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<document ID-DOI="10.15560/15.4.651" ID-ISSN="1809-127X" approvalRequired="239" approvalRequired_for_taxonomicNames="94" approvalRequired_for_textStreams="143" approvalRequired_for_treatments="2" checkinTime="1630357287411" checkinUser="marcus" docAuthor="Ortiz, Orlando Oriel, de Stapf, María Sánchez, Baldini, Riccardo Maria &amp; Croat, Thomas Bernard" docDate="2019" docId="03F487E39D5CFFEAFC87F8DD3319F9B8" docLanguage="en" docName="CheckList.2019.15.4.651-689.pdf" docOrigin="Check List 15 (4)" docSource="http://dx.doi.org/10.15560/15.4.651" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Philodendron verrucosum Schott" docType="treatment" docVersion="1" lastPageId="33" lastPageNumber="684" masterDocId="FFCDFF9B9D7CFFCBFFAAFFEE313BFFD1" masterDocTitle="Synopsis of aroids (Alismatales, Araceae) from Cerro Pirre (Darién Province, Panama)" masterLastPageNumber="689" masterPageNumber="651" pageId="32" pageNumber="683" updateTime="1630357298930" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synopsis of aroids (Alismatales, Araceae) from Cerro Pirre (Darién Province, Panama)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortiz, Orlando Oriel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Stapf, María Sánchez</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baldini, Riccardo Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Croat, Thomas Bernard</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>15</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>651</mods:start>
<mods:end>689</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/15.4.651</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/15.4.651</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F487E39D5CFFEAFC87F8DD3319F9B8" httpUri="http://treatment.plazi.org/id/03F487E39D5CFFEAFC87F8DD3319F9B8" lastPageId="33" lastPageNumber="684" pageId="32" pageNumber="683">
<subSubSection box="[813,1338,1843,1868]" pageId="32" pageNumber="683" type="nomenclature">
<paragraph blockId="32.[813,1338,1843,1868]" box="[813,1338,1843,1868]" pageId="32" pageNumber="683">
<heading box="[813,1338,1843,1868]" fontSize="10" level="2" pageId="32" pageNumber="683" reason="2">
<taxonomicName authority="L. Mathieu ex Schott" authorityName="Schott" box="[813,1338,1843,1868]" class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="32" pageNumber="683" phylum="Tracheophyta" rank="species" species="verrucosum">
<emphasis box="[813,1338,1843,1868]" inLineHeading="true" pageId="32" pageNumber="683" reason="1">
<emphasis bold="true" box="[813,1104,1843,1867]" italics="true" pageId="32" pageNumber="683">Philodendron verrucosum</emphasis>
L. Mathieu ex Schott
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="33" lastPageNumber="684" pageId="32" pageNumber="683" type="materials_examined">
<paragraph blockId="32.[813,1421,1893,1984]" pageId="32" pageNumber="683">
<emphasis bold="true" box="[813,1040,1893,1917]" pageId="32" pageNumber="683">Material examined.</emphasis>
Serranía de Pirre, trail
<quantity box="[1309,1394,1893,1917]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" metricValueMax="6.437376" metricValueMin="1.609344" pageId="32" pageNumber="683" unit="mi" value="2.5" valueMax="4.0" valueMin="1.0">14 mi.</quantity>
<collectionCode box="[1402,1421,1894,1917]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="32" pageNumber="683" type="Herbarium">N</collectionCode>
of Cerro Pirre on main ridge;
<geoCoordinate box="[1132,1223,1925,1950]" degrees="07" direction="north" minutes="56" orientation="latitude" pageId="32" pageNumber="683" precision="925" value="7.9333334">07°56N</geoCoordinate>
,
<geoCoordinate box="[1233,1341,1925,1950]" degrees="077" direction="west" minutes="42" orientation="longitude" pageId="32" pageNumber="683" precision="925" value="-77.7">077°42W</geoCoordinate>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="32" pageNumber="683" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000 1100 m</quantity>
;
<emphasis box="[905,1114,1960,1984]" italics="true" pageId="32" pageNumber="683">
<collectionCode box="[905,925,1961,1984]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="32" pageNumber="683">R</collectionCode>
.
<collectionCode box="[928,947,1961,1984]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="32" pageNumber="683" type="Herbarium">L</collectionCode>
. Hartman 4829
</emphasis>
(
<collectionCode box="[1129,1176,1960,1984]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="32" pageNumber="683" type="Herbarium">MO</collectionCode>
).
</paragraph>
<paragraph blockId="32.[813,1420,2009,2067]" pageId="32" pageNumber="683">
<emphasis bold="true" box="[813,974,2009,2033]" pageId="32" pageNumber="683">Identification.</emphasis>
This species is characterized by having short internodes, subterete petioles (almost as long as
</paragraph>
<paragraph blockId="33.[167,775,160,351]" pageId="33" pageNumber="684">
the blade), broadly ovate-cordate blades, glossy above (giving a velvety appearance), and especially by having scaly-setose indumentum in the stems, cataphylls, petioles, and inflorescences.
<taxonomicName box="[455,744,260,284]" class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="33" pageNumber="684" phylum="Tracheophyta" rank="species" species="verrucosum">
<emphasis box="[455,744,260,284]" italics="true" pageId="33" pageNumber="684">Philodendron verrucosum</emphasis>
</taxonomicName>
is the only species of the genus having a scaly indumentum reported in the study area.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="684" type="distribution">
<paragraph blockId="33.[168,778,376,500]" pageId="33" pageNumber="684">
<emphasis bold="true" box="[168,457,376,400]" pageId="33" pageNumber="684">Distribution and ecology.</emphasis>
<collectingCountry box="[462,570,376,400]" name="Colombia" pageId="33" pageNumber="684">Colombia</collectingCountry>
,
<collectingCountry box="[580,701,376,400]" name="Costa Rica" pageId="33" pageNumber="684">Costa Rica</collectingCountry>
,
<collectingCountry name="Ecuador" pageId="33" pageNumber="684">Ecuador</collectingCountry>
,
<collectingCountry box="[218,332,410,434]" name="Nicaragua" pageId="33" pageNumber="684">Nicaragua</collectingCountry>
,
<collectingCountry box="[345,433,410,433]" name="Panama" pageId="33" pageNumber="684">Panama</collectingCountry>
, and
<collectingCountry box="[493,546,410,433]" name="Peru" pageId="33" pageNumber="684">Peru</collectingCountry>
. At the study site,
<taxonomicName class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="33" pageNumber="684" phylum="Tracheophyta" rank="species" species="verrucosum">
<emphasis italics="true" pageId="33" pageNumber="684">P. verrucosum</emphasis>
</taxonomicName>
has only been reported in submontane forests, at
<quantity box="[248,396,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="33" pageNumber="684" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">10001100 m</quantity>
.
</paragraph>
<paragraph blockId="33.[168,754,536,594]" pageId="33" pageNumber="684">
<taxonomicName authority="Croat &amp; Grayum" authorityName="Croat &amp; Grayum" class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="33" pageNumber="684" phylum="Tracheophyta" rank="variety" species="wilburii" variety="longipedunculatum">
<emphasis box="[168,754,536,560]" inLineHeading="true" pageId="33" pageNumber="684" reason="1">
<emphasis bold="true" box="[168,416,536,560]" italics="true" pageId="33" pageNumber="684">Philodendron wilburii</emphasis>
var.
<emphasis bold="true" box="[469,688,536,560]" italics="true" pageId="33" pageNumber="684">longipedunculatum</emphasis>
Croat
</emphasis>
&amp; Grayum
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="684" type="materials_examined">
<paragraph blockId="33.[166,776,619,976]" pageId="33" pageNumber="684">
<emphasis bold="true" box="[168,399,619,643]" pageId="33" pageNumber="684">Material examined.</emphasis>
Parque Nacional
<collectingRegion box="[607,684,619,643]" country="Panama" name="Darien" pageId="33" pageNumber="684">Darien</collectingRegion>
, middle slopes on
<collectionCode box="[285,310,653,676]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="33" pageNumber="684" type="Herbarium">W</collectionCode>
side of Cerro Pirre;
<geoCoordinate box="[557,648,651,676]" degrees="07" direction="north" minutes="56" orientation="latitude" pageId="33" pageNumber="684" precision="925" value="7.9333334">07°56N</geoCoordinate>
,
<geoCoordinate box="[663,772,651,676]" degrees="077" direction="west" minutes="43" orientation="longitude" pageId="33" pageNumber="684" precision="925" value="-77.71667">077°43W</geoCoordinate>
;
<quantity box="[168,303,685,709]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="10.5" metricValueMin="8.0" pageId="33" pageNumber="684" unit="m" value="925.0" valueMax="1050.0" valueMin="800.0">8001050 m</quantity>
;
<date box="[313,448,686,710]" pageId="33" pageNumber="684" value="1988-06-30">30 Jun. 1988</date>
;
<emphasis box="[459,641,686,710]" italics="true" pageId="33" pageNumber="684">
<collectionCode box="[459,477,686,709]" name="Tavera, Department of Geology and Geophysics" pageId="33" pageNumber="684">T</collectionCode>
.
<collectionCode box="[480,498,686,709]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="33" pageNumber="684" type="Herbarium">B</collectionCode>
. Croat 69001
</emphasis>
(
<collectionCode box="[653,700,686,710]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="33" pageNumber="684" type="Herbarium">MO</collectionCode>
). Ibid., Cerro Pirre, Rancho Frío;
<geoCoordinate box="[450,577,718,743]" degrees="08" direction="north" minutes="01" orientation="latitude" pageId="33" pageNumber="684" precision="15" seconds="14" value="8.0205555">08°0114″N</geoCoordinate>
,
<geoCoordinate box="[587,729,718,743]" degrees="077" direction="west" minutes="43" orientation="longitude" pageId="33" pageNumber="684" precision="15" seconds="41" value="-77.72806">077°4341″W</geoCoordinate>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.43" pageId="33" pageNumber="684" unit="m" value="143.0">143 m</quantity>
;
<date box="[202,344,752,777]" pageId="33" pageNumber="684" value="2016-04-13">13 Apr. 2016</date>
;
<emphasis box="[359,532,752,776]" italics="true" pageId="33" pageNumber="684">
<collectionCode box="[359,381,753,776]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
.
<collectionCode box="[385,407,753,776]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
. Ortiz 2556
</emphasis>
(
<collectionCode box="[546,609,752,776]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="33" pageNumber="684">PMA</collectionCode>
). Ibid., Rancho Plástico, camino hacia la cima del Cerro;
<geoCoordinate box="[644,772,784,810]" degrees="07" direction="north" minutes="59" orientation="latitude" pageId="33" pageNumber="684" precision="15" seconds="49" value="7.996945">07°5949″N</geoCoordinate>
,
<geoCoordinate box="[167,310,818,843]" degrees="077" direction="west" minutes="42" orientation="longitude" pageId="33" pageNumber="684" precision="15" seconds="45" value="-77.712494">077°4245″W</geoCoordinate>
;
<quantity box="[327,404,819,843]" metricMagnitude="2" metricUnit="m" metricValue="6.36" pageId="33" pageNumber="684" unit="m" value="636.0">636 m</quantity>
;
<date box="[421,573,819,843]" pageId="33" pageNumber="684" value="2016-04-15">15 Apr. 2016</date>
;
<emphasis box="[590,775,819,843]" italics="true" pageId="33" pageNumber="684">
<collectionCode box="[590,612,819,842]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
.
<collectionCode box="[616,638,819,842]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
. Ortiz 2582
</emphasis>
(
<collectionCode box="[176,239,852,876]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="33" pageNumber="684">PMA</collectionCode>
). Ibid.;
<geoCoordinate box="[319,447,851,876]" degrees="07" direction="north" minutes="59" orientation="latitude" pageId="33" pageNumber="684" precision="15" seconds="13" value="7.9869447">07°5913″N</geoCoordinate>
,
<geoCoordinate box="[457,602,851,876]" degrees="077" direction="west" minutes="42" orientation="longitude" pageId="33" pageNumber="684" precision="15" seconds="28" value="-77.70777">077°4228″W</geoCoordinate>
;
<quantity box="[613,693,852,876]" metricMagnitude="3" metricUnit="m" metricValue="1.127" pageId="33" pageNumber="684" unit="m" value="1127.0">1127 m</quantity>
;
<date pageId="33" pageNumber="684" value="2016-07-30">30 Jul. 2016</date>
;
<emphasis box="[233,410,886,910]" italics="true" pageId="33" pageNumber="684">
<collectionCode box="[233,255,886,909]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
.
<collectionCode box="[259,281,886,909]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
. Ortiz 2648
</emphasis>
(
<collectionCode box="[425,488,886,910]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="33" pageNumber="684">PMA</collectionCode>
). Ibid., campamento cerca del segundo mirador;
<geoCoordinate box="[410,537,918,943]" degrees="07" direction="north" minutes="59" orientation="latitude" pageId="33" pageNumber="684" precision="15" seconds="49" value="7.996945">07°5949″N</geoCoordinate>
,
<geoCoordinate box="[547,690,918,943]" degrees="077" direction="west" minutes="42" orientation="longitude" pageId="33" pageNumber="684" precision="15" seconds="43" value="-77.711945">077°4243″W</geoCoordinate>
;
<quantity box="[702,770,919,943]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="33" pageNumber="684" unit="m" value="610.0">610 m</quantity>
;
<date box="[168,297,952,976]" pageId="33" pageNumber="684" value="2016-12-03">3 Dec. 2016</date>
;
<emphasis box="[308,478,952,976]" italics="true" pageId="33" pageNumber="684">
<collectionCode box="[308,330,953,976]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
.
<collectionCode box="[334,356,953,976]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="33" pageNumber="684" type="Herbarium">O</collectionCode>
. Ortiz 2717
</emphasis>
(
<collectionCode box="[492,555,952,976]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="33" pageNumber="684">PMA</collectionCode>
).
</paragraph>
<paragraph blockId="33.[168,776,1001,1459]" pageId="33" pageNumber="684">
<emphasis bold="true" box="[168,330,1001,1025]" pageId="33" pageNumber="684">Identification.</emphasis>
This taxon is characterized by having elongated internodes, intact and deciduous cataphylls in most of the upper nodes, subteretes petioles, typically triangular-ovate blades, usually reddish when dry, primary lateral veins three to four per side, and long pedunculate inflorescences (peduncles usually longer than the spathe). This taxon differs from the typical variety
<taxonomicName authority="var. wilburii" class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="33" pageNumber="684" phylum="Tracheophyta" rank="variety" species="wilburii" variety="wilburii">
<emphasis italics="true" pageId="33" pageNumber="684">P. wilburii</emphasis>
var.
<emphasis box="[312,398,1235,1259]" italics="true" pageId="33" pageNumber="684">wilburii</emphasis>
</taxonomicName>
(absent on Cerro Pirre), mainly in having greenish blades (when dry) and peduncles usually of equal size or smaller than spathe. In the herbar- ium, specimens of this taxon tend to confuse them with those of
<taxonomicName authority="K. Krause" authorityName="K. Krause" box="[259,654,1368,1392]" class="Liliopsida" family="Araceae" genus="Philodendron" kingdom="Plantae" order="Alismatales" pageId="33" pageNumber="684" phylum="Tracheophyta" rank="species" species="panamense">
<emphasis box="[259,535,1368,1392]" italics="true" pageId="33" pageNumber="684">Philodendron panamense</emphasis>
K. Krause
</taxonomicName>
, which differs by having ovate blades (blackish when dry) with four to seven primary lateral veins per side.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="684" type="distribution">
<paragraph blockId="33.[168,776,1484,1642]" pageId="33" pageNumber="684">
<emphasis bold="true" box="[168,455,1484,1508]" pageId="33" pageNumber="684">Distribution and ecology.</emphasis>
<collectingCountry box="[460,577,1484,1508]" name="Costa Rica" pageId="33" pageNumber="684">Costa Rica</collectingCountry>
and
<collectingCountry box="[627,716,1485,1508]" name="Panama" pageId="33" pageNumber="684">Panama</collectingCountry>
. This taxon is distributed throughout the elevation gradient of Cerro Pirre. It can ocurrs both in semideciduous forests (to a lesser extent), as well as in evergreen, submontane, and montane forests at
<quantity box="[418,542,1618,1642]" metricMagnitude="2" metricUnit="m" metricValue="6.95" metricValueMax="13.0" metricValueMin="0.9" pageId="33" pageNumber="684" unit="m" value="695.0" valueMax="1300.0" valueMin="90.0">901300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>