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<mods:title id="111A1BAB3FF1BC627961926D1EC670E7">A new species of Characidium (Characiformes: Crenuchidae) from the upper rio São Francisco basin, Minas Gerais, Brazil</mods:title>
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<mods:namePart id="9395068BF4CED15A04018BA0A88F0CF8">Zanata, Angela M.</mods:namePart>
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(
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sp.: 
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Vieira 
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<bibRefCitation id="EFC509D2FFD1FF96FEF5FA81FDAE014A" author="Alves, C. B. M. &amp; Pompeu, P. S." box="[332,621,1331,1356]" pageId="1" pageNumber="506" pagination="167 - 189" refId="ref5124" refString="Alves, C. B. M. &amp; Pompeu, P. S. (2010) A Fauna de peixes da bacia do Rio das Velhas no final do seculo XX. In: Alves, C. B. M. &amp; Pompeu, P. S. (Eds.), Peixes do Rio das Velhas: passado e presente. ARGVMENTVM, Belo Horizonte, pp. 167 - 189." type="book chapter" year="2010">Alves &amp; Pompeu, 2010: 186</bibRefCitation>
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, rio Cipó, Serra do Cipó National Park, listed).
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. Penido &amp; 
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<materialsCitation id="3B3C7E7EFFD1FF96FF7EFA54FE120222" collectorName="das Velhas" country="Brazil" county="All" location="Serra do Cipo National Park" municipality="Jaboticatubas" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="B920A831FFD1FF96FF7EFA54FE850206" bold="true" box="[199,326,1510,1536]" pageId="1" pageNumber="506">
<typeStatus id="54EFCA81FFD1FF96FF7EFA54FE820206" box="[199,321,1510,1536]" pageId="1" pageNumber="506" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="628A0CAFFFD1FF96FEF5FA54FEB30206" box="[332,368,1510,1536]" pageId="1" pageNumber="506">All</collectingCounty>
from 
<collectingCountry id="F34334B3FFD1FF96FE0FFA54FE3E0206" box="[438,509,1510,1536]" name="Brazil" pageId="1" pageNumber="506">Brazil</collectingCountry>
, 
<collectingRegion id="4990BAC1FFD1FF96FDB0FA54FD600206" box="[521,675,1510,1536]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="506">Minas Gerais</collectingRegion>
, 
<collectingMunicipality id="6B8FEE59FFD1FF96FD17FA54FC8B0206" box="[686,840,1510,1536]" pageId="1" pageNumber="506">Jaboticatubas</collectingMunicipality>
, 
<location id="8E8B22F8FFD1FF96FCEDFA54FB540206" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FCEDFA54FB540206" box="[852,1175,1510,1536]" country="Brazil" county="All" municipality="Jaboticatubas" name="Serra do Cipo National Park" pageId="1" pageNumber="506" stateProvince="Minas Gerais">Serra do Cipó National Park</location>
, rio 
<collectorName id="26A111F5FFD1FF96FB70FA54FA820206" box="[1225,1345,1510,1536]" pageId="1" pageNumber="506">das Velhas</collectorName>
sub-basin, 
<location id="8E8B22F8FFD1FF96FF7FF9B8FE490222" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FF7FF9B8FE490222" box="[198,394,1546,1572]" country="Brazil" county="All" municipality="Jaboticatubas" name="rio Sao Francisco" pageId="1" pageNumber="506" stateProvince="Minas Gerais">rio São Francisco</location>
basin
</materialsCitation>
: 
<materialsCitation id="3B3C7E7EFFD1FF96FE65F9B8FCC5024E" collectingDate="2018-09-25" collectionCode="CICMG" country="Brazil" county="rio Cipo" elevation="785" latitude="-19.34714" location="rio Mascates" longLatPrecision="1" longitude="-43.605694" municipality="rio Bocaina" pageId="1" pageNumber="506" specimenCode="CICMG 93, 1" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FE65F9B8FD9E0222" box="[476,605,1546,1572]" collectionCode="CICMG" pageId="1" pageNumber="506">CICMG 93</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FDD0F9B8FDB40222" box="[617,631,1546,1572]" collectionCode="CICMG" pageId="1" pageNumber="506">1</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FD3CF9B8FD2A0223" box="[645,745,1546,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="1" pageNumber="506" unit="mm" value="26.1">26.1 mm</quantity>
SL, 
<collectingCounty id="628A0CAFFFD1FF96FCA6F9B8FCB80222" box="[799,891,1546,1572]" pageId="1" pageNumber="506">rio Cipó</collectingCounty>
below meeting of 
<collectingMunicipality id="6B8FEE59FFD1FF96FBF7F9B8FB130222" box="[1102,1232,1546,1572]" pageId="1" pageNumber="506">rio Bocaina</collectingMunicipality>
and 
<location id="8E8B22F8FFD1FF96FABEF9B8FA5B0223" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FABEF9B8FA5B0223" box="[1287,1432,1546,1573]" country="Brazil" county="rio Cipo" latitude="-19.34714" longLatPrecision="1" longitude="-43.605694" municipality="rio Bocaina" name="rio Mascates" pageId="1" pageNumber="506" stateProvince="Minas Gerais">rio Mascates</location>
, 
<geoCoordinate id="EE6012E4FFD1FF96FF2EF99CFEF2024E" box="[151,305,1582,1608]" degrees="19" direction="south" minutes="20" orientation="latitude" pageId="1" pageNumber="506" precision="1" seconds="49.7" value="-19.34714">19°20’49.7”S</geoCoordinate>
<geoCoordinate id="EE6012E4FFD1FF96FE81F99FFE1F024E" box="[312,476,1581,1608]" degrees="43" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="506" precision="1" seconds="20.5" value="-43.605694">43°36’20.5”W</geoCoordinate>
, 
<elevation id="00799310FFD1FF96FE5EF99FFDA8024E" box="[487,619,1581,1609]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="1" pageNumber="506" unit="m" value="785.0">
<quantity id="4CACD9C6FFD1FF96FE5EF99FFDED024F" box="[487,558,1581,1609]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="1" pageNumber="506" unit="m" value="785.0">785 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFEA52E3FFD1FF96FDCEF99FFCC5024E" box="[631,774,1581,1608]" pageId="1" pageNumber="506" value="2018-09-25">
<collectingDate id="EFAEAB0BFFD1FF96FDCEF99FFCC5024E" box="[631,774,1581,1608]" pageId="1" pageNumber="506" value="2018-09-25">25 Sep 2018</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B3C7E7EFFD1FF96FCA9F99DFBC3024F" box="[784,1024,1582,1609]" collectionCode="T" collectorName="Ribeiro &amp; F. Porto" country="Brazil" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectionCode id="ED45ECE6FFD1FF96FCA9F99DFCE1024F" box="[784,802,1583,1609]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="506">T</collectionCode>
. 
<collectorName id="26A111F5FFD1FF96FC94F99CFC42024E" box="[813,897,1582,1608]" pageId="1" pageNumber="506">Ribeiro</collectorName>
&amp; 
<collectorName id="26A111F5FFD1FF96FC1CF99DFBC3024F" box="[933,1024,1583,1609]" pageId="1" pageNumber="506">F. Porto</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B3C7E7EFFD1FF96FBB2F99CFC8A026A" collectingDate="2018-08-23" collectionCode="CICMG" country="Brazil" elevation="789" latitude="-19.345028" location="rio Bocaina" longLatPrecision="1" longitude="-43.605804" pageId="1" pageNumber="506" specimenCode="CICMG 95, 1" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FBB2F99CFB48024E" box="[1035,1163,1581,1608]" collectionCode="CICMG" pageId="1" pageNumber="506">CICMG 95</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FB2EF99CFB66024E" box="[1175,1189,1582,1608]" collectionCode="CICMG" pageId="1" pageNumber="506">1</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FB0AF99CFAD4024F" box="[1203,1303,1582,1609]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="1" pageNumber="506" unit="mm" value="26.8">26.8 mm</quantity>
SL, 
<location id="8E8B22F8FFD1FF96FAF5F99CFF16026A" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FAF5F99CFF16026A" country="Brazil" latitude="-19.345028" longLatPrecision="1" longitude="-43.605804" name="rio Bocaina" pageId="1" pageNumber="506" stateProvince="Minas Gerais">rio Bocaina</location>
, 
<geoCoordinate id="EE6012E4FFD1FF96FF66F9E0FEBA026A" box="[223,377,1618,1644]" degrees="19" direction="south" minutes="20" orientation="latitude" pageId="1" pageNumber="506" precision="1" seconds="42.1" value="-19.345028">19°20’42.1”S</geoCoordinate>
<geoCoordinate id="EE6012E4FFD1FF96FEC7F9E0FDE1026A" box="[382,546,1618,1644]" degrees="43" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="506" precision="1" seconds="20.9" value="-43.605804">43°36’20.9”W</geoCoordinate>
, 
<elevation id="00799310FFD1FF96FD94F9E0FD6E026A" box="[557,685,1618,1645]" metricMagnitude="2" metricUnit="m" metricValue="7.89" pageId="1" pageNumber="506" unit="m" value="789.0">
<quantity id="4CACD9C6FFD1FF96FD94F9E0FDB1026B" box="[557,626,1618,1645]" metricMagnitude="2" metricUnit="m" metricValue="7.89" pageId="1" pageNumber="506" unit="m" value="789.0">789 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFEA52E3FFD1FF96FD01F9E0FC8A026A" box="[696,841,1618,1644]" pageId="1" pageNumber="506" value="2018-08-23">
<collectingDate id="EFAEAB0BFFD1FF96FD01F9E0FC8A026A" box="[696,841,1618,1644]" pageId="1" pageNumber="506" value="2018-08-23">23 Aug 2018</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B3C7E7EFFD1FF96FCEAF9E1FBFE026B" box="[851,1085,1618,1645]" collectionCode="T" collectorName="Ribeiro &amp; F. Porto" country="Brazil" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectionCode id="ED45ECE6FFD1FF96FCEAF9E1FCA6026B" box="[851,869,1619,1645]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="506">T</collectionCode>
. 
<collectorName id="26A111F5FFD1FF96FCD7F9E0FC01026A" box="[878,962,1618,1644]" pageId="1" pageNumber="506">Ribeiro</collectorName>
&amp; 
<collectorName id="26A111F5FFD1FF96FC5AF9E1FBFE026B" box="[995,1085,1619,1645]" pageId="1" pageNumber="506">F. Porto</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B3C7E7EFFD1FF96FBFEF9E0FD660297" collectionCode="CICMG" country="Brazil" elevation="120" location="Lagoa Rasgada" pageId="1" pageNumber="506" specimenCode="CICMG 154, 3" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FBFEF9E0FB17026A" box="[1095,1236,1617,1644]" collectionCode="CICMG" pageId="1" pageNumber="506">CICMG 154</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FB66F9E0FB2E026A" box="[1247,1261,1618,1644]" collectionCode="CICMG" pageId="1" pageNumber="506">3</specimenCode>
c&amp;s, 
<quantity id="4CACD9C6FFD1FF96FA95F9E0FF000297" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.41" metricValueMin="1.7899999999999998" pageId="1" pageNumber="506" unit="mm" value="21.0" valueMax="24.1" valueMin="17.9">17.9–24.1 mm</quantity>
SL, 
<location id="8E8B22F8FFD1FF96FF43F9C5FE6A0296" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FF43F9C5FE6A0296" box="[250,425,1654,1681]" country="Brazil" name="Lagoa Rasgada" pageId="1" pageNumber="506" stateProvince="Minas Gerais">Lagoa Rasgada</location>
, approximately 
<quantity id="4CACD9C6FFD1FF96FDE7F9C4FD660297" box="[606,677,1654,1681]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="506" unit="m" value="120.0">
<elevation id="00799310FFD1FF96FDE7F9C4FD660297" box="[606,677,1654,1681]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="506" unit="m" value="120.0">120 m</elevation>
</quantity>
</materialsCitation>
<materialsCitation id="3B3C7E7EFFD1FF96FD14F9C4FEA402B2" collectingDate="2019-06-20" country="Brazil" elevation="791" latitude="-19.34664" location="rio Cipo" longLatPrecision="1" longitude="-43.61122" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
of the left margin of 
<location id="8E8B22F8FFD1FF96FC21F9C4FC340296" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FC21F9C4FC340296" box="[920,1015,1654,1680]" country="Brazil" latitude="-19.34664" longLatPrecision="1" longitude="-43.61122" name="rio Cipo" pageId="1" pageNumber="506" stateProvince="Minas Gerais">rio Cipó</location>
, 
<geoCoordinate id="EE6012E4FFD1FF96FBBAF9C4FB5E0296" box="[1027,1181,1654,1680]" degrees="19" direction="south" minutes="20" orientation="latitude" pageId="1" pageNumber="506" precision="1" seconds="47.9" value="-19.34664">19°20’47.9”S</geoCoordinate>
<geoCoordinate id="EE6012E4FFD1FF96FB1DF9C4FA8A0296" box="[1188,1353,1654,1680]" degrees="43" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="506" precision="1" seconds="40.4" value="-43.61122">43°36’40.4”W</geoCoordinate>
, 
<elevation id="00799310FFD1FF96FAECF9C4FF0E02B2" metricMagnitude="2" metricUnit="m" metricValue="7.909999999999999" pageId="1" pageNumber="506" unit="m" value="791.0">
<quantity id="4CACD9C6FFD1FF96FAECF9C4FA5F0297" box="[1365,1436,1654,1681]" metricMagnitude="2" metricUnit="m" metricValue="7.909999999999999" pageId="1" pageNumber="506" unit="m" value="791.0">791 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFEA52E3FFD1FF96FF63F928FEA402B2" box="[218,359,1690,1717]" pageId="1" pageNumber="506" value="2019-06-20">
<collectingDate id="EFAEAB0BFFD1FF96FF63F928FEA402B2" box="[218,359,1690,1717]" pageId="1" pageNumber="506" value="2019-06-20">20 Jun 2019</collectingDate>
</date>
</materialsCitation>
, F. Porto &amp; M. Kadri. 
<materialsCitation id="3B3C7E7EFFD1FF96FDDFF928FC3802B2" box="[614,1019,1689,1717]" collectionCode="CICMG" country="Brazil" pageId="1" pageNumber="506" specimenCode="CICMG 157, 10, 20.4" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FDDFF928FD3602B2" box="[614,757,1689,1716]" collectionCode="CICMG" pageId="1" pageNumber="506">CICMG 157</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FCBBF928FCE302B2" box="[770,800,1690,1716]" collectionCode="CICMG" pageId="1" pageNumber="506">10</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FC95F928FC9E02B2" box="[812,861,1690,1716]" collectionCode="CICMG" pageId="1" pageNumber="506">20.4</specimenCode>
–27.0 mm SL
</materialsCitation>
; 
<materialsCitation id="3B3C7E7EFFD1FF96FBBEF928FDE402DF" collectionCode="MCNIP" country="Brazil" elevation="130" location="Lagoa do Boi" pageId="1" pageNumber="506" specimenCode="MCNIP 4530, 6" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FBBEF928FB6102B2" box="[1031,1186,1689,1716]" collectionCode="MCNIP" pageId="1" pageNumber="506">MCNIP 4530</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FB14F928FB7E02B2" box="[1197,1213,1690,1716]" collectionCode="MCNIP" pageId="1" pageNumber="506">6</specimenCode>
, 21.0– 
<quantity id="4CACD9C6FFD1FF96FAB0F92BFAAE02B3" box="[1289,1389,1689,1717]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="506" unit="mm" value="25.4">25.4 mm</quantity>
SL, 
<location id="8E8B22F8FFD1FF96FF2EF90DFEF302DE" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FF2EF90DFEF302DE" box="[151,304,1726,1753]" country="Brazil" name="Lagoa do Boi" pageId="1" pageNumber="506" stateProvince="Minas Gerais">Lagoa do Boi</location>
, approximately 
<quantity id="4CACD9C6FFD1FF96FE58F90CFDE402DF" box="[481,551,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="506" unit="m" value="130.0">
<elevation id="00799310FFD1FF96FE58F90CFDE402DF" box="[481,551,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="506" unit="m" value="130.0">130 m</elevation>
</quantity>
</materialsCitation>
<materialsCitation id="3B3C7E7EFFD1FF96FD95F90CFF1102FA" collectingDate="2019-05-15" country="Brazil" elevation="803" latitude="-19.348307" location="rio Cipo" longLatPrecision="1" longitude="-43.60822" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
from left margin of 
<location id="8E8B22F8FFD1FF96FCBCF90CFCA002DE" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD1FF96FCBCF90CFCA002DE" box="[773,867,1726,1752]" country="Brazil" latitude="-19.348307" longLatPrecision="1" longitude="-43.60822" name="rio Cipo" pageId="1" pageNumber="506" stateProvince="Minas Gerais">rio Cipó</location>
, 
<geoCoordinate id="EE6012E4FFD1FF96FCD4F90FFBC402DE" box="[877,1031,1725,1752]" degrees="19" direction="south" minutes="20" orientation="latitude" pageId="1" pageNumber="506" precision="1" seconds="53.9" value="-19.348307">19°20’53.9”S</geoCoordinate>
<geoCoordinate id="EE6012E4FFD1FF96FBB4F90CFB7202DE" box="[1037,1201,1726,1752]" degrees="43" direction="west" minutes="36" orientation="longitude" pageId="1" pageNumber="506" precision="1" seconds="29.6" value="-43.60822">43°36’29.6”W</geoCoordinate>
, 
<elevation id="00799310FFD1FF96FB02F90CFAFF02DE" box="[1211,1340,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="8.03" pageId="1" pageNumber="506" unit="m" value="803.0">
<quantity id="4CACD9C6FFD1FF96FB02F90CFAC202DF" box="[1211,1281,1726,1753]" metricMagnitude="2" metricUnit="m" metricValue="8.03" pageId="1" pageNumber="506" unit="m" value="803.0">803 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFEA52E3FFD1FF96FAFEF90FFF1102FA" pageId="1" pageNumber="506" value="2019-05-15">
<collectingDate id="EFAEAB0BFFD1FF96FAFEF90FFF1102FA" pageId="1" pageNumber="506" value="2019-05-15">15 May 2019</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B3C7E7EFFD1FF96FF67F951FD0402FA" box="[222,711,1762,1789]" collectionCode="T" collectorName="Pessali, F &amp; Porto, I &amp; Penido &amp; Santana" country="Brazil" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectionCode id="ED45ECE6FFD1FF96FF67F951FF3302FB" box="[222,240,1763,1789]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="506">T</collectionCode>
. 
<collectorName id="26A111F5FFD1FF96FF45F950FEA902FB" box="[252,362,1762,1789]" pageId="1" pageNumber="506">Pessali, F</collectorName>
. 
<collectorName id="26A111F5FFD1FF96FECCF951FE0E02FB" box="[373,461,1763,1789]" pageId="1" pageNumber="506">Porto, I</collectorName>
. 
<collectorName id="26A111F5FFD1FF96FE60F950FDE402FA" box="[473,551,1762,1788]" pageId="1" pageNumber="506">Penido</collectorName>
&amp; S. 
<collectorName id="26A111F5FFD1FF96FDD2F950FD0402FA" box="[619,711,1762,1788]" pageId="1" pageNumber="506">Santana</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B3C7E7EFFD1FF96FD6AF950FB9F02FA" box="[723,1116,1761,1789]" collectionCode="CICMG" country="Brazil" pageId="1" pageNumber="506" specimenCode="CICMG 158, 8" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FD6AF950FCA102FA" box="[723,866,1761,1788]" collectionCode="CICMG" pageId="1" pageNumber="506">CICMG 158</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FCD6F950FCBD02FA" box="[879,894,1762,1788]" collectionCode="CICMG" pageId="1" pageNumber="506">8</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FC35F953FBF202FB" box="[908,1073,1761,1789]" metricMagnitude="-2" metricUnit="m" metricValue="3.075" metricValueMax="3.62" metricValueMin="2.53" pageId="1" pageNumber="506" unit="mm" value="30.75" valueMax="36.2" valueMin="25.3">25.3–36.2 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B3C7E7EFFD1FF96FBD0F950FF330326" collectionCode="MCNIP" country="Brazil" pageId="1" pageNumber="506" specimenCode="MCNIP 4528, 2" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FBD0F950FAC002FA" box="[1129,1283,1761,1788]" collectionCode="MCNIP" pageId="1" pageNumber="506">MCNIP 4528</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FAB6F950FADC02FA" box="[1295,1311,1762,1788]" collectionCode="MCNIP" pageId="1" pageNumber="506">2</specimenCode>
, 29.0– 
<quantity id="4CACD9C6FFD1FF96FAD5F950FF000327" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="1" pageNumber="506" unit="mm" value="36.8">36.8 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B3C7E7EFFD1FF96FF47F8B4FD7B0326" box="[254,696,1797,1825]" collectionCode="MZUSP" country="Brazil" pageId="1" pageNumber="506" specimenCode="MZUSP 125766, 3" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FF47F8B4FE790326" box="[254,442,1797,1824]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="506">MZUSP 125766</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FE71F8B4FE140326" box="[456,471,1798,1824]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="506">3</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FE5FF8B4FD480327" box="[486,651,1798,1825]" metricMagnitude="-2" metricUnit="m" metricValue="3.2049999999999996" metricValueMax="3.62" metricValueMin="2.79" pageId="1" pageNumber="506" unit="mm" value="32.05" valueMax="36.2" valueMin="27.9">27.9–36.2 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B3C7E7EFFD1FF96FD7FF8B4FA860326" box="[710,1349,1797,1825]" collectionCode="UFBA" country="Brazil" pageId="1" pageNumber="506" specimenCode="UFBA 9056, 8" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FD7FF8B4FC900326" box="[710,851,1797,1824]" collectionCode="UFBA" pageId="1" pageNumber="506">UFBA 9056</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FCD9F8B4FCAC0326" box="[864,879,1798,1824]" collectionCode="UFBA" pageId="1" pageNumber="506">8</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FCC6F8B7FBE70327" box="[895,1060,1797,1825]" metricMagnitude="-2" metricUnit="m" metricValue="3.075" metricValueMax="3.62" metricValueMin="2.53" pageId="1" pageNumber="506" unit="mm" value="30.75" valueMax="36.2" valueMin="25.3">25.3–36.2 mm</quantity>
SL, 1 c&amp;s, 
<quantity id="4CACD9C6FFD1FF96FB0BF8B7FADA0327" box="[1202,1305,1797,1825]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="1" pageNumber="506" unit="mm" value="35.5">35.5 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B3C7E7EFFD1FF96FAEAF8B4FC170342" collectingDate="2019-05-14" collectionCode="ZUEC" collectorName="das Velhas &amp; Pessali, F &amp; Porto, I &amp; S. Santana." country="Brazil" county="Jaboticatubas" elevation="783" latitude="-19.34339" location="Serra do Cipo National Park" longLatPrecision="1" longitude="-43.608444" municipality="rio Cipo" pageId="1" pageNumber="506" specimenCode="ZUEC 17227, 4" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD1FF96FAEAF8B4FF230342" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="1" pageNumber="506">ZUEC 17227</specimenCode>
, 
<specimenCode id="DBF2DC58FFD1FF96FF52F898FF380342" box="[235,251,1834,1860]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="1" pageNumber="506">4</specimenCode>
, 
<quantity id="4CACD9C6FFD1FF96FEBEF89BFE690343" box="[263,426,1833,1861]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" metricValueMax="3.86" metricValueMin="2.56" pageId="1" pageNumber="506" unit="mm" value="32.1" valueMax="38.6" valueMin="25.6">25.6–38.6 mm</quantity>
SL, 1 c&amp;s, 
<quantity id="4CACD9C6FFD1FF96FD89F89BFD570343" box="[560,660,1833,1861]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="506" unit="mm" value="27.5">27.5 mm</quantity>
SL, collected with holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E27A8FFD1FF95FF7EF8FFFE78058A" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506" type="diagnosis">
<paragraph id="8BEB7423FFD1FF95FF7EF8FFFE78058A" blockId="1.[151,1437,1401,2041]" lastBlockId="2.[151,1437,153,647]" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506">
<emphasis id="B920A831FFD1FF96FF7EF8FFFE81036E" bold="true" box="[199,322,1869,1896]" pageId="1" pageNumber="506">Diagnosis.</emphasis>
<taxonomicName id="4C540FA0FFD1FF96FEF7F8FCFDF1036E" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[334,562,1870,1896]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD1FF96FEF7F8FCFDF1036E" box="[334,562,1870,1896]" italics="true" pageId="1" pageNumber="506">Characidium cacah</emphasis>
</taxonomicName>
can be distinguished from all congeners, except 
<taxonomicName id="4C540FA0FFD1FF96FB3AF8FDFED8038A" authority="Graca, Ota &amp; Domingues" authorityName="Graca, Ota &amp; Domingues" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="chicoi">
<emphasis id="B920A831FFD1FF96FB3AF8FDFB2E036E" box="[1155,1261,1870,1897]" italics="true" pageId="1" pageNumber="506">C. chicoi</emphasis>
Graça, Ota &amp; Domingues
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD1FF96FE9FF8C1FC62038A" authority="Zanata, Sarmento-Soares &amp; Martins-Pinheiro" authorityName="Zanata, Sarmento-Soares &amp; Martins-Pinheiro" authorityYear="2015" box="[294,929,1906,1933]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="helmeri">
<emphasis id="B920A831FFD1FF96FE9FF8C1FE5E038A" box="[294,413,1906,1933]" italics="true" pageId="1" pageNumber="506">C. helmeri</emphasis>
Zanata, Sarmento-Soares &amp; Martins-Pinheiro
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD1FF96FC16F8C1FF7003B7" authority="Netto-Ferreira, Birindelli &amp; Buckup" authorityName="Netto-Ferreira, Birindelli &amp; Buckup" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="mirim">
<emphasis id="B920A831FFD1FF96FC16F8C1FBD0038A" box="[943,1043,1907,1933]" italics="true" pageId="1" pageNumber="506">C. mirim</emphasis>
Netto-Ferreira, Birindelli &amp; Buckup
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD1FF96FF79F825FD9403B6" authority="Mendonca &amp; Netto-Ferreira" authorityName="Mendonca &amp; Netto-Ferreira" authorityYear="2015" box="[192,599,1942,1969]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="nana">
<emphasis id="B920A831FFD1FF96FF79F825FED403B6" box="[192,279,1943,1969]" italics="true" pageId="1" pageNumber="506">C. nana</emphasis>
Mendonça &amp; Netto-Ferreira
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD1FF96FDD8F825FBCE03B6" authority="Graca, Pavanelli &amp; Buckup" authorityName="Graca, Pavanelli &amp; Buckup" box="[609,1037,1942,1969]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="nupelia">
<emphasis id="B920A831FFD1FF96FDD8F825FD1703B6" box="[609,724,1942,1969]" italics="true" pageId="1" pageNumber="506">C. nupelia</emphasis>
Graça, Pavanelli &amp; Buckup
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD1FF96FBAEF825FAA903B6" authority="Melo &amp; Buckup" authorityName="Melo &amp; Buckup" authorityYear="2002" box="[1047,1386,1942,1969]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="stigmosum">
<emphasis id="B920A831FFD1FF96FBAEF825FB6D03B6" box="[1047,1198,1943,1969]" italics="true" pageId="1" pageNumber="506">C. stigmosum</emphasis>
Melo &amp; Buckup
</taxonomicName>
, and 
<taxonomicName id="4C540FA0FFD1FF96FF2EF809FD8803D2" authority="Graca, Pavanelli &amp; Buckup" authorityName="Graca, Pavanelli &amp; Buckup" box="[151,587,1978,2005]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="xavante">
<emphasis id="B920A831FFD1FF96FF2EF809FECD03D2" box="[151,270,1979,2005]" italics="true" pageId="1" pageNumber="506">C. xavante</emphasis>
Graça, Pavanelli &amp; Buckup
</taxonomicName>
, by the presence of an incomplete lateral line and the absence of the adipose fin. The new species can be distinguished from 
<taxonomicName id="4C540FA0FFD1FF96FD11F86DFCDD03FE" authorityName="Zanata, Sarmento-Soares &amp; Martins-Pinheiro" authorityYear="2015" box="[680,798,2014,2041]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="helmeri">
<emphasis id="B920A831FFD1FF96FD11F86DFCDD03FE" box="[680,798,2014,2041]" italics="true" pageId="1" pageNumber="506">C. helmeri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C540FA0FFD1FF96FCEAF86DFC2803FE" authorityName="Melo &amp; Buckup" authorityYear="2002" box="[851,1003,2015,2041]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="506" phylum="Chordata" rank="species" species="stigmosum">
<emphasis id="B920A831FFD1FF96FCEAF86DFC2803FE" box="[851,1003,2015,2041]" italics="true" pageId="1" pageNumber="506">C. stigmosum</emphasis>
</taxonomicName>
by having 12 circumpeduncular scales (
<emphasis id="B920A831FFD2FF95FF18FF28FF7C04B5" box="[161,191,154,179]" italics="true" pageId="2" pageNumber="507">vs.</emphasis>
13 or 14), from 
<taxonomicName id="4C540FA0FFD2FF95FEC2FF2BFE2304B5" box="[379,480,153,179]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="chicoi">
<emphasis id="B920A831FFD2FF95FEC2FF2BFE2304B5" box="[379,480,153,179]" italics="true" pageId="2" pageNumber="507">C. chicoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD2FF95FE56FF2BFD9004B5" box="[495,595,153,179]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="mirim">
<emphasis id="B920A831FFD2FF95FE56FF2BFD9004B5" box="[495,595,153,179]" italics="true" pageId="2" pageNumber="507">C. mirim</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C540FA0FFD2FF95FD33FF2BFD2704B5" authorityName="Mendonca &amp; Netto-Ferreira" authorityYear="2015" box="[650,740,153,179]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="nana">
<emphasis id="B920A831FFD2FF95FD33FF2BFD2704B5" box="[650,740,153,179]" italics="true" pageId="2" pageNumber="507">C. nana</emphasis>
</taxonomicName>
by the presence of a thin, inconspicuous or dashed midlateral dark stripe (
<emphasis id="B920A831FFD2FF95FEA7FF0CFEFE04D1" box="[286,317,190,215]" italics="true" pageId="2" pageNumber="507">vs.</emphasis>
presence of conspicuous dark midlateral stripe) and from 
<emphasis id="B920A831FFD2FF95FC71FF0FFB6004D1" box="[968,1187,189,215]" italics="true" pageId="2" pageNumber="507">
<taxonomicName id="4C540FA0FFD2FF95FC71FF0FFBE004D1" authorityName="Mendonca &amp; Netto-Ferreira" authorityYear="2015" box="[968,1059,189,215]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="nana">C. nana</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD2FF95FB97FF0FFB6004D1" box="[1070,1187,189,215]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="nupelia">C. nupelia</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C540FA0FFD2FF95FB60FF0FFA9304D1" box="[1241,1360,189,215]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="xavante">
<emphasis id="B920A831FFD2FF95FB60FF0FFA9304D1" box="[1241,1360,189,215]" italics="true" pageId="2" pageNumber="507">C. xavante</emphasis>
</taxonomicName>
by the absence of a peduncular blotch (
<emphasis id="B920A831FFD2FF95FDB3FF50FDEA04FD" box="[522,553,226,251]" italics="true" pageId="2" pageNumber="507">vs.</emphasis>
presence). It can be further distinguished from 
<taxonomicName id="4C540FA0FFD2FF95FBF3FF53FB0004FD" authorityName="Zanata, Sarmento-Soares &amp; Martins-Pinheiro" authorityYear="2015" box="[1098,1219,225,251]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="helmeri">
<emphasis id="B920A831FFD2FF95FBF3FF53FB0004FD" box="[1098,1219,225,251]" italics="true" pageId="2" pageNumber="507">C. helmeri</emphasis>
</taxonomicName>
by having a single series of dentary teeth (
<emphasis id="B920A831FFD2FF95FE1AFEB4FE790519" box="[419,442,262,287]" italics="true" pageId="2" pageNumber="507">vs</emphasis>
. two series of dentary teeth), a well-developed supraorbital (
<emphasis id="B920A831FFD2FF95FBD6FEB4FB450519" box="[1135,1158,262,287]" italics="true" pageId="2" pageNumber="507">vs</emphasis>
. supraorbital reduced or absent) and isthmus completely covered by scales (
<emphasis id="B920A831FFD2FF95FD5EFE98FCC50545" box="[743,774,298,323]" italics="true" pageId="2" pageNumber="507">vs.</emphasis>
anteriormost portion of isthmus naked). The new species further differs from 
<taxonomicName id="4C540FA0FFD2FF95FEC0FEFFFE1C0561" box="[377,479,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="chicoi">
<emphasis id="B920A831FFD2FF95FEC0FEFFFE1C0561" box="[377,479,333,359]" italics="true" pageId="2" pageNumber="507">C. chicoi</emphasis>
</taxonomicName>
by the presence of a dark rounded inconspicuous humeral blotch, shaped like an upside-down acute triangle).
</paragraph>
</subSubSection>
<subSubSection id="C34E27A8FFD2FF92FF7EFE26FD3506BD" lastPageId="5" lastPageNumber="510" pageId="2" pageNumber="507" type="description">
<paragraph id="8BEB7423FFD2FF95FF7EFE26FB430681" blockId="2.[151,1437,153,647]" pageId="2" pageNumber="507">
<emphasis id="B920A831FFD2FF95FF7EFE26FE9A05A9" bold="true" box="[199,345,404,431]" pageId="2" pageNumber="507">Description.</emphasis>
Morphometric data of 
<typeStatus id="54EFCA81FFD2FF95FDE5FE27FD7D05A9" box="[604,702,405,431]" pageId="2" pageNumber="507" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D52BFAAFFD2FF95FD4DFE27FC4105A9" box="[756,898,405,431]" pageId="2" pageNumber="507" type="generic" typeStatus="paratypes">24 paratypes</specimenCount>
in 
<tableCitation id="C6D64198FFD2FF95FC1CFE27FC3A05A9" box="[933,1017,405,431]" captionStart="TABLE 1" captionStartId="2.[151,239,694,718]" captionText="TABLE 1. Morphometric data of holotype and paratypes of Characidium cacah (n = 25), range includes the holotype. SD = standard deviation." pageId="2" pageNumber="507">Table 1</tableCitation>
. Body elongate and moderately compressed. Greatest body depth at vertical through dorsal-fin origin. Dorsal profile convex from upper lip to interorbital area, slightly convex from this point to dorsal-fin origin, convex and posteroventrally inclined along dorsal-fin base, almost straight between dorsal-fin terminus to anteriormost caudal-fin procurrent ray. Ventral profile slightly convex near dentary symphysis, straight or slightly convex from that area to pelvic-fin origin, straight from latter point to anal-fin origin, slightly concave along anal-fin base, and straight from anal-fin terminus to anteriormost ventral caudal-fin procurrent ray. Snout triangular in lateral view, rounded in dorsal view.
</paragraph>
<caption id="DF2B24ABFFD2FF95FF2EFD04FE6206F5" pageId="2" pageNumber="507">
<paragraph id="8BEB7423FFD2FF95FF2EFD04FE6206F5" blockId="2.[151,1437,694,755]" pageId="2" pageNumber="507">
<emphasis id="B920A831FFD2FF95FF2EFD04FEC806C9" bold="true" box="[151,267,694,719]" pageId="2" pageNumber="507">TABLE 1.</emphasis>
Morphometric data of holotype and paratypes of 
<taxonomicName id="4C540FA0FFD2FF95FC91FD05FC3E06C9" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[808,1021,695,719]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="507" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD2FF95FC91FD05FC3E06C9" box="[808,1021,695,719]" italics="true" pageId="2" pageNumber="507">Characidium cacah</emphasis>
</taxonomicName>
(n = 25), range includes the holotype. SD = standard deviation.
</paragraph>
</caption>
<paragraph id="8BEB7423FFD2FF95FD47FCB0FAF102BE" pageId="2" pageNumber="507">
<table id="F9548683FFD20068FF1FFCB0FAF102BF" box="[166,1330,770,1721]" gridcols="5" gridrows="24" pageId="2" pageNumber="507">
<tr id="35647661FFD20068FF1FFCB0FAF1071D" box="[166,1330,770,795]" gridrow="0" pageId="2" pageNumber="507" rowspan-0="1">
<th id="76B51F1DFFD20068FD47FCB0FC9F071D" box="[766,860,770,795]" gridcol="1" gridrow="0" pageId="2" pageNumber="507">Holotype</th>
<th id="76B51F1DFFD20068FC1DFCB0FBCA071D" box="[932,1033,770,795]" gridcol="2" gridrow="0" pageId="2" pageNumber="507">Range</th>
<th id="76B51F1DFFD20068FB3DFCB0FB7D071D" box="[1156,1214,770,795]" gridcol="3" gridrow="0" pageId="2" pageNumber="507">Mean</th>
<th id="76B51F1DFFD20068FAABFCB0FAF1071D" box="[1298,1330,770,795]" gridcol="4" gridrow="0" pageId="2" pageNumber="507">SD</th>
</tr>
<tr id="35647661FFD20068FF1FFC98FAF10745" box="[166,1330,810,835]" gridrow="1" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFC98FE780745" box="[166,443,810,835]" gridcol="0" gridrow="1" pageId="2" pageNumber="507">Total length (mm)</th>
<td id="76B51F1DFFD20068FD47FC98FC9F0745" box="[766,860,810,835]" gridcol="1" gridrow="1" pageId="2" pageNumber="507">45.0</td>
<td id="76B51F1DFFD20068FC1DFC98FBCA0745" box="[932,1033,810,835]" gridcol="2" gridrow="1" pageId="2" pageNumber="507">31.8–49.7</td>
<td id="76B51F1DFFD20068FB3DFC98FB7D0745" box="[1156,1214,810,835]" gridcol="3" gridrow="1" pageId="2" pageNumber="507">-</td>
<td id="76B51F1DFFD20068FAABFC98FAF10745" box="[1298,1330,810,835]" gridcol="4" gridrow="1" pageId="2" pageNumber="507">-</td>
</tr>
<tr id="35647661FFD20068FF1FFCE1FAF1076A" box="[166,1330,851,876]" gridrow="2" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFCE1FE78076A" box="[166,443,851,876]" gridcol="0" gridrow="2" pageId="2" pageNumber="507">Standard length (mm)</th>
<td id="76B51F1DFFD20068FD47FCE1FC9F076A" box="[766,860,851,876]" gridcol="1" gridrow="2" pageId="2" pageNumber="507">36.5</td>
<td id="76B51F1DFFD20068FC1DFCE1FBCA076A" box="[932,1033,851,876]" gridcol="2" gridrow="2" pageId="2" pageNumber="507">25.4–39.7</td>
<td id="76B51F1DFFD20068FB3DFCE1FB7D076A" box="[1156,1214,851,876]" gridcol="3" gridrow="2" pageId="2" pageNumber="507">-</td>
<td id="76B51F1DFFD20068FAABFCE1FAF1076A" box="[1298,1330,851,876]" gridcol="4" gridrow="2" pageId="2" pageNumber="507">-</td>
</tr>
<tr id="35647661FFD20068FF1FFCC8FAF10795" box="[166,1330,890,915]" gridrow="3" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFCC8FAF10795" box="[166,1330,890,915]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="2" pageNumber="507">
<emphasis id="B920A831FFD2FF95FF1FFCC9FE0C0795" bold="true" box="[166,463,890,915]" pageId="2" pageNumber="507">Percents of standard length</emphasis>
</th>
</tr>
<tr id="35647661FFD20068FF1FFC11FAF107BA" box="[166,1330,931,956]" gridrow="4" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFC11FE7807BA" box="[166,443,931,956]" gridcol="0" gridrow="4" pageId="2" pageNumber="507">Depth at dorsal-fin origin</th>
<td id="76B51F1DFFD20068FD47FC11FC9F07BA" box="[766,860,931,956]" gridcol="1" gridrow="4" pageId="2" pageNumber="507">27.4</td>
<td id="76B51F1DFFD20068FC1DFC11FBCA07BA" box="[932,1033,931,956]" gridcol="2" gridrow="4" pageId="2" pageNumber="507">25.0–28.3</td>
<td id="76B51F1DFFD20068FB3DFC11FB7D07BA" box="[1156,1214,931,956]" gridcol="3" gridrow="4" pageId="2" pageNumber="507">26.3</td>
<td id="76B51F1DFFD20068FAABFC11FAF107BA" box="[1298,1330,931,956]" gridcol="4" gridrow="4" pageId="2" pageNumber="507">1.1</td>
</tr>
<tr id="35647661FFD20068FF1FFC79FAF107E2" box="[166,1330,971,996]" gridrow="5" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFC79FE7807E2" box="[166,443,971,996]" gridcol="0" gridrow="5" pageId="2" pageNumber="507">Depth at anal-fin origin</th>
<td id="76B51F1DFFD20068FD47FC79FC9F07E2" box="[766,860,971,996]" gridcol="1" gridrow="5" pageId="2" pageNumber="507">18.6</td>
<td id="76B51F1DFFD20068FC1DFC79FBCA07E2" box="[932,1033,971,996]" gridcol="2" gridrow="5" pageId="2" pageNumber="507">15.4–19.0</td>
<td id="76B51F1DFFD20068FB3DFC79FB7D07E2" box="[1156,1214,971,996]" gridcol="3" gridrow="5" pageId="2" pageNumber="507">17.5</td>
<td id="76B51F1DFFD20068FAABFC79FAF107E2" box="[1298,1330,971,996]" gridcol="4" gridrow="5" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FFC46FAF1000B" box="[166,1330,1012,1037]" gridrow="6" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFC46FE78000B" box="[166,443,1012,1037]" gridcol="0" gridrow="6" pageId="2" pageNumber="507">Caudal-peduncle depth</th>
<td id="76B51F1DFFD20068FD47FC46FC9F000B" box="[766,860,1012,1037]" gridcol="1" gridrow="6" pageId="2" pageNumber="507">12.1</td>
<td id="76B51F1DFFD20068FC1DFC46FBCA000B" box="[932,1033,1012,1037]" gridcol="2" gridrow="6" pageId="2" pageNumber="507">10.7–13.0</td>
<td id="76B51F1DFFD20068FB3DFC46FB7D000B" box="[1156,1214,1012,1037]" gridcol="3" gridrow="6" pageId="2" pageNumber="507">11.9</td>
<td id="76B51F1DFFD20068FAABFC46FAF1000B" box="[1298,1330,1012,1037]" gridcol="4" gridrow="6" pageId="2" pageNumber="507">0.5</td>
</tr>
<tr id="35647661FFD20068FF1FFBAEFAF10033" box="[166,1330,1052,1077]" gridrow="7" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFBAEFE780033" box="[166,443,1052,1077]" gridcol="0" gridrow="7" pageId="2" pageNumber="507">Caudal-peduncle length</th>
<td id="76B51F1DFFD20068FD47FBAEFC9F0033" box="[766,860,1052,1077]" gridcol="1" gridrow="7" pageId="2" pageNumber="507">16.7</td>
<td id="76B51F1DFFD20068FC1DFBAEFBCA0033" box="[932,1033,1052,1077]" gridcol="2" gridrow="7" pageId="2" pageNumber="507">15.9–18.9</td>
<td id="76B51F1DFFD20068FB3DFBAEFB7D0033" box="[1156,1214,1052,1077]" gridcol="3" gridrow="7" pageId="2" pageNumber="507">17.5</td>
<td id="76B51F1DFFD20068FAABFBAEFAF10033" box="[1298,1330,1052,1077]" gridcol="4" gridrow="7" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FFBF6FAF1005B" box="[166,1330,1092,1117]" gridrow="8" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFBF6FE78005B" box="[166,443,1092,1117]" gridcol="0" gridrow="8" pageId="2" pageNumber="507">Snout to dorsal-fin origin</th>
<td id="76B51F1DFFD20068FD47FBF6FC9F005B" box="[766,860,1092,1117]" gridcol="1" gridrow="8" pageId="2" pageNumber="507">51.8</td>
<td id="76B51F1DFFD20068FC1DFBF6FBCA005B" box="[932,1033,1092,1117]" gridcol="2" gridrow="8" pageId="2" pageNumber="507">49.6–53.2</td>
<td id="76B51F1DFFD20068FB3DFBF6FB7D005B" box="[1156,1214,1092,1117]" gridcol="3" gridrow="8" pageId="2" pageNumber="507">51.0</td>
<td id="76B51F1DFFD20068FAABFBF6FAF1005B" box="[1298,1330,1092,1117]" gridcol="4" gridrow="8" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FFBDEFAF10083" box="[166,1330,1132,1157]" gridrow="9" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFBDEFE780083" box="[166,443,1132,1157]" gridcol="0" gridrow="9" pageId="2" pageNumber="507">Snout to pectoral-fin origin</th>
<td id="76B51F1DFFD20068FD47FBDEFC9F0083" box="[766,860,1132,1157]" gridcol="1" gridrow="9" pageId="2" pageNumber="507">28.2</td>
<td id="76B51F1DFFD20068FC1DFBDEFBCA0083" box="[932,1033,1132,1157]" gridcol="2" gridrow="9" pageId="2" pageNumber="507">26.2–31.1</td>
<td id="76B51F1DFFD20068FB3DFBDEFB7D0083" box="[1156,1214,1132,1157]" gridcol="3" gridrow="9" pageId="2" pageNumber="507">28.3</td>
<td id="76B51F1DFFD20068FAABFBDEFAF10083" box="[1298,1330,1132,1157]" gridcol="4" gridrow="9" pageId="2" pageNumber="507">1.0</td>
</tr>
<tr id="35647661FFD20068FF1FFB27FAF100A8" box="[166,1330,1173,1198]" gridrow="10" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFB27FE7800A8" box="[166,443,1173,1198]" gridcol="0" gridrow="10" pageId="2" pageNumber="507">Snout to pelvic-fin origin</th>
<td id="76B51F1DFFD20068FD47FB27FC9F00A8" box="[766,860,1173,1198]" gridcol="1" gridrow="10" pageId="2" pageNumber="507">53.4</td>
<td id="76B51F1DFFD20068FC1DFB27FBCA00A8" box="[932,1033,1173,1198]" gridcol="2" gridrow="10" pageId="2" pageNumber="507">52.4–55.7</td>
<td id="76B51F1DFFD20068FB3DFB27FB7D00A8" box="[1156,1214,1173,1198]" gridcol="3" gridrow="10" pageId="2" pageNumber="507">53.7</td>
<td id="76B51F1DFFD20068FAABFB27FAF100A8" box="[1298,1330,1173,1198]" gridcol="4" gridrow="10" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FFB0FFAF100D0" box="[166,1330,1213,1238]" gridrow="11" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFB0FFE7800D0" box="[166,443,1213,1238]" gridcol="0" gridrow="11" pageId="2" pageNumber="507">Snout to anal-fin origin</th>
<td id="76B51F1DFFD20068FD47FB0FFC9F00D0" box="[766,860,1213,1238]" gridcol="1" gridrow="11" pageId="2" pageNumber="507">77.5</td>
<td id="76B51F1DFFD20068FC1DFB0FFBCA00D0" box="[932,1033,1213,1238]" gridcol="2" gridrow="11" pageId="2" pageNumber="507">75.5–78.8</td>
<td id="76B51F1DFFD20068FB3DFB0FFB7D00D0" box="[1156,1214,1213,1238]" gridcol="3" gridrow="11" pageId="2" pageNumber="507">77.1</td>
<td id="76B51F1DFFD20068FAABFB0FFAF100D0" box="[1298,1330,1213,1238]" gridcol="4" gridrow="11" pageId="2" pageNumber="507">0.8</td>
</tr>
<tr id="35647661FFD20068FF1FFB57FAF100F8" box="[166,1330,1253,1278]" gridrow="12" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFB57FE7800F8" box="[166,443,1253,1278]" gridcol="0" gridrow="12" pageId="2" pageNumber="507">Anal-apex distance</th>
<td id="76B51F1DFFD20068FD47FB57FC9F00F8" box="[766,860,1253,1278]" gridcol="1" gridrow="12" pageId="2" pageNumber="507">93.4</td>
<td id="76B51F1DFFD20068FC1DFB57FBCA00F8" box="[932,1033,1253,1278]" gridcol="2" gridrow="12" pageId="2" pageNumber="507">92.1–97.2</td>
<td id="76B51F1DFFD20068FB3DFB57FB7D00F8" box="[1156,1214,1253,1278]" gridcol="3" gridrow="12" pageId="2" pageNumber="507">94.2</td>
<td id="76B51F1DFFD20068FAABFB57FAF100F8" box="[1298,1330,1253,1278]" gridcol="4" gridrow="12" pageId="2" pageNumber="507">1.2</td>
</tr>
<tr id="35647661FFD20068FF1FFABFFAF10120" box="[166,1330,1293,1318]" gridrow="13" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFABFFE780120" box="[166,443,1293,1318]" gridcol="0" gridrow="13" pageId="2" pageNumber="507">Anus to anal-fin origin</th>
<td id="76B51F1DFFD20068FD47FABFFC9F0120" box="[766,860,1293,1318]" gridcol="1" gridrow="13" pageId="2" pageNumber="507">6.3</td>
<td id="76B51F1DFFD20068FC1DFABFFBCA0120" box="[932,1033,1293,1318]" gridcol="2" gridrow="13" pageId="2" pageNumber="507">4.7–7.1</td>
<td id="76B51F1DFFD20068FB3DFABFFB7D0120" box="[1156,1214,1293,1318]" gridcol="3" gridrow="13" pageId="2" pageNumber="507">6.1</td>
<td id="76B51F1DFFD20068FAABFABFFAF10120" box="[1298,1330,1293,1318]" gridcol="4" gridrow="13" pageId="2" pageNumber="507">0.6</td>
</tr>
<tr id="35647661FFD20068FF1FFA87FAF10148" box="[166,1330,1333,1358]" gridrow="14" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFA87FE780148" box="[166,443,1333,1358]" gridcol="0" gridrow="14" pageId="2" pageNumber="507">Body width</th>
<td id="76B51F1DFFD20068FD47FA87FC9F0148" box="[766,860,1333,1358]" gridcol="1" gridrow="14" pageId="2" pageNumber="507">15.3</td>
<td id="76B51F1DFFD20068FC1DFA87FBCA0148" box="[932,1033,1333,1358]" gridcol="2" gridrow="14" pageId="2" pageNumber="507">12.7–16.3</td>
<td id="76B51F1DFFD20068FB3DFA87FB7D0148" box="[1156,1214,1333,1358]" gridcol="3" gridrow="14" pageId="2" pageNumber="507">14.8</td>
<td id="76B51F1DFFD20068FAABFA87FAF10148" box="[1298,1330,1333,1358]" gridcol="4" gridrow="14" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FFAECFAF10171" box="[166,1330,1374,1399]" gridrow="15" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFAECFE780171" box="[166,443,1374,1399]" gridcol="0" gridrow="15" pageId="2" pageNumber="507">Head length</th>
<td id="76B51F1DFFD20068FD47FAECFC9F0171" box="[766,860,1374,1399]" gridcol="1" gridrow="15" pageId="2" pageNumber="507">28.3</td>
<td id="76B51F1DFFD20068FC1DFAECFBCA0171" box="[932,1033,1374,1399]" gridcol="2" gridrow="15" pageId="2" pageNumber="507">25.8–29.0</td>
<td id="76B51F1DFFD20068FB3DFAECFB7D0171" box="[1156,1214,1374,1399]" gridcol="3" gridrow="15" pageId="2" pageNumber="507">27.4</td>
<td id="76B51F1DFFD20068FAABFAECFAF10171" box="[1298,1330,1374,1399]" gridcol="4" gridrow="15" pageId="2" pageNumber="507">1.0</td>
</tr>
<tr id="35647661FFD20068FF1FFA34FAF10199" box="[166,1330,1414,1439]" gridrow="16" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFA34FAF10199" box="[166,1330,1414,1439]" colspan="5" colspanRight="4" gridcol="0" gridrow="16" pageId="2" pageNumber="507">
<emphasis id="B920A831FFD2FF95FF1FFA34FE610198" bold="true" box="[166,418,1414,1439]" pageId="2" pageNumber="507">Percents of head length</emphasis>
</th>
</tr>
<tr id="35647661FFD20068FF1FFA1CFAF101C1" box="[166,1330,1454,1479]" gridrow="17" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFA1CFE7801C1" box="[166,443,1454,1479]" gridcol="0" gridrow="17" pageId="2" pageNumber="507">Horizontal eye diameter</th>
<td id="76B51F1DFFD20068FD47FA1CFC9F01C1" box="[766,860,1454,1479]" gridcol="1" gridrow="17" pageId="2" pageNumber="507">26.2</td>
<td id="76B51F1DFFD20068FC1DFA1CFBCA01C1" box="[932,1033,1454,1479]" gridcol="2" gridrow="17" pageId="2" pageNumber="507">25.5–29.9</td>
<td id="76B51F1DFFD20068FB3DFA1CFB7D01C1" box="[1156,1214,1454,1479]" gridcol="3" gridrow="17" pageId="2" pageNumber="507">27.3</td>
<td id="76B51F1DFFD20068FAABFA1CFAF101C1" box="[1298,1330,1454,1479]" gridcol="4" gridrow="17" pageId="2" pageNumber="507">1.3</td>
</tr>
<tr id="35647661FFD20068FF1FFA64FAF101E9" box="[166,1330,1494,1519]" gridrow="18" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFA64FE7801E9" box="[166,443,1494,1519]" gridcol="0" gridrow="18" pageId="2" pageNumber="507">Snout length</th>
<td id="76B51F1DFFD20068FD47FA64FC9F01E9" box="[766,860,1494,1519]" gridcol="1" gridrow="18" pageId="2" pageNumber="507">24.3</td>
<td id="76B51F1DFFD20068FC1DFA64FBCA01E9" box="[932,1033,1494,1519]" gridcol="2" gridrow="18" pageId="2" pageNumber="507">19.7–24.3</td>
<td id="76B51F1DFFD20068FB3DFA64FB7D01E9" box="[1156,1214,1494,1519]" gridcol="3" gridrow="18" pageId="2" pageNumber="507">21.9</td>
<td id="76B51F1DFFD20068FAABFA64FAF101E9" box="[1298,1330,1494,1519]" gridcol="4" gridrow="18" pageId="2" pageNumber="507">1.1</td>
</tr>
<tr id="35647661FFD20068FF1FFA4DFAF1021E" box="[166,1330,1535,1560]" gridrow="19" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FFA4DFE78021E" box="[166,443,1535,1560]" gridcol="0" gridrow="19" pageId="2" pageNumber="507">Snout to maxillary tip</th>
<td id="76B51F1DFFD20068FD47FA4DFC9F021E" box="[766,860,1535,1560]" gridcol="1" gridrow="19" pageId="2" pageNumber="507">26.2</td>
<td id="76B51F1DFFD20068FC1DFA4DFBCA021E" box="[932,1033,1535,1560]" gridcol="2" gridrow="19" pageId="2" pageNumber="507">23.4–26.4</td>
<td id="76B51F1DFFD20068FB3DFA4DFB7D021E" box="[1156,1214,1535,1560]" gridcol="3" gridrow="19" pageId="2" pageNumber="507">24.8</td>
<td id="76B51F1DFFD20068FAABFA4DFAF1021E" box="[1298,1330,1535,1560]" gridcol="4" gridrow="19" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FF995FAF10246" box="[166,1330,1575,1600]" gridrow="20" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FF995FE780246" box="[166,443,1575,1600]" gridcol="0" gridrow="20" pageId="2" pageNumber="507">Anterior naris to orbit</th>
<td id="76B51F1DFFD20068FD47F995FC9F0246" box="[766,860,1575,1600]" gridcol="1" gridrow="20" pageId="2" pageNumber="507">10.7</td>
<td id="76B51F1DFFD20068FC1DF995FBCA0246" box="[932,1033,1575,1600]" gridcol="2" gridrow="20" pageId="2" pageNumber="507">8.0–11.3</td>
<td id="76B51F1DFFD20068FB3DF995FB7D0246" box="[1156,1214,1575,1600]" gridcol="3" gridrow="20" pageId="2" pageNumber="507">9.7</td>
<td id="76B51F1DFFD20068FAABF995FAF10246" box="[1298,1330,1575,1600]" gridcol="4" gridrow="20" pageId="2" pageNumber="507">0.9</td>
</tr>
<tr id="35647661FFD20068FF1FF9FDFAF1026E" box="[166,1330,1615,1640]" gridrow="21" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FF9FDFE78026E" box="[166,443,1615,1640]" gridcol="0" gridrow="21" pageId="2" pageNumber="507">Posterior naris to orbit</th>
<td id="76B51F1DFFD20068FD47F9FDFC9F026E" box="[766,860,1615,1640]" gridcol="1" gridrow="21" pageId="2" pageNumber="507">4.9</td>
<td id="76B51F1DFFD20068FC1DF9FDFBCA026E" box="[932,1033,1615,1640]" gridcol="2" gridrow="21" pageId="2" pageNumber="507">3.0–5.5</td>
<td id="76B51F1DFFD20068FB3DF9FDFB7D026E" box="[1156,1214,1615,1640]" gridcol="3" gridrow="21" pageId="2" pageNumber="507">4.3</td>
<td id="76B51F1DFFD20068FAABF9FDFAF1026E" box="[1298,1330,1615,1640]" gridcol="4" gridrow="21" pageId="2" pageNumber="507">0.8</td>
</tr>
<tr id="35647661FFD20068FF1FF9C5FAF10296" box="[166,1330,1655,1680]" gridrow="22" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FF9C5FE780296" box="[166,443,1655,1680]" gridcol="0" gridrow="22" pageId="2" pageNumber="507">Cheek depth</th>
<td id="76B51F1DFFD20068FD47F9C5FC9F0296" box="[766,860,1655,1680]" gridcol="1" gridrow="22" pageId="2" pageNumber="507">6.8</td>
<td id="76B51F1DFFD20068FC1DF9C5FBCA0296" box="[932,1033,1655,1680]" gridcol="2" gridrow="22" pageId="2" pageNumber="507">5.3–7.9</td>
<td id="76B51F1DFFD20068FB3DF9C5FB7D0296" box="[1156,1214,1655,1680]" gridcol="3" gridrow="22" pageId="2" pageNumber="507">6.6</td>
<td id="76B51F1DFFD20068FAABF9C5FAF10296" box="[1298,1330,1655,1680]" gridcol="4" gridrow="22" pageId="2" pageNumber="507">0.7</td>
</tr>
<tr id="35647661FFD20068FF1FF912FAF102BF" box="[166,1330,1696,1721]" gridrow="23" pageId="2" pageNumber="507">
<th id="76B51F1DFFD20068FF1FF912FE7802BF" box="[166,443,1696,1721]" gridcol="0" gridrow="23" pageId="2" pageNumber="507">Least interorbital width</th>
<td id="76B51F1DFFD20068FD47F912FC9F02BF" box="[766,860,1696,1721]" gridcol="1" gridrow="23" pageId="2" pageNumber="507">20.4</td>
<td id="76B51F1DFFD20068FC1DF912FBCA02BF" box="[932,1033,1696,1721]" gridcol="2" gridrow="23" pageId="2" pageNumber="507">17.3–20.6</td>
<td id="76B51F1DFFD20068FB3DF912FB7D02BF" box="[1156,1214,1696,1721]" gridcol="3" gridrow="23" pageId="2" pageNumber="507">19.1</td>
<td id="76B51F1DFFD20068FAABF912FAF102BF" box="[1298,1330,1696,1721]" gridcol="4" gridrow="23" pageId="2" pageNumber="507">1.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEB7423FFD2FF93FF7EF95AFE940519" blockId="2.[151,1437,1768,1975]" lastBlockId="4.[151,1437,153,863]" lastPageId="4" lastPageNumber="509" pageId="2" pageNumber="507">Mouth subterminal, aligned or slightly lower than ventral margin of orbit. Distal tip of maxilla not reaching vertical through anterior margin of orbit. Orbit approximately round, larger than snout length. Cheek thin, depth approximately a fourth to a fifth of orbit diameter. Nares separated, without distinctly raised margins; posterior naris considerably closer to orbit than to anterior naris. Supraorbital well developed, inner border convex and outer border slightly concave. Nasal bones restricted to the ossified canal. Parietal fontanel limited anteriorly by frontals. Parietal branch of supraorbital canal absent. Dentary teeth in single row with 8 (1), 9 (4) or 10 (3) uni- or bicuspid teeth; teeth decreasing in size from symphysis. Premaxilla with single row of 6 (1), 7* (18) or 8 (6) unicuspid triangular teeth, decreasing in size from symphysis. Maxillary edentulous. Ectopterygoid with a patch of about 6–10 (2) teeth. Mesopterygoid teeth absent. Branchiostegal rays 4 (2), 3 connected to anterior ceratohyal, 1 connected to the posterior ceratohyal.</paragraph>
<caption id="DF2B24ABFFD3FF94FF2EF8C4FE9303FD" ID-DOI="http://doi.org/10.5281/zenodo.3891281" ID-Zenodo-Dep="3891281" httpUri="https://zenodo.org/record/3891281/files/figure.png" pageId="3" pageNumber="508" startId="3.[151,250,1910,1935]" targetBox="[215,1373,179,1882]" targetPageId="3">
<paragraph id="8BEB7423FFD3FF94FF2EF8C4FE9303FD" blockId="3.[151,1437,1910,2043]" pageId="3" pageNumber="508">
<emphasis id="B920A831FFD3FF94FF2EF8C4FED70389" bold="true" box="[151,276,1910,1935]" pageId="3" pageNumber="508">FIGURE 1.</emphasis>
<taxonomicName id="4C540FA0FFD3FF94FEA3F8C4FE200388" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[282,483,1910,1934]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="508" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD3FF94FEA3F8C4FE200388" box="[282,483,1910,1934]" italics="true" pageId="3" pageNumber="508">Characidium cacah</emphasis>
</taxonomicName>
: (a) holotype, MZUSP 125765, 36.5 mm SL; (b) paratype, UFBA 9056, 36.2 mm SL, in life; (c) paratype, ZUEC 17227, 36.0 mm SL; (d) paratype, ZUEC 17227, 25.6 mm SL; (e) paratype, CICMG 158, 25.3 mm SL; (f) paratype, MCNIP 4530, 21.0 mm SL. All from Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub-basin, rio Cipó drainage.
</paragraph>
</caption>
<paragraph id="8BEB7423FFD4FF93FF7EFE9BFA92061D" blockId="4.[151,1437,153,863]" pageId="4" pageNumber="509">
Scales cycloid; 
<emphasis id="B920A831FFD4FF93FECFFE9BFE7D0545" box="[374,446,297,323]" italics="true" pageId="4" pageNumber="509">circuli</emphasis>
absent on exposed portion of scales; up to 12 divergent 
<emphasis id="B920A831FFD4FF93FB88FE9BFBA40545" box="[1073,1127,297,323]" italics="true" pageId="4" pageNumber="509">radii</emphasis>
present on exposed portion of scales. Lateral line reduced; perforated scales 6 (1), 7* (17), or 8 (7); total scales along lateral line series 34 (14) or 35* (11). Horizontal scale rows above lateral line 4* (24) or 5 (1); horizontal scale rows below lateral line 5* (22) or 6 (3). Predorsal scales 9 (1), 10* (8), 11 (10) or irregularly arranged (6). Scale rows around caudal peduncle 12 (25). Isthmus completely covered with scales. Pseudotympanum present as a muscular hiatus at vertical through anterior portion of swimbladder and mostly situated between ribs of the fifth and sixth vertebrae, with large opening anterior to the sixth vertebrae; rib of fifth vertebrae largely evident and a small part of sixth evident (
<figureCitation id="136F68A6FFD4FF93FB46FDB3FA87061D" box="[1279,1348,513,539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[151,1437,885,1777]" captionTargetId="figure@4.[151,1437,885,1777]" captionTargetPageId="4" captionText="FIGURE 2. Pseudotympanum of Characidium cacah, paratype CICMG 158, 35.6 mm SL, lateral view, right side." figureDoi="http://doi.org/10.5281/zenodo.3891283" httpUri="https://zenodo.org/record/3891283/files/figure.png" pageId="4" pageNumber="509">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BEB7423FFD4FF93FF7EFD97FAC30759" blockId="4.[151,1437,153,863]" pageId="4" pageNumber="509">Dorsal-fin rays ii,8 (1), ii,9* (23) or iii,9 (1); distal margin of dorsal fin rounded. Adipose fin absent. Pectoral-fin rays 10–11 total rays; unbranched rays ii (1), iii (19) or iv* (5), and posterior total rays 5 (1), 6* (7), 7 (16) or 8 (1) branched rays; first and second branched pectoral-fin rays usually longest; posterior tip of pectoral fin reaching pelvicfin origin in specimens up to 30.0 mm SL and larger specimens with pectoral-fin tip falling short of pelvic-fin insertion. Pelvic-fin rays i,6,i* (14) or i,7 (11); second and third branched rays longest; posterior tip of pelvic fin falling short of anal-fin origin.Anal-fin rays iii,5 (1) or iii,6* (24); posterior margin of anal fin slightly rounded, with second branched ray usually longest; last ray adnate* (24) or simple (1). Caudal-fin rays i,8,8,i* (22), i,8,7,i (2) or i,9,8,i (1). Dorsal procurrent caudal-fin rays 8 (2); ventral procurrent caudal-fin rays 6 (2). Total vertebrae 33 (2); precaudal vertebrae 18 (2); caudal vertebrae 15 (2). Supraneural bones 5 (1) or 6 (1). Epural bones 2 (2). Uroneural bone 1 (2).</paragraph>
<caption id="DF2B24ABFFD4FF93FF2EF8B8FAE10325" ID-DOI="http://doi.org/10.5281/zenodo.3891283" ID-Zenodo-Dep="3891283" box="[151,1314,1802,1827]" httpUri="https://zenodo.org/record/3891283/files/figure.png" pageId="4" pageNumber="509" startId="4.[151,250,1802,1827]" targetBox="[151,1437,885,1777]" targetPageId="4">
<paragraph id="8BEB7423FFD4FF93FF2EF8B8FAE10325" blockId="4.[151,1314,1802,1827]" box="[151,1314,1802,1827]" pageId="4" pageNumber="509">
<emphasis id="B920A831FFD4FF93FF2EF8B8FED00325" bold="true" box="[151,275,1802,1827]" pageId="4" pageNumber="509">FIGURE 2.</emphasis>
Pseudotympanum of 
<taxonomicName id="4C540FA0FFD4FF93FE48F8B9FD7A0325" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[497,697,1803,1827]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="509" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD4FF93FE48F8B9FD7A0325" box="[497,697,1803,1827]" italics="true" pageId="4" pageNumber="509">Characidium cacah</emphasis>
</taxonomicName>
, paratype CICMG 158, 35.6 mm SL, lateral view, right side.
</paragraph>
</caption>
<paragraph id="8BEB7423FFD4FF92FF7EF8E2FC880655" blockId="4.[151,1436,1872,2044]" lastBlockId="5.[151,1437,153,2026]" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
<emphasis id="B920A831FFD4FF93FF7EF8E2FE51036D" bold="true" box="[199,402,1872,1899]" pageId="4" pageNumber="509">Color in alcohol.</emphasis>
Ground color of head and body pale brown (
<figureCitation id="136F68A6FFD4FF93FC1BF8E3FBCE036A" box="[930,1037,1873,1900]" captionStart="FIGURE 1" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[215,1373,179,1882]" captionTargetId="figure@3.[215,1373,179,1882]" captionTargetPageId="3" captionText="FIGURE 1. Characidium cacah: (a) holotype, MZUSP 125765, 36.5 mm SL; (b) paratype, UFBA 9056, 36.2 mm SL, in life; (c) paratype, ZUEC 17227, 36.0 mm SL; (d) paratype, ZUEC 17227, 25.6 mm SL; (e) paratype, CICMG 158, 25.3 mm SL; (f) paratype, MCNIP 4530, 21.0 mm SL. All from Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub-basin, rio Cipó drainage." figureDoi="http://doi.org/10.5281/zenodo.3891281" httpUri="https://zenodo.org/record/3891281/files/figure.png" pageId="4" pageNumber="509">Fig. 1a, 1</figureCitation>
c-f). Head with tiny melanophores distributed over dorsal, lateral and ventral surfaces, not forming blotches; dorsum darker than lateral surface and ventral surface clearest. Concentration of melanophores forming dark area from tip of snout to eye, not forming a conspicuous dark stripe. Opercle usually darker than surrounding areas, with higher concentration of melanophores on its central portion or dorsal half. Humeral region with round humeral blotch, variably marked. Midlateral pig- mentation forming a continuous narrow dark stripe in some specimens or a series of dark dashes from area posterior to the humeral blotch to caudal peduncle in other specimens, but not reaching end of caudal peduncle. Scales on flanks with dark borders, forming an overall reticulate pattern; pattern more conspicuous on dorsal half of body. Most specimens with up to 14 somewhat zigzag-shaped narrow bars and less pigmented specimens almost without dark spots or vertical bars. Variation in pattern of bars apparently not related to ontogeny. When present, bars usually reaching at least the horizontal through pectoral-fin origin ventrally; bars located posteriorly to the dorsal fin connected dorsally in some specimens. Ventral portion of the body less pigmented, with sparse tiny melanophores, not forming reticulate pattern or spots. Basicaudal spot variably marked, well visible in some specimens and inconspicuous or absent in others. All fins with melanophores concentrated over rays, interradial membranes mostly hyaline. Proximal portion of dorsal-fin rays and interradial membranes dark, forming a distinctly dark basal area; rays with dark dashes sparsely distributed, usually not forming bands. Caudal fin with dark dashes on rays, usually not forming bands; some specimens with dashes roughly arranged in up to three vertical bands crossing fin. Paired fins and anal fin slightly less pigmented, without dark dashes.
</paragraph>
<paragraph id="8BEB7423FFD5FF92FF7EFDE9FD3506BD" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EFDE9FE990670" bold="true" box="[199,346,603,630]" pageId="5" pageNumber="510">Color in life</emphasis>
. Ground color varying from pale yellow to pale brown, slightly olivaceous dorsally and white ventrally (
<figureCitation id="136F68A6FFD5FF92FEB3FDCCFE9E069F" box="[266,349,638,665]" captionStart="FIGURE 1" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[215,1373,179,1882]" captionTargetId="figure@3.[215,1373,179,1882]" captionTargetPageId="3" captionText="FIGURE 1. Characidium cacah: (a) holotype, MZUSP 125765, 36.5 mm SL; (b) paratype, UFBA 9056, 36.2 mm SL, in life; (c) paratype, ZUEC 17227, 36.0 mm SL; (d) paratype, ZUEC 17227, 25.6 mm SL; (e) paratype, CICMG 158, 25.3 mm SL; (f) paratype, MCNIP 4530, 21.0 mm SL. All from Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub-basin, rio Cipó drainage." figureDoi="http://doi.org/10.5281/zenodo.3891281" httpUri="https://zenodo.org/record/3891281/files/figure.png" pageId="5" pageNumber="510">Fig. 1b</figureCitation>
). Parts of iris, infraorbitals, and opercle silvery. Fins pale yellow. Dark marks and spots on body and fins similar to the pattern of preserved specimens.
</paragraph>
</subSubSection>
<subSubSection id="C34E27A8FFD5FF92FF7EFD76FEC7078D" pageId="5" pageNumber="510" type="etymology">
<paragraph id="8BEB7423FFD5FF92FF7EFD76FEAC0743" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EFD76FE8D06D8" bold="true" box="[199,334,708,734]" pageId="5" pageNumber="510">Etymology.</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FEEFFD76FDF606D8" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[342,565,708,734]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FEEFFD76FDF606D8" box="[342,565,708,734]" italics="true" pageId="5" pageNumber="510">Characidium cacah</emphasis>
</taxonomicName>
is named in honor of the ichthyologist Carlos B. M. Alves (Projeto Manuel-zão—UFMG), known to his friends as “Cacá”, for his great contribution to the knowledge of the ichthyofauna of the rio das Velhas basin and for being one of the collectors of the first specimens of the species described herein. A noun in apposition.
</paragraph>
<paragraph id="8BEB7423FFD5FF92FF7EFCFFFEC7078D" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EFCFFFE79076E" bold="true" box="[199,442,845,872]" pageId="5" pageNumber="510">Sexual dimorphism.</emphasis>
Dissected specimens were not mature sexually and no sexually dimorphic traits were observed.
</paragraph>
</subSubSection>
<subSubSection id="C34E27A8FFD5FF92FF7EFC21FACC02E8" pageId="5" pageNumber="510" type="distribution">
<paragraph id="8BEB7423FFD5FF92FF7EFC21FB9007D6" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EFC21FEA207A8" bold="true" box="[199,353,915,942]" pageId="5" pageNumber="510">Distribution.</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FEDEFC26FD8607A8" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[359,581,916,942]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FEDEFC26FD8607A8" box="[359,581,916,942]" italics="true" pageId="5" pageNumber="510">Characidium cacah</emphasis>
</taxonomicName>
is only known to occur in the rio Cipó drainage, a tributary of the middle rio das Velhas sub-basin, rio São Francisco basin, 
<collectingRegion id="4990BAC1FFD5FF92FD1BFC04FCB907D6" box="[674,890,950,976]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="510">Minas Gerais State</collectingRegion>
, 
<collectingCountry id="F34334B3FFD5FF92FC3CFC04FC0A07D6" box="[901,969,950,976]" name="Brazil" pageId="5" pageNumber="510">Brazil</collectingCountry>
(
<figureCitation id="136F68A6FFD5FF92FC60FC04FB8507D6" box="[985,1094,950,976]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1956,1981]" captionStartId-1="7.[151,250,1925,1950]" captionTargetBox-0="[141,1445,603,1924]" captionTargetBox-1="[188,1400,181,1900]" captionTargetId-0="figure@6.[136,1451,597,1931]" captionTargetId-1="figure@7.[188,1400,181,1900]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. Map of the rio das Velhas sub-basin, rio São Francisco basin, showing the collection sites of Characidium cacah (red star); symbol represents more than one sampling point." captionText-1="FIGURE 4. Sampling localities of Characidium cacah, Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub basin, rio Cipó drainage: (a) satellite image of a stretch of rio Cipó and surrounding lagoons (from Google Earth); (b) rio Cipó just after the meeting of rio Bocaina and rio Mascates; (c) rio Cipó at type locality; d) Lagoa do Boi." figureDoi-0="http://doi.org/10.5281/zenodo.3891285" figureDoi-1="http://doi.org/10.5281/zenodo.3891287" httpUri-0="https://zenodo.org/record/3891285/files/figure.png" httpUri-1="https://zenodo.org/record/3891287/files/figure.png" pageId="5" pageNumber="510">Figs. 3–4</figureCitation>
).
</paragraph>
<paragraph id="8BEB7423FFD5FF92FF7EFC6AFB8B01FD" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EFC6AFDE507F5" bold="true" box="[199,550,984,1011]" pageId="5" pageNumber="510">Habitat and ecological notes.</emphasis>
The rio Cipó is formed by the meeting of the rio Mascates and rio Bocaina, which have their headsprings within the Serra do Cipó National Park, possessing several temporary and permanent marginal lagoons mainly located at lower altitudes (
<figureCitation id="136F68A6FFD5FF92FD53FBACFCFC003E" box="[746,831,1054,1080]" captionStart="FIGURE 4" captionStartId="7.[151,250,1925,1950]" captionTargetBox="[188,1400,181,1900]" captionTargetId="figure@7.[188,1400,181,1900]" captionTargetPageId="7" captionText="FIGURE 4. Sampling localities of Characidium cacah, Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub basin, rio Cipó drainage: (a) satellite image of a stretch of rio Cipó and surrounding lagoons (from Google Earth); (b) rio Cipó just after the meeting of rio Bocaina and rio Mascates; (c) rio Cipó at type locality; d) Lagoa do Boi." figureDoi="http://doi.org/10.5281/zenodo.3891287" httpUri="https://zenodo.org/record/3891287/files/figure.png" pageId="5" pageNumber="510">Fig. 4a</figureCitation>
). Specimens of 
<taxonomicName id="4C540FA0FFD5FF92FC4EFBADFB9E003E" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[1015,1117,1054,1081]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FC4EFBADFB9E003E" box="[1015,1117,1054,1081]" italics="true" pageId="5" pageNumber="510">C. cacah</emphasis>
</taxonomicName>
were caught near submerse macrophytes and marginal grasses in lotic and lentic environments of the rio Cipó sub-basin, at relatively high elevations (783–803 meters a.s.l.). The stretches of the rio Cipó (
<figureCitation id="136F68A6FFD5FF92FC31FBD1FC0E0078" box="[904,973,1123,1150]" captionStart="FIGURE 4" captionStartId="7.[151,250,1925,1950]" captionTargetBox="[188,1400,181,1900]" captionTargetId="figure@7.[188,1400,181,1900]" captionTargetPageId="7" captionText="FIGURE 4. Sampling localities of Characidium cacah, Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub basin, rio Cipó drainage: (a) satellite image of a stretch of rio Cipó and surrounding lagoons (from Google Earth); (b) rio Cipó just after the meeting of rio Bocaina and rio Mascates; (c) rio Cipó at type locality; d) Lagoa do Boi." figureDoi="http://doi.org/10.5281/zenodo.3891287" httpUri="https://zenodo.org/record/3891287/files/figure.png" pageId="5" pageNumber="510">Fig. 4</figureCitation>
b-c) and rio Bocaina sampled vary from 9–15 meters wide, up to 
<quantity id="4CACD9C6FFD5FF92FE13FB34FE1600A7" box="[426,469,1158,1185]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="510" unit="m" value="1.0">1 m</quantity>
deep, possessing sandy substrate, riparian forest and slow to moderate water flow, and transparent water with a brown tinge due to the presence of humic and fulvic acids. Syntopic species in lotic sites include 
<taxonomicName id="4C540FA0FFD5FF92FF49FB7EFDE900E0" authority="(Cuvier)" baseAuthorityName="Cuvier" baseAuthorityYear="1819" box="[240,554,1227,1254]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B920A831FFD5FF92FF49FB7EFE0100E0" box="[240,450,1228,1254]" italics="true" pageId="5" pageNumber="510">Astyanax fasciatus</emphasis>
(Cuvier)
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FD8DFB7EFCDC00E0" authority="(Lutken)" baseAuthorityName="Lutken" baseAuthorityYear="1875" box="[564,799,1227,1254]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="B920A831FFD5FF92FD8DFB7EFD7000E0" box="[564,691,1228,1254]" italics="true" pageId="5" pageNumber="510">A. lacustris</emphasis>
(Lütken)
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FC93FB7EFC5200E0" box="[810,913,1228,1254]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD5FF92FC93FB7EFC5200E0" box="[810,913,1228,1254]" italics="true" pageId="5" pageNumber="510">Astyanax</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C540FA0FFD5FF92FC7CFB7EFACF00E3" authority="Ottoni" authorityName="Ottoni" authorityYear="2012" box="[965,1292,1227,1254]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="mattosi">
<emphasis id="B920A831FFD5FF92FC7CFB7EFB7800E3" box="[965,1211,1228,1254]" italics="true" pageId="5" pageNumber="510">Australoheros mattosi</emphasis>
Ottoni
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FAA1FB7EFEF9010E" authority="Gunther" authorityName="Gunther" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B920A831FFD5FF92FAA1FB7EFF17010E" italics="true" pageId="5" pageNumber="510">Brycon nattereri</emphasis>
Günther
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FEFFFB5CFCDA010F" authority=", Gymnotus" authorityName="Gymnotus" box="[326,793,1262,1289]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lagosantense">
<emphasis id="B920A831FFD5FF92FEFFFB5CFE19010E" box="[326,474,1262,1288]" italics="true" pageId="5" pageNumber="510">Characidium</emphasis>
cf. 
<emphasis id="B920A831FFD5FF92FDBFFB5CFCDA010F" box="[518,793,1262,1289]" italics="true" pageId="5" pageNumber="510">lagosantense, Gymnotus</emphasis>
</taxonomicName>
cf. 
<emphasis id="B920A831FFD5FF92FCFCFB5DFC57010E" box="[837,916,1263,1288]" italics="true" pageId="5" pageNumber="510">carapo</emphasis>
Linnaeus, 
<taxonomicName id="4C540FA0FFD5FF92FBABFB5DFB51010E" authorityName="Ellis" authorityYear="1911" box="[1042,1170,1263,1288]" class="Actinopterygii" family="Characidae" genus="Hasemania" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD5FF92FBABFB5DFB51010E" box="[1042,1170,1263,1288]" italics="true" pageId="5" pageNumber="510">Hasemania</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C540FA0FFD5FF92FB70FB5DFAAD010E" authorityName="Gill" authorityYear="1858" box="[1225,1390,1263,1288]" class="Actinopterygii" family="Characidae" genus="Hemigrammus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD5FF92FB70FB5DFAAD010E" box="[1225,1390,1263,1288]" italics="true" pageId="5" pageNumber="510">Hemigrammus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C540FA0FFD5FF92FF2EFAA3FE32012A" authority="(Gunther)" baseAuthorityName="Gunther" box="[151,497,1297,1324]" class="Actinopterygii" family="Erythrinidae" genus="Hoplias" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B920A831FFD5FF92FF2EFAA3FEB8012D" box="[151,379,1297,1323]" italics="true" pageId="5" pageNumber="510">Hoplias intermedius</emphasis>
(Günther)
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FE46FAA3FD96012D" authorityName="Eigenmann" authorityYear="1911" box="[511,597,1297,1323]" class="Actinopterygii" family="Characidae" genus="Knodus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD5FF92FE46FAA3FD96012D" box="[511,597,1297,1323]" italics="true" pageId="5" pageNumber="510">Knodus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C540FA0FFD5FF92FD30FAA3FBB0012D" authority="Lehmann A. &amp; Reis" authorityName="Lehmann A. &amp; Reis" authorityYear="2012" box="[649,1139,1297,1323]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B920A831FFD5FF92FD30FAA3FC48012D" box="[649,907,1297,1323]" italics="true" pageId="5" pageNumber="510">Parotocinclus robustus</emphasis>
Lehmann A. &amp; Reis
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FB38FAA3FA56012D" authority="Lucinda" authorityName="Lucinda" authorityYear="2008" box="[1153,1429,1297,1323]" class="Actinopterygii" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="uai">
<emphasis id="B920A831FFD5FF92FB38FAA3FAF0012D" box="[1153,1331,1297,1323]" italics="true" pageId="5" pageNumber="510">Phalloceros uai</emphasis>
Lucinda
</taxonomicName>
, and 
<taxonomicName id="4C540FA0FFD5FF92FF71FA86FDC9014B" authority="Reinhardt." authorityName="Reinhardt." box="[200,522,1331,1358]" class="Actinopterygii" family="Characidae" genus="Piabina" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="argentea">
<emphasis id="B920A831FFD5FF92FF71FA86FE4E014B" box="[200,397,1332,1358]" italics="true" pageId="5" pageNumber="510">Piabina argentea</emphasis>
Reinhardt.
</taxonomicName>
<taxonomicName id="4C540FA0FFD5FF92FDAAFA86FD310148" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[531,754,1332,1358]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FDAAFA86FD310148" box="[531,754,1332,1358]" italics="true" pageId="5" pageNumber="510">Characidium cacah</emphasis>
</taxonomicName>
was also sampled in the Lagoa do Boi (
<figureCitation id="136F68A6FFD5FF92FB78FA81FAD60148" box="[1217,1301,1331,1358]" captionStart="FIGURE 4" captionStartId="7.[151,250,1925,1950]" captionTargetBox="[188,1400,181,1900]" captionTargetId="figure@7.[188,1400,181,1900]" captionTargetPageId="7" captionText="FIGURE 4. Sampling localities of Characidium cacah, Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub basin, rio Cipó drainage: (a) satellite image of a stretch of rio Cipó and surrounding lagoons (from Google Earth); (b) rio Cipó just after the meeting of rio Bocaina and rio Mascates; (c) rio Cipó at type locality; d) Lagoa do Boi." figureDoi="http://doi.org/10.5281/zenodo.3891287" httpUri="https://zenodo.org/record/3891287/files/figure.png" pageId="5" pageNumber="510">Fig. 4d</figureCitation>
) and Lagoa Rasgada, the former a natural lagoon and the latter a man-rectified lagoon with a flooded swampy area. Both possess dark water, muddy bottom and are surrounded by cattle pastures and marginal macrophytes. In these relatively shallow (0.2–1.0 m deep) lentic sites, only small juveniles of 
<taxonomicName id="4C540FA0FFD5FF92FCEDFA2EFC7B01B0" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[852,952,1436,1462]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FCEDFA2EFC7B01B0" box="[852,952,1436,1462]" italics="true" pageId="5" pageNumber="510">C. cacah</emphasis>
</taxonomicName>
were caught (
<quantity id="4CACD9C6FFD5FF92FBE3FA2EFB3C01B0" box="[1114,1279,1436,1462]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" metricValueMax="2.79" metricValueMin="1.7899999999999998" pageId="5" pageNumber="510" unit="mm" value="22.9" valueMax="27.9" valueMin="17.9">17.9–27.9 mm</quantity>
SL), whereas the larger specimens were sampled only in the lotic sites. In the lagoons, 
<taxonomicName id="4C540FA0FFD5FF92FC06FA0DFBE101DE" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[959,1058,1470,1497]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FC06FA0DFBE101DE" box="[959,1058,1470,1497]" italics="true" pageId="5" pageNumber="510">C. cacah</emphasis>
</taxonomicName>
was sampled syntopically with 
<taxonomicName id="4C540FA0FFD5FF92FA3DFA0DFF3401FD" baseAuthorityName="Lutken" baseAuthorityYear="1875" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="B920A831FFD5FF92FA3DFA0DFF3401FD" italics="true" pageId="5" pageNumber="510">A. lacustris</emphasis>
</taxonomicName>
, 
<emphasis id="B920A831FFD5FF92FEBFFA53FEBD01FD" box="[262,382,1505,1531]" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C540FA0FFD5FF92FEBFFA53FEB901FD" authorityName="Ottoni" authorityYear="2012" box="[262,378,1505,1531]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="mattosi">A. mattosi</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FE3CFA53FD9F01FD" box="[389,604,1505,1531]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lagosantense">
<emphasis id="B920A831FFD5FF92FE3CFA53FE5C01FD" box="[389,415,1505,1531]" italics="true" pageId="5" pageNumber="510">C.</emphasis>
cf. 
<emphasis id="B920A831FFD5FF92FE73FA53FD9F01FD" box="[458,604,1505,1531]" italics="true" pageId="5" pageNumber="510">lagosantense</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FDD3FA50FD2901FD" authorityName="Ellis" authorityYear="1911" box="[618,746,1506,1531]" class="Actinopterygii" family="Characidae" genus="Hasemania" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD5FF92FDD3FA50FD2901FD" box="[618,746,1506,1531]" italics="true" pageId="5" pageNumber="510">Hasemania</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C540FA0FFD5FF92FCA7FA50FC0701FD" box="[798,964,1505,1531]" class="Actinopterygii" family="Erythrinidae" genus="Hoplias" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B920A831FFD5FF92FCA7FA50FC0701FD" box="[798,964,1505,1531]" italics="true" pageId="5" pageNumber="510">H. intermedius</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C540FA0FFD5FF92FBBBFA50FB8101FD" authorityName="Lucinda" authorityYear="2008" box="[1026,1090,1506,1531]" class="Actinopterygii" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="uai">
<emphasis id="B920A831FFD5FF92FBBBFA50FB8101FD" box="[1026,1090,1506,1531]" italics="true" pageId="5" pageNumber="510">P. uai</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEB7423FFD5FF92FF7EF9B1FACC02E8" blockId="5.[151,1437,153,2026]" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EF9B1FE780218" bold="true" box="[199,443,1539,1566]" pageId="5" pageNumber="510">Conservation status.</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FE7BF9B6FD630218" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[450,672,1540,1566]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FE7BF9B6FD630218" box="[450,672,1540,1566]" italics="true" pageId="5" pageNumber="510">Characidium cacah</emphasis>
</taxonomicName>
had been collected in five nearby localities in the rio Cipó drainage, rio das Velhas sub-basin. Several anthropogenic impacts are reported for the rio das Velhas sub-basin, among them pollution by sewage effluents and siltation (
<emphasis id="B920A831FFD5FF92FDC5F9F8FD680265" box="[636,683,1610,1635]" italics="true" pageId="5" pageNumber="510">e.g.,</emphasis>
<bibRefCitation id="EFC509D2FFD5FF92FD09F9FBFC740265" author="Pompeu, P. S. &amp; Alves, C. B. M. &amp; Callisto, M." box="[688,951,1608,1635]" pageId="5" pageNumber="510" pagination="11 - 22" refId="ref6184" refString="Pompeu, P. S., Alves, C. B. M. &amp; Callisto, M. (2005) The effects of urbanization on biodiversity and water quality in the Rio das Velhas basin, Brazil. American Fisheries Society Symposium, 47, 11 - 22." type="journal article" year="2005">Alves &amp; Pompeu, 2005</bibRefCitation>
; 
<bibRefCitation id="EFC509D2FFD5FF92FC7BF9FBFB600265" author="Pompeu, P. S. &amp; Alves, C. B. M. &amp; Callisto, M." box="[962,1187,1608,1635]" pageId="5" pageNumber="510" pagination="11 - 22" refId="ref6184" refString="Pompeu, P. S., Alves, C. B. M. &amp; Callisto, M. (2005) The effects of urbanization on biodiversity and water quality in the Rio das Velhas basin, Brazil. American Fisheries Society Symposium, 47, 11 - 22." type="journal article" year="2005">
Pompeu 
<emphasis id="B920A831FFD5FF92FB9DF9F8FB970265" box="[1060,1108,1609,1635]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
., 2005
</bibRefCitation>
; 
<bibRefCitation id="EFC509D2FFD5FF92FB17F9FBFAA00265" author="Leal, C. G. &amp; Junqueira, N. T. &amp; Pompeu, P. S." box="[1198,1379,1609,1635]" pageId="5" pageNumber="510" pagination="143 - 157" refId="ref5717" refString="Leal, C. G., Junqueira, N. T. &amp; Pompeu, P. S. (2011) Morphology and habitat use by fishes of the Rio das Velhas basin in southeastern Brazil. Environmental Biology of Fishes, 90, 143 - 157. https: // doi. org / 10.1007 / s 10641 - 010 - 9726 - 6" type="journal article" year="2011">
Leal 
<emphasis id="B920A831FFD5FF92FB5CF9F8FAD50265" box="[1253,1302,1609,1635]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
., 2011
</bibRefCitation>
; 
<bibRefCitation id="EFC509D2FFD5FF92FAD7F9FBFE920280" author="Ferreira, W. R. &amp; Rodrigues, D. N. &amp; Alves, C. B. M. &amp; Callisto, M." pageId="5" pageNumber="510" pagination="253 - 259" refId="ref5477" refString="Ferreira, W. R., Rodrigues, D. N., Alves, C. B. M. &amp; Callisto, M. (2012) Biomonitoramento de longo prazo da bacia do rio das Velhas atraves de um indice multimetrico bentonico. Revista Brasileira de Recursos Hidricos, 17 (3), 253 - 259. https: // doi. org / 10.21168 / rbrh. v 17 n 3. p 253 - 259" type="journal article" year="2012">
Ferreira 
<emphasis id="B920A831FFD5FF92FF69F9DEFEC20280" box="[208,257,1644,1670]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
., 2012
</bibRefCitation>
). However, 
<taxonomicName id="4C540FA0FFD5FF92FE6CF9DEFDF40280" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[469,567,1644,1670]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FE6CF9DEFDF40280" box="[469,567,1644,1670]" italics="true" pageId="5" pageNumber="510">C. cacah</emphasis>
</taxonomicName>
occurs within the Serra do Cipó National Park, a conservation unit. According to 
<bibRefCitation id="EFC509D2FFD5FF92FF0DF93DFE5C02AE" author="Pompeu, P. S. &amp; Alves, C. B. M. &amp; Callisto, M." box="[180,415,1677,1705]" pageId="5" pageNumber="510" pagination="11 - 22" refId="ref6184" refString="Pompeu, P. S., Alves, C. B. M. &amp; Callisto, M. (2005) The effects of urbanization on biodiversity and water quality in the Rio das Velhas basin, Brazil. American Fisheries Society Symposium, 47, 11 - 22." type="journal article" year="2005">
Pompeu 
<emphasis id="B920A831FFD5FF92FEAFF93DFE8402AE" box="[278,327,1678,1704]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
. (2005)
</bibRefCitation>
and 
<bibRefCitation id="EFC509D2FFD5FF92FE6DF93CFD0E02AE" author="Carvalho, D. R. &amp; Flecker, A. S. &amp; Alves, C. B. M. &amp; Sparks, J. P. &amp; Pompeu, P. S." box="[468,717,1678,1705]" pageId="5" pageNumber="510" pagination="503 - 515" refId="ref5356" refString="Carvalho, D. R., Flecker, A. S., Alves, C. B. M., Sparks, J. P. &amp; Pompeu, P. S. (2019) Trophic responses to aquatic pollution of native and exotic livebearer fishes. Science of The Total Environment, 681, 503 - 515. https: // doi. org / 10.1016 / j. scitotenv. 2019.05.092" type="journal article" year="2019">
Carvalho 
<emphasis id="B920A831FFD5FF92FDF8F93DFDB102AE" box="[577,626,1678,1704]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
. (2019)
</bibRefCitation>
, the rio Cipó is one of the best-preserved major tributaries of the rio das Velhas watershed. We thus consider that 
<taxonomicName id="4C540FA0FFD5FF92FD00F903FCDD02CD" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[697,798,1713,1739]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FD00F903FCDD02CD" box="[697,798,1713,1739]" italics="true" pageId="5" pageNumber="510">C. cacah</emphasis>
</taxonomicName>
could be classified as Least Concern (LC), according to the International Union for Conservation of Nature Standards and Petitions Committee (IUCN, 2019).
</paragraph>
</subSubSection>
<subSubSection id="C34E27A8FFD5FF9FFF7EF94AFE930545" lastPageId="8" lastPageNumber="513" pageId="5" pageNumber="510" type="discussion">
<paragraph id="8BEB7423FFD5FF91FF7EF94AFC820639" blockId="5.[151,1437,153,2026]" lastBlockId="6.[151,1437,153,575]" lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510">
<emphasis id="B920A831FFD5FF92FF7EF94AFEF80314" bold="true" box="[199,315,1784,1810]" pageId="5" pageNumber="510">Remarks.</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FEFFF945FDE40317" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[326,551,1783,1809]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD5FF92FEFFF945FDE40317" box="[326,551,1783,1809]" italics="true" pageId="5" pageNumber="510">Characidium cacah</emphasis>
</taxonomicName>
shares the absence of the parietal branch of the supraorbital canal, used by 
<bibRefCitation id="EFC509D2FFD5FF92FF2EF8A9FE8D0330" author="Buckup, P. A." box="[151,334,1819,1846]" pageId="5" pageNumber="510" pagination="305 - 341" refId="ref5247" refString="Buckup, P. A. (1993 b) Phylogenetic interrelationships and reductive evolution in neotropical characidiin fishes (Characiformes, Ostariophysi). Cladistics, 9 (3), 305 - 341. https: // doi. org / 10.1111 / j. 1096 - 0031.1993. tb 00227. x." type="journal article" year="1993">Buckup (1993b)</bibRefCitation>
to diagnose the Clade C4 group of species that includes 
<emphasis id="B920A831FFD5FF92FC71F8AEFB900333" box="[968,1107,1819,1846]" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C540FA0FFD5FF92FC71F8AEFB8C0333" authorityName="Almeida" authorityYear="1971" box="[968,1103,1819,1846]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="bahiense">C. bahiense</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FBE3F8AEFAAE0333" authority="Pellegrin" authorityName="Pellegrin" authorityYear="1909" box="[1114,1389,1819,1846]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="interruptum">
<emphasis id="B920A831FFD5FF92FBE3F8AEFAC20333" box="[1114,1281,1820,1846]" italics="true" pageId="5" pageNumber="510">C. interruptum</emphasis>
Pellegrin
</taxonomicName>
and 
<taxonomicName id="4C540FA0FFD5FF92FF2EF88DFE6D035F" authority="Travassos. More" authorityName="Travassos. More" authorityYear="1967" box="[151,430,1855,1881]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lanei">
<emphasis id="B920A831FFD5FF92FF2EF88DFF2C035F" box="[151,239,1855,1881]" italics="true" pageId="5" pageNumber="510">C. lanei</emphasis>
Travassos. More
</taxonomicName>
recently, the taxonomic limits of Clade C4 were expanded with the addition of 
<emphasis id="B920A831FFD5FF92FAA2F88DFF14037B" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C540FA0FFD5FF92FAA2F88DFF10037B" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="lagosantense">C. lagosantense</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FF67F8D1FE260378" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1895" box="[222,485,1891,1918]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="laterale">
<emphasis id="B920A831FFD5FF92FF67F8D1FE96037B" box="[222,341,1891,1917]" italics="true" pageId="5" pageNumber="510">C. laterale</emphasis>
(Boulenger)
</taxonomicName>
, 
<emphasis id="B920A831FFD5FF92FE56F8D1FD19037B" box="[495,730,1891,1917]" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C540FA0FFD5FF92FE56F8D1FD96037B" box="[495,597,1891,1917]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="mirim">C. mirim</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FDE6F8D1FD15037B" box="[607,726,1891,1917]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="nupelia">C. nupelia</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FD59F8D1FBF7037B" authority="Buckup &amp; Reis" authorityName="Buckup &amp; Reis" authorityYear="1997" box="[736,1076,1891,1917]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="occidentale">
<emphasis id="B920A831FFD5FF92FD59F8D1FC42037B" box="[736,897,1891,1917]" italics="true" pageId="5" pageNumber="510">C. occidentale</emphasis>
Buckup &amp; Reis
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FB86F8D1FAB5037B" authority="Buckup &amp; Reis" authorityName="Buckup &amp; Reis" authorityYear="1997" box="[1087,1398,1891,1917]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="orientale">
<emphasis id="B920A831FFD5FF92FB86F8D1FB07037B" box="[1087,1220,1891,1917]" italics="true" pageId="5" pageNumber="510">C. orientale</emphasis>
Buckup &amp; Reis
</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD5FF92FA3AF8D1FE8003A7" authority="Regan" authorityName="Regan" authorityYear="1913" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="rachovii">
<emphasis id="B920A831FFD5FF92FA3AF8D1FF3703A7" italics="true" pageId="5" pageNumber="510">C. rachovii</emphasis>
Regan
</taxonomicName>
, 
<emphasis id="B920A831FFD5FF92FEE9F835FE2C03A7" box="[336,495,1927,1953]" italics="true" pageId="5" pageNumber="510">
<taxonomicName id="4C540FA0FFD5FF92FEE9F835FE2F03A7" authorityName="Melo &amp; Buckup" authorityYear="2002" box="[336,492,1927,1953]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="stigmosum">C. stigmosum</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD5FF92FE4FF835FC9D03A7" authority="Buckup &amp; Hahn" authorityName="Buckup &amp; Hahn" authorityYear="2000" box="[502,862,1927,1953]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="vestigipinne">
<emphasis id="B920A831FFD5FF92FE4FF835FD5E03A7" box="[502,669,1927,1953]" italics="true" pageId="5" pageNumber="510">C. vestigipinne</emphasis>
Buckup &amp; Hahn
</taxonomicName>
, and 
<taxonomicName id="4C540FA0FFD5FF92FC21F835FBCC03A7" box="[920,1039,1927,1953]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="510" phylum="Chordata" rank="species" species="xavante">
<emphasis id="B920A831FFD5FF92FC21F835FBCC03A7" box="[920,1039,1927,1953]" italics="true" pageId="5" pageNumber="510">C. xavante</emphasis>
</taxonomicName>
, and the putative monophyly of the clade supported also by the presence of more than 12 bars on body and the presence of a single row of dentary teeth (
<bibRefCitation id="EFC509D2FFD5FF92FF27F87DFE0903EF" author="Netto-Ferreira, A. L. &amp; Birindelli, J. L. O. &amp; Buckup, P. A." box="[158,458,1999,2025]" pageId="5" pageNumber="510" pagination="361 - 368" refId="ref6112" refString="Netto-Ferreira, A. L., Birindelli, J. L. O. &amp; Buckup, P. A. (2013) A new miniature species of Characidium Reinhardt (Ostariophysi: Characiformes: Crenuchidae) from the headwaters of the rio Araguaia, Brazil. Zootaxa, 3664 (3), 361 - 368. https: // doi. org / 10.11646 / zootaxa. 3664.3.6" type="journal article" year="2013">
Netto-Ferreira 
<emphasis id="B920A831FFD5FF92FEF1F862FEBA03EF" box="[328,377,1999,2025]" italics="true" pageId="5" pageNumber="510">et al</emphasis>
., 2013
</bibRefCitation>
). However, according to 
<bibRefCitation id="EFC509D2FFD5FF92FD5BF87DFBC603EC" author="Melo, M. R. S. &amp; Espindola, V. C." box="[738,1029,1999,2026]" pageId="5" pageNumber="510" pagination="552 - 568" refId="ref6039" refString="Melo, M. R. S. &amp; Espindola, V. C. (2016) Description of a new species of Characidium Reinhardt, 1867 (Characiformes: Crenuchidae) from the Chapada Diamantina, Bahia, and redescription of Characidium bimaculatum Fowler, 1941. Zootaxa, 4196 (4), 552 - 568. https: // doi. org / 10.11646 / zootaxa. 4196.4.5." type="journal article" year="2016">Melo &amp; Espíndola (2016)</bibRefCitation>
, these characters were not evaluated correctly in some of the species included in the group and are not useful to diagnose the clade C4 as recently proposed. At any rate, 
<taxonomicName id="4C540FA0FFD6FF91FECAFF0FFE1B04D1" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[371,472,189,215]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD6FF91FECAFF0FFE1B04D1" box="[371,472,189,215]" italics="true" pageId="6" pageNumber="511">C. cacah</emphasis>
</taxonomicName>
shares the three features proposed up to date to circumscribe the Clade C4 and other reductive characters with subgroups of the cited species. The new species shares an incomplete lateral line, the absence of the adipose fin, and the absence of the parietal branch of the supraorbital canal exclusively with 
<emphasis id="B920A831FFD6FF91FA89FEB7FEB30545" italics="true" pageId="6" pageNumber="511">
<taxonomicName id="4C540FA0FFD6FF91FA89FEB7FA5B0519" box="[1328,1432,261,287]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="chicoi">C. chicoi</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD6FF91FF2EFE9BFF310545" authorityName="Mendonca &amp; Netto-Ferreira" authorityYear="2015" box="[151,242,297,323]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="nana">C. nana</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD6FF91FF45FE9BFEB30545" box="[252,368,297,323]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="nupelia">C. nupelia</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C540FA0FFD6FF91FEC7FE9BFDD50545" authorityName="Melo &amp; Buckup" authorityYear="2002" box="[382,534,297,323]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="stigmosum">
<emphasis id="B920A831FFD6FF91FEC7FE9BFDD50545" box="[382,534,297,323]" italics="true" pageId="6" pageNumber="511">C. stigmosum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C540FA0FFD6FF91FDE8FE9BFD0B0545" box="[593,712,297,323]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="xavante">
<emphasis id="B920A831FFD6FF91FDE8FE9BFD0B0545" box="[593,712,297,323]" italics="true" pageId="6" pageNumber="511">C. xavante</emphasis>
</taxonomicName>
, and shares the presence of only four branchiostegal rays with 
<emphasis id="B920A831FFD6FF91FA3AFE9BFD520561" italics="true" pageId="6" pageNumber="511">
<taxonomicName id="4C540FA0FFD6FF91FA3AFE9BFF3E0561" authorityName="Almeida" authorityYear="1971" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="bahiense">C. bahiense</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD6FF91FEB0FEFFFEA70561" authorityName="Mendonca &amp; Netto-Ferreira" authorityYear="2015" box="[265,356,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="nana">C. nana</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD6FF91FED6FEFFFE240561" box="[367,487,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="nupelia">C. nupelia</taxonomicName>
, 
<taxonomicName id="4C540FA0FFD6FF91FE4BFEFFFD4D0561" authorityName="Melo &amp; Buckup" authorityYear="2002" box="[498,654,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="stigmosum">C. stigmosum</taxonomicName>
,
</emphasis>
<taxonomicName id="4C540FA0FFD6FF91FD21FEFFFC0D0561" authority="Zanata &amp; Ohara" authorityName="Zanata &amp; Ohara" authorityYear="2015" box="[664,974,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="summus">
<emphasis id="B920A831FFD6FF91FD21FEFFFCD00561" box="[664,787,333,359]" italics="true" pageId="6" pageNumber="511">C. summus</emphasis>
Zanata &amp; Ohara
</taxonomicName>
and 
<taxonomicName id="4C540FA0FFD6FF91FBBCFEFFFBBE0561" box="[1029,1149,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="xavante">
<emphasis id="B920A831FFD6FF91FBBCFEFFFBBE0561" box="[1029,1149,333,359]" italics="true" pageId="6" pageNumber="511">C. xavante</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C540FA0FFD6FF91FB32FEFFFAAB0561" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[1163,1384,333,359]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD6FF91FB32FEFFFAAB0561" box="[1163,1384,333,359]" italics="true" pageId="6" pageNumber="511">Characidium cacah</emphasis>
</taxonomicName>
also shares with species from the C4 clade and some other congeners an overall shape of body and fins related to species dwelling in slow to moderate water current, and does not possess adaptations of congeners that inhabit fast water current environments, as a streamlined body and paired-fins modifications, as discussed by several authors (
<emphasis id="B920A831FFD6FF91FAD4FE08FA4D05D5" box="[1389,1422,442,467]" italics="true" pageId="6" pageNumber="511">e.g</emphasis>
., 
<bibRefCitation id="EFC509D2FFD6FF91FF2EFE6FFEB605F1" author="Buckup, P. A. &amp; Zamprogno, C. &amp; Vieira, F. &amp; Teixeira, R. L." box="[151,373,477,503]" pageId="6" pageNumber="511" pagination="273 - 278" refId="ref5303" refString="Buckup, P. A., Zamprogno, C., Vieira, F. &amp; Teixeira, R. L. (2000) Waterfall climbing in Characidium (Crenuchidae, Characidiinae) from eastern Brazil. Ichthyological Exploration of Freshwaters, 11 (3), 273 - 278." type="journal article" year="2000">
Buckup 
<emphasis id="B920A831FFD6FF91FF4DFE6CFEE605F1" box="[244,293,477,503]" italics="true" pageId="6" pageNumber="511">et al</emphasis>
., 2000
</bibRefCitation>
; 
<bibRefCitation id="EFC509D2FFD6FF91FEC7FE6FFDBA05F1" author="Zanata, A. M. &amp; Ohara, W. M." box="[382,633,476,503]" pageId="6" pageNumber="511" pagination="368 - 376" refId="ref6428" refString="Zanata, A. M. &amp; Ohara, W. M. (2015) A new species of Characidium Reinhardt (Ostariophysi: Characiformes: Crenuchidae) from headwaters of rio Pacaas Novos, rio Madeira basin, Rondonia, Brazil. Zootaxa, 4021 (2), 368 - 376. https: // doi. org / 10.11646 / zootaxa. 4021.2.7" type="journal article" year="2015">Zanata &amp; Ohara, 2015</bibRefCitation>
; Zanata 
<emphasis id="B920A831FFD6FF91FD6FFE6CFCC505F1" box="[726,774,477,503]" italics="true" pageId="6" pageNumber="511">et al</emphasis>
., 2015; 
<bibRefCitation id="EFC509D2FFD6FF91FCD9FE6FFBF005F1" author="Zanata, A. M. &amp; Ramos, T. P. A. &amp; Oliveira-Silva, L." box="[864,1075,477,503]" pageId="6" pageNumber="511" pagination="77 - 86" refId="ref6495" refString="Zanata, A. M., Ramos, T. P. A. &amp; Oliveira-Silva, L. (2018) A new species of Characidium (Characiformes: Crenuchidae) from the rio Parnaiba basin, northeastern Brazil. Zootaxa, 4514 (1), 77 - 86. https: // doi. org / 10.11646 / zootaxa. 4514.1.6" type="journal article" year="2018">
Zanata 
<emphasis id="B920A831FFD6FF91FC0BFE6CFC2005F1" box="[946,995,477,503]" italics="true" pageId="6" pageNumber="511">et al</emphasis>
., 2018
</bibRefCitation>
). A comprehensive phylogenetic study of 
<taxonomicName id="4C540FA0FFD6FF91FF41FDB3FE4F061D" authorityName="Reinhardt" authorityYear="1867" box="[248,396,513,539]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="genus">
<emphasis id="B920A831FFD6FF91FF41FDB3FE4F061D" box="[248,396,513,539]" italics="true" pageId="6" pageNumber="511">Characidium</emphasis>
</taxonomicName>
is necessary to properly evaluate the evolution of the morphological traits shared by 
<taxonomicName id="4C540FA0FFD6FF91FA80FDB3FA5F061D" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[1337,1436,513,539]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD6FF91FA80FDB3FA5F061D" box="[1337,1436,513,539]" italics="true" pageId="6" pageNumber="511">C. cacah</emphasis>
</taxonomicName>
with congeners and to define the relationships of the species.
</paragraph>
<caption id="DF2B24ABFFD6FF91FF2EF816FD3603E7" ID-DOI="http://doi.org/10.5281/zenodo.3891285" ID-Zenodo-Dep="3891285" httpUri="https://zenodo.org/record/3891285/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,250,1956,1981]" targetBox="[141,1445,603,1924]" targetPageId="6">
<paragraph id="8BEB7423FFD6FF91FF2EF816FD3603E7" blockId="6.[151,1436,1956,2017]" pageId="6" pageNumber="511">
<emphasis id="B920A831FFD6FF91FF2EF816FED703BB" bold="true" box="[151,276,1956,1981]" pageId="6" pageNumber="511">FIGURE 3.</emphasis>
Map of the rio das Velhas sub-basin, rio São Francisco basin, showing the collection sites of 
<taxonomicName id="4C540FA0FFD6FF91FB6AF816FA5F03BA" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[1235,1436,1956,1980]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="511" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD6FF91FB6AF816FA5F03BA" box="[1235,1436,1956,1980]" italics="true" pageId="6" pageNumber="511">Characidium cacah</emphasis>
</taxonomicName>
(red star); symbol represents more than one sampling point.
</paragraph>
</caption>
<caption id="DF2B24ABFFD7FF90FF2EF837FB5203E0" ID-DOI="http://doi.org/10.5281/zenodo.3891287" ID-Zenodo-Dep="3891287" httpUri="https://zenodo.org/record/3891287/files/figure.png" pageId="7" pageNumber="512" startId="7.[151,250,1925,1950]" targetBox="[188,1400,181,1900]" targetPageId="7">
<paragraph id="8BEB7423FFD7FF90FF2EF837FB5203E0" blockId="7.[151,1436,1925,2022]" pageId="7" pageNumber="512">
<emphasis id="B920A831FFD7FF90FF2EF837FED00398" bold="true" box="[151,275,1925,1950]" pageId="7" pageNumber="512">FIGURE 4.</emphasis>
Sampling localities of 
<taxonomicName id="4C540FA0FFD7FF90FE47F834FD050398" authorityName="Zanata &amp; Ribeiro &amp; Araújo-Porto &amp; Pessali &amp; Oliveira-Silva" authorityYear="2020" box="[510,710,1926,1950]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="512" phylum="Chordata" rank="species" species="cacah">
<emphasis id="B920A831FFD7FF90FE47F834FD050398" box="[510,710,1926,1950]" italics="true" pageId="7" pageNumber="512">Characidium cacah</emphasis>
</taxonomicName>
, Brazil, Minas Gerais State, rio São Francisco basin, rio das Velhas sub basin, rio Cipó drainage: (a) satellite image of a stretch of rio Cipó and surrounding lagoons (from Google Earth); (b) rio Cipó just after the meeting of rio Bocaina and rio Mascates; (c) rio Cipó at type locality; d) Lagoa do Boi.
</paragraph>
</caption>
<paragraph id="8BEB7423FFD8FF9FFF7EFF2AFE930545" blockId="8.[151,1436,152,323]" pageId="8" pageNumber="513">
<emphasis id="B920A831FFD8FF9FFF7EFF2AFC3304B5" bold="true" box="[199,1008,152,179]" pageId="8" pageNumber="513">
Comparative material examined (additional to 
<bibRefCitation id="EFC509D2FFD8FF9FFD46FF2BFC2104B5" author="Zanata, A. M. &amp; Ramos, T. P. A. &amp; Oliveira-Silva, L." box="[767,994,152,179]" pageId="8" pageNumber="513" pagination="77 - 86" refId="ref6495" refString="Zanata, A. M., Ramos, T. P. A. &amp; Oliveira-Silva, L. (2018) A new species of Characidium (Characiformes: Crenuchidae) from the rio Parnaiba basin, northeastern Brazil. Zootaxa, 4514 (1), 77 - 86. https: // doi. org / 10.11646 / zootaxa. 4514.1.6" type="journal article" year="2018">
Zanata 
<emphasis id="B920A831FFD8FF9FFCE4FF2BFC4C04B5" bold="true" box="[861,911,152,179]" italics="true" pageId="8" pageNumber="513">et al</emphasis>
., 2018
</bibRefCitation>
):
</emphasis>
<taxonomicName id="4C540FA0FFD8FF9FFC41FF2BFB1404B5" box="[1016,1239,153,179]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="513" phylum="Chordata" rank="species" species="mirim">
<emphasis id="B920A831FFD8FF9FFC41FF2BFB1404B5" box="[1016,1239,153,179]" italics="true" pageId="8" pageNumber="513">Characidium mirim</emphasis>
</taxonomicName>
: 
<materialsCitation id="3B3C7E7EFFD8FF9FFB5EFF2BFB8904D1" collectingDate="23-11-11" collectionCode="MZUSP" collectorName="das Mortes." country="Brazil" location="Mato Grosso" pageId="8" pageNumber="513" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<collectionCode id="ED45ECE6FFD8FF9FFB5EFF2BFA8004B5" box="[1255,1347,153,179]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="513">MZUSP</collectionCode>
<date id="FFEA52E3FFD8FF9FFAF2FF2BFA5F04B5" box="[1355,1436,153,179]" pageId="8" pageNumber="513" value="23-11-11">
<collectingDate id="EFAEAB0BFFD8FF9FFAF2FF2BFA5F04B5" box="[1355,1436,153,179]" pageId="8" pageNumber="513" value="23-11-11">111123</collectingDate>
</date>
(
<typeStatus id="54EFCA81FFD8FF9FFF26FF0FFEC604D1" box="[159,261,189,215]" pageId="8" pageNumber="513" type="holotype">holotype</typeStatus>
, 
<quantity id="4CACD9C6FFD8FF9FFEADFF0FFEB804D1" box="[276,379,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="8" pageNumber="513" unit="mm" value="20.2">20.2 mm</quantity>
SL), 
<collectingCountry id="F34334B3FFD8FF9FFE79FF0FFDC404D1" box="[448,519,189,215]" name="Brazil" pageId="8" pageNumber="513">Brazil</collectingCountry>
, 
<collectingRegion id="4990BAC1FFD8FF9FFDAFFF0FFD6D04D1" box="[534,686,189,215]" country="Brazil" name="Mato Grosso" pageId="8" pageNumber="513">Mato Grosso</collectingRegion>
, rio 
<collectorName id="26A111F5FFD8FF9FFD5FFF0FFCAF04D1" box="[742,876,189,215]" pageId="8" pageNumber="513">das Mortes.</collectorName>
<taxonomicName id="4C540FA0FFD8FF9FFCCFFF0FFB8904D1" authorityName="Melo &amp; Oyakawa" authorityYear="2015" box="[886,1098,189,215]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="513" phylum="Chordata" rank="species" species="satoi">
<emphasis id="B920A831FFD8FF9FFCCFFF0FFB8904D1" box="[886,1098,189,215]" italics="true" pageId="8" pageNumber="513">Characidium satoi</emphasis>
</taxonomicName>
</materialsCitation>
: 
<materialsCitation id="3B3C7E7EFFD8FF9FFBE4FF0FFE070519" collectionCode="MZUSP" collectorName="Corrego Curral das Eguas" country="Brazil" location="rio Sao Francisco" municipality="rio Abaete" pageId="8" pageNumber="513" specimenCode="MZUSP 115059" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD8FF9FFBE4FF0FFAD504D1" box="[1117,1302,188,215]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="513">MZUSP 115059</specimenCode>
(
<typeStatus id="54EFCA81FFD8FF9FFA91FF0FFA5B04D1" box="[1320,1432,189,215]" pageId="8" pageNumber="513" type="paratype">paratypes</typeStatus>
, 17, 
<quantity id="4CACD9C6FFD8FF9FFF7BFF53FEA604FD" box="[194,357,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="3.525" metricValueMax="4.43" metricValueMin="2.62" pageId="8" pageNumber="513" unit="mm" value="35.25" valueMax="44.3" valueMin="26.2">26.2–44.3 mm</quantity>
SL), 
<collectingCountry id="F34334B3FFD8FF9FFE1CFF53FE2F04FD" box="[421,492,225,251]" name="Brazil" pageId="8" pageNumber="513">Brazil</collectingCountry>
, 
<collectingRegion id="4990BAC1FFD8FF9FFE4EFF53FD5204FD" box="[503,657,225,251]" country="Brazil" name="Minas Gerais" pageId="8" pageNumber="513">Minas Gerais</collectingRegion>
, 
<collectorName id="26A111F5FFD8FF9FFD25FF53FC0104FD" box="[668,962,224,251]" pageId="8" pageNumber="513">Córrego Curral das Éguas</collectorName>
, tributary of 
<collectingMunicipality id="6B8FEE59FFD8FF9FFBECFF53FB0F04FD" box="[1109,1228,225,251]" pageId="8" pageNumber="513">rio Abaeté</collectingMunicipality>
, 
<location id="8E8B22F8FFD8FF9FFB6EFF53FA5F04FD" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD8FF9FFB6EFF53FA5F04FD" box="[1239,1436,225,251]" country="Brazil" municipality="rio Abaete" name="rio Sao Francisco" pageId="8" pageNumber="513" stateProvince="Minas Gerais">rio São Francisco</location>
basin. 
<taxonomicName id="4C540FA0FFD8FF9FFF58FEB7FE070519" authorityName="Zanata, Ramos &amp; Oliveira-Silva" authorityYear="2018" box="[225,452,261,287]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="513" phylum="Chordata" rank="species" species="tapuia">
<emphasis id="B920A831FFD8FF9FFF58FEB7FE070519" box="[225,452,261,287]" italics="true" pageId="8" pageNumber="513">Characidium tapuia</emphasis>
</taxonomicName>
</materialsCitation>
: 
<materialsCitation id="3B3C7E7EFFD8FF9FFE6AFEB7FE8F0545" collectionCode="UFBA" country="Brazil" location="rio Balsas" pageId="8" pageNumber="513" specimenCode="UFBA 8511" specimenCount="1" stateProvince="Maranhao" typeStatus="paratype">
<specimenCode id="DBF2DC58FFD8FF9FFE6AFEB7FD980519" box="[467,603,260,287]" collectionCode="UFBA" pageId="8" pageNumber="513">UFBA 8511</specimenCode>
(
<typeStatus id="54EFCA81FFD8FF9FFDD3FEB7FD190519" box="[618,730,261,287]" pageId="8" pageNumber="513" type="paratype">paratypes</typeStatus>
, 3, 
<quantity id="4CACD9C6FFD8FF9FFCBBFEB7FC660519" box="[770,933,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.095" metricValueMax="3.21" metricValueMin="2.98" pageId="8" pageNumber="513" unit="mm" value="30.95" valueMax="32.1" valueMin="29.8">29.8–32.1 mm</quantity>
SL, 1 c&amp;s, 
<quantity id="4CACD9C6FFD8FF9FFB94FEB7FB520519" box="[1069,1169,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" pageId="8" pageNumber="513" unit="mm" value="30.8">30.8 mm</quantity>
SL), 
<collectingCountry id="F34334B3FFD8FF9FFB68FEB7FADB0519" box="[1233,1304,261,287]" name="Brazil" pageId="8" pageNumber="513">Brazil</collectingCountry>
, 
<collectingRegion id="4990BAC1FFD8FF9FFA9DFEB7FA5B0519" box="[1316,1432,261,287]" country="Brazil" name="Maranhao" pageId="8" pageNumber="513">Maranhão</collectingRegion>
, 
<location id="8E8B22F8FFD8FF9FFF2EFE9BFEC50545" LSID="urn:lsid:plazi:treatment:03FDC535FFD1FF9FFF2EFB1FFE930545:8E8B22F8FFD8FF9FFF2EFE9BFEC50545" box="[151,262,297,323]" country="Brazil" name="rio Balsas" pageId="8" pageNumber="513" stateProvince="Maranhao">rio Balsas</location>
basin
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>