358 lines
47 KiB
XML
358 lines
47 KiB
XML
<document id="75CEAD2BF22BC3044BAAFF04D47DE98E" ID-DOI="10.1206/0003-0082(2006)3539[1:RCIBAN]2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5385443" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630301771567" checkinUser="carolina" docAuthor="ARILLO, ANTONIO & ENGEL, MICHAEL S" docDate="2006" docId="14292535A642FFBCC0F9C5A0FB45FD2A" docLanguage="en" docName="N3539.pdf" docOrigin="American Museum Novitates 3539 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2006)3539%5B1%3ARCIBAN%5D2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Adicophasma grylloblattoides ARILLO & ENGEL 2006, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="E8105D4DA640FFBAC061C06FFF89FFC0" masterDocTitle="Rock Crawlers in Baltic Amber (Notoptera: Mantophasmatodea)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="3" updateTime="1699220750920" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="C32BD0CAD693AD60129B4F5D0FB5CB20" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="53E669845C25C3DB4DFE6098BF4D544C">
|
||
<mods:title id="403FB1D47D07CC3A2F4E77EC54A12D75">Rock Crawlers in Baltic Amber (Notoptera: Mantophasmatodea)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E3B21C8FEF4C55F8A82844E65DD6A5D6" type="personal">
|
||
<mods:role id="A5E72FAD4918A2D535A4901FF824531D">
|
||
<mods:roleTerm id="89B8D1B4565B24143A26270F35739505">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CB839DA19B28CC7FAEC6F88BAB6A25D4">ARILLO, ANTONIO</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2FC063CCCD2A6FFEA3033D96D5237AAB" type="personal">
|
||
<mods:role id="080F9A70160EFFB77A76AA472BEA70AB">
|
||
<mods:roleTerm id="C24764D29FE3900539C08AD13EB18672">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0A36042E43C2BBB67CF861199B28AB7D">ENGEL, MICHAEL S</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="2402F9E63237266DE33F0F4C0E998159">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2A85DEA21450960DD44CDC5E9FFDC4C4" type="host">
|
||
<mods:titleInfo id="49F3F64E733F5983860FA77AA7E8771E">
|
||
<mods:title id="E4ABD52E2BF1796D11C06FB896454AD1">American Museum Novitates</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CD14FE7B53167D5638FFEDA648FFC788">
|
||
<mods:date id="64CD41343562669834CE5C8052089568">2006</mods:date>
|
||
<mods:detail id="7E9EBC1A0F24E018898FB9222E5F6097" type="pubDate">
|
||
<mods:number id="55F5DE0AAD63886B6456AD003B15D0C5">2006-12-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CBD56C226A20AA1C772E13C728572295" type="volume">
|
||
<mods:number id="5E889367DC79AC18A3288A69C6110AFE">3539</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FE724FE224C302820050BEAD48AFAF11" type="issue">
|
||
<mods:number id="F7DEFF58F3D96E77FCE246958FFDA4CA">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A6A2CA214C31E9795E579F2C8D4723B7" unit="page">
|
||
<mods:start id="8DE9A8BF8288AF8AE8269289A128C6B5">1</mods:start>
|
||
<mods:end id="BB923135331579364D50AD21ACA8820E">10</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="13E70349B7B61D16DB1A9AB89C9FBBA9">
|
||
<mods:url id="3F949F212A717E7FFC813B786985C51F">http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2006)3539%5B1%3ARCIBAN%5D2.0.CO%3B2</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="7B36ADF62744E56D4F4B0214E10C51E9">journal article</mods:classification>
|
||
<mods:identifier id="74D848BE4F2399E484D2B3F6123DE5F8" type="DOI">10.1206/0003-0082(2006)3539[1:RCIBAN]2.0.CO;2</mods:identifier>
|
||
<mods:identifier id="B938951B0588E350B1D7F4D6E004A367" type="ISSN">0003-0082</mods:identifier>
|
||
<mods:identifier id="EFB724DD011C5A9A02051F4A113F501B" type="Zenodo-Dep">5385443</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="14292535A642FFBCC0F9C5A0FB45FD2A" LSID="urn:lsid:plazi:treatment:14292535A642FFBCC0F9C5A0FB45FD2A" httpUri="http://treatment.plazi.org/id/14292535A642FFBCC0F9C5A0FB45FD2A" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
|
||
<subSubSection id="D49AC7A8A642FFB8C0F9C5A0FE4CF9D7" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="9C3F9423A642FFB8C0F9C5A0FE4CF9D7" blockId="2.[152,629,1487,1511]" lastBlockId="2.[328,453,1535,1559]" pageId="2" pageNumber="3">
|
||
<heading id="C777234FA642FFB8C0F9C5A0FDFCFA27" box="[152,629,1487,1511]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="3" reason="8">
|
||
<taxonomicName id="5B80EFA0A642FFB8C0F9C5A0FE68FA27" ID-CoL="64T2Q" authority="ARILLO & ENGEL, 2006" authorityName="ARILLO & ENGEL" authorityYear="2006" box="[152,481,1487,1511]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A642FFB8C0F9C5A0FE68FA27" bold="true" box="[152,481,1487,1511]" italics="true" pageId="2" pageNumber="3">Adicophasma grylloblattoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B5C7F54AA642FFB8C190C5A0FDFCFA27" box="[497,629,1487,1511]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
|
||
</heading>
|
||
<figureCitation id="04BB88A6A642FFB8C129C590FE4CF9D7" box="[328,453,1535,1559]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[112,150,716,738]" captionStartId-1="3.[112,150,1705,1727]" captionStartId-2="4.[151,189,692,714]" captionStartId-3="4.[151,189,1705,1727]" captionTargetBox-0="[88,1191,210,682]" captionTargetBox-1="[105,1169,1326,1670]" captionTargetBox-2="[306,1049,211,658]" captionTargetBox-3="[205,1150,749,1670]" captionTargetId-0="figure-220@3.[88,1191,209,682]" captionTargetId-1="figure-237@3.[104,1170,1326,1671]" captionTargetId-2="figure-1@4.[306,1049,211,658]" captionTargetId-3="figure-19@4.[205,1150,749,1670]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="Fig. 1. Photomicrograph of holotype of Adicophasma grylloblattoides Arillo and Engel, new species" captionText-1="Fig. 2. Illustration of holotype of Adicophasma grylloblattoides Arillo and Engel, new species" captionText-2="Fig. 3. Photomicrograph of holotype foreleg of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." captionText-3="Fig. 4. Illustration of head and pro- and mesothoracic segments of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." figureDoi-0="http://doi.org/10.5281/zenodo.5385445" figureDoi-1="http://doi.org/10.5281/zenodo.5385447" figureDoi-2="http://doi.org/10.5281/zenodo.5385449" figureDoi-3="http://doi.org/10.5281/zenodo.5385451" httpUri-0="https://zenodo.org/record/5385445/files/figure.png" httpUri-1="https://zenodo.org/record/5385447/files/figure.png" httpUri-2="https://zenodo.org/record/5385449/files/figure.png" httpUri-3="https://zenodo.org/record/5385451/files/figure.png" pageId="2" pageNumber="3">figures 1–4</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A642FFB8C0FBC641FBD5FB26" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="9C3F9423A642FFB8C0FBC641FBD5FB26" blockId="2.[128,654,1582,1754]" lastBlockId="2.[701,1228,1201,1754]" pageId="2" pageNumber="3">
|
||
DIAGNOSIS: The new species differs from its congener,
|
||
<emphasis id="AEF44831A642FFB8C178C623FEA5F9A4" box="[281,300,1612,1636]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A642FFB8C15AC623FE29F9A4" box="[315,416,1612,1636]" italics="true" pageId="2" pageNumber="3">spinosum</emphasis>
|
||
Engel and Grimaldi (see appendix 1), by the shortened pedicel that is about as long as wide, the absence of mesofemoral spination, the absence of spines on the cardines, the proportions of the thoracic nota, and the more elongate abdomen (see also Key to Baltic Amber
|
||
<taxonomicName id="5B80EFA0A642FFB8C3E1C4A1FC73FB26" authorityName="Crampton" authorityYear="1915" box="[896,1018,1230,1254]" order="Notoptera" pageId="2" pageNumber="10" rank="order">Notoptera</taxonomicName>
|
||
, below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A642FFBFC2B9C483FF1EFB27" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="9C3F9423A642FFB8C2B9C483FB42F91A" blockId="2.[701,1228,1201,1754]" pageId="2" pageNumber="3">
|
||
DESCRIPTION:
|
||
<emphasis id="AEF44831A642FFB8C3E7C483FC57FAC4" bold="true" box="[902,990,1260,1284]" pageId="2" pageNumber="3">Female.</emphasis>
|
||
Total body length (excluding antennae) 6.0 mm (additional metrics provided in
|
||
<bibRefCitation id="F811E9D2A642FFB8C3AAC548FB49FAFF" author="Arillo, A. & V. M. Ortuno & A. Nel" box="[971,1216,1319,1343]" pageId="2" pageNumber="3" pagination="11 - 14" refId="ref3855" refString="Arillo, A., V. M. Ortuno, and A. Nel. 1997. Description of an enigmatic insect from Baltic amber. Bulletin de la Societe Entomologique de France 102 (1): 11 - 14." type="journal article" year="1997">Arillo et al., 1997</bibRefCitation>
|
||
). Coloration uniformly light brown. Head transverse. Face with sparse, simple setae; integument impunctate. Compound eyes large, extending entire length of head, encompassing almost entire lateral area of head; ommatidia small and numerous. Vertex straight. Gena impunctate. Malar space exceedingly short. Antennal socket apparently just below facial midlength and near compound eye; scape about equal in size to pedicel; pedicel about as long as wide; 16 flagellomeres elongate and with sparse setae. Maxillary palp five-segmented, basalmost palpomere smallest, approximately equal in
|
||
</paragraph>
|
||
<caption id="C8FFC4ABA643FFB9C011C2A3FB2AFD22" ID-DOI="http://doi.org/10.5281/zenodo.5385445" ID-Zenodo-Dep="5385445" box="[112,1187,716,738]" httpUri="https://zenodo.org/record/5385445/files/figure.png" pageId="3" pageNumber="4" startId="3.[112,150,716,738]" targetBox="[88,1191,210,682]" targetPageId="3">
|
||
<paragraph id="9C3F9423A643FFB9C011C2A3FB2AFD22" blockId="3.[88,1187,716,765]" box="[112,1187,716,738]" pageId="3" pageNumber="4">
|
||
Fig. 1. Photomicrograph of holotype of
|
||
<taxonomicName id="5B80EFA0A643FFB9C250C2A3FB95FD22" authority="Arillo and Engel" authorityName="Arillo and Engel" authorityYear="2006" box="[561,1052,716,738]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A643FFB9C250C2A3FCD5FD22" bold="true" box="[561,860,716,738]" italics="true" pageId="3" pageNumber="4">Adicophasma grylloblattoides</emphasis>
|
||
Arillo and Engel
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B5C7F54AA643FFB9C44BC2A3FB2AFD22" box="[1066,1187,716,738]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9C3F9423A643FFB9C039C288FE8BFD3D" blockId="3.[88,1187,716,765]" box="[88,258,743,765]" pageId="3" pageNumber="4">(MCNA 10686).</paragraph>
|
||
<paragraph id="9C3F9423A643FFB9C039C372FB2AFAC9" blockId="3.[88,614,797,1289]" lastBlockId="3.[662,1188,797,1289]" pageId="3" pageNumber="4">
|
||
size to second palpomere; third, fourth, and fifth maxillary palpomeres elongate; third palpomere longest; combined lengths of fourth and fifth palpomeres slightly longer than length of third palpomere; distinct setae scattered on apical palpomeres; stipes broad; cardo without spines. Coxae elongate; trochanters short; pro- and mesotibiae slightly longer than respective femora; metatibia longer than tibiae of preceding segments. Procoxa with three stout spines; profemur noticeably swollen, without dorsal carinae, with two longitudinal rows of stout spines ventrally (distinct ventral surface formed between rows of spines as in
|
||
<emphasis id="AEF44831A643FFB9C1A9C4D8FE52FB0F" box="[456,475,1207,1231]" italics="true" pageId="3" pageNumber="4">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A643FFB9C191C4D8FDDCFB0F" box="[496,597,1207,1231]" italics="true" pageId="3" pageNumber="4">spinosum</emphasis>
|
||
); inner surface of protibia with similar stiff, elongate, simple spines as well as elongate setae on outer surface (
|
||
<figureCitation id="04BB88A6A643FFB9C3D5C372FB90FCF5" box="[948,1049,797,821]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[112,150,1705,1727]" captionStartId-1="4.[151,189,692,714]" captionStartId-2="4.[151,189,1705,1727]" captionTargetBox-0="[105,1169,1326,1670]" captionTargetBox-1="[306,1049,211,658]" captionTargetBox-2="[205,1150,749,1670]" captionTargetId-0="figure-237@3.[104,1170,1326,1671]" captionTargetId-1="figure-1@4.[306,1049,211,658]" captionTargetId-2="figure-19@4.[205,1150,749,1670]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionText-0="Fig. 2. Illustration of holotype of Adicophasma grylloblattoides Arillo and Engel, new species" captionText-1="Fig. 3. Photomicrograph of holotype foreleg of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." captionText-2="Fig. 4. Illustration of head and pro- and mesothoracic segments of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." figureDoi-0="http://doi.org/10.5281/zenodo.5385447" figureDoi-1="http://doi.org/10.5281/zenodo.5385449" figureDoi-2="http://doi.org/10.5281/zenodo.5385451" httpUri-0="https://zenodo.org/record/5385447/files/figure.png" httpUri-1="https://zenodo.org/record/5385449/files/figure.png" httpUri-2="https://zenodo.org/record/5385451/files/figure.png" pageId="3" pageNumber="4">figs. 2–4</figureCitation>
|
||
); tarsi pentamerous, basal four tarsomeres with euplantulae; tarsomere IV weakly bilobed such that origin of tarsomere V slightly recessed, tarsomere V elongate and slender, bearing simple pretarsal claws and large, broad arolium (
|
||
<figureCitation id="04BB88A6A643FFB9C2FCC3A2FD55FC25" box="[669,732,973,997]" captionStart="Fig" captionStartId="4.[151,189,1705,1727]" captionTargetBox="[205,1150,749,1670]" captionTargetId="figure-19@4.[205,1150,749,1670]" captionTargetPageId="4" captionText="Fig. 4. Illustration of head and pro- and mesothoracic segments of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." figureDoi="http://doi.org/10.5281/zenodo.5385451" httpUri="https://zenodo.org/record/5385451/files/figure.png" pageId="3" pageNumber="4">fig. 4</figureCitation>
|
||
); arolium without fans of elongate setae. Middle and hind legs similar in construction and with sparsely scattered, simple setae; mesofemur not as swollen as profemur and lacking spination. Thoracic nota longer than wide and each of approximately equal length and width, with exceedingly sparse, simple setae; integument impunctate; thorax about as broad as posterior width of head; pronotum only slightly longer than mesonotum. Abdomen elongate with sides relatively
|
||
</paragraph>
|
||
<caption id="C8FFC4ABA643FFB9C011C6C6FB2AF97F" ID-DOI="http://doi.org/10.5281/zenodo.5385447" ID-Zenodo-Dep="5385447" box="[112,1187,1705,1727]" httpUri="https://zenodo.org/record/5385447/files/figure.png" pageId="3" pageNumber="4" startId="3.[112,150,1705,1727]" targetBox="[105,1169,1326,1670]" targetPageId="3">
|
||
<paragraph id="9C3F9423A643FFB9C011C6C6FB2AF97F" blockId="3.[88,1187,1705,1753]" box="[112,1187,1705,1727]" pageId="3" pageNumber="4">
|
||
Fig. 2. Illustration of holotype of
|
||
<taxonomicName id="5B80EFA0A643FFB9C26CC6C6FB99F97F" authority="Arillo and Engel" authorityName="Arillo and Engel" authorityYear="2006" box="[525,1040,1705,1727]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A643FFB9C26CC6C6FCB7F97F" bold="true" box="[525,830,1705,1727]" italics="true" pageId="3" pageNumber="4">Adicophasma grylloblattoides</emphasis>
|
||
Arillo and Engel
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B5C7F54AA643FFB9C445C6C6FB2AF97F" box="[1060,1187,1705,1727]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9C3F9423A643FFB9C039C6ACFE8BF919" blockId="3.[88,1187,1705,1753]" box="[88,258,1731,1753]" pageId="3" pageNumber="4">(MCNA 10686).</paragraph>
|
||
<caption id="C8FFC4ABA644FFBEC0F6C2DBFEFEFD25" ID-DOI="http://doi.org/10.5281/zenodo.5385449" ID-Zenodo-Dep="5385449" httpUri="https://zenodo.org/record/5385449/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,189,692,714]" targetBox="[306,1049,211,658]" targetPageId="4">
|
||
<paragraph id="9C3F9423A644FFBEC0F6C2DBFEFEFD25" blockId="4.[128,1228,692,741]" pageId="4" pageNumber="5">
|
||
Fig. 3. Photomicrograph of holotype foreleg of
|
||
<taxonomicName id="5B80EFA0A644FFBEC2CBC2DBFB1DFD0A" authority="Arillo and Engel" authorityName="Arillo and Engel" authorityYear="2006" box="[682,1172,692,714]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A644FFBEC2CBC2DBFC5DFD0A" bold="true" box="[682,980,692,714]" italics="true" pageId="4" pageNumber="5">Adicophasma grylloblattoides</emphasis>
|
||
Arillo and Engel
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B5C7F54AA644FFBEC4C2C2DBFF4CFD25" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
|
||
(MCNA 10686).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C8FFC4ABA644FFBEC0F6C6C6FD96F919" ID-DOI="http://doi.org/10.5281/zenodo.5385451" ID-Zenodo-Dep="5385451" httpUri="https://zenodo.org/record/5385451/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,189,1705,1727]" targetBox="[205,1150,749,1670]" targetPageId="4">
|
||
<paragraph id="9C3F9423A644FFBEC0F6C6C6FD96F919" blockId="4.[128,1227,1705,1753]" pageId="4" pageNumber="5">
|
||
Fig. 4. Illustration of head and pro- and mesothoracic segments of
|
||
<taxonomicName id="5B80EFA0A644FFBEC301C6C6FF62F919" authority="Arillo and Engel" authorityName="Arillo and Engel" authorityYear="2006" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A644FFBEC301C6C6FB0EF97F" bold="true" box="[864,1159,1705,1727]" italics="true" pageId="4" pageNumber="5">Adicophasma grylloblattoides</emphasis>
|
||
Arillo and Engel
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B5C7F54AA644FFBEC097C6ACFEE5F919" box="[246,364,1731,1753]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
|
||
(MCNA 10686).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C8FFC4ABA645FFBFC011C3A3FCE6FBD7" ID-DOI="http://doi.org/10.5281/zenodo.5385454" ID-Zenodo-Dep="5385454" httpUri="https://zenodo.org/record/5385454/files/figure.png" pageId="5" pageNumber="6" startId="5.[112,150,972,994]" targetBox="[158,1117,210,938]" targetPageId="5">
|
||
<paragraph id="9C3F9423A645FFBFC011C3A3FCE6FBD7" blockId="5.[88,1187,972,1047]" pageId="5" pageNumber="6">
|
||
Fig. 5. Photomicrograph of holotype of
|
||
<emphasis id="AEF44831A645FFBFC246C3A3FC99FC21" box="[551,784,972,993]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="5B80EFA0A645FFBFC246C3A3FD25FC21" authorityName="Engel & Grimaldi" authorityYear="2004" box="[551,684,972,993]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Adicophasma</taxonomicName>
|
||
spinosum
|
||
</emphasis>
|
||
Engel and Grimaldi showing detail of forelegs, particularly the presence of profemoral and protibial spination (similar, albeit shorter and less stout, spines are also present on the midlegs; see
|
||
<bibRefCitation id="F811E9D2A645FFBFC234C46EFCE8FBD7" author="Engel, M. S. & D. A. Grimaldi" box="[597,865,1025,1047]" pageId="5" pageNumber="6" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi, 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9C3F9423A645FFBFC039C452FF1EFB27" blockId="5.[88,614,1085,1754]" pageId="5" pageNumber="6">parallel, segments short, terga and sterna broader than long; terga and sterna with scattered, simple setae. Cerci short, unsegmented, gently tapering to apices, covered with numerous erect to suberect elongate setae.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A645FFBFC013C482FE9BFA9D" pageId="5" pageNumber="6" type="materials_examined">
|
||
<paragraph id="9C3F9423A645FFBFC013C482FE9BFA9D" blockId="5.[88,614,1085,1754]" pageId="5" pageNumber="6">
|
||
<materialsCitation id="2CE89E7EA645FFBFC013C482FE9BFA9D" collectionCode="MCNA" country="Spain" location="Museo de Ciencias Naturales de Alava" municipality="Vitoria" pageId="5" pageNumber="6" specimenCode="MCNA 10686" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="433B2A81A645FFBFC013C482FF6AFAC3" box="[114,227,1261,1285]" pageId="5" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
|
||
: Female, Baltic amber (middle Eocene),
|
||
<specimenCode id="CC263C58A645FFBFC0A7C565FEE2FAE2" box="[198,363,1290,1314]" collectionCode="MCNA" country="Spain" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="5" pageNumber="6">MCNA 10686</specimenCode>
|
||
(
|
||
<figureCitation id="04BB88A6A645FFBFC1E1C565FE33FAE2" box="[384,442,1290,1314]" captionStart="Fig" captionStartId="3.[112,150,716,738]" captionTargetBox="[88,1191,210,682]" captionTargetId="figure-220@3.[88,1191,209,682]" captionTargetPageId="3" captionText="Fig. 1. Photomicrograph of holotype of Adicophasma grylloblattoides Arillo and Engel, new species" figureDoi="http://doi.org/10.5281/zenodo.5385445" httpUri="https://zenodo.org/record/5385445/files/figure.png" pageId="5" pageNumber="6">fig. 1</figureCitation>
|
||
), deposited in the
|
||
<location id="995FC2F8A645FFBFC0E8C548FDEFFA80" LSID="urn:lsid:plazi:treatment:14292535A642FFBCC0F9C5A0FB45FD2A:995FC2F8A645FFBFC0E8C548FDEFFA80" box="[137,614,1314,1344]" country="Spain" municipality="Vitoria" name="Museo de Ciencias Naturales de Alava" pageId="5" pageNumber="6">Museo de Ciencias Naturales de Álava</location>
|
||
(
|
||
<collectingMunicipality id="7C5B0E59A645FFBFC03EC52AFF3DFA9D" box="[95,180,1349,1373]" pageId="5" pageNumber="6">Vitoria</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="E497D4B3A645FFBFC0A0C52AFE8FFA9D" box="[193,262,1349,1373]" name="Spain" pageId="5" pageNumber="6">Spain</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A645FFBFC013C50CFE97FA12" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="9C3F9423A645FFBFC013C50CFE97FA12" blockId="5.[88,614,1085,1754]" pageId="5" pageNumber="6">
|
||
ETYMOLOGY: The specific epithet is a reference to the similarity and phylogenetic affinity of rock crawlers to the ice crawlers (suborder
|
||
<taxonomicName id="5B80EFA0A645FFBFC039C5D5FE9AFA12" baseAuthorityName="Walker" baseAuthorityYear="1914" box="[88,275,1466,1490]" class="Insecta" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="order">Grylloblattodea</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A645FFBCC013C5B7FE90FAF3" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="9C3F9423A645FFBFC013C5B7FE9EF91A" blockId="5.[88,614,1085,1754]" pageId="5" pageNumber="6">
|
||
COMMENTS: The new species, like all fossil
|
||
<taxonomicName id="5B80EFA0A645FFBFC039C59AFF5BF9CD" authorityName="Crampton" authorityYear="1915" box="[88,210,1525,1549]" order="Notoptera" pageId="5" pageNumber="10" rank="order">Notoptera</taxonomicName>
|
||
, lack the setal fan on the arolium, a feature synapomorphic for the modern, African species of
|
||
<taxonomicName id="5B80EFA0A645FFBFC120C65FFDA3F988" box="[321,554,1584,1608]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
|
||
. like
|
||
<taxonomicName id="5B80EFA0A645FFBFC039C622FE65F9A5" authority="Zompro" authorityName="Zompro" authorityYear="2001" box="[88,492,1613,1637]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="AEF44831A645FFBFC039C622FEF1F9A5" box="[88,376,1613,1637]" italics="true" pageId="5" pageNumber="6">Raptophasma kerneggeri</emphasis>
|
||
Zompro
|
||
</taxonomicName>
|
||
the new species lacks mesofemoral spination but otherwise has all of the features of the genus
|
||
<taxonomicName id="5B80EFA0A645FFBFC039C6CAFF65F97D" authorityName="Engel & Grimaldi" authorityYear="2004" box="[88,236,1701,1725]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEF44831A645FFBFC039C6CAFF65F97D" box="[88,236,1701,1725]" italics="true" pageId="5" pageNumber="6">Adicophasma</emphasis>
|
||
</taxonomicName>
|
||
as characterized by
|
||
<bibRefCitation id="F811E9D2A645FFBFC18CC6CAFE9AF91A" author="Engel, M. S. & D. A. Grimaldi" pageId="5" pageNumber="6" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi (2004)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A645FFBCC2D1C452FE90FAF3" blockId="5.[662,1188,1085,1754]" lastBlockId="6.[128,654,211,1331]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<bibRefCitation id="F811E9D2A645FFBFC2D1C452FCE9FB95" author="Zompro, O." box="[688,864,1085,1109]" pageId="5" pageNumber="6" pagination="85 - 116" refId="ref4643" refString="Zompro, O. 2005. Inter- and intra-ordinal relationships of the Mantophasmatodea, with comments on the phylogeny of polyneopteran orders (Insecta: Polyneoptera). Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg 89: 85 - 116." type="journal article" year="2005">Zompro (2005)</bibRefCitation>
|
||
considered the
|
||
<typeStatus id="433B2A81A645FFBFC47CC452FB08FB95" box="[1053,1153,1085,1109]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="AEF44831A645FFBFC2F7C434FD20FBB3" box="[662,681,1115,1139]" italics="true" pageId="5" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A645FFBFC2DBC434FC96FBB3" box="[698,799,1115,1139]" italics="true" pageId="5" pageNumber="6">spinosum</emphasis>
|
||
as a nymph solely on its smaller body size. However,
|
||
<emphasis id="AEF44831A645FFBFC3FAC417FC27FB50" box="[923,942,1144,1168]" italics="true" pageId="5" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A645FFBFC3A4C417FBA3FB50" box="[965,1066,1144,1168]" italics="true" pageId="5" pageNumber="6">spinosum</emphasis>
|
||
has fully developed genitalia indicative of an adult and has numerous significant features distinguishing it from
|
||
<taxonomicName id="5B80EFA0A645FFBFC353C4BFFC59FB28" authorityName="Zompro" authorityYear="2001" box="[818,976,1232,1256]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="AEF44831A645FFBFC353C4BFFCCCFB28" box="[818,837,1232,1256]" italics="true" pageId="5" pageNumber="6">R</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A645FFBFC33DC4BFFC59FB28" box="[860,976,1232,1256]" italics="true" pageId="5" pageNumber="6">kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
as noted in the original description (
|
||
<bibRefCitation id="F811E9D2A645FFBFC3FCC482FD44FAE2" author="Engel, M. S. & D. A. Grimaldi" pageId="5" pageNumber="6" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi, 2004</bibRefCitation>
|
||
) and in the following key. Perhaps most notably the adult of
|
||
<taxonomicName id="5B80EFA0A645FFBFC3E0C547FB90FA80" authorityName="Zompro" authorityYear="2001" box="[897,1049,1320,1344]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEF44831A645FFBFC3E0C547FB90FA80" box="[897,1049,1320,1344]" italics="true" pageId="5" pageNumber="6">Raptophasma</emphasis>
|
||
</taxonomicName>
|
||
is supposed to lack femoral spines, while they are exceptionally well developed in
|
||
<taxonomicName id="5B80EFA0A645FFBFC469C50CFB15FABB" authorityName="Engel & Grimaldi" authorityYear="2004" box="[1032,1180,1379,1403]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEF44831A645FFBFC469C50CFB15FABB" box="[1032,1180,1379,1403]" italics="true" pageId="5" pageNumber="6">Adicophasma</emphasis>
|
||
</taxonomicName>
|
||
. Zompro’s statement to the contrary is entirely erroneous as the photograph in
|
||
<bibRefCitation id="F811E9D2A645FFBFC448C5F2FCD8FA12" author="Engel, M. S. & D. A. Grimaldi" pageId="5" pageNumber="6" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi (2004)</bibRefCitation>
|
||
was not taken in the focal plane of the legs, yet the out-of-focus spines on at least one leg can nonetheless be discerned; similar, albeit slightly weaker, spines are obvious on
|
||
<taxonomicName id="5B80EFA0A645FFBFC3A9C65FFB2DF988" authorityName="Arillo and Engel" authorityYear="2006" box="[968,1188,1583,1608]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grylloblattoides">
|
||
<emphasis id="AEF44831A645FFBFC3A9C65FFC52F988" box="[968,987,1584,1608]" italics="true" pageId="5" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A645FFBFC399C65FFB2DF988" box="[1016,1188,1584,1608]" italics="true" pageId="5" pageNumber="6">grylloblattoides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="04BB88A6A645FFBFC2FCC622FD52F9A5" box="[669,731,1613,1637]" captionStart="Fig" captionStartId="4.[151,189,692,714]" captionTargetBox="[306,1049,211,658]" captionTargetId="figure-1@4.[306,1049,211,658]" captionTargetPageId="4" captionText="Fig. 3. Photomicrograph of holotype foreleg of Adicophasma grylloblattoides Arillo and Engel, new species (MCNA 10686)." figureDoi="http://doi.org/10.5281/zenodo.5385449" httpUri="https://zenodo.org/record/5385449/files/figure.png" pageId="5" pageNumber="6">fig. 3</figureCitation>
|
||
). That the spines are indeed present in
|
||
<emphasis id="AEF44831A645FFBFC2F7C605FD20F942" box="[662,681,1642,1666]" italics="true" pageId="5" pageNumber="6">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A645FFBFC2D6C605FC95F942" box="[695,796,1642,1666]" italics="true" pageId="5" pageNumber="6">spinosum</emphasis>
|
||
is evidenced by a new photograph of the
|
||
<typeStatus id="433B2A81A645FFBFC281C6E8FCCDF95F" box="[736,836,1671,1695]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
taken in the focal plane of the legs (
|
||
<figureCitation id="04BB88A6A645FFBFC2B0C6CAFC87F97D" box="[721,782,1701,1725]" captionStart="Fig" captionStartId="5.[112,150,972,994]" captionTargetBox="[158,1117,210,938]" captionTargetId="figure-313@5.[158,1117,210,938]" captionTargetPageId="5" captionText="Fig. 5. Photomicrograph of holotype of Adicophasma spinosum Engel and Grimaldi showing detail of forelegs, particularly the presence of profemoral and protibial spination (similar, albeit shorter and less stout, spines are also present on the midlegs; see Engel and Grimaldi, 2004)." figureDoi="http://doi.org/10.5281/zenodo.5385454" httpUri="https://zenodo.org/record/5385454/files/figure.png" pageId="5" pageNumber="6">fig. 5</figureCitation>
|
||
), demonstrating that statements as to the absence of these structures are meant to mislead. Nymphal forms of rock crawlers can have spines in the latest of nymphal stages, but only when such features are similarly present in the adult (i.e., the latest nymphal stages acquire adult-like femoral spination). Thus, even if the
|
||
<typeStatus id="433B2A81A646FFBCC178C108FEF4FEBF" box="[281,381,359,383]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="AEF44831A646FFBCC1D6C108FE43FEBF" box="[439,458,359,383]" italics="true" pageId="6" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC183C108FDCEFEBF" box="[482,583,359,383]" italics="true" pageId="6" pageNumber="7">spinosum</emphasis>
|
||
were a nymph (which the terminal structures indicate is not the case), it could scarcely be considered conspecific with
|
||
<taxonomicName id="5B80EFA0A646FFBCC182C1AFFD03FE18" authorityName="Zompro" authorityYear="2001" box="[483,650,448,472]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="AEF44831A646FFBCC182C1AFFE7FFE18" box="[483,502,448,472]" italics="true" pageId="6" pageNumber="7">R</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC273C1AFFD03FE18" box="[530,650,448,472]" italics="true" pageId="6" pageNumber="7">kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AEF44831A646FFBCC2E8C1AFFD04FE18" box="[649,653,448,472]" italics="true" pageId="6" pageNumber="7">,</emphasis>
|
||
which is supposed to lack such features entirely in the adult stage [as evidenced by the fact that the adult
|
||
<typeStatus id="433B2A81A646FFBCC1FEC276FD8AFDF1" box="[415,515,537,561]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
male lacks such characters, as do, apparently, the
|
||
<typeStatus id="433B2A81A646FFBCC230C259FF37FDAC" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
; since spines in at least
|
||
<specimenCount id="8A865FAAA646FFBCC193C23BFD04FDAC" box="[498,653,596,620]" pageId="6" pageNumber="7" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
were entirely overlooked (
|
||
<figureCitation id="04BB88A6A646FFBCC1A0C21DFE77FD4A" box="[449,510,626,650]" captionStart="Fig" captionStartId="5.[112,150,972,994]" captionTargetBox="[158,1117,210,938]" captionTargetId="figure-313@5.[158,1117,210,938]" captionTargetPageId="5" captionText="Fig. 5. Photomicrograph of holotype of Adicophasma spinosum Engel and Grimaldi showing detail of forelegs, particularly the presence of profemoral and protibial spination (similar, albeit shorter and less stout, spines are also present on the midlegs; see Engel and Grimaldi, 2004)." figureDoi="http://doi.org/10.5281/zenodo.5385454" httpUri="https://zenodo.org/record/5385454/files/figure.png" pageId="6" pageNumber="7">fig. 5</figureCitation>
|
||
), the series should be re-examined]. The same can be said for the dorsal profemoral carinae present in
|
||
<taxonomicName id="5B80EFA0A646FFBCC212C2C2FF7DFD23" authorityName="Zompro" authorityYear="2001" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="AEF44831A646FFBCC212C2C2FD0FFD05" box="[627,646,685,709]" italics="true" pageId="6" pageNumber="7">R</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC0E1C2A4FF7DFD23" box="[128,244,715,739]" italics="true" pageId="6" pageNumber="7">kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
but lacking in
|
||
<emphasis id="AEF44831A646FFBCC1FCC2A4FDB0FD23" box="[413,569,715,739]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="5B80EFA0A646FFBCC1FCC2A4FDBCFD23" authorityName="Engel & Grimaldi" authorityYear="2004" box="[413,565,715,739]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Adicophasma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
among other generic features (see
|
||
<bibRefCitation id="F811E9D2A646FFBCC264C287FEBFFCDE" author="Engel, M. S. & D. A. Grimaldi" pageId="6" pageNumber="7" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi, 2004</bibRefCitation>
|
||
). Most descriptions and figures of Baltic amber rock crawlers are inadequate (e.g.,
|
||
<bibRefCitation id="F811E9D2A646FFBCC159C32EFE6AFC99" author="Zompro, O." box="[312,483,833,857]" pageId="6" pageNumber="7" pagination="229 - 261" refId="ref4601" refString="Zompro, O. 2001. The Phasmatodea and Raptophasma n. gen., Orthoptera incertae sedis, in Baltic amber (Insecta: Orthoptera). Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg 85: 229 - 261." type="journal article" year="2001">Zompro, 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F811E9D2A646FFBCC195C32EFDA2FC99" author="Zompro, O." box="[500,555,833,857]" pageId="6" pageNumber="7" pagination="85 - 116" refId="ref4643" refString="Zompro, O. 2005. Inter- and intra-ordinal relationships of the Mantophasmatodea, with comments on the phylogeny of polyneopteran orders (Insecta: Polyneoptera). Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg 89: 85 - 116." type="journal article" year="2005">2005</bibRefCitation>
|
||
) as evidenced by the oversight of obvious features such as the presence/absence of femoral spination among
|
||
<typeStatus id="433B2A81A646FFBCC134C3F5FE4AFC72" box="[341,451,922,946]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
and should not be relied upon. For more extensive descriptions and figures refer to
|
||
<bibRefCitation id="F811E9D2A646FFBCC1C9C3BAFF4FFBCB" author="Engel, M. S. & D. A. Grimaldi" pageId="6" pageNumber="7" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi (2004)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="F811E9D2A646FFBCC160C39CFDB1FBCB" author="Grimaldi, D. & M. S. Engel" box="[257,568,1011,1035]" pageId="6" pageNumber="7" refId="ref4073" refString="Grimaldi, D., and M. S. Engel. 2005. Evolution of the insects. Cambridge: Cambridge University Press, xv + 755 pp." type="book" year="2005">Grimaldi and Engel (2005)</bibRefCitation>
|
||
. As the subfamily
|
||
<taxonomicName id="5B80EFA0A646FFBCC09AC47FFE5AFBE8" authorityName="Zompro" authorityYear="2005" box="[251,467,1040,1064]" class="Insecta" family="Mantophasmatidae" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Raptophasmatinae">Raptophasmatinae</taxonomicName>
|
||
and the family
|
||
<taxonomicName id="5B80EFA0A646FFBCC0E1C441FEE6FB86" authorityName="Zompro" authorityYear="2005" box="[128,367,1070,1094]" class="Insecta" family="Ensiferophasmatidae" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Ensiferophasmatidae</taxonomicName>
|
||
are based entirely on plesiomorphies, they are herein newly synonymized with
|
||
<taxonomicName id="5B80EFA0A646FFBCC175C406FE7BFB41" box="[276,498,1129,1153]" class="Insecta" family="Mantophasmatidae" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Mantophasmatidae</taxonomicName>
|
||
(
|
||
<emphasis id="AEF44831A646FFBCC265C406FF38FB5F" bold="true" pageId="6" pageNumber="7">new synonymies</emphasis>
|
||
). The
|
||
<taxonomicName id="5B80EFA0A646FFBCC167C4E8FE6EFB5F" authorityName="Zompro" authorityYear="2005" box="[262,487,1159,1183]" class="Insecta" family="Mantophasmatidae" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Tyrannophasmatini">Tyrannophasmatini</taxonomicName>
|
||
is also suspicious and thus is only tentatively retained (table 1). Indeed, the recognition of multiple subfamilies or tribes within this remarkably small and relatively homogeneous family is unwarranted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A646FFBCC0A2C516FD07F9C1" pageId="6" pageNumber="7" type="key">
|
||
<paragraph id="9C3F9423A646FFBCC0A2C516FDC3FA4F" blockId="6.[195,586,1401,1425]" box="[195,586,1401,1425]" pageId="6" pageNumber="7">
|
||
KEY TO BALTIC AMBER
|
||
<taxonomicName id="5B80EFA0A646FFBCC1ABC516FDC3FA4F" authorityName="Crampton" authorityYear="1915" box="[458,586,1401,1425]" order="Notoptera" pageId="6" pageNumber="10" rank="order">NOTOPTERA</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A646FFBCC0E3C5DAFD07F9C1" blockId="6.[130,654,1461,1754]" pageId="6" pageNumber="7">1. Compound eyes occupying major portion of head; antennae shorter than body; arolium projecting beyond pretarsal claws...... 2</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D49AC7A8A646FFBCC0E3C669FB45FD2A" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="9C3F9423A646FFBCC0E3C669FD04F948" blockId="6.[130,654,1461,1754]" pageId="6" pageNumber="7">
|
||
_ Compound eyes placed in posterior half of head; antennae several times longer than body; arolium not projecting beyond pretarsal claws (Genus
|
||
<emphasis id="AEF44831A646FFBCC1F0C638FDB9F9AC" box="[401,560,1623,1644]" italics="true" pageId="6" pageNumber="7">Ensiferophasma</emphasis>
|
||
)....................
|
||
<taxonomicName id="5B80EFA0A646FFBCC1FDC61DFD04F948" authority="Zompro" authorityName="Zompro" authorityYear="2005" box="[412,653,1650,1672]" class="Insecta" family="Ensiferophasmatidae" genus="Ensiferophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="velociraptor">
|
||
<emphasis id="AEF44831A646FFBCC1FDC61DFDB8F947" box="[412,561,1650,1671]" italics="true" pageId="6" pageNumber="7">E. velociraptor</emphasis>
|
||
Zompro
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A646FFBCC0E3C6E1FD07F91A" blockId="6.[130,654,1461,1754]" pageId="6" pageNumber="7">
|
||
2. At least some femora with distinct, ventral spination; profemur without dorsal carinae (Genus
|
||
<taxonomicName id="5B80EFA0A646FFBCC170C6ABFE1FF919" authorityName="Engel & Grimaldi" authorityYear="2004" box="[273,406,1732,1753]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEF44831A646FFBCC170C6ABFE1FF919" box="[273,406,1732,1753]" italics="true" pageId="6" pageNumber="7">Adicophasma</emphasis>
|
||
</taxonomicName>
|
||
)............... 3
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A646FFBCC2A1C0BDFB42FEDE" blockId="6.[704,1228,210,746]" pageId="6" pageNumber="7">
|
||
_ Femora without spination; profemur with paired, dorsal, longitudinal carinae (Genus
|
||
<taxonomicName id="5B80EFA0A646FFBCC29CC167FC0CFEDD" authorityName="Zompro" authorityYear="2001" box="[765,901,264,285]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEF44831A646FFBCC29CC167FC0CFEDD" box="[765,901,264,285]" italics="true" pageId="6" pageNumber="7">Raptophasma</emphasis>
|
||
</taxonomicName>
|
||
).....
|
||
<taxonomicName id="5B80EFA0A646FFBCC386C167FB42FEDE" authority="Zompro" authorityName="Zompro" authorityYear="2001" box="[999,1227,264,286]" class="Insecta" family="Mantophasmatidae" genus="Raptophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="AEF44831A646FFBCC386C167FC71FEDD" box="[999,1016,264,285]" italics="true" pageId="6" pageNumber="7">R</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC467C167FBE6FEDD" box="[1030,1135,264,285]" italics="true" pageId="6" pageNumber="7">kerneggeri</emphasis>
|
||
Zompro
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A646FFBCC2A1C14BFB42FDEC" blockId="6.[704,1228,210,746]" pageId="6" pageNumber="7">
|
||
3. Mesofemur with distinct, ventral spination; pedicel elongate, at least three times longer than wide; cardo with two stout, apical spines; pronotum slightly longer than wide, meso- and metanota each wider than long; lateral margins of abdomen convex (i.e., abdomen roughly ovoid); compound eyes with large ommatidial facets (see
|
||
<bibRefCitation id="F811E9D2A646FFBCC400C18FFC13FDD1" author="Engel, M. S. & D. A. Grimaldi" pageId="6" pageNumber="7" pagination="1 - 11" refId="ref4029" refString="Engel, M. S., and D. A. Grimaldi. 2004. A new rock crawler in Baltic amber, with comments on the order (Mantophasmatodea: Mantophasmatidae). American Museum Novitates 3431: 1 - 11." type="journal article" year="2004">Engel and Grimaldi, 2004</bibRefCitation>
|
||
)............................
|
||
<emphasis id="AEF44831A646FFBCC31AC278FC05FDEC" box="[891,908,535,556]" italics="true" pageId="6" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC3FAC278FC7FFDEC" box="[923,1014,535,556]" italics="true" pageId="6" pageNumber="7">spinosum</emphasis>
|
||
Engel and Grimaldi
|
||
</paragraph>
|
||
<paragraph id="9C3F9423A646FFBCC2A1C25DFB45FD2A" blockId="6.[704,1228,210,746]" pageId="6" pageNumber="7">
|
||
_ Mesofemur without ventral spination; pedicel short, about as long as wide; cardo without spines; all thoracic nota longer than wide and each of approximately equal size; abdomen elongate; compound eyes with smaller and more numerous ommatidial facets.....................
|
||
<taxonomicName id="5B80EFA0A646FFBCC3B3C2BBFB06FD29" authority="ARILLO & ENGEL, 2006" authorityName="ARILLO & ENGEL" authorityYear="2006" box="[978,1167,724,746]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grylloblattoides" status="sp. nov.">
|
||
<emphasis id="AEF44831A646FFBCC3B3C2BBFC6AFD29" box="[978,995,724,745]" italics="true" pageId="6" pageNumber="7">A</emphasis>
|
||
.
|
||
<emphasis id="AEF44831A646FFBCC393C2BBFB06FD29" box="[1010,1167,724,745]" italics="true" pageId="6" pageNumber="7">grylloblattoides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AEF44831A646FFBCC4EEC2BBFB1AFD29" box="[1167,1171,724,745]" italics="true" pageId="6" pageNumber="7">,</emphasis>
|
||
<taxonomicNameLabel id="B5C7F54AA646FFBCC4FAC2BBFB45FD2A" box="[1179,1228,724,746]" pageId="6" pageNumber="7" rank="species">n.sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |