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<document id="115168D527D8E233925D6202A76848D6" ID-CLB-Dataset="30267" ID-DOI="10.11646/zootaxa.4407.4.4" ID-GBIF-Dataset="a850a99f-acfa-4f8e-b098-e7e783587f92" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221080" ID-ZooBank="D112CD45-7907-4F42-9D32-0DC7DC3A39EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524143490108" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2018" docId="FD5D6D7EFFD5FF979D80618BFC27858E" docLanguage="en" docName="zootaxa.4407.4.4.pdf" docOrigin="Zootaxa 4407 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia maculata Shavrin 2018, sp.n." docType="treatment" docVersion="7" lastPageNumber="528" masterDocId="01641506FFD3FF909D176341FFB48743" masterDocTitle="Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan" masterLastPageNumber="532" masterPageNumber="521" pageNumber="527" updateTime="1698537035611" updateUser="ExternalLinkService">
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<mods:title id="CED8DBE6A180E1C0FEC4BB02215ED174">Four new species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae) from south-western China and Taiwan</mods:title>
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<mods:namePart id="7C8493311BEA60C33C84FADDF859B51D">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="7CFDF00F8A1484F7264670A43A11DFAC">2018</mods:date>
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<treatment id="FD5D6D7EFFD5FF979D80618BFC27858E" ID-DOI="http://doi.org/10.5281/zenodo.5949832" ID-GBIF-Taxon="143605183" ID-Zenodo-Dep="5949832" LSID="urn:lsid:plazi:treatment:FD5D6D7EFFD5FF979D80618BFC27858E" httpUri="http://treatment.plazi.org/id/FD5D6D7EFFD5FF979D80618BFC27858E" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527">
<subSubSection id="3DEE8FE3FFD5FF969D80618BFED18445" pageId="6" pageNumber="527" type="nomenclature">
<paragraph id="754BDC68FFD5FF969D80618BFDC385A7" blockId="6.[151,631,714,774]" box="[151,631,714,740]" pageId="6" pageNumber="527">
<heading id="2E036B04FFD5FF969D80618BFDC385A7" bold="true" box="[151,631,714,740]" fontSize="11" level="1" pageId="6" pageNumber="527" reason="1">
<emphasis id="4780007AFFD5FF969D80618BFDC385A7" bold="true" box="[151,631,714,740]" pageId="6" pageNumber="527">
<taxonomicName id="B2F4A7EBFFD5FF969D80618BFD9B85A7" ID-CoL="3XVMM" authority="Shavrin" authorityName="Shavrin" authorityYear="2018" box="[151,559,714,740]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
<emphasis id="4780007AFFD5FF969D80618BFE7785A7" bold="true" box="[151,451,714,740]" italics="true" pageId="6" pageNumber="527">Mannerheimia maculata</emphasis>
Shavrin
</taxonomicName>
,
<taxonomicNameLabel id="5CB3BD01FFD5FF969F2B618BFDC385A7" box="[572,631,714,740]" pageId="6" pageNumber="527" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="754BDC68FFD5FF969D8061ACFED18445" blockId="6.[151,631,714,774]" box="[151,357,749,774]" pageId="6" pageNumber="527">
(
<figureCitation id="EDCFC0EDFFD5FF969D8861ACFF598445" box="[159,237,749,774]" captionStart="FIGURES 14" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="6" pageNumber="527">Figs. 3</figureCitation>
,
<figureCitation id="EDCFC0EDFFD5FF969DED61AFFEBE8445" box="[250,266,750,774]" captionStart="FIGURES 910" captionStartId="7.[151,264,1329,1352]" captionTargetBox="[201,1360,771,1291]" captionTargetId="figure@7.[187,1400,756,1330]" captionTargetPageId="7" captionText="FIGURES 910. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri="https://zenodo.org/record/1221086/files/figure.png" pageId="6" pageNumber="527">9</figureCitation>
,
<figureCitation id="EDCFC0EDFFD5FF969C0161AFFEEA8445" box="[278,350,750,774]" captionStart="FIGURES 1215" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 1215. Aedeagus of Mannerheimia maculata (1213: paratype) and M. taiwanica (1415: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="6" pageNumber="527">1213</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3DEE8FE3FFD5FF969D806074FB6B832D" pageId="6" pageNumber="527" type="materials_examined">
<paragraph id="754BDC68FFD5FF969D806074FB5C84D5" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">
<emphasis id="4780007AFFD5FF969D806074FE73840D" bold="true" box="[151,455,821,846]" pageId="6" pageNumber="527">
<typeStatus id="AA4F62CAFFD5FF969D806074FF66840D" box="[151,210,821,846]" pageId="6" pageNumber="527">Type</typeStatus>
material examined:
</emphasis>
<typeStatus id="AA4F62CAFFD5FF969CC56074FD8E840D" box="[466,570,821,846]" pageId="6" pageNumber="527" type="holotype">Holotype</typeStatus>
<specimenCount id="63F217E1FFD5FF969F536077FDED840E" box="[580,601,822,845]" pageId="6" pageNumber="527" type="male"></specimenCount>
[specimen dissected]:
<collectingCountry id="0DE39CF8FFD5FF969E706077FC73840D" box="[871,967,822,846]" name="China" pageId="6" pageNumber="527">CHINA</collectingCountry>
,
<collectingRegion id="B730128AFFD5FF969EC16077FB9B840E" box="[982,1071,822,845]" country="China" name="Yunnan" pageId="6" pageNumber="527">Yunnan</collectingRegion>
,
<quantity id="B20C718DFFD5FF96992B6074FBC9840D" box="[1084,1149,821,846]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="527" unit="km" value="15.0">15km</quantity>
| W Deqin, Mingyong, |
<geoCoordinate id="10C0BAAFFFD5FF969D80601BFEAD8431" box="[151,281,858,882]" direction="north" orientation="latitude" pageId="6" pageNumber="527" precision="15" value="28.461945">N28°27'43''</geoCoordinate>
<geoCoordinate id="10C0BAAFFFD5FF969C366018FE168431" box="[289,418,857,882]" direction="east" orientation="longitude" pageId="6" pageNumber="527" precision="15" value="98.781395">E98°46'53''</geoCoordinate>
, |
<date id="014AFAA8FFD5FF969CAC6018FD938431" box="[443,551,857,882]" pageId="6" pageNumber="527" value="2012-06-08">8.vi.2012</date>
, 2546m, | sift18, V. Grebennikov &lt;rectangular label, printed&gt;,
<typeStatus id="AA4F62CAFFD5FF9699E1601BFA3B8431" box="[1270,1423,858,882]" pageId="6" pageNumber="527" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="B2F4A7EBFFD5FF969D80603CFE8E84D5" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[151,314,893,918]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
|
<taxonomicName id="B2F4A7EBFFD5FF969C59603CFE0784D5" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[334,435,893,918]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">maculata</taxonomicName>
<taxonomicNameLabel id="5CB3BD01FFD5FF969CAD603FFE5B84D6" box="[442,495,894,917]" pageId="6" pageNumber="527" rank="species">
<emphasis id="4780007AFFD5FF969CAD603FFE5B84D6" bold="true" box="[442,495,894,917]" pageId="6" pageNumber="527">sp.n.</emphasis>
</taxonomicNameLabel>
| Shavrin A.V. des. 2017 &lt;red rectangular label, printed&gt; (
<collectionCode id="13E544ADFFD5FF96999F603FFB6884D5" box="[1160,1244,894,918]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="527">NSMT</collectionCode>
).
</paragraph>
<paragraph id="754BDC68FFD5FF969DD060E3FB6B832D" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">
Paratypes (6 specimens): 2 ♂, 2 ♀ [both specimens without right antennomeres 311]: same label as the holotype (1 ♂, 1 ♀: CS; 1 ♂, 1 ♀: NSMT); 1 ♂: CHINA, Yunnan,
<quantity id="B20C718DFFD5FF969EA36084FC40849D" box="[948,1012,965,990]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="527" unit="km" value="15.0">15km</quantity>
| W Deqin, Mingyong, | N28°27'29'' E98°45'28'', |
<date id="014AFAA8FFD5FF969C2D60A8FE128341" box="[314,422,1001,1026]" pageId="6" pageNumber="527" value="2012-06-07">7.vi.2012</date>
, 3289m, | sift16, V. Grebennikov &lt;rectangular label, printed&gt; (CNC); 1 ♂ [specimen (without right elytron) dissected]: CHINA, Yunnan,
<quantity id="B20C718DFFD5FF969E12674CFCF18365" box="[773,837,1037,1062]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="527" unit="km" value="15.0">15km</quantity>
| W Deqin, Mingyong, | N28°27'39'' E98°46'19'', |
<date id="014AFAA8FFD5FF969D806770FEB78309" box="[151,259,1073,1098]" pageId="6" pageNumber="527" value="2012-06-08">8.vi.2012</date>
, 2735m, | sift17, V. Grebennikov &lt;rectangular label, printed&gt; (NSMT). All paratypes with additional red rectangular printed label: PARATYPE |
<taxonomicName id="B2F4A7EBFFD5FF969F486714FCB6832D" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[607,770,1109,1134]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="genus" status="sp. nov.">Mannerheimia</taxonomicName>
|
<taxonomicName id="B2F4A7EBFFD5FF969E016714FCCF832D" authority="Shavrin, 2018" authorityName="Shavrin" authorityYear="2018" box="[790,891,1109,1134]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">maculata</taxonomicName>
<taxonomicNameLabel id="5CB3BD01FFD5FF969E956717FC0C832E" box="[898,952,1110,1133]" pageId="6" pageNumber="527" rank="species">
<emphasis id="4780007AFFD5FF969E956717FC0C832E" bold="true" box="[898,952,1110,1133]" pageId="6" pageNumber="527">sp.n.</emphasis>
</taxonomicNameLabel>
| Shavrin A.V. des. 2017.
</paragraph>
</subSubSection>
<subSubSection id="3DEE8FE3FFD5FF979DD06738FBB7857F" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527" type="description">
<paragraph id="754BDC68FFD5FF969DD06738FB8A8399" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">
<emphasis id="4780007AFFD5FF969DD06738FEE583D1" bold="true" box="[199,337,1145,1170]" pageId="6" pageNumber="527">Description</emphasis>
. Measurements (n=7): HW: 0.480.51; HL: 0.260.28; AL (
<typeStatus id="AA4F62CAFFD5FF96993C6738FB2083D1" box="[1067,1172,1145,1170]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
): 0.61; OL: 0.13; PL: 0.370.38; PW: 0.700.75; EL: 0.76; EW: 0.941.07; AW: 0.891.05; MTbL (
<typeStatus id="AA4F62CAFFD5FF96990267DCFBCA83F5" box="[1045,1150,1181,1206]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
): 0.41; MTrL (
<typeStatus id="AA4F62CAFFD5FF96983067DCFA2483F5" box="[1319,1424,1181,1206]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
): 0.24 (MTrL 14: 0.15; MTrL 5: 0.09); AedL: 0.37; TL: 2.222.57 (
<typeStatus id="AA4F62CAFFD5FF969E9E6780FC458399" box="[905,1009,1217,1242]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
: 2.25).
</paragraph>
<paragraph id="754BDC68FFD5FF969DD067A4FC8781C9" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">
Body moderately elongate, convex. Yellow-brown to reddish-brown, head and abdomen (except paratergites) brown in some; antennomeres 611 yellow brown; mouthparts, antennomeres 15, sometimes lateral and basal margins of pronotum, latero-basal and apical portions of elytra, legs, paratergites and apical abdominal tergites yellow. Body shiny, without microsculpture except postocular portions of head and neck with very fine and indistinct transverse microsculpture and abdominal tergites with distinct isodiametric microscuilpture. Head with irregular, moderately dense and deep punctation, distinctly larger and deeper on posterolateral portions of head and between ocelli, sometimes with impunctate transverse portion in middle at level of posterior third of eyes; punctation of pronotum as that of head, but very dense and more or less regular, with small impunctate portion in medio-basal third; scutellum with several very small punctures; punctation of elytra as that of pronotum, each elytron with six to seven tangled and vague longitudinal rows of punctures, remaining elytral punctures forming indistinct and tangled diagonal rows and becoming irregularly scattered laterally; abdominal tergites with very indistinct and sparse, small punctation. Habitus as in
<figureCitation id="EDCFC0EDFFD5FF969FFC6530FC9A81C9" box="[747,814,1649,1674]" captionStart="FIGURES 14" captionStartId="3.[151,264,1963,1986]" captionTargetBox="[242,1406,245,1886]" captionTargetId="figure@3.[151,1436,235,1895]" captionTargetPageId="3" captionText="FIGURES 14. Habitus of Mannerheimia: 1—M. grandilobata (paratype), 2—M. emeishanensis (paratype), 3—M. maculata (paratype), 4—M. taiwanica (holotype).Scale bar:1.0 mm." httpUri="https://zenodo.org/record/1221082/files/figure.png" pageId="6" pageNumber="527">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="754BDC68FFD5FF969DD065D4FAC1807D" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">
Head 1.8 times as wide as long, without occipital furrow between ocelli, with indistinct short grooves in front of ocelli. Ocelli moderately large, slightly convex; distance between ocelli one and a half to two times as long as distance between ocellus and posterior margin of eye. Antenna with antennomeres gradually widened toward apex, antennomere 4 1.5× as long as wide; length × width of antennomeres (
<typeStatus id="AA4F62CAFFD5FF969EA76440FBAD8059" box="[944,1049,1793,1818]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
): 1: 0.06 × 0.05; 2: 0.06 × 0.04; 3: 0.07 × 0.03; 4: 0.02 × 0.03; 5: 0.03 × 0.04; 6: 0.03 × 0.05; 78: 0.05 × 0.05; 910: 0.06 × 0.06; 11: 0.12 × 0.06.
</paragraph>
<paragraph id="754BDC68FFD5FF969DD0640BFD3380E9" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">Pronotum transverse, 1.81.9 times as wide as long, 1.4 times as wide as head, widest in middle, gradually slightly narrowed anterad and posterad; anterior and posterior angles widely rounded; lateral portions narrow, indistinctly explanate, without pit at middle.</paragraph>
<paragraph id="754BDC68FFD5FF969DD064F4FE7480B1" blockId="6.[151,1437,821,2034]" pageId="6" pageNumber="527">Elytra 1.2 times as wide as long, twice as long as pronotum; latero-apical margins of elytra with very indistinct and smoothed crenulation.</paragraph>
<paragraph id="754BDC68FFD4FF979DD063D9FD2887F3" blockId="7.[151,1436,151,717]" box="[199,668,151,176]" pageId="7" pageNumber="528">Metatarsus 1.7 times as long as metatibia.</paragraph>
<paragraph id="754BDC68FFD4FF979DD063FDFB3F8797" blockId="7.[151,1436,151,717]" box="[199,1163,188,213]" pageId="7" pageNumber="528">Abdomen with a pair of moderately large oval tomentose spots on middle of tergite V.</paragraph>
<paragraph id="754BDC68FFD4FF979DD0639EFF5A86CB" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="528">
Male. Protarsomeres 14 slightly widened. Apical margin of abdominal tergite VIII and sternite VIII widely and moderately deeply sinuate. Aedeagus (
<figureCitation id="EDCFC0EDFFD4FF979F966245FD61865F" box="[641,725,260,285]" captionStart="FIGURES 1215" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 1215. Aedeagus of Mannerheimia maculata (1213: paratype) and M. taiwanica (1415: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="7" pageNumber="528">Fig. 12</figureCitation>
) with wide basal part of the median lobe gradually narrowing toward apex, with very narrow apical projection; parameres wide, slightly exceeding apex of aedeagus; internal sac moderately long and narrow, with two fields of sclerites consisting of numerous thorns. Aedeagus laterally as in
<figureCitation id="EDCFC0EDFFD4FF979D80622EFF5D86CB" box="[151,233,367,392]" captionStart="FIGURES 1215" captionStartId="9.[151,264,1980,2003]" captionTargetBox="[363,1366,219,1902]" captionTargetId="figure@9.[176,1411,193,1959]" captionTargetPageId="9" captionText="FIGURES 1215. Aedeagus of Mannerheimia maculata (1213: paratype) and M. taiwanica (1415: paratype): 12, 14—parameral view, 13, 15—lateral view.Scale bar:0.1 mm." httpUri="https://zenodo.org/record/1221092/files/figure.png" pageId="7" pageNumber="528">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="754BDC68FFD4FF979DD062D5FA3A86EE" blockId="7.[151,1436,151,717]" box="[199,1422,404,429]" pageId="7" pageNumber="528">Female. Protarsomeres 14 not widened. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
<paragraph id="754BDC68FFD4FF979DD062F6FBB7857F" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="528">
<emphasis id="4780007AFFD4FF979DD062F6FE058693" bold="true" box="[199,433,439,464]" pageId="7" pageNumber="528">Comparative notes.</emphasis>
Based on the body size, shape of the pronotum and antennomeres, by the very indistinct punctation of the abdomen and general shape of the aedeagus, the new species is most similar to the Himalayan
<taxonomicName id="B2F4A7EBFFD4FF97986A629CFE49855B" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Mannerheimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="4780007AFFD4FF97986A629CFF6F8554" italics="true" pageId="7" pageNumber="528">M. vicina</emphasis>
Coiffait, 1982 (Kashmir)
</taxonomicName>
, from which it differs by paler coloration of the body with yellow latero-basal and apical portions of elytra, the wider aedeagus, and wider and longer parameres.
</paragraph>
</subSubSection>
<subSubSection id="3DEE8FE3FFD4FF979DD06106FADC85C6" pageId="7" pageNumber="528" type="distribution">
<paragraph id="754BDC68FFD4FF979DD06106FB548523" blockId="7.[151,1436,151,717]" box="[199,1248,583,608]" pageId="7" pageNumber="528">
<emphasis id="4780007AFFD4FF979DD06106FED48523" bold="true" box="[199,352,583,608]" pageId="7" pageNumber="528">Distribution.</emphasis>
The new species is known the Meili Xue Shan range in
<collectingRegion id="B730128AFFD4FF979ECF6109FB85851C" box="[984,1073,584,607]" country="China" name="Yunnan" pageId="7" pageNumber="528">Yunnan</collectingRegion>
,
<collectingCountry id="0DE39CF8FFD4FF97992C6106FBCA8523" box="[1083,1150,583,608]" name="China" pageId="7" pageNumber="528">China</collectingCountry>
(
<figureCitation id="EDCFC0EDFFD4FF97999A6106FB668523" box="[1165,1234,583,608]" captionStart="FIGURES 910" captionStartId="7.[151,264,1329,1352]" captionTargetBox="[201,1360,771,1291]" captionTargetId="figure@7.[187,1400,756,1330]" captionTargetPageId="7" captionText="FIGURES 910. Distribution of Mannerheimia in Yunnan and Sichuan, south-western China (9—M. grandilobata (black circles), M.emeishanensis (white circle), M.maculata (black triangle)) and Taiwan (10—M.taiwanica (black circles))." httpUri="https://zenodo.org/record/1221086/files/figure.png" pageId="7" pageNumber="528">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="754BDC68FFD4FF979DD0612DFADC85C6" blockId="7.[151,1436,151,717]" box="[199,1384,620,645]" pageId="7" pageNumber="528">
<emphasis id="4780007AFFD4FF979DD0612DFEFE85C6" bold="true" box="[199,330,620,645]" pageId="7" pageNumber="528">Bionomics.</emphasis>
All specimens were taken by sifting forest floor litter at elevations from
<quantity id="B20C718DFFD4FF97996B612DFA9885C7" box="[1148,1324,620,645]" metricMagnitude="3" metricUnit="m" metricValue="2.9175" metricValueMax="3.289" metricValueMin="2.546" pageId="7" pageNumber="528" unit="m" value="2917.5" valueMax="3289.0" valueMin="2546.0">2546 to 3289 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="3DEE8FE3FFD4FF979DD061CEFC27858E" pageId="7" pageNumber="528" type="etymology">
<paragraph id="754BDC68FFD4FF979DD061CEFC27858E" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="528">
<emphasis id="4780007AFFD4FF979DD061CEFEFA85EB" bold="true" box="[199,334,655,680]" pageId="7" pageNumber="528">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="4780007AFFD4FF979E3161D1FC2D85EB" box="[806,921,656,680]" italics="true" pageId="7" pageNumber="528">maculatus</emphasis>
, -
<emphasis id="4780007AFFD4FF979EA761D0FC0A85EB" box="[944,958,657,680]" italics="true" pageId="7" pageNumber="528">a</emphasis>
, -
<emphasis id="4780007AFFD4FF979EC261D0FC4385EB" box="[981,1015,657,680]" italics="true" pageId="7" pageNumber="528">um</emphasis>
(blotched, maculated). It is alludes to the presence of yellow latero-basal and lateral portions of the elytra.
</paragraph>
</subSubSection>
</treatment>
</document>