pageNumber="55">York</collectingRegion>
<location id="8EF5C0DEFFA58D7CA014FE0156AF90DA" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA014FE0156AF90DA" box="[857,966,439,464]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Peninsula" pageId="14" pageNumber="55" stateProvince="York">Peninsula</location>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA097FE0E513090DA" box="[986,1113,440,464]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.666667">11°40'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA72AFE01519E90DA" box="[1127,1271,439,464]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.75">142°45'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA646FE0150F190DA" box="[1291,1432,439,464]" pageId="14" pageNumber="55" value="1975-07-04" valueMax="1975-07-09" valueMin="1975-07-04">
<collectingDate id="EFD0492DFFA58D7CA646FE0150F190DA" box="[1291,1432,439,464]" pageId="14" pageNumber="55" value="1975-07-04" valueMax="1975-07-09" valueMin="1975-07-04">49.vii.1975</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA3DAFE6A545A90FF" box="[151,307,476,501]" pageId="14" pageNumber="55">G.B. Monteith</collectorName>
(25-015558
<date id="FF94B0C5FFA58D7CA2F2FE6A54B390FE" box="[447,474,476,500]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA2F2FE6A54B390FE" box="[447,474,476,500]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♀ in QM also as T50228); “ABRS
<location id="8EF5C0DEFFA58D7CA02FFE6A56FE90FE" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA02FFE6A56FE90FE" box="[866,919,476,500]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Area" pageId="14" pageNumber="55" stateProvince="York">Area</location>
3”,
<location id="8EF5C0DEFFA58D7CA088FE6A511F90FF" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA088FE6A511F90FF" box="[965,1142,476,501]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Dividing Range" pageId="14" pageNumber="55" stateProvince="York">Dividing Range</location>
,
<locationDeviation id="9A278F27FFA58D7CA7CDFE6A55B29312" location="Captain Billy Creek" pageId="14" pageNumber="55">
<quantity id="4CD23BE0FFA58D7CA7CDFE6A51AC90FF" box="[1152,1221,476,501]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="55" unit="km" value="15.0">15 km</quantity>
W of Captain Billy Creek
</locationDeviation>
,
<location id="8EF5C0DEFFA58D7CA3AAFDB654769312" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA3AAFDB654769312" box="[231,287,512,536]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Cape" pageId="14" pageNumber="55" stateProvince="York">Cape</location>
<collectingRegion id="49EE58E7FFA58D7CA265FE4954379312" box="[296,350,511,536]" country="United Kingdom" name="York" pageId="14" pageNumber="55">York</collectingRegion>
<location id="8EF5C0DEFFA58D7CA22BFE4954BA9312" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA22BFE4954BA9312" box="[358,467,511,536]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Peninsula" pageId="14" pageNumber="55" stateProvince="York">Peninsula</location>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA2AAFDB6570F9312" box="[487,614,512,536]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.666667">11°40'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA13EFE49566A9312" box="[627,771,511,536]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.75">142°45'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA05AFE4956CA9312" box="[791,931,511,536]" pageId="14" pageNumber="55" value="1975-07-04" valueMax="1975-07-09" valueMin="1975-07-04">
<collectingDate id="EFD0492DFFA58D7CA05AFE4956CA9312" box="[791,931,511,536]" pageId="14" pageNumber="55" value="1975-07-04" valueMax="1975-07-09" valueMin="1975-07-04">49.vii.1975</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA0FDFDB651279312" box="[944,1102,511,536]" pageId="14" pageNumber="55">G.B. Monteith</collectorName>
(25-015556
<date id="FF94B0C5FFA58D7CA793FDB651909312" box="[1246,1273,512,536]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA793FDB651909312" box="[1246,1273,512,536]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂, 25-015622
<date id="FF94B0C5FFA58D7CA3DAFD9255DB9336" box="[151,178,548,572]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA3DAFD9255DB9336" box="[151,178,548,572]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂,
<date id="FF94B0C5FFA58D7CA3A9FD9255969336" box="[228,255,548,572]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA3A9FD9255969336" box="[228,255,548,572]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
damaged in QM also as T50226, T49910T49911); “ABRS
<location id="8EF5C0DEFFA58D7CA08FFD92569E9336" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA08FFD92569E9336" box="[962,1015,548,572]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Area" pageId="14" pageNumber="55" stateProvince="York">Area</location>
6”,
<location id="8EF5C0DEFFA58D7CA763FD92507D9337" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA763FD92507D9337" box="[1070,1300,548,573]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Captain Billy Creek" pageId="14" pageNumber="55" stateProvince="York">Captain Billy Creek</location>
,
<location id="8EF5C0DEFFA58D7CA66EFD9250329336" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA66EFD9250329336" box="[1315,1371,548,572]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Cape" pageId="14" pageNumber="55" stateProvince="York">Cape</location>
<collectingRegion id="49EE58E7FFA58D7CA62AFD9250F59337" box="[1383,1436,548,573]" country="United Kingdom" name="York" pageId="14" pageNumber="55">York</collectingRegion>
<location id="8EF5C0DEFFA58D7CA3DAFDF1546D936A" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA3DAFDF1546D936A" box="[151,260,583,608]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Peninsula" pageId="14" pageNumber="55" stateProvince="York">Peninsula</location>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA256FDFE54F2936A" box="[283,411,584,608]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.666667">11°40'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA2E1FDF15755936A" box="[428,572,583,608]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.83333">142°50'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA11EFDF15787936A" box="[595,750,583,608]" pageId="14" pageNumber="55" value="1975-07-09" valueMax="1975-07-13" valueMin="1975-07-09">
<collectingDate id="EFD0492DFFA58D7CA11EFDF15787936A" box="[595,750,583,608]" pageId="14" pageNumber="55" value="1975-07-09" valueMax="1975-07-13" valueMin="1975-07-09">913.vii.1975</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA1B3FDFE56C9936A" box="[766,928,583,608]" pageId="14" pageNumber="55">G.B. Monteith</collectorName>
(25-015557
<date id="FF94B0C5FFA58D7CA775FDFE513A936A" box="[1080,1107,584,608]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA775FDFE513A936A" box="[1080,1107,584,608]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♀ in QM also as T50227);
<location id="8EF5C0DEFFA58D7CA3DAFDDA547C938F" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA3DAFDDA547C938F" box="[151,277,620,645]" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Heathlands" pageId="14" pageNumber="55" stateProvince="York">Heathlands</location>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA264FDDA54C1938F" box="[297,424,620,645]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-11.75">11°45'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA2FBFDDA572E938F" box="[438,583,620,645]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.58333">142°35'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA116FDDA57AC938F" box="[603,709,620,645]" pageId="14" pageNumber="55" value="1992-01-22">
<collectingDate id="EFD0492DFFA58D7CA116FDDA57AC938F" box="[603,709,620,645]" pageId="14" pageNumber="55" value="1992-01-22">22.i.1992</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA19EFDDA5629938F" box="[723,832,620,645]" pageId="14" pageNumber="55">T.A. Weir</collectorName>
&amp;
<collectorName id="26DFF3D3FFA58D7CA024FDDB5161938E" box="[873,1032,621,644]" pageId="14" pageNumber="55">I.D. Naumann</collectorName>
(25-023702
<date id="FF94B0C5FFA58D7CA7D7FDDA51DC938E" box="[1178,1205,620,644]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA7D7FDDA51DC938E" box="[1178,1205,620,644]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂ in ANIC);
<location id="8EF5C0DEFFA58D7CA614FDDA541293A2" LSID="urn:lsid:plazi:treatment:03832713FFA28D7CA3DAF944567297EA:8EF5C0DEFFA58D7CA614FDDA541293A2" county="Area" latitude="-11.85" longLatPrecision="21" longitude="142.63333" municipality="Dividing Range" name="Green Hill Thursday Island" pageId="14" pageNumber="55" stateProvince="York">Green Hill Thursday Island</location>
, [
<geoCoordinate id="EE1EF0C2FFA58D7CA2C3FD39576793A2" box="[398,526,655,680]" direction="south" orientation="latitude" pageId="14" pageNumber="55" precision="15" value="-10.583333">10°35'00&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FFA58D7CA154FD2657C093A2" box="[537,681,656,680]" direction="east" orientation="longitude" pageId="14" pageNumber="55" precision="15" value="142.21666">142°13'00&quot;E</geoCoordinate>
],
<date id="FF94B0C5FFA58D7CA1F1FD26563B93A2" box="[700,850,656,680]" pageId="14" pageNumber="55" value="2003-05-20" valueMax="2003-05-23" valueMin="2003-05-20">
<collectingDate id="EFD0492DFFA58D7CA1F1FD26563B93A2" box="[700,850,656,680]" pageId="14" pageNumber="55" value="2003-05-20" valueMax="2003-05-23" valueMin="2003-05-20">2023.v.2003</collectingDate>
</date>
,
<collectorName id="26DFF3D3FFA58D7CA013FD26569393A2" box="[862,1018,655,680]" pageId="14" pageNumber="55">G.B. Monteith</collectorName>
(T169468
<date id="FF94B0C5FFA58D7CA73EFD2651E793A2" box="[1139,1166,656,680]" pageId="14" pageNumber="55">
<collectingDate id="EFD0492DFFA58D7CA73EFD2651E793A2" box="[1139,1166,656,680]" pageId="14" pageNumber="55">x1</collectingDate>
</date>
♂ in QM).
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA58D7CA38AFD0257A694EF" pageId="14" pageNumber="55" type="description">
<paragraph id="8B959605FFA58D7CA38AFD0251A193C7" blockId="14.[151,1438,223,1760]" box="[199,1224,692,717]" pageId="14" pageNumber="55">
<emphasis id="B95E4A17FFA58D7CA38AFD02543093C7" bold="true" box="[199,345,692,717]" pageId="14" pageNumber="55">Description.</emphasis>
Head and pronotum dark brown and elytra light brown. Antennal clubs white.
</paragraph>
<paragraph id="8B959605FFA58D7CA38AFD61519B93E5" blockId="14.[151,1438,223,1760]" box="[199,1266,727,752]" pageId="14" pageNumber="55">
Total length:
<quantity id="4CD23BE0FFA58D7CA211FD6E548F93E5" box="[348,486,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="14" pageNumber="55" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
.
<typeStatus id="549128A7FFA58D7CA2BDFD61573193FA" box="[496,600,727,752]" pageId="14" pageNumber="55" type="holotype">Holotype</typeStatus>
measurements: Total length 2.0 mm, elytral width
<quantity id="4CD23BE0FFA58D7CA7D8FD6E518793E5" box="[1173,1262,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="55" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FFA58D7CA38AFD4A51DD928A" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">Male: Head: Broad U-shaped between clypeal teeth, which are short and very slightly upturned. Head reticulate between clypeal suture and anterior edge. Margin slightly angulate at clypeogenal suture. Genal angle rounded. Basal carina absent. Dorsal part of eyes moderate in size, separated by interocular space approximately 10 times eye width (29:3); eye canthus nearly dividing eye. Head 1.25 × wider than long (35:28).</paragraph>
<paragraph id="8B959605FFA58D7CA38AFC3A519E92C2" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">Prothorax: Basal margin slightly angulate. Hypomeral surface smooth with light punctation and long setae. Hypomeral striae about half the length of the hypomeron. Pronotum 1.75 × wider than long (56:32).</paragraph>
<paragraph id="8B959605FFA58D7CA38AFC6256FF9552" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">Elytra: Surface reticulate at apical half and at the sides. Intervals finely punctate, and with very fine, straight setae longer than the space between punctures. Elytral striae distinct but slightly impressed on apical half. Striae 1 7 double with very fine, punctate edges. All striae on disc equal width. 8th striae almost extends to the apex of the elytra forming a slight flange. Elytra 1.05 × wider than long (68:65).</paragraph>
<paragraph id="8B959605FFA58D7CA38AFBD350F795AA" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">Legs: Front edge of protibiae deeply angulate; apical digit somewhat elongate and sharp; two teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws minute and simple.</paragraph>
<paragraph id="8B959605FFA58D7CA38AFB1A566B943A" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">Abdomen: Pygidium with U-shaped depression containing a faint tubercle that is a raised line but may be absent. Depression occupies medial third of pygidial disc and extends under the angulate upper edge of pygidium. Edges of depression are only moderately sharp, bottom edge is slightly pointed. Abdominal surface reticulate right across. Segment 6 slightly gibbous and finely punctate.</paragraph>
<paragraph id="8B959605FFA58D7CA38AFA8A518294CA" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">
Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite punctate with fine reticulation. Meso-metaventral suture very slightly arched but with a fine short carina extending backwards from middle of meso-metaventral suture as in
<figureCitation id="13118A80FFA58D7CA768FA3251149496" box="[1061,1149,1412,1437]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="14" pageNumber="55">Fig. 4C</figureCitation>
. Mesoventrite with slight notch anteriorly. Mesoventrite smooth. Mesepimeron finely reticulate. Metanepisternum reticulate.
</paragraph>
<paragraph id="8B959605FFA58D7CA38AFA7A57A694EF" blockId="14.[151,1438,223,1760]" box="[199,719,1484,1509]" pageId="14" pageNumber="55">
Female: As per the
<taxonomicName id="4C2AED86FFA58D7CA2EFFA7B575994EE" authorityName=": Matthews" authorityYear="1974" box="[418,560,1485,1508]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FFA58D7CA2EFFA7B54D894EE" box="[418,433,1485,1508]" italics="true" pageId="14" pageNumber="55">L</emphasis>
.
<emphasis id="B95E4A17FFA58D7CA28DFA7B575994EE" box="[448,560,1485,1508]" italics="true" pageId="14" pageNumber="55">pygmaeus</emphasis>
</taxonomicName>
species group
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA58D7CA38AFA5951F69727" pageId="14" pageNumber="55" type="etymology">
<paragraph id="8B959605FFA58D7CA38AFA5951F69727" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">
<emphasis id="B95E4A17FFA58D7CA38AFA5954279702" bold="true" box="[199,334,1519,1544]" pageId="14" pageNumber="55">Etymology.</emphasis>
Named for A. Chace and Josie Anderson for their generous support of The Cleveland Museum of Natural History for this research, and encouragement of women pursuing careers in science.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA58D7CA38AF981507F977F" pageId="14" pageNumber="55" type="distribution">
<paragraph id="8B959605FFA58D7CA38AF981507F977F" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">
<emphasis id="B95E4A17FFA58D7CA38AF9815409975A" bold="true" box="[199,352,1591,1616]" pageId="14" pageNumber="55">Distribution.</emphasis>
Known only from closed forests of northern Cape
<collectingRegion id="49EE58E7FFA58D7CA0D5F98156A6975A" box="[920,975,1591,1616]" country="United Kingdom" name="York" pageId="14" pageNumber="55">York</collectingRegion>
Peninsula and Thursday Island (
<figureCitation id="13118A80FFA58D7CA671F98150E6975A" box="[1340,1423,1591,1616]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="14" pageNumber="55">Fig. 14</figureCitation>
). So far only
<specimenCount id="9D2C5D8CFFA58D7CA257F9EA54DB977F" box="[282,434,1628,1653]" pageId="14" pageNumber="55" type="generic">15 specimens</specimenCount>
are known and the species may be more widespread than the CYP bioregion.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFFA58D7CA38AF9C9567297EA" pageId="14" pageNumber="55" type="discussion">
<paragraph id="8B959605FFA58D7CA38AF9C9567297EA" blockId="14.[151,1438,223,1760]" pageId="14" pageNumber="55">
<emphasis id="B95E4A17FFA58D7CA38AF9C954269792" bold="true" box="[199,335,1663,1688]" pageId="14" pageNumber="55">Comments.</emphasis>
<taxonomicName id="4C2AED86FFA58D7CA211F93757059792" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[348,620,1663,1688]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="species" species="andersonorum">
<emphasis id="B95E4A17FFA58D7CA211F93757059792" box="[348,620,1663,1688]" italics="true" pageId="14" pageNumber="55">Lepanus andersonorum</emphasis>
</taxonomicName>
was previously referred to as “
<taxonomicName id="4C2AED86FFA58D7CA0BEF937513B9792" authorityName=", Matthews" authorityYear="1974" box="[1011,1106,1665,1688]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FFA58D7CA0BEF937513B9792" box="[1011,1106,1665,1688]" italics="true" pageId="14" pageNumber="55">Lepanus</emphasis>
</taxonomicName>
CYP7” under the informal nomenclature sensu
<bibRefCitation id="EFBBEBF4FFA58D7CA236F913573F97B6" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[379,598,1700,1725]" pageId="14" pageNumber="55" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FFA58D7CA286F913576A97B7" box="[459,515,1700,1725]" italics="true" pageId="14" pageNumber="55">et al.</emphasis>
(2011)
</bibRefCitation>
. Specimens have been collected using a variety of methods including dung baited pitfall traps, berlesed leaf-litter and intercept traps.
</paragraph>
</subSubSection>
</treatment>
</document>