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<document id="B7EFF406A1BA7D350E7A91D89C2CFFFD" ID-CLB-Dataset="8640" ID-DOI="10.11646/zootaxa.4909.1.1" ID-GBIF-Dataset="fa031765-b474-48aa-9d7e-b62dd092cfa4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4448697" ID-ZooBank="92ECCF97-7315-4A1A-A94A-537D0F331EBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611041096358" checkinUser="plazi" docAuthor="Skartveit, John &amp; Wedmann, Sonja" docDate="2021" docId="035E87B2FFBDFF90FF023346FB45D2C3" docLanguage="en" docName="zootaxa.4909.1.1.pdf" docOrigin="Zootaxa 4909 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Penthetria aestimata Statz. LACMIP 1943" docType="treatment" docVersion="12" lastPageNumber="15" masterDocId="FF67FFCAFFB0FF9EFF953571FFBBD142" masterDocTitle="A Revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany" masterLastPageNumber="77" masterPageNumber="1" pageNumber="14" updateTime="1698924763774" updateUser="ExternalLinkService">
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<mods:title id="1A5FB114E81F239263C900687A3CB2AA">A Revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany</mods:title>
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<mods:namePart id="020E0986FFF492D2401F98F7338BE909">Skartveit, John</mods:namePart>
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<mods:namePart id="8EC30884D76277132C6A0F0E26674E09">Wedmann, Sonja</mods:namePart>
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<mods:title id="92D452DC085B39A775A38EFC9D74145F">Zootaxa</mods:title>
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<treatment id="035E87B2FFBDFF90FF023346FB45D2C3" ID-DOI="http://doi.org/10.5281/zenodo.4464928" ID-GBIF-Taxon="177132508" ID-Zenodo-Dep="4464928" LSID="urn:lsid:plazi:treatment:035E87B2FFBDFF90FF023346FB45D2C3" httpUri="http://treatment.plazi.org/id/035E87B2FFBDFF90FF023346FB45D2C3" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C3ED652FFFBDFF93FF023346FD40D710" box="[151,763,1591,1618]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B4836A4FFBDFF93FF023346FD40D710" blockId="13.[151,763,1591,1618]" box="[151,763,1591,1618]" pageId="13" pageNumber="14">
<heading id="D00081C8FFBDFF93FF023346FD40D710" bold="true" box="[151,763,1591,1618]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B983EAB6FFBDFF93FF023346FD40D710" bold="true" box="[151,763,1591,1618]" pageId="13" pageNumber="14">
<taxonomicName id="4CF74D27FFBDFF93FF023346FD99D710" ID-CoL="8D8K3" authority="Statz, 1943" authorityName="Statz. LACMIP" authorityYear="1943" box="[151,546,1591,1618]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aestimata">
<emphasis id="B983EAB6FFBDFF93FF023346FE29D710" bold="true" box="[151,402,1591,1618]" italics="true" pageId="13" pageNumber="14">Penthetria aestimata</emphasis>
<bibRefCitation id="EF664B55FFBDFF93FE0C3349FD99D710" author="Statz, G." box="[409,546,1592,1618]" pageId="13" pageNumber="14" pagination="1 - 65" refId="ref51587" refString="Statz, G. (1943) Neue Dipteren (Nematocera) aus dem Oberoligozan von Rott. I. Familie Bibionidae (Haarmucken). Palaeontographica, Series A, 95, 1 - 65." type="journal article" year="1943">Statz, 1943</bibRefCitation>
</taxonomicName>
(figs. 67, 9194)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFBDFF93FF02330DFEC2D79F" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8B4836A4FFBDFF93FF02330DFDAAD7D7" blockId="13.[151,529,1660,1685]" box="[151,529,1660,1685]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="ABE7118AFFBDFF93FF02330DFDAAD7D7" box="[151,529,1660,1685]" pageId="13" pageNumber="14">
<treatmentCitation id="0A5610B5FFBDFF93FF02330DFDB6D7D7" author="Statz, G." box="[151,525,1660,1685]" page="39" pageId="13" pageNumber="14" year="1943">
<taxonomicName id="4CF74D27FFBDFF93FF02330DFDB6D7D7" ID-CoL="8D8K3" authority="Statz, 1943: 39" authorityName="Statz. LACMIP" authorityPageNumber="39" authorityYear="1943" box="[151,525,1660,1685]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aestimata">
<emphasis id="B983EAB6FFBDFF93FF02330DFED0D7D6" box="[151,363,1660,1684]" italics="true" pageId="13" pageNumber="14">Penthetria aestimata</emphasis>
<bibRefCitation id="EF664B55FFBDFF93FEE4330CFDB6D7D7" author="Statz, G." box="[369,525,1661,1685]" pageId="13" pageNumber="14" pagination="1 - 65" refId="ref51587" refString="Statz, G. (1943) Neue Dipteren (Nematocera) aus dem Oberoligozan von Rott. I. Familie Bibionidae (Haarmucken). Palaeontographica, Series A, 95, 1 - 65." type="journal article" year="1943">Statz, 1943: 39</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B4836A4FFBDFF93FF5233B3FEC2D79F" blockId="13.[151,1436,1730,2009]" box="[199,377,1730,1757]" pageId="13" pageNumber="14">
<emphasis id="B983EAB6FFBDFF93FF5233B3FE81D79E" bold="true" box="[199,314,1730,1756]" pageId="13" pageNumber="14">Localities</emphasis>
: Rott
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFBDFF93FF523397FAF8D608" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8B4836A4FFBDFF93FF523397FCAED667" blockId="13.[151,1436,1730,2009]" pageId="13" pageNumber="14">
<emphasis id="B983EAB6FFBDFF93FF523397FE12D642" bold="true" box="[199,425,1766,1792]" pageId="13" pageNumber="14">Material examined</emphasis>
:
<typeStatus id="544C8806FFBDFF93FE2D3396FDA4D643" box="[440,543,1767,1793]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
LACMIP 2533.249 / LACMIP Type 3379 (female) (fig. 6), LACMIP 2533.250 / LACMIP Type 3380 (male, part and counterpart, fig. 7)
</paragraph>
<paragraph id="8B4836A4FFBDFF93FF52325EFAF8D608" blockId="13.[151,1436,1730,2009]" box="[199,1347,1838,1866]" pageId="13" pageNumber="14">
Additional material: LACMIP 2533.251/LACMIP
<typeStatus id="544C8806FFBDFF93FC95325EFC83D60B" box="[768,824,1839,1865]" pageId="13" pageNumber="14">Type</typeStatus>
3381 (female), LACMIP 2533.1260ab (male).
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFBDFF90FF523223FB45D2C3" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8B4836A4FFBDFF93FF523223FEE1D62F" blockId="13.[151,1436,1730,2009]" box="[199,346,1874,1901]" pageId="13" pageNumber="14">
<emphasis id="B983EAB6FFBDFF93FF523223FEEAD62E" bold="true" box="[199,337,1874,1900]" pageId="13" pageNumber="14">Description</emphasis>
:
</paragraph>
<paragraph id="8B4836A4FFBDFF93FF523206FD84D6D3" blockId="13.[151,1436,1730,2009]" box="[199,575,1910,1937]" pageId="13" pageNumber="14">Female (N=2, figs. 6, 91, 9394):</paragraph>
<paragraph id="8B4836A4FFBDFF93FF5232EAFDB8D6F7" blockId="13.[151,1436,1730,2009]" box="[199,515,1947,1973]" pageId="13" pageNumber="14">
Total length
<quantity id="4C0F9B41FFBDFF93FEC132EAFE0CD6F7" box="[340,439,1947,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="13" pageNumber="14" unit="mm" value="11.2">11.2 mm</quantity>
(N=1)
</paragraph>
<paragraph id="8B4836A4FFBDFF90FF5232CEFF60D197" blockId="13.[151,1436,1730,2009]" lastBlockId="14.[151,1437,151,897]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Head (fig. 93): Width
<quantity id="4C0F9B41FFBDFF93FE5132CEFDA5D69B" box="[452,542,1983,2009]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="14" unit="mm" value="1.2">1.2 mm</quantity>
. Brownish-black, rather short and wide. Antenna nearly as long as head width, flagellum 1.0 mm long,
<quantity id="4C0F9B41FFBEFF90FE2635E6FDA1D1F3" box="[435,538,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="mm" value="0.23">0.23 mm</quantity>
wide, conical, 8-segmented, ultimate flagellomere small. Complex eye rather small.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF5235AEFB6ED1BB" blockId="14.[151,1437,151,897]" box="[199,1237,223,249]" pageId="14" pageNumber="15">
Thorax: Length
<quantity id="4C0F9B41FFBEFF90FEE835AEFE6ED1BB" box="[381,469,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="15" unit="mm" value="2.9">2.9 mm</quantity>
, width
<quantity id="4C0F9B41FFBEFF90FDBD35AEFD3AD1BB" box="[552,641,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="15" unit="mm" value="2.2">2.2 mm</quantity>
. Brownish-black, humeri brown, haltere light brown.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF523472FDAED003" blockId="14.[151,1437,151,897]" pageId="14" pageNumber="15">
Legs: Brown, slender. Fore femur
<quantity id="4C0F9B41FFBEFF90FDDF3472FD1AD05F" box="[586,673,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="mm" value="2.3">2.3 mm</quantity>
long,
<quantity id="4C0F9B41FFBEFF90FD7D3472FCF6D05F" box="[744,845,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="15" unit="mm" value="0.38">0.38 mm</quantity>
wide, hind femur
<quantity id="4C0F9B41FFBEFF90FB893472FBC8D05F" box="[1052,1139,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="15" unit="mm" value="2.9">2.9 mm</quantity>
long,
<quantity id="4C0F9B41FFBEFF90FB2F3472FAA4D05F" box="[1210,1311,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="15" unit="mm" value="0.44">0.44 mm</quantity>
wide, hind tibia
<quantity id="4C0F9B41FFBEFF90FF453457FE9DD003" box="[208,294,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="mm" value="3.5">3.5 mm</quantity>
long,
<quantity id="4C0F9B41FFBEFF90FEF93456FE6BD003" box="[364,464,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="0.3">0.30 mm</quantity>
wide.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF52343AFBE0D37F" blockId="14.[151,1437,151,897]" pageId="14" pageNumber="15">
Wing (fig. 91): Length
<quantity id="4C0F9B41FFBEFF90FE58343AFD9ED027" box="[461,549,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="15" unit="mm" value="9.8">9.8 mm</quantity>
, width
<quantity id="4C0F9B41FFBEFF90FDED343AFD6BD027" box="[632,720,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="15" unit="mm" value="3.4">3.4 mm</quantity>
, length/width = 2.9. Oval overall shape, light brownish fumose, veins brown throughout, distinctive, R-veins strong, posterior veins considerably finer. Pterostigma oval, brown, not highly distinctive. Costal cell slightly darkened. Humeral vein present, oblique. Subcosta prominent, long and straight. R
<subScript id="177334E1FFBEFF90FE9E34B8FE9FD095" attach="left" box="[267,292,457,471]" fontSize="6" pageId="14" pageNumber="15">2+3</subScript>
long, horizontal, sharply bent near base, slightly sinuous.
<collectingRegion id="4933F846FFBEFF90FC3734C6FC59D093" box="[930,994,439,465]" country="New Zealand" name="West Coast" pageId="14" pageNumber="15">Costa</collectingRegion>
extends to apex of R
<subScript id="177334E1FFBEFF90FB5C34B8FB53D095" attach="both" box="[1225,1256,457,471]" fontSize="6" pageId="14" pageNumber="15">4+5,</subScript>
which is evenly, gently curved. Crossvein R-M short and vertical, much closer to R
<subScript id="177334E1FFBEFF90FC17349CFC20D0B9" attach="left" box="[898,923,493,507]" fontSize="6" pageId="14" pageNumber="15">2+3</subScript>
than to R
<subScript id="177334E1FFBEFF90FB9F349CFBA9D0B9" attach="left" box="[1034,1042,493,507]" fontSize="6" pageId="14" pageNumber="15">1</subScript>
. Fork of M a little rounded. Crossvein bM-Cu joins M far basad to R-M. CuA-veins fine, CuA
<subScript id="177334E1FFBEFF90FCC33760FCE5D35D" attach="left" box="[854,862,529,543]" fontSize="6" pageId="14" pageNumber="15">2</subScript>
apically a little bent basad. CuP fine and straight, reaches wing margin. A
<subScript id="177334E1FFBEFF90FE343744FE12D301" attach="left" box="[417,425,565,579]" fontSize="6" pageId="14" pageNumber="15">1</subScript>
visible basally only. Vein measurements are given in
<tableCitation id="C675031FFFBEFF90FB943752FBEDD37F" box="[1025,1110,547,573]" captionStart="TABLE 3" captionStartId="15.[151,239,1088,1114]" captionText="TABLE 3. Wing vein measurements, genus Penthetria" pageId="14" pageNumber="15">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF523736FE02D3C7" blockId="14.[151,1437,151,897]" pageId="14" pageNumber="15">
Abdomen (fig. 94): Length
<quantity id="4C0F9B41FFBEFF90FD943737FDE1D323" box="[513,602,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="15" unit="mm" value="8.5">8.5 mm</quantity>
, width
<quantity id="4C0F9B41FFBEFF90FD3B3736FCB3D323" box="[686,776,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="mm" value="2.3">2.3 mm</quantity>
. Dark brown, conical. Cerci two-segmented, terminal segment slender and pointed.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF5237FEFE69D3EB" blockId="14.[151,1437,151,897]" box="[199,466,655,681]" pageId="14" pageNumber="15">Male (N=2, figs. 7, 92):</paragraph>
<paragraph id="8B4836A4FFBEFF90FF5237C2FB3DD38F" blockId="14.[151,1437,151,897]" box="[199,1158,691,717]" pageId="14" pageNumber="15">
Head: Width
<quantity id="4C0F9B41FFBEFF90FEC937C2FE0ED38F" box="[348,437,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="15" unit="mm" value="1.3">1.3 mm</quantity>
. Dark brown, antennal flagellum
<quantity id="4C0F9B41FFBEFF90FCB837C2FC2AD38F" box="[813,913,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="14" pageNumber="15" unit="mm" value="0.67">0.67 mm</quantity>
long,
<quantity id="4C0F9B41FFBEFF90FC4237C2FB80D38F" box="[983,1083,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="mm" value="0.16">0.16 mm</quantity>
wide..
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF5237A6FCC5D3B3" blockId="14.[151,1437,151,897]" box="[199,894,727,753]" pageId="14" pageNumber="15">
Thorax: Length
<quantity id="4C0F9B41FFBEFF90FEE837A6FE6DD3B3" box="[381,470,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="14" pageNumber="15" unit="mm" value="3.1">3.1 mm</quantity>
. Brownish-black, haltere light brown.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF52378AFAE0D257" blockId="14.[151,1437,151,897]" box="[199,1371,763,789]" pageId="14" pageNumber="15">
Legs: Brown, slender. Fore femur 2.0 mm long,
<quantity id="4C0F9B41FFBEFF90FD71378AFCF3D257" box="[740,840,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="0.3">0.30 mm</quantity>
wide, hind femur 3.0 mm long,
<quantity id="4C0F9B41FFBEFF90FB27378AFAADD257" box="[1202,1302,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="15" unit="mm" value="0.42">0.42 mm</quantity>
wide.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF52366EFC0FD21F" blockId="14.[151,1437,151,897]" pageId="14" pageNumber="15">
Wing (fig. 92): Length 8.0 mm, width
<quantity id="4C0F9B41FFBEFF90FD13366EFD5BD27B" box="[646,736,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="15" unit="mm" value="2.4">2.4 mm</quantity>
, length/width = 3.4. Yellowish-brown fumose, darker than in female. Venation as in female, vein measurements are given in
<tableCitation id="C675031FFFBEFF90FCCF3632FC15D21F" box="[858,942,835,861]" captionStart="TABLE 3" captionStartId="15.[151,239,1088,1114]" captionText="TABLE 3. Wing vein measurements, genus Penthetria" pageId="14" pageNumber="15">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8B4836A4FFBEFF90FF523616FB45D2C3" blockId="14.[151,1437,151,897]" box="[199,1278,870,897]" pageId="14" pageNumber="15">
Abdomen: Width
<quantity id="4C0F9B41FFBEFF90FE053617FE52D2C3" box="[400,489,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="mm" value="1.5">1.5 mm</quantity>
. Basally rather wide, tapering in the middle. Terminalia not preserved.
</paragraph>
</subSubSection>
</treatment>
</document>