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<document id="D570D300F898BB0107A6A63A587C230E" ID-CLB-Dataset="8640" ID-DOI="10.11646/zootaxa.4909.1.1" ID-GBIF-Dataset="fa031765-b474-48aa-9d7e-b62dd092cfa4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4448697" ID-ZooBank="92ECCF97-7315-4A1A-A94A-537D0F331EBF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611041096358" checkinUser="plazi" docAuthor="Skartveit, John &amp; Wedmann, Sonja" docDate="2021" docId="035E87B2FFACFF83FF0230D0FAE0D553" docLanguage="en" docName="zootaxa.4909.1.1.pdf" docOrigin="Zootaxa 4909 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Plecia luctuosa Statz 1943" docType="treatment" docVersion="12" lastPageNumber="30" masterDocId="FF67FFCAFFB0FF9EFF953571FFBBD142" masterDocTitle="A Revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany" masterLastPageNumber="77" masterPageNumber="1" pageNumber="29" updateTime="1698924763774" updateUser="ExternalLinkService">
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<mods:title id="40B37004509B261F5E05B0DD6E3FDDE7">A Revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany</mods:title>
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<mods:namePart id="3F7A280E4DCD96DEB27CDD4DBD83FAD1">Skartveit, John</mods:namePart>
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<mods:namePart id="70E578D599217E7489451A9992AB90B2">Wedmann, Sonja</mods:namePart>
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<mods:title id="B6F37D47123C6281A7116A4D0DAC5AFD">Zootaxa</mods:title>
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<mods:date id="8D0E69FDEF17358524EF13FC59F5DF00">2021</mods:date>
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<mods:number id="225AD51B24724F720CA6998107269B96">4909</mods:number>
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<mods:identifier id="D55F8975EC049F81DA9113830936B647" type="DOI">10.11646/zootaxa.4909.1.1</mods:identifier>
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<treatment id="035E87B2FFACFF83FF0230D0FAE0D553" ID-DOI="http://doi.org/10.5281/zenodo.4472322" ID-GBIF-Taxon="177132519" ID-Zenodo-Dep="4472322" LSID="urn:lsid:plazi:treatment:035E87B2FFACFF83FF0230D0FAE0D553" httpUri="http://treatment.plazi.org/id/035E87B2FFACFF83FF0230D0FAE0D553" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C3ED652FFFACFF82FF0230D0FC0BD4F9" box="[151,944,1441,1468]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B4836A4FFACFF82FF0230D0FC0BD4F9" blockId="28.[151,944,1441,1468]" box="[151,944,1441,1468]" pageId="28" pageNumber="29">
<heading id="D00081C8FFACFF82FF0230D0FC0BD4F9" bold="true" box="[151,944,1441,1468]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B983EAB6FFACFF82FF0230D0FC0BD4F9" bold="true" box="[151,944,1441,1468]" pageId="28" pageNumber="29">
<taxonomicName id="4CF74D27FFACFF82FF0230D0FD27D4F9" ID-CoL="8DFGC" authority="(Heyden &amp; Heyden, 1865)" authorityName="Statz" authorityYear="1943" baseAuthorityName="Heyden &amp; Heyden" baseAuthorityYear="1865" box="[151,668,1441,1468]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="B983EAB6FFACFF82FF0230D0FEEBD4FE" bold="true" box="[151,336,1441,1468]" italics="true" pageId="28" pageNumber="29">Plecia luctuosa</emphasis>
(
<bibRefCitation id="EF664B55FFACFF82FEF430D0FD28D4F9" author="Heyden, C. H. G. von &amp; Heyden, L. F. J. D. von" box="[353,659,1441,1467]" pageId="28" pageNumber="29" pagination="19 - 30" refId="ref50399" refString="Heyden, C. H. G. von &amp; Heyden, L. F. J. D. von (1865) Bibioniden aus der rheinischen Braunkohle von Rott. Palaeontographica, 14, 19 - 30." type="journal article" year="1865">Heyden &amp; Heyden, 1865</bibRefCitation>
)
</taxonomicName>
(figs. 41–42, 130–137)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFACFF82FF023097FEEAD7C6" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8B4836A4FFACFF82FF023097FD33D4BC" blockId="28.[151,681,1510,1597]" box="[151,648,1510,1534]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="ABE7118AFFACFF82FF023097FD33D4BC" box="[151,648,1510,1534]" pageId="28" pageNumber="29">
<treatmentCitation id="0A5610B5FFACFF82FF023097FD3FD4BC" author="Heyden, C. H. G. von &amp; Heyden, L. F. J. D. von" box="[151,644,1510,1534]" page="22" pageId="28" pageNumber="29" year="1865">
<taxonomicName id="4CF74D27FFACFF82FF023097FD3FD4BC" ID-CoL="8DHLJ" authority="Heyden &amp; Heyden 1865: 22" authorityName="Heyden &amp; Heyden" authorityPageNumber="22" authorityYear="1865" box="[151,644,1510,1534]" class="Insecta" family="Bibionidae" genus="Protomyia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="B983EAB6FFACFF82FF023097FEE5D4BC" box="[151,350,1510,1534]" italics="true" pageId="28" pageNumber="29">Protomyia luctuosa</emphasis>
<bibRefCitation id="EF664B55FFACFF82FEF13097FD3FD4BC" author="Heyden, C. H. G. von &amp; Heyden, L. F. J. D. von" box="[356,644,1510,1534]" pageId="28" pageNumber="29" pagination="19 - 30" refId="ref50399" refString="Heyden, C. H. G. von &amp; Heyden, L. F. J. D. von (1865) Bibioniden aus der rheinischen Braunkohle von Rott. Palaeontographica, 14, 19 - 30." type="journal article" year="1865">Heyden &amp; Heyden 1865: 22</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B4836A4FFACFF82FF023375FD12D75E" blockId="28.[151,681,1510,1597]" box="[151,681,1540,1565]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="ABE7118AFFACFF82FF023375FD12D75E" box="[151,681,1540,1565]" pageId="28" pageNumber="29">
<treatmentCitation id="0A5610B5FFACFF82FF023375FD1ED75E" author="Statz, G." box="[151,677,1540,1565]" page="15" pageId="28" pageNumber="29" year="1943">
<taxonomicName id="4CF74D27FFACFF82FF023375FD1ED75E" ID-CoL="8DFGC" authority="(Heyden &amp; Heyden) Statz 1943: 15" authorityName="Statz" authorityPageNumber="15" authorityYear="1943" baseAuthorityName="Heyden &amp; Heyden" baseAuthorityYear="1865" box="[151,677,1540,1565]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="B983EAB6FFACFF82FF023375FE8FD75E" box="[151,308,1540,1564]" italics="true" pageId="28" pageNumber="29">Plecia luctuosa</emphasis>
(Heyden &amp; Heyden)
<bibRefCitation id="EF664B55FFACFF82FD853374FD1ED75E" author="Statz, G." box="[528,677,1540,1565]" pageId="28" pageNumber="29" pagination="1 - 65" refId="ref51587" refString="Statz, G. (1943) Neue Dipteren (Nematocera) aus dem Oberoligozan von Rott. I. Familie Bibionidae (Haarmucken). Palaeontographica, Series A, 95, 1 - 65." type="journal article" year="1943">Statz 1943: 15</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B4836A4FFACFF82FF023352FE4BD779" blockId="28.[151,681,1510,1597]" box="[151,496,1571,1597]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="ABE7118AFFACFF82FF023352FE4BD779" box="[151,496,1571,1597]" pageId="28" pageNumber="29">
<taxonomicName id="4CF74D27FFACFF82FF023352FF63D779" ID-CoL="9CJT2" box="[151,216,1571,1595]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B983EAB6FFACFF82FF023352FF63D779" box="[151,216,1571,1595]" italics="true" pageId="28" pageNumber="29">Plecia</emphasis>
</taxonomicName>
♂ Nr. 4
<treatmentCitation id="0A5610B5FFACFF82FEA43352FE57D779" author="Statz, G." box="[305,492,1571,1595]" page="30" pageId="28" pageNumber="29" year="1943">
<bibRefCitation id="EF664B55FFACFF82FEA43352FE57D779" author="Statz, G." box="[305,492,1571,1595]" pageId="28" pageNumber="29" pagination="1 - 65" refId="ref51587" refString="Statz, G. (1943) Neue Dipteren (Nematocera) aus dem Oberoligozan von Rott. I. Familie Bibionidae (Haarmucken). Palaeontographica, Series A, 95, 1 - 65." type="journal article" year="1943">Statz 1943: 30–31</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B4836A4FFACFF82FF023318FEEAD7C6" blockId="28.[151,1436,1641,2028]" box="[151,337,1641,1668]" pageId="28" pageNumber="29">
<emphasis id="B983EAB6FFACFF82FF023318FEB1D7C1" bold="true" box="[151,266,1641,1667]" pageId="28" pageNumber="29">Localities</emphasis>
: Rott.
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFACFF82FF5233FCFC95D7B2" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B4836A4FFACFF82FF5233FCFC7ED7E5" blockId="28.[151,1436,1641,2028]" box="[199,965,1677,1704]" pageId="28" pageNumber="29">
<emphasis id="B983EAB6FFACFF82FF5233FCFE12D7E5" bold="true" box="[199,425,1677,1703]" pageId="28" pageNumber="29">Material examined</emphasis>
:
<materialsCitation id="3B9F3CF9FFACFF82FE2D33FCFC7ED7E5" ID-GBIF-Occurrence="3026467321" box="[440,965,1677,1704]" collectionCode="NHMUK" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="544C8806FFACFF82FE2D33FCFD9BD7E5" box="[440,544,1677,1703]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
(female)
<collectionCode id="EDE6AE61FFACFF82FD1833FFFD4CD7EA" box="[653,759,1678,1704]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="28" pageNumber="29" type="Museum">NHMUK</collectionCode>
In 58792 (fig. 41)
</materialsCitation>
</paragraph>
<paragraph id="8B4836A4FFACFF82FF5233C0FC95D7B2" blockId="28.[151,1436,1641,2028]" pageId="28" pageNumber="29">
Additional material: Male: LACMIP 2533.260/LACMIP
<typeStatus id="544C8806FFACFF82FCDB33C3FC3DD78E" box="[846,902,1714,1740]" pageId="28" pageNumber="29">Type</typeStatus>
3390 (fig. 42), Females: LACMIP 2533.281ab LACMIP
<typeStatus id="544C8806FFACFF82FE9333A7FE85D7B2" box="[262,318,1750,1776]" pageId="28" pageNumber="29">Type</typeStatus>
3413, 2533.1234, 2533.1293, 2533.4187ab.
</paragraph>
</subSubSection>
<subSubSection id="C3ED652FFFACFF83FF523388FAE0D553" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="8B4836A4FFACFF82FF523388FEE1D656" blockId="28.[151,1436,1641,2028]" box="[199,346,1785,1812]" pageId="28" pageNumber="29">
<emphasis id="B983EAB6FFACFF82FF523388FEEAD651" bold="true" box="[199,337,1785,1811]" pageId="28" pageNumber="29">Description</emphasis>
:
</paragraph>
<paragraph id="8B4836A4FFACFF82FF52326CFD84D675" blockId="28.[151,1436,1641,2028]" box="[199,575,1821,1848]" pageId="28" pageNumber="29">Female (N=5, figs. 41, 130–133):</paragraph>
<paragraph id="8B4836A4FFACFF82FF523230FE60D61E" blockId="28.[151,1436,1641,2028]" box="[199,475,1857,1884]" pageId="28" pageNumber="29">
Total length 8.0–
<quantity id="4C0F9B41FFACFF82FE133230FE60D61E" box="[390,475,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="28" pageNumber="29" unit="mm" value="9.5">9.5 mm</quantity>
</paragraph>
<paragraph id="8B4836A4FFACFF82FF523214FF74D6E1" blockId="28.[151,1436,1641,2028]" pageId="28" pageNumber="29">Head (fig. 132): Dark brown. Antenna robust, pointed, flagellum 6-segmented? Complex eye rather large, oval.</paragraph>
<paragraph id="8B4836A4FFACFF82FF5232DCFD74D685" blockId="28.[151,1436,1641,2028]" box="[199,719,1965,1992]" pageId="28" pageNumber="29">Thorax (fig. 132): Brown, haltere light brown.</paragraph>
<paragraph id="8B4836A4FFACFF82FF5232A3FCE7D6A9" blockId="28.[151,1436,1641,2028]" box="[199,860,2001,2028]" pageId="28" pageNumber="29">Legs: Brown, slender, femora basally pale. Hind tibia pale.</paragraph>
<paragraph id="8B4836A4FFADFF83FF5235E6FBA2D003" blockId="29.[151,1437,151,1041]" pageId="29" pageNumber="30">
Wing (fig. 133): Brown fumose, veins brown. Costal cell considerably darkened. Humeral vein absent. Subcosta relatively strong, straight, running close to but well separated from R. Pterostigma brown, not very distinctive.
<collectingRegion id="4933F846FFADFF83FF0235AEFF6CD1BB" box="[151,215,223,249]" country="New Zealand" name="West Coast" pageId="29" pageNumber="30">Costa</collectingRegion>
extends to a little beyond apex of R
<subScript id="177334E1FFADFF83FDEC3580FD28D1BD" attach="left" box="[633,659,241,255]" fontSize="6" pageId="29" pageNumber="30">4+5</subScript>
. R
<subScript id="177334E1FFADFF83FD203580FD75D1BD" attach="left" box="[693,718,241,255]" fontSize="6" pageId="29" pageNumber="30">2+3</subScript>
short, steep, evenly curved. R
<subScript id="177334E1FFADFF83FBB23580FBFAD1BD" attach="left" box="[1063,1089,241,255]" fontSize="6" pageId="29" pageNumber="30">4+5</subScript>
apically strongly curved. R-M vertical. M- and CuA-veins apically straight. CuP straight, reaching wing margin, running more or less parallel to CuA and well separated from this vein. Vein measurements are given in
<tableCitation id="C675031FFFADFF83FC553456FBAED003" box="[960,1045,295,321]" captionStart="TABLE 4" captionStartId="21.[151,239,588,614]" captionText="TABLE 4. Wing vein measurements, genus Plecia" pageId="29" pageNumber="30">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8B4836A4FFADFF83FF52343AFCBBD027" blockId="29.[151,1437,151,1041]" box="[199,768,331,357]" pageId="29" pageNumber="30">Abdomen: dark brown, cylindrical. Cerci rounded.</paragraph>
<paragraph id="8B4836A4FFADFF83FF52341EFD9CD0CB" blockId="29.[151,1437,151,1041]" box="[199,551,367,393]" pageId="29" pageNumber="30">Male (N=1, figs. 42, 134–137):</paragraph>
<paragraph id="8B4836A4FFADFF83FF5234E2FE11D0EF" blockId="29.[151,1437,151,1041]" box="[199,426,403,429]" pageId="29" pageNumber="30">Total length 7.0 mm</paragraph>
<paragraph id="8B4836A4FFADFF83FF5234C6FBF5D0B7" blockId="29.[151,1437,151,1041]" pageId="29" pageNumber="30">
Head (fig.136): Length
<quantity id="4C0F9B41FFADFF83FE4434C6FD8CD093" box="[465,567,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="29" pageNumber="30" unit="mm" value="1.04">1.04 mm</quantity>
, width
<quantity id="4C0F9B41FFADFF83FD1F34C6FD4AD093" box="[650,753,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="29" pageNumber="30" unit="mm" value="1.18">1.18 mm</quantity>
. Blackish-brown. Antenna relatively slender, cylindrical, flagellum
<quantity id="4C0F9B41FFADFF83FF7934AAFEEBD0B7" box="[236,336,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="29" pageNumber="30" unit="mm" value="0.42">0.42 mm</quantity>
long,
<quantity id="4C0F9B41FFADFF83FE0334AAFE41D0B7" box="[406,506,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="29" pageNumber="30" unit="mm" value="0.09">0.09 mm</quantity>
wide, possibly 8-segmented. Palpus blackish-brown.
</paragraph>
<paragraph id="8B4836A4FFADFF83FF52348EFC54D35B" blockId="29.[151,1437,151,1041]" box="[199,1007,511,538]" pageId="29" pageNumber="30">
Thorax (fig. 136): Length
<quantity id="4C0F9B41FFADFF83FE7B348EFDFCD35B" box="[494,583,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="mm" value="2.1">2.1 mm</quantity>
. Blackish-brown, haltere light brown.
</paragraph>
<paragraph id="8B4836A4FFADFF83FF523752FE60D3EB" blockId="29.[151,1437,151,1041]" pageId="29" pageNumber="30">
Legs: Slender. Fore- and mid-femora pale yellowish with distal fourth blackish-brown, hind femur pale yellowish with extreme tip dark. All tibiae and tarsi blackish-brown. Fore femur
<quantity id="4C0F9B41FFADFF83FC773736FB82D323" box="[994,1081,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="mm" value="1.8">1.8 mm</quantity>
long,
<quantity id="4C0F9B41FFADFF83FB173736FB53D323" box="[1154,1256,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="29" pageNumber="30" unit="mm" value="0.33">0.33 mm</quantity>
wide, fore tibia
<quantity id="4C0F9B41FFADFF83FF02371AFF56D3C7" box="[151,237,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="mm" value="2.1">2.1 mm</quantity>
long,
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wide, fore first tarsomere
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long, hind femur
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long,
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wide, hind tibia
<quantity id="4C0F9B41FFADFF83FF0237FEFF56D3EB" box="[151,237,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="mm" value="2.6">2.6 mm</quantity>
long,
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wide,
</paragraph>
<paragraph id="8B4836A4FFADFF83FF5237C2FEE9D2E7" blockId="29.[151,1437,151,1041]" pageId="29" pageNumber="30">
Wing (fig. 135): length
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, width
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, length/width = 2.56. Relatively narrow, apically rounded, brown fumose, costal cell somewhat darkened. Veins dark brown, relatively fine but distinctive throughout. Pterostigma broadly oval, brown, not very distinctive. Humeral vein present, oblique. Subcosta straight, basally running very close to R and difficult to separate from this vein.
<collectingRegion id="4933F846FFADFF83FD51366EFCBFD27B" box="[708,772,799,825]" country="New Zealand" name="West Coast" pageId="29" pageNumber="30">Costa</collectingRegion>
extends to a little beyond apex of R
<subScript id="177334E1FFADFF83FB063640FB16D27D" attach="left" box="[1171,1197,817,831]" fontSize="6" pageId="29" pageNumber="30">4+5</subScript>
. R
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relatively short, oblique, strongly and evenly curved. R
<subScript id="177334E1FFADFF83FDC63624FDD6D221" attach="left" box="[595,621,853,867]" fontSize="6" pageId="29" pageNumber="30">4+5</subScript>
apically strongly curved. R-M vertical. Fork of M almost symmetrical, veins gently curved. CuA-veins strong, apically gently curved. CuP rather strong, straight. Vein measurements are given in
<tableCitation id="C675031FFFADFF83FF6C36FAFEF5D2E7" box="[249,334,907,933]" captionStart="TABLE 4" captionStartId="21.[151,239,588,614]" captionText="TABLE 4. Wing vein measurements, genus Plecia" pageId="29" pageNumber="30">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8B4836A4FFADFF83FF5236DEFB62D288" blockId="29.[151,1437,151,1041]" box="[199,1241,942,970]" pageId="29" pageNumber="30">
Abdomen: Length
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, width
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. Blackish-brown, conical (basally relatively wide).
</paragraph>
<paragraph id="8B4836A4FFADFF83FF5236A2FAE0D553" blockId="29.[151,1437,151,1041]" pageId="29" pageNumber="30">
Terminalia (fig. 137): Hypopygium
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wide, rectangular in outline. Gonostylus
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long, attached laterally, straight and relatively long, blunt. Epandrium relatively small, rectangular, apically gently concave.
</paragraph>
</subSubSection>
</treatment>
</document>
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