224 lines
28 KiB
XML
224 lines
28 KiB
XML
<document id="1EA7FF8CDA67F3C2D1CC2EBC150FC4B7" ID-CLB-Dataset="30267" ID-DOI="10.11646/zootaxa.4407.4.4" ID-GBIF-Dataset="a850a99f-acfa-4f8e-b098-e7e783587f92" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221080" ID-ZooBank="D112CD45-7907-4F42-9D32-0DC7DC3A39EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524143490108" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2018" docId="FD5D6D7EFFD7FF969D806158FF5C85C7" docLanguage="en" docName="zootaxa.4407.4.4.pdf" docOrigin="Zootaxa 4407 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia emeishanensis Shavrin 2018, sp.n." docType="treatment" docVersion="7" lastPageNumber="527" masterDocId="01641506FFD3FF909D176341FFB48743" masterDocTitle="Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan" masterLastPageNumber="532" masterPageNumber="521" pageNumber="525" updateTime="1698537035611" updateUser="ExternalLinkService">
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<mods:title id="2A496E27893C3FF059F4746263942325">Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan</mods:title>
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<mods:name id="7A974324D5172AAE4264CA701204B0C7" type="personal">
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<mods:namePart id="C09B0261602BFE9C2F1AD7760A6BE554">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="6B045401AE1C1FEDEFB4346DE1DEAFB9">Zootaxa</mods:title>
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<mods:part id="ACE2CDC7318CD9EFD71D6030E31B7E3C">
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<mods:date id="14AA193C0B0D653BF6E2B8786BF99F43">2018</mods:date>
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<mods:number id="3B1771381244F41AC6395D43AC8E2286">2018-04-12</mods:number>
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<mods:number id="CD430CD7791F3938098F465FC35DDC78">4407</mods:number>
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<treatment id="FD5D6D7EFFD7FF969D806158FF5C85C7" ID-DOI="http://doi.org/10.5281/zenodo.5949830" ID-GBIF-Taxon="143605182" ID-Zenodo-Dep="5949830" LSID="urn:lsid:plazi:treatment:FD5D6D7EFFD7FF969D806158FF5C85C7" httpUri="http://treatment.plazi.org/id/FD5D6D7EFFD7FF969D806158FF5C85C7" lastPageId="6" lastPageNumber="527" pageId="4" pageNumber="525">
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<subSubSection id="3DEE8FE3FFD7FF949D806158FE9A8517" pageId="4" pageNumber="525" type="nomenclature">
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<paragraph id="754BDC68FFD7FF949D806158FD038570" blockId="4.[151,695,537,597]" box="[151,695,537,563]" pageId="4" pageNumber="525">
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<heading id="2E036B04FFD7FF949D806158FD038570" bold="true" box="[151,695,537,563]" fontSize="11" level="1" pageId="4" pageNumber="525" reason="1">
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<emphasis id="4780007AFFD7FF949D806158FD038570" bold="true" box="[151,695,537,563]" pageId="4" pageNumber="525">
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<taxonomicName id="B2F4A7EBFFD7FF949D806158FDDB8570" ID-CoL="3XVMD" authority="Shavrin" authorityName="Shavrin" authorityYear="2018" box="[151,623,537,563]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="emeishanensis" status="sp. nov.">
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<emphasis id="4780007AFFD7FF949D806158FDB78570" bold="true" box="[151,515,537,563]" italics="true" pageId="4" pageNumber="525">Mannerheimia emeishanensis</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="5CB3BD01FFD7FF949F6B6158FD038570" box="[636,695,537,563]" pageId="4" pageNumber="525" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="754BDC68FFD7FF949D80617DFE9A8517" blockId="4.[151,695,537,597]" box="[151,302,572,597]" pageId="4" pageNumber="525">
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(
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<figureCitation id="EDCFC0EDFFD7FF949D88617DFF5A8517" box="[159,238,572,597]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="4" pageNumber="525">Figs. 2</figureCitation>
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,
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<figureCitation id="EDCFC0EDFFD7FF949DED617DFE928517" box="[250,294,572,596]" captionStart-0="FIGURES 5–8" captionStart-1="FIGURES 9–10" captionStartId-0="5.[151,264,1837,1860]" captionStartId-1="7.[151,264,1329,1352]" captionTargetBox-0="[342,1327,200,1779]" captionTargetBox-1="[201,1360,771,1291]" captionTargetId-0="figure@5.[210,1374,181,1827]" captionTargetId-1="figure@7.[187,1400,756,1330]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURES 5–8. Aedeagus of Mannerheimia grandilobata (5–6: paratype) and M. emeishanensis (7–8: paratype): 5, 7—parameral view, 6, 8—lateral view.Scale bar: 0.1 mm." captionText-1="FIGURES 9–10. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri-0="https://zenodo.org/record/1221084/files/figure.png" httpUri-1="https://zenodo.org/record/1221086/files/figure.png" pageId="4" pageNumber="525">7–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="3DEE8FE3FFD7FF949D8061C5FAF2830F" pageId="4" pageNumber="525" type="materials_examined">
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<paragraph id="754BDC68FFD7FF949D8061C5FF4B844B" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">
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<emphasis id="4780007AFFD7FF949D8061C5FE7E85DE" bold="true" box="[151,458,644,669]" pageId="4" pageNumber="525">
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<typeStatus id="AA4F62CAFFD7FF949D8061C5FF6685DE" box="[151,210,644,669]" pageId="4" pageNumber="525">Type</typeStatus>
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material examined:
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</emphasis>
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<typeStatus id="AA4F62CAFFD7FF949CC161C5FD8A85DE" box="[470,574,644,669]" pageId="4" pageNumber="525" type="holotype">Holotype</typeStatus>
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<specimenCount id="63F217E1FFD7FF949F5D61C4FDEB85DF" box="[586,607,645,668]" pageId="4" pageNumber="525" type="male">♂</specimenCount>
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[specimen dissected]: ‘R.
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<collectingCountry id="0DE39CF8FFD7FF949E8C61C5FC4385DF" box="[923,1015,644,668]" name="China" pageId="4" pageNumber="525">CHINA</collectingCountry>
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,
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<collectingRegion id="B730128AFFD7FF94991061C5FBD085DE" box="[1031,1124,644,669]" country="China" name="Sichuan" pageId="4" pageNumber="525">Sichuan</collectingRegion>
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, | Emei Shan,
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<geoCoordinate id="10C0BAAFFFD7FF94980161C5FA3F85DE" box="[1302,1419,644,669]" direction="north" orientation="latitude" pageId="4" pageNumber="525" precision="15" value="29.551666">N29°33'6''</geoCoordinate>
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<geoCoordinate id="10C0BAAFFFD7FF949D8061E6FEAC8583" box="[151,280,679,704]" direction="east" orientation="longitude" pageId="4" pageNumber="525" precision="15" value="103.335">E103°20'6''</geoCoordinate>
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,
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<date id="014AFAA8FFD7FF949C3C61E6FE198583" box="[299,429,679,704]" pageId="4" pageNumber="525">27.vi.-5.vii.</date>
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| 2009,
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<quantity id="B20C718DFFD7FF949F0D61E9FD188583" box="[538,684,680,704]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="4" pageNumber="525" unit="m" value="2100.0" valueMax="2400.0" valueMin="1800.0">1800-2400m</quantity>
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, sifti- | ng11-17, V. Grebennikov’ <rectangular label, printed>, ‘
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<typeStatus id="AA4F62CAFFD7FF949D8C618DFE8085A7" box="[155,308,716,740]" pageId="4" pageNumber="525" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="B2F4A7EBFFD7FF949C58618DFE4685A6" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[335,498,716,741]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
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<taxonomicName id="B2F4A7EBFFD7FF949F1A618DFD1B85A6" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[525,687,716,741]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="emeishanensis" status="sp. nov.">emeishanensis</taxonomicName>
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<taxonomicNameLabel id="5CB3BD01FFD7FF949FAD618DFD5B85A0" box="[698,751,716,739]" pageId="4" pageNumber="525" rank="species">
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<emphasis id="4780007AFFD7FF949FAD618DFD5B85A0" bold="true" box="[698,751,716,739]" pageId="4" pageNumber="525">sp.n.</emphasis>
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</taxonomicNameLabel>
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| Shavrin A.V. des. 2017’ <red rectangular label, printed> (
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<collectionCode id="13E544ADFFD7FF949D8861B1FF47844B" box="[159,243,752,776]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="4" pageNumber="525">NSMT</collectionCode>
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).
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</paragraph>
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<paragraph id="754BDC68FFD7FF949DD06054FAF2830F" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">
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Paratypes (17 specimens): 1 ♀: same label as the holotype (CS); 1 ♂, 1 ♀: ‘P.R. CHINA, Sichuan, | Emei Shan, N29°33'51'' | E103°20'47'',
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<date id="014AFAA8FFD7FF949CDF6079FD838413" box="[456,567,824,848]" pageId="4" pageNumber="525" value="2011-05-23">23.v.2011</date>
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, |
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<quantity id="B20C718DFFD7FF949F586079FD2B8413" box="[591,671,824,848]" metricMagnitude="3" metricUnit="m" metricValue="1.779" pageId="4" pageNumber="525" unit="m" value="1779.0">1779m</quantity>
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, | sift03, V. Grebennikov’ <rectangular label, printed> (CS, NSMT); 1 ♀: ‘P.R. CHINA, Sichuan, | Emei Shan, N29°34'46'' | E103°22'04'',
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<date id="014AFAA8FFD7FF949E88601DFBBB8437" box="[927,1039,860,884]" pageId="4" pageNumber="525" value="2011-05-27">27.v.2011</date>
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, |
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<quantity id="B20C718DFFD7FF949931601DFBC98437" box="[1062,1149,860,885]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="4" pageNumber="525" unit="m" value="1463.0">1463 m</quantity>
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, | sift07, V. Grebennikov’ <rectangular label, printed> (CNC); 1 ♂, 4 ♀ [one female without antennae]: ‘P.R. CHINA, Sichuan, | Emei Shan, N29°33'04'' | E103°21'19'',
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<date id="014AFAA8FFD7FF949CDD60E5FD8D84FE" box="[458,569,932,957]" pageId="4" pageNumber="525" value="2011-05-25">25.v.2011</date>
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, |
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<quantity id="B20C718DFFD7FF949F4460E5FD1784FF" box="[595,675,932,956]" metricMagnitude="3" metricUnit="m" metricValue="1.7289999999999999" pageId="4" pageNumber="525" unit="m" value="1729.0">1729m</quantity>
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, | sift05, V. Grebennikov’ <rectangular label, printed> (CNC); 7 ♀: ‘P.R. CHINA, Sichuan, | Emei Shan, N29°33'56'' | E103°21'24'',
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<date id="014AFAA8FFD7FF949E9A6086FC4984A3" box="[909,1021,967,992]" pageId="4" pageNumber="525" value="2011-05-26">26.v.2011</date>
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, |
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<quantity id="B20C718DFFD7FF94990B6089FBD884A3" box="[1052,1132,968,992]" metricMagnitude="3" metricUnit="m" metricValue="1.829" pageId="4" pageNumber="525" unit="m" value="1829.0">1829m</quantity>
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, | sift06, V. Grebennikov’ <rectangular label, printed> (NSMT); 1 ♀: ‘P.R. CHINA, Sichuan, | Emei Shan, N29°33'39'' | E103°20'42'',
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<date id="014AFAA8FFD7FF949D806751FEB2836B" box="[151,262,1040,1064]" pageId="4" pageNumber="525" value="2011-05-23">23.v.2011</date>
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, |
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||
<quantity id="B20C718DFFD7FF949C36674EFEC6836B" box="[289,370,1039,1064]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="4" pageNumber="525" unit="m" value="1852.0">1852m</quantity>
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, | sift04, V. Grebennikov’ <rectangular label, printed> (NSMT). All paratypes with additional red rectangular printed label: ‘PARATYPE |
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<taxonomicName id="B2F4A7EBFFD7FF949F9D6775FC99830E" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[650,813,1076,1101]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
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|
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<taxonomicName id="B2F4A7EBFFD7FF949E566775FC57830E" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[833,995,1076,1101]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="emeishanensis" status="sp. nov.">emeishanensis</taxonomicName>
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<taxonomicNameLabel id="5CB3BD01FFD7FF949EFE6775FBAA8308" box="[1001,1054,1076,1099]" pageId="4" pageNumber="525" rank="species">
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<emphasis id="4780007AFFD7FF949EFE6775FBAA8308" bold="true" box="[1001,1054,1076,1099]" pageId="4" pageNumber="525">sp.n.</emphasis>
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</taxonomicNameLabel>
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| Shavrin A.V. des. 2017’.
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</paragraph>
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</subSubSection>
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<subSubSection id="3DEE8FE3FFD7FF969DD06716FE9A8693" lastPageId="6" lastPageNumber="527" pageId="4" pageNumber="525" type="description">
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<paragraph id="754BDC68FFD7FF949DD06716FB7B83FB" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">
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<emphasis id="4780007AFFD7FF949DD06716FEE58333" bold="true" box="[199,337,1111,1136]" pageId="4" pageNumber="525">Description</emphasis>
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. Measurements (n=18): HW: 0.46–0.50; HL: 0.25–0.29; AL (
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<typeStatus id="AA4F62CAFFD7FF9499116716FBDB8333" box="[1030,1135,1111,1136]" pageId="4" pageNumber="525" type="holotype">holotype</typeStatus>
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): 0.60; OL: 0.12–0.14; PL: 0.37–0.42; PW: 0.71–0.82; EL: 0.78–0.86; EW: 0.94–1.12; AW: 0.87–1.04; MTbL (
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<typeStatus id="AA4F62CAFFD7FF949998673DFB4C83D6" box="[1167,1272,1148,1173]" pageId="4" pageNumber="525" type="holotype">holotype</typeStatus>
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): 0.37; MTrL (
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<typeStatus id="AA4F62CAFFD7FF949D8867DEFEBC83FB" box="[159,264,1183,1208]" pageId="4" pageNumber="525" type="holotype">holotype</typeStatus>
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): 0.20 (MTrL 1–4: 0.12; MTrL 5: 0.080); AedL: 0.30; TL: 1.75–2.30 (
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<typeStatus id="AA4F62CAFFD7FF94990C67DEFB3783FB" box="[1051,1155,1183,1208]" pageId="4" pageNumber="525" type="holotype">holotype</typeStatus>
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: 2.10).
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</paragraph>
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<paragraph id="754BDC68FFD7FF949DD06785FDFD8163" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">
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Body very wide and convex. Yellow-brown to reddish-brown, sometimes head darker and pronotum paler, with yellow margins; mouthparts, antennae, legs, paratergites and apical abdominal tergites yellow. Forebody very shiny, without microsculpture; abdomen with indistinct or distinct isodiametric microsculpture. Head with irregular and sparse, small and moderately deep punctation, distinctly denser on median portion between eyes, sometimes medio-posterior portion of head between posterior third length of eyes impunctate, some
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<typeStatus id="AA4F62CAFFD7FF9499686614FB5F822E" box="[1151,1259,1365,1389]" pageId="4" pageNumber="525" type="paratype">paratypes</typeStatus>
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with transverse row of punctures between ocelli; punctation of pronotum sparse, with wide impunctate portion in medio-basal third; scutellum with several small punctures; punctation of elytra as that in pronotum, each elytron with very tangled and vague six longitudinal rows of punctures, remaining elytral punctures forming indistinct to distinct diagonal rows and becoming irregularly scattered laterally; abdominal tergites with indistinct, very small and sparse punctation. Habitus as in
|
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<figureCitation id="EDCFC0EDFFD7FF949F166546FDF28163" box="[513,582,1543,1568]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="4" pageNumber="525">Fig. 2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD7FF949DD0656DFE29805E" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">
|
||
Head 1.7–1.8 times as wide as long, without occipital furrow between and grooves in front of ocelli; anterior portion of head between antennal insertion and anterior margin of eye slightly concave. Ocelli indistinct, flattened and large, apical margin of ocelli situated at or slightly behind level of posterior margin of eyes; distance between ocelli about one and a half times as long as distance between ocellus and posterior margin of eye. Antenna with antennomeres gradually widened toward apex, antennomere 4 as long as wide; length × width of antennomeres (
|
||
<typeStatus id="AA4F62CAFFD7FF949D88659EFEBC81BB" box="[159,264,1759,1784]" pageId="4" pageNumber="525" type="holotype">holotype</typeStatus>
|
||
): 1: 0.08 × 0.03; 2: 0.07 × 0.02; 3: 0.06 × 0.02; 4: 0.02 × 0.02; 5: 0.03 × 0.03; 6: 0.03 × 0.04; 7–10: 0.05 × 0.05; 11: 0.11 × 0.05.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD7FF949DD06468FBC680CB" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">Pronotum transverse, 1.9 times as wide as long, 1.5–1.6 times as wide as head, widest in middle, gradually slightly narrowed posterad and anterad; anterior angles widely rounded, indistinctly protruded anterad, posterior angles obtuse; lateral portions very narrow and slightly explanate, without pit at middle.</paragraph>
|
||
<paragraph id="754BDC68FFD7FF949DD064D5FD6680B6" blockId="4.[151,1437,644,2037]" pageId="4" pageNumber="525">Elytra 1.2–1.3 times as wide as long, significantly widened apicad, two to two and a half times as long as pronotum and sometimes covering first five abdominal tergites; lateral portions of elytra narrow, as in pronotum; latero-apical margins of elytra without crenulation.</paragraph>
|
||
<caption id="218B8CE0FFD6FF959D80646CFD518020" httpUri="https://zenodo.org/record/1221084/files/figure.png" pageId="5" pageNumber="526" startId="5.[151,264,1837,1860]" targetBox="[342,1327,200,1779]" targetPageId="5">
|
||
<paragraph id="754BDC68FFD6FF959D80646CFD518020" blockId="5.[151,1436,1837,1891]" pageId="5" pageNumber="526">
|
||
<emphasis id="4780007AFFD6FF959D80646CFEF78000" bold="true" box="[151,323,1837,1860]" pageId="5" pageNumber="526">FIGURES 5–8.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="B2F4A7EBFFD6FF959CFF646FFCBB8007" authorityName="Shavrin" authorityYear="2018" box="[488,783,1838,1860]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="grandilobata">
|
||
<emphasis id="4780007AFFD6FF959CFF646FFCBB8007" box="[488,783,1838,1860]" italics="true" pageId="5" pageNumber="526">Mannerheimia grandilobata</emphasis>
|
||
</taxonomicName>
|
||
(5–6: paratype) and
|
||
<taxonomicName id="B2F4A7EBFFD6FF959913646EFB758007" authorityName="Shavrin" authorityYear="2018" box="[1028,1217,1838,1860]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="emeishanensis">
|
||
<emphasis id="4780007AFFD6FF959913646EFB758007" box="[1028,1217,1838,1860]" italics="true" pageId="5" pageNumber="526">M. emeishanensis</emphasis>
|
||
</taxonomicName>
|
||
(7–8: paratype): 5, 7—parameral view, 6, 8—lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="754BDC68FFD6FF959DD064D1FD1C80EB" blockId="5.[151,1436,1935,2032]" box="[199,680,1935,1960]" pageId="5" pageNumber="526">Meatatarsus 1.8 times as long as metatibia.</paragraph>
|
||
<paragraph id="754BDC68FFD6FF959DD064F2FF5580B3" blockId="5.[151,1436,1935,2032]" pageId="5" pageNumber="526">Abdomen slightly narrower than elytra; apical margin of abdominal tergite VIII with very narrow palisade fringe.</paragraph>
|
||
<paragraph id="754BDC68FFD5FF969DD063D6FA84865F" blockId="6.[151,1436,151,644]" pageId="6" pageNumber="527">
|
||
Male. Protarsomeres 1–4 slightly widened. Apical margin of abdominal tergite VIII straight or slightly rounded. Apical margin of abdominal sternite VIII widely and moderately deep sinuate. Aedeagus (
|
||
<figureCitation id="EDCFC0EDFFD5FF96980463FDFAEC8797" box="[1299,1368,188,213]" captionStart="FIGURES 5–8" captionStartId="5.[151,264,1837,1860]" captionTargetBox="[342,1327,200,1779]" captionTargetId="figure@5.[210,1374,181,1827]" captionTargetPageId="5" captionText="FIGURES 5–8. Aedeagus of Mannerheimia grandilobata (5–6: paratype) and M. emeishanensis (7–8: paratype): 5, 7—parameral view, 6, 8—lateral view.Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/1221084/files/figure.png" pageId="6" pageNumber="527">Fig. 7</figureCitation>
|
||
) with very wide basal part and narrow median lobe gradually narrowing toward rounded apex; parameres distinctly exceeding apex of median lobe; internal sac narrow and moderately long. Aedeagus laterally as in
|
||
<figureCitation id="EDCFC0EDFFD5FF9699F06245FA9F865F" box="[1255,1323,260,285]" captionStart="FIGURES 5–8" captionStartId="5.[151,264,1837,1860]" captionTargetBox="[342,1327,200,1779]" captionTargetId="figure@5.[210,1374,181,1827]" captionTargetPageId="5" captionText="FIGURES 5–8. Aedeagus of Mannerheimia grandilobata (5–6: paratype) and M. emeishanensis (7–8: paratype): 5, 7—parameral view, 6, 8—lateral view.Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/1221084/files/figure.png" pageId="6" pageNumber="527">Fig. 8</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD5FF969DD06266FA3A8603" blockId="6.[151,1436,151,644]" box="[199,1422,295,320]" pageId="6" pageNumber="527">Female. Protarsomeres 1–4 not widened. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
|
||
<paragraph id="754BDC68FFD5FF969DD0620DFE9A8693" blockId="6.[151,1436,151,644]" pageId="6" pageNumber="527">
|
||
<emphasis id="4780007AFFD5FF969DD0620DFE008626" bold="true" box="[199,436,332,357]" pageId="6" pageNumber="527">Comparative notes.</emphasis>
|
||
Based on the body size, coloration, shape of the pronotum and general shape of the aedeagus,
|
||
<taxonomicName id="B2F4A7EBFFD5FF969C1B6230FE6786CB" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[268,467,368,392]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="emeishanensis" status="sp. nov.">
|
||
<emphasis id="4780007AFFD5FF969C1B6230FE6786CB" box="[268,467,368,392]" italics="true" pageId="6" pageNumber="527">M. emeishanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CB3BD01FFD5FF969CCB622EFDA786CB" box="[476,531,367,392]" pageId="6" pageNumber="527" rank="species">
|
||
<emphasis id="4780007AFFD5FF969CCB622EFDA786CB" bold="true" box="[476,531,367,392]" pageId="6" pageNumber="527">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most similar to Japanese
|
||
<taxonomicName id="B2F4A7EBFFD5FF969E456230FB2586CB" authority="Watanabe, 1990" authorityName="Watanabe" authorityYear="1990" box="[850,1169,367,392]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="tsurumii">
|
||
<emphasis id="4780007AFFD5FF969E456230FC6186C4" box="[850,981,368,392]" italics="true" pageId="6" pageNumber="527">M. tsurumii</emphasis>
|
||
<bibRefCitation id="1165A199FFD5FF969ECA622EFB2586CB" box="[989,1169,367,392]" pageId="6" pageNumber="527" refId="ref8184" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article">Watanabe, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
(central Honshu), from which it differs by the narrower elytra and apical portions of parameres and wider apical part of the median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DEE8FE3FFD5FF969DD0629DFADC857E" pageId="6" pageNumber="527" type="distribution">
|
||
<paragraph id="754BDC68FFD5FF969DD0629DFF56855B" blockId="6.[151,1436,151,644]" pageId="6" pageNumber="527">
|
||
<emphasis id="4780007AFFD5FF969DD0629DFED486B6" bold="true" box="[199,352,476,501]" pageId="6" pageNumber="527">Distribution.</emphasis>
|
||
The new species is known from several locations in the Emei Shan range (
|
||
<figureCitation id="EDCFC0EDFFD5FF9699D2629DFABF86B7" box="[1221,1291,476,501]" captionStart="FIGURES 9–10" captionStartId="7.[151,264,1329,1352]" captionTargetBox="[201,1360,771,1291]" captionTargetId="figure@7.[187,1400,756,1330]" captionTargetPageId="7" captionText="FIGURES 9–10. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri="https://zenodo.org/record/1221086/files/figure.png" pageId="6" pageNumber="527">Fig. 9</figureCitation>
|
||
) in
|
||
<collectingRegion id="B730128AFFD5FF96982C629DFA2C86B6" box="[1339,1432,476,501]" country="China" name="Sichuan" pageId="6" pageNumber="527">Sichuan</collectingRegion>
|
||
,
|
||
<collectingCountry id="0DE39CF8FFD5FF969D8062BEFF6A855B" box="[151,222,511,536]" name="China" pageId="6" pageNumber="527">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="754BDC68FFD5FF969DD06165FADC857E" blockId="6.[151,1436,151,644]" box="[199,1384,548,573]" pageId="6" pageNumber="527">
|
||
<emphasis id="4780007AFFD5FF969DD06165FEFE857E" bold="true" box="[199,330,548,573]" pageId="6" pageNumber="527">Bionomics.</emphasis>
|
||
All specimens were taken by sifting forest floor litter at elevations from
|
||
<quantity id="B20C718DFFD5FF96996B6165FA98857F" box="[1148,1324,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.9315" metricValueMax="2.4" metricValueMin="1.463" pageId="6" pageNumber="527" unit="m" value="1931.5" valueMax="2400.0" valueMin="1463.0">
|
||
<date id="014AFAA8FFD5FF96996B6165FABB857F" box="[1148,1295,548,573]" pageId="6" pageNumber="527" value="1463" valueMax="2400">1463 to 2400</date>
|
||
m
|
||
</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DEE8FE3FFD5FF969DD06106FF5C85C7" pageId="6" pageNumber="527" type="etymology">
|
||
<paragraph id="754BDC68FFD5FF969DD06106FF5C85C7" blockId="6.[151,1436,151,644]" pageId="6" pageNumber="527">
|
||
<emphasis id="4780007AFFD5FF969DD06106FEFA8523" bold="true" box="[199,334,583,608]" pageId="6" pageNumber="527">Etymology.</emphasis>
|
||
The species epitet is the Latinized adjective of Emei Shan, a mountain range, where the species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |