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<document ID-DOI="10.15560/15.4.651" ID-ISSN="1809-127X" approvalRequired="239" approvalRequired_for_taxonomicNames="94" approvalRequired_for_textStreams="143" approvalRequired_for_treatments="2" checkinTime="1630357287411" checkinUser="marcus" docAuthor="Ortiz, Orlando Oriel, de Stapf, María Sánchez, Baldini, Riccardo Maria &amp; Croat, Thomas Bernard" docDate="2019" docId="03F487E39D6EFFDFFC87F8B530BDFC30" docLanguage="en" docName="CheckList.2019.15.4.651-689.pdf" docOrigin="Check List 15 (4)" docSource="http://dx.doi.org/10.15560/15.4.651" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Anthurium scandens subsp. pusillum R. Sheffer" docType="treatment" docVersion="1" lastPageId="20" lastPageNumber="671" masterDocId="FFCDFF9B9D7CFFCBFFAAFFEE313BFFD1" masterDocTitle="Synopsis of aroids (Alismatales, Araceae) from Cerro Pirre (Darién Province, Panama)" masterLastPageNumber="689" masterPageNumber="651" pageId="18" pageNumber="669" updateTime="1630357298930" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synopsis of aroids (Alismatales, Araceae) from Cerro Pirre (Darién Province, Panama)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortiz, Orlando Oriel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Stapf, María Sánchez</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baldini, Riccardo Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Croat, Thomas Bernard</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>15</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>651</mods:start>
<mods:end>689</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/15.4.651</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/15.4.651</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F487E39D6EFFDFFC87F8B530BDFC30" httpUri="http://treatment.plazi.org/id/03F487E39D6EFFDFFC87F8B530BDFC30" lastPageId="20" lastPageNumber="671" pageId="18" pageNumber="669">
<subSubSection box="[813,1420,1883,1908]" pageId="18" pageNumber="669" type="nomenclature">
<paragraph blockId="18.[813,1420,1883,1908]" box="[813,1420,1883,1908]" pageId="18" pageNumber="669">
<heading box="[813,1420,1883,1908]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="669" reason="2">
<emphasis box="[813,1420,1883,1908]" inLineHeading="true" pageId="18" pageNumber="669" reason="1">
<taxonomicName authority="R. Sheffer" authorityName="R. Sheffer" box="[813,1321,1883,1908]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="18" pageNumber="669" phylum="Tracheophyta" rank="subSpecies" species="scandens" subSpecies="pusillum">
<emphasis bold="true" box="[813,1039,1883,1907]" italics="true" pageId="18" pageNumber="669">Anthurium scandens</emphasis>
subsp.
<emphasis bold="true" box="[1115,1210,1883,1907]" italics="true" pageId="18" pageNumber="669">pusillum</emphasis>
R. Sheffer
</taxonomicName>
(
<figureCitation box="[1334,1412,1883,1908]" captionStart="Figure 5" captionStartId="19.[168,231,1904,1924]" captionTargetBox="[168,1419,215,1879]" captionTargetId="figure-6@19.[168,1419,215,1879]" captionTargetPageId="19" captionText="Figure 5. Some aroids from Cerro Pirre. A. Anthurium rubrifructum (inflorescence). B. Anthurium salvinii (inflorescence). C. Anthurium scandens (infructescence). D. Anthurium talamancae (inflorescence). E. Anthurium terryae (infructescence). F. Anthurium terryae (inflorescence). G. Anthurium tonduzii (inflorescence). H. Anthurium trilobum (inflorescence). I. Anthurium sp. 2 (infructescence)." pageId="18" pageNumber="669">Fig. 5C</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="671" pageId="18" pageNumber="669" type="materials_examined">
<paragraph blockId="18.[812,1419,1933,2091]" lastBlockId="20.[167,777,160,320]" lastPageId="20" lastPageNumber="671" pageId="18" pageNumber="669">
<emphasis bold="true" box="[813,1046,1933,1957]" pageId="18" pageNumber="669">Material examined.</emphasis>
Parque Nacional
<collectingRegion box="[1261,1338,1933,1957]" country="Panama" name="Darien" pageId="18" pageNumber="669">Darién</collectingRegion>
, Cerro Pirre, Rancho Plástico;
<geoCoordinate box="[1059,1188,1965,1991]" degrees="07" direction="north" minutes="59" orientation="latitude" pageId="18" pageNumber="669" precision="15" seconds="43" value="7.995278">07°5943″N</geoCoordinate>
,
<geoCoordinate box="[1194,1339,1965,1991]" degrees="077" direction="west" minutes="42" orientation="longitude" pageId="18" pageNumber="669" precision="15" seconds="39" value="-77.71083">077°4239″W</geoCoordinate>
;
<quantity box="[1347,1415,1967,1991]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="669" unit="m" value="700.0">700 m</quantity>
;
<date box="[813,941,2000,2024]" pageId="18" pageNumber="669" value="2016-07-18">18 Jul. 2016</date>
;
<emphasis box="[951,1094,2000,2024]" italics="true" pageId="18" pageNumber="669">
<collectionCode box="[951,963,2001,2024]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="18" pageNumber="669" type="Herbarium">J</collectionCode>
Aranda s.n.
</emphasis>
(
<collectionCode box="[1108,1171,2000,2024]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="18" pageNumber="669">PMA</collectionCode>
). Cerro Pirre;
<geoCoordinate box="[1324,1415,1999,2024]" degrees="07" direction="north" minutes="52" orientation="latitude" pageId="18" pageNumber="669" precision="925" value="7.866667">07°52N</geoCoordinate>
,
<geoCoordinate box="[812,922,2032,2058]" degrees="077" direction="west" minutes="44" orientation="longitude" pageId="18" pageNumber="669" precision="925" value="-77.73333">077°44W</geoCoordinate>
;
<date box="[937,1080,2033,2058]" pageId="18" pageNumber="669" value="1967-08-04">4 Aug. 1967</date>
;
<emphasis box="[1096,1273,2034,2058]" italics="true" pageId="18" pageNumber="669">
<collectionCode box="[1096,1117,2034,2057]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="18" pageNumber="669" type="Herbarium">N</collectionCode>
. Bristán 1242
</emphasis>
(
<collectionCode box="[1293,1340,2034,2058]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="18" pageNumber="669" type="Herbarium">MO</collectionCode>
). Ibid., cloud forest and/or mossy forest;
<geoCoordinate box="[1198,1290,2065,2090]" degrees="07" direction="north" minutes="50" orientation="latitude" pageId="18" pageNumber="669" precision="925" value="7.8333335">07°50N</geoCoordinate>
,
<geoCoordinate box="[1306,1416,2065,2090]" degrees="077" direction="west" minutes="44" orientation="longitude" pageId="18" pageNumber="669" precision="925" value="-77.73333">077°44W</geoCoordinate>
;
<quantity box="[168,308,160,184]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.3" metricValueMin="0.75" pageId="20" pageNumber="671" unit="m" value="1025.0" valueMax="1300.0" valueMin="750.0">7501300 m</quantity>
;
<date box="[322,502,160,185]" pageId="20" pageNumber="671" value="1967-08-09" valueMax="1967-08-10" valueMin="1967-08-09">910 Aug. 1967</date>
;
<emphasis box="[516,698,160,184]" italics="true" pageId="20" pageNumber="671">
<collectionCode box="[516,532,161,184]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="20" pageNumber="671" type="Herbarium">J</collectionCode>
.
<collectionCode box="[535,554,161,184]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="20" pageNumber="671" type="Herbarium">A</collectionCode>
. Duke 13701
</emphasis>
(
<collectionCode box="[717,765,160,184]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="20" pageNumber="671" type="Herbarium">MO</collectionCode>
). Serranía Pirre, on main ridge from intersection with trail down to Rancho Frío to
<quantity box="[427,501,228,252]" metricMagnitude="3" metricUnit="m" metricValue="2.414016" pageId="20" pageNumber="671" unit="mi" value="1.5">1.5 mi.</quantity>
<collectionCode box="[507,526,228,251]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="20" pageNumber="671" type="Herbarium">N</collectionCode>
of that point;
<geoCoordinate box="[676,769,227,252]" degrees="08" direction="north" minutes="00" orientation="latitude" pageId="20" pageNumber="671" precision="925" value="8.0">08°00N</geoCoordinate>
,
<geoCoordinate box="[168,275,260,285]" degrees="077" direction="west" minutes="43" orientation="longitude" pageId="20" pageNumber="671" precision="925" value="-77.71667">077°43W</geoCoordinate>
;
<quantity box="[286,413,262,286]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="20" pageNumber="671" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
;
<date box="[424,556,262,286]" pageId="20" pageNumber="671" value="1977-07-12">12 Jul. 1977</date>
;
<emphasis box="[566,775,262,286]" italics="true" pageId="20" pageNumber="671">
<collectionCode box="[566,586,262,285]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="671">R</collectionCode>
.
<collectionCode box="[590,609,262,285]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="20" pageNumber="671" type="Herbarium">L</collectionCode>
. Hartman 4547
</emphasis>
(
<collectionCode box="[176,223,296,320]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="20" pageNumber="671" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[233,294,296,320]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="20" pageNumber="671">PMA</collectionCode>
).
</paragraph>
<caption pageId="19" pageNumber="670" startId="19.[168,231,1904,1924]" targetBox="[168,1419,215,1879]" targetPageId="19">
<paragraph blockId="19.[168,1421,1904,1983]" pageId="19" pageNumber="670">
<emphasis bold="true" box="[168,253,1904,1924]" pageId="19" pageNumber="670">Figure 5.</emphasis>
Some aroids from Cerro Pirre.
<emphasis bold="true" box="[531,552,1904,1924]" pageId="19" pageNumber="670">A.</emphasis>
<taxonomicName box="[556,768,1904,1924]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="rubrifructum">
<emphasis box="[556,768,1904,1924]" italics="true" pageId="19" pageNumber="670">Anthurium rubrifructum</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[916,935,1904,1924]" pageId="19" pageNumber="670">B.</emphasis>
<taxonomicName box="[940,1101,1904,1924]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="salvinii">
<emphasis box="[940,1101,1904,1924]" italics="true" pageId="19" pageNumber="670">Anthurium salvinii</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[1249,1268,1904,1924]" pageId="19" pageNumber="670">C.</emphasis>
<taxonomicName class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="scandens">
<emphasis italics="true" pageId="19" pageNumber="670">Anthurium scandens</emphasis>
</taxonomicName>
(infructescence).
<emphasis bold="true" box="[370,390,1933,1953]" pageId="19" pageNumber="670">D.</emphasis>
<taxonomicName box="[395,600,1933,1953]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="talamancae">
<emphasis box="[395,600,1933,1953]" italics="true" pageId="19" pageNumber="670">Anthurium talamancae</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[748,766,1933,1953]" pageId="19" pageNumber="670">E.</emphasis>
<taxonomicName box="[771,932,1933,1953]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="terryae">
<emphasis box="[771,932,1933,1953]" italics="true" pageId="19" pageNumber="670">Anthurium terryae</emphasis>
</taxonomicName>
(infructescence).
<emphasis bold="true" box="[1094,1110,1933,1953]" pageId="19" pageNumber="670">F.</emphasis>
<taxonomicName box="[1114,1276,1933,1953]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="terryae">
<emphasis box="[1114,1276,1933,1953]" italics="true" pageId="19" pageNumber="670">Anthurium terryae</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[168,189,1963,1983]" pageId="19" pageNumber="670">G.</emphasis>
<taxonomicName box="[194,365,1963,1983]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis box="[194,365,1963,1983]" italics="true" pageId="19" pageNumber="670">Anthurium tonduzii</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[514,534,1963,1983]" pageId="19" pageNumber="670">H.</emphasis>
<taxonomicName box="[539,715,1963,1983]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="trilobum">
<emphasis box="[539,715,1963,1983]" italics="true" pageId="19" pageNumber="670">Anthurium trilobum</emphasis>
</taxonomicName>
(inflorescence).
<emphasis bold="true" box="[864,876,1963,1983]" pageId="19" pageNumber="670">I.</emphasis>
<taxonomicName box="[881,1022,1963,1983]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="19" pageNumber="670" phylum="Tracheophyta" rank="species" species="undefined-2">
<emphasis box="[881,976,1963,1983]" italics="true" pageId="19" pageNumber="670">Anthurium</emphasis>
sp. 2
</taxonomicName>
(infructescence).
</paragraph>
</caption>
<paragraph blockId="20.[167,775,345,741]" pageId="20" pageNumber="671">
<emphasis bold="true" box="[168,330,345,369]" pageId="20" pageNumber="671">Identification.</emphasis>
This taxon is characterized by having an epiphytic-climbing habit, short stems and internodes, fibrous-persistent cataphylls, black glandular-punctate blades, usually pendent inflorescences, small spathes and spadices (&lt;1.0 cm long) and whitish berries.
<taxonomicName class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="20" pageNumber="671" phylum="Tracheophyta" rank="subSpecies" species="scandens" subSpecies="pusillum">
<emphasis italics="true" pageId="20" pageNumber="671">Anthurium scandens</emphasis>
subsp.
<emphasis box="[409,502,514,538]" italics="true" pageId="20" pageNumber="671">pusillum</emphasis>
</taxonomicName>
differs from the typical subspecies (
<taxonomicName authority="subsp. scandens" box="[298,606,548,572]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="20" pageNumber="671" phylum="Tracheophyta" rank="subSpecies" species="scandens" subSpecies="scandens">
<emphasis box="[298,428,548,572]" italics="true" pageId="20" pageNumber="671">A. scandens</emphasis>
subsp.
<emphasis box="[508,606,548,572]" italics="true" pageId="20" pageNumber="671">scandens</emphasis>
</taxonomicName>
) in having spadices with fewer flowers (&lt;13 flowers per spadix), small leaves, and very small spadices (up to 1.0 cm long). In the study area it could be confused with
<taxonomicName authority="Engl." authorityName="Engl." box="[582,771,649,673]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="20" pageNumber="671" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis box="[582,703,649,673]" italics="true" pageId="20" pageNumber="671">A. tonduzii</emphasis>
Engl.
</taxonomicName>
, but the latter differs in having intact and deciduous cata- phylls, markedly ovate blades and reddish spadices.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="671" type="distribution">
<paragraph blockId="20.[168,775,767,994]" pageId="20" pageNumber="671">
<emphasis bold="true" box="[168,457,767,791]" pageId="20" pageNumber="671">Distribution and ecology.</emphasis>
<collectingCountry box="[462,570,767,791]" name="Colombia" pageId="20" pageNumber="671">Colombia</collectingCountry>
,
<collectingCountry box="[580,701,767,791]" name="Costa Rica" pageId="20" pageNumber="671">Costa Rica</collectingCountry>
,
<collectingCountry name="Ecuador" pageId="20" pageNumber="671">Ecuador</collectingCountry>
,
<collectingCountry box="[215,323,801,825]" name="Honduras" pageId="20" pageNumber="671">Honduras</collectingCountry>
,
<collectingCountry box="[333,448,801,825]" name="Nicaragua" pageId="20" pageNumber="671">Nicaragua</collectingCountry>
,
<collectingCountry box="[458,547,801,824]" name="Panama" pageId="20" pageNumber="671">Panama</collectingCountry>
, and
<collectingCountry box="[602,717,801,825]" name="Venezuela" pageId="20" pageNumber="671">Venezuela</collectingCountry>
. This taxon generally ocurrs in montane and submontane forests, above
<quantity box="[294,363,868,892]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="671" unit="m" value="600.0">600 m</quantity>
(
<bibRefCitation author="Croat TB" box="[377,513,868,892]" pageId="20" pageNumber="671" refId="ref39425" refString="Croat TB (1986 b) A revision of genus Anthurium (Araceae) of Mexico and Central America. Part II: Panama. Missouri Botanical Garden Press, St Louis, 204 pp." type="book" year="1986">Croat 1986b</bibRefCitation>
). On Cerro Pirre, it represents a relatively rare species. Along the elevation gradient, it can only be observed in the montane cloud forest above
<quantity box="[305,386,970,994]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="671" unit="m" value="1100.0">1100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>