197 lines
19 KiB
XML
197 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.986.56276" ID-GBIF-Dataset="b2e2aca7-d96d-4fb3-8d67-bb0a1adcb9e3" ID-PMC="PMC7661483" ID-Pensoft-Pub="1313-2970-986-1" ID-Pensoft-UUID="2A3CD185EDDF52BDB0AD1E5DA924299B" ID-PubMed="33223879" ID-ZooBank="CDFA38CA1B6F424D85246B540E63E954" ModsDocID="1313-2970-986-1" checkinTime="1604647765603" checkinUser="pensoft" docAuthor="Hrivniak, Ľubos, Sroka, Pavel, Bojkova, Jindriska & Godunko, Roman J." docDate="2020" docId="FA6CD7B919575AE8996221A1CB78E2E5" docLanguage="en" docName="ZooKeys 986: 1-53" docOrigin="ZooKeys 986" docSource="http://dx.doi.org/10.3897/zookeys.986.56276" docTitle="Epeorus (Caucasiron) shargi Hrivniak & Sroka 2020" docType="treatment" docVersion="4" id="2A3CD185EDDF52BDB0AD1E5DA924299B" lastPageNumber="1" masterDocId="2A3CD185EDDF52BDB0AD1E5DA924299B" masterDocTitle="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="1" updateTime="1668169768372" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hrivniak, Ľubos</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Faculty of Sciences, University of South Bohemia, Branisovska 31, 37005 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="email">lubos.hrivniak@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sroka, Pavel</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4367-6564</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bojkova, Jindriska</mods:namePart>
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<mods:affiliation>Department of Botany and Zoology, Masaryk University, Kotlarska 2, 61137 Brno, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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<mods:affiliation>Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005 Ceske Budejovice, Czech Republic & Department of Invertebrate Zoology and Hydrobiology, University of Lodz, Banacha 12 / 16, 90237 Lodz, Poland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-2196-3327</mods:nameIdentifier>
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<mods:nameIdentifier type="email">godunko@seznam.cz</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>986</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.986.56276</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.986.56276</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-986-1</mods:identifier>
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<mods:identifier type="ZooBank">CDFA38CA1B6F424D85246B540E63E954</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2A3CD185EDDF52BDB0AD1E5DA924299B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168815722" LSID="urn:lsid:plazi:treatment:FA6CD7B919575AE8996221A1CB78E2E5" httpUri="http://treatment.plazi.org/id/FA6CD7B919575AE8996221A1CB78E2E5" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="FA6CD7B9-1957-5AE8-9962-21A1CB78E2E5" authority="Hrivniak & Sroka, 2020" authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) shargi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">Epeorus (Caucasiron) shargi Hrivniak & Sroka, 2020</taxonomicName>
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Geographical (left) and vertical (right) distribution of Epeorus (Caucasiron) shargi." figureDoi="10.3897/zookeys.986.56276.figure42" httpUri="https://binary.pensoft.net/fig/471965" pageId="0" pageNumber="1">Figs 42</figureCitation>
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">, 43</figureCitation>
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Epeorus (Caucasiron) shargi, larva: A labrum (left half in dorsal view right half in ventral view) B incisors of left mandible C incisors of right mandible D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J abdominal segments VIII-X in lateral view K sternum IX of female with observed variability." figureDoi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1">, 44</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Iran, Golestan Province, Shirinabad village, unnamed river;
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<geoCoordinate degrees="36" direction="north" minutes="48" orientation="latitude" precision="1" seconds="01.44" value="36.8004">36°48'01.44"N</geoCoordinate>
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/
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<geoCoordinate degrees="55" direction="east" minutes="01" orientation="longitude" precision="1" seconds="05.78" value="55.018272">55°01'05.78"E</geoCoordinate>
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(locality no. 108); 740 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Northern Iran. Known only from three sites in the eastern Alborz (Fig.
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Geographical (left) and vertical (right) distribution of Epeorus (Caucasiron) shargi." figureDoi="10.3897/zookeys.986.56276.figure42" httpUri="https://binary.pensoft.net/fig/471965" pageId="0" pageNumber="1">42</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="habitat">
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<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Larvae inhabit streams at middle altitude, 740-1450 m a.s.l. (Fig.
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Geographical (left) and vertical (right) distribution of Epeorus (Caucasiron) shargi." figureDoi="10.3897/zookeys.986.56276.figure42" httpUri="https://binary.pensoft.net/fig/471965" pageId="0" pageNumber="1">42</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="main morphological diagnostics of larvae">
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<paragraph pageId="0" pageNumber="1">Main morphological diagnostics of larvae.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(i) abdominal terga V-VII with triangular or T-shaped medial macula (Fig.
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">43I-K</figureCitation>
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); (ii) abdominal sterna without coloration pattern (Fig.
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">43B, L</figureCitation>
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); (iii) tergum X without postero-lateral projections (Fig.
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Epeorus (Caucasiron) shargi, larva: A labrum (left half in dorsal view right half in ventral view) B incisors of left mandible C incisors of right mandible D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J abdominal segments VIII-X in lateral view K sternum IX of female with observed variability." figureDoi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1">44J</figureCitation>
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); (iv) femora with medial hypodermal spot (Fig.
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">43F-H</figureCitation>
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); (v) gill plates VII (in natural position from ventral view) wide (Figs
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">43M</figureCitation>
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,
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Epeorus (Caucasiron) shargi, larva: A labrum (left half in dorsal view right half in ventral view) B incisors of left mandible C incisors of right mandible D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J abdominal segments VIII-X in lateral view K sternum IX of female with observed variability." figureDoi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1">44H, I</figureCitation>
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); (vi) setae on abdominal terga hair-like (Fig.
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Epeorus (Caucasiron) shargi, larva: A labrum (left half in dorsal view right half in ventral view) B incisors of left mandible C incisors of right mandible D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J abdominal segments VIII-X in lateral view K sternum IX of female with observed variability." figureDoi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1">44E</figureCitation>
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); (vii) gill plates III with well-developed projection (Fig.
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<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Epeorus (Caucasiron) shargi, larva: A labrum (left half in dorsal view right half in ventral view) B incisors of left mandible C incisors of right mandible D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J abdominal segments VIII-X in lateral view K sternum IX of female with observed variability." figureDoi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1">44G</figureCitation>
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); (viii) shape of head of male oval trapezoidal (Fig.
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<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Epeorus (Caucasiron) shargi, larva: A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F-H middle leg in dorsal view I-K abdominal terga L abdominal sterna II-VI M gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1">43D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Taxonomy</emphasis>
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.
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</emphasis>
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This species was described based on larvae collected from Alborz Mts. (
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.947.51259" author="Hrivniak, Ľ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="71 - 102" refId="B21" refString="Hrivniak, Ľ, Sroka, P, Bojkova, J, Godunko, RJ, Imanpour Namin, J, Bagheri, S, Nejat, F, Abdoli, A, Staniczek, AH, 2020a. Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) in Iran, with descriptions of three new species. ZooKeys 947: 71 - 102, DOI: https://doi.org/10.3897/zookeys.947.51259" title="Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) in Iran, with descriptions of three new species." url="https://doi.org/10.3897/zookeys.947.51259" volume="947" year="2020 a">Hrivniak et al. 2020a</bibRefCitation>
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). Imagines not described. The type series is currently deposited in SMNS, IECA, and MMTT_DOE.
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</paragraph>
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<caption doi="10.3897/zookeys.986.56276.figure42" httpUri="https://binary.pensoft.net/fig/471965" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
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Geographical (left) and vertical (right) distribution of
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<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) shargi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">Epeorus (Caucasiron) shargi</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.986.56276.figure43" httpUri="https://binary.pensoft.net/fig/471966" pageId="0" pageNumber="1" start="Figure 43" startId="F43">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 43.</emphasis>
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<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) shargi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">Epeorus (Caucasiron) shargi</taxonomicName>
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, larva:
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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habitus in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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habitus in ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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habitus in lateral view
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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head of male in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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head of female in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">F-H</emphasis>
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middle leg in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">I-K</emphasis>
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abdominal terga
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<emphasis bold="true" pageId="0" pageNumber="1">L</emphasis>
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abdominal sterna II-VI
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<emphasis bold="true" pageId="0" pageNumber="1">M</emphasis>
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gills VII (in natural position from ventral view).
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.986.56276.figure44" httpUri="https://binary.pensoft.net/fig/471967" pageId="0" pageNumber="1" start="Figure 44" startId="F44">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 44.</emphasis>
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<taxonomicName authorityName="Hrivniak & Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus (Caucasiron) shargi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">Epeorus (Caucasiron) shargi</taxonomicName>
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, larva:
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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labrum (left half in dorsal view right half in ventral view)
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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incisors of left mandible
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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incisors of right mandible
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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setae on dorsal surface of femora
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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surface and posterior margin of abdominal tergum VII
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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gill I
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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gill III
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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gill VII (flattened on slide)
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<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
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gill VII (in natural position from ventral view)
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<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
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abdominal segments VIII-X in lateral view
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<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
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sternum IX of female with observed variability.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |