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<document id="2BB5F0BA7F5B89DE0C892C109262D455" ID-DOI="10.11646/zootaxa.4032.4.1" ID-GBIF-Dataset="1459e41f-b82d-4c88-be4d-ee86e329f9c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="243485" ID-ZooBank="AFEF8567-B7C6-4F54-AC51-4C990389BF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461294283733" checkinUser="plazi" docAuthor="Kustov, Semen &amp; Zherebilo, Danil" docDate="2015" docId="03F487D4FF8A1A71FF04F96DB863FC90" docLanguage="en" docName="zt04032p369.pdf" docOrigin="Zootaxa 4032 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wiedemannia (Philolutra) pseudovaillanti Joost" docType="treatment" docVersion="9" lastPageNumber="365" masterDocId="FFCDFFACFF871A7FFF93FFA0BD01FFCD" masterDocTitle="New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species" masterLastPageNumber="369" masterPageNumber="351" pageNumber="364" updateTime="1698616197258" updateUser="plazi">
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<mods:title id="B953F4F7B6311B2EDE8985BD7A193423">New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species</mods:title>
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<mods:namePart id="BB6AFBA42CA767CCB864A183E8B853B4">Kustov, Semen</mods:namePart>
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<mods:date id="8450C0F51939E186B3992CBC2774857F">2015</mods:date>
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<treatment id="03F487D4FF8A1A71FF04F96DB863FC90" ID-DOI="http://doi.org/10.5281/zenodo.5691039" ID-GBIF-Taxon="119642275" ID-Zenodo-Dep="5691039" LSID="urn:lsid:plazi:treatment:03F487D4FF8A1A71FF04F96DB863FC90" httpUri="http://treatment.plazi.org/id/03F487D4FF8A1A71FF04F96DB863FC90" lastPageId="14" lastPageNumber="365" pageId="13" pageNumber="364">
<subSubSection id="C3476549FF8A1A72FF04F96DBDF8F8C5" pageId="13" pageNumber="364" type="nomenclature">
<paragraph id="8BE236C2FF8A1A72FF04F96DBFDDF92A" blockId="13.[151,732,1741,1801]" box="[151,732,1741,1767]" pageId="13" pageNumber="364">
<heading id="D0AA81AEFF8A1A72FF04F96DBFDDF92A" bold="true" box="[151,732,1741,1767]" fontSize="11" level="1" pageId="13" pageNumber="364" reason="1">
<taxonomicName id="4C5D4D41FF8A1A72FF04F96DBFDDF92A" ID-CoL="8FM7V" authority="Joost" authorityName="Joost" box="[151,732,1741,1767]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="pseudovaillanti" subGenus="Philolutra">
<emphasis id="B929EAD0FF8A1A72FF04F96DBFDDF92A" bold="true" box="[151,732,1741,1767]" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF04F96DBC3FF92A" bold="true" box="[151,318,1741,1767]" italics="true" pageId="13" pageNumber="364">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF8A1A72FEDDF96DBCCBF92A" bold="true" box="[334,458,1741,1767]" italics="true" pageId="13" pageNumber="364">Philolutra</emphasis>
)
<emphasis id="B929EAD0FF8A1A72FE48F96DBF92F92A" bold="true" box="[475,659,1741,1767]" italics="true" pageId="13" pageNumber="364">pseudovaillanti</emphasis>
Joost
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE236C2FF8A1A72FF04F950BDF8F8C5" blockId="13.[151,732,1741,1801]" box="[151,249,1776,1801]" pageId="13" pageNumber="364">
(
<figureCitation id="13662A47FF8A1A72FF0CF950BDF1F8C5" box="[159,240,1776,1801]" captionStart="FIGURES 12 17" captionStartId="12.[151,263,1937,1959]" captionTargetBox="[247,1322,214,1871]" captionTargetId="figure@12.[229,1337,193,1891]" captionTargetPageId="12" captionText="FIGURES 12 17. Male terminalia of Wiedemannia, lateral view. 12, W. pseguashae sp. nov.; 13, W. pseudovaillanti Joost; 14, W. shamshevi Kustov &amp; Zherebilo; 15, W. sinclairi Kustov &amp; Zherebilo; 16, W. vaillanti Joost; 17, W. zetterstedti (Fallén). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/243488/files/figure.png" pageId="13" pageNumber="364">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A72FF04F898BFF7F883" box="[151,758,1848,1870]" pageId="13" pageNumber="364" type="reference_group">
<paragraph id="8BE236C2FF8A1A72FF04F898BFF7F883" blockId="13.[151,758,1848,1870]" box="[151,758,1848,1870]" pageId="13" pageNumber="364">
<treatmentCitationGroup id="AB4D11ECFF8A1A72FF04F898BFF7F883" box="[151,758,1848,1870]" pageId="13" pageNumber="364">
<taxonomicName id="4C5D4D41FF8A1A72FF04F898BFF3F883" ID-CoL="8FM7V" authority="Joost, 1981: 188" authorityName="Joost" authorityPageNumber="188" authorityYear="1981" box="[151,754,1848,1870]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="pseudovaillanti" subGenus="Philolutra">
<emphasis id="B929EAD0FF8A1A72FF04F898BC20F883" box="[151,289,1848,1870]" italics="true" pageId="13" pageNumber="364">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF8A1A72FEBCF898BC96F883" box="[303,407,1848,1870]" italics="true" pageId="13" pageNumber="364">Philolutra</emphasis>
)
<emphasis id="B929EAD0FF8A1A72FE35F898BF43F883" box="[422,578,1848,1870]" italics="true" pageId="13" pageNumber="364">pseudovaillanti</emphasis>
<treatmentCitation id="0AFC10D3FF8A1A72FDDAF899BFF3F883" author="Joost" box="[585,754,1848,1870]" page="188" pageId="13" pageNumber="364" year="1981">
<bibRefCitation id="EFCC4B33FF8A1A72FDDAF899BFBEF883" author="Joost" box="[585,703,1848,1870]" pageId="13" pageNumber="364" refString="Joost, W. (1981) Beitrag zur Kenntnis der Hemerodromiiniae des Kaukasus (I) (Diptera, Empididae). Reichenbachia, 19 (31), 183 - 191." type="journal article" year="1981">Joost, 1981</bibRefCitation>
: 188
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF8A1A71FF04F8DBB836FE39" lastPageId="14" lastPageNumber="365" pageId="13" pageNumber="364" type="materials_examined">
<paragraph id="8BE236C2FF8A1A71FF04F8DBB836FE39" blockId="13.[151,1436,1915,2012]" lastBlockId="14.[151,1437,151,861]" lastPageId="14" lastPageNumber="365" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FF04F8DBBC78F859" bold="true" box="[151,377,1915,1940]" pageId="13" pageNumber="364">Material examined</emphasis>
.
<collectingCountry id="F34A7652FF8A1A72FE1BF8DBBCEEF859" box="[392,495,1915,1940]" name="Russia" pageId="13" pageNumber="364">
<emphasis id="B929EAD0FF8A1A72FE1BF8DBBCEEF859" bold="true" box="[392,495,1915,1940]" pageId="13" pageNumber="364">RUSSIA</emphasis>
</collectingCountry>
: Karachay-Cherkess Republic, env. Rozhkao (13),
<date id="FFE31002FF8A1A72FBA3F8DBB99CF859" box="[1072,1181,1915,1940]" pageId="13" pageNumber="364" value="2013-06-08">8.vi.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF8A1A72FADBF8DBB874F85E" box="[1352,1397,1915,1939]" pageId="13" pageNumber="364" type="generic">2 ♂</specimenCount>
, 11 ♀); Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (5),
<date id="FFE31002FF8A1A72FC7DF83EB96EF87A" box="[1006,1135,1950,1975]" pageId="13" pageNumber="364" value="2013-06-24">24.06.2013</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF8A1A72FAB7F83FB851F87A" box="[1316,1360,1951,1975]" pageId="13" pageNumber="364" type="generic">3 ♂</specimenCount>
, 9 ♀); same data,
<date id="FFE31002FF8A1A72FE89F863BC7CF811" box="[282,381,1987,2012]" pageId="13" pageNumber="364" value="2015-05-03">3.v.2015</date>
, S.Yu. Kustov, D. Zherebilo (2 ♀); Nature Reserve “Kamyshanova Polyana” (7),
<date id="FFE31002FF8A1A72FAA6F863B896F816" box="[1333,1431,1987,2011]" pageId="13" pageNumber="364" value="2014-05-02">2.v.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF891A71FED6FF38BC74FF62" box="[325,373,152,176]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
); Nature Reserve “Kamyshanova Polyana” (8),
<date id="FFE31002FF891A71FC22FF38B915FF7D" box="[945,1044,152,176]" pageId="14" pageNumber="365" value="2014-05-04">4.v.2014</date>
, S.Yu. Kustov (3 ♀); same data,
<date id="FFE31002FF891A71FF04FF1CBC18FF18" box="[151,281,188,213]" pageId="14" pageNumber="365" value="2014-07-17">17.vii.2014</date>
, V.V. Gladun (
<specimenCount id="9D5BFD4BFF891A71FE53FF1CBCEDFF19" box="[448,492,188,212]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
, 3 ♀); same data,
<date id="FFE31002FF891A71FD2DFF1CBE41FF18" box="[702,832,188,213]" pageId="14" pageNumber="365" value="2014-07-19">19.vii.2014</date>
, V.V. Gladun (
<specimenCount id="9D5BFD4BFF891A71FC79FF1CB912FF19" box="[1002,1043,188,212]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
, 2 ♀); same data,
<date id="FFE31002FF891A71FB75FF1CB852FF18" box="[1254,1363,188,213]" pageId="14" pageNumber="365" value="2014-09-06">6.ix.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF891A71FF6DFF40BC2CFF3A" box="[254,301,224,248]" pageId="14" pageNumber="365" type="generic">3 ♂</specimenCount>
, 2 ♀); same data,
<date id="FFE31002FF891A71FD84FF7FBF7BFF35" box="[535,634,223,248]" pageId="14" pageNumber="365" value="2015-05-02">2.v.2015</date>
, S.Yu. Kustov, D. Zherebilo (6 ♀); Nature Reserve “Kamyshanova Polyana” (6),
<date id="FFE31002FF891A71FED3FEA4BCADFED0" box="[320,428,260,285]" pageId="14" pageNumber="365" value="2014-06-01">1.vi.2014</date>
, S.Yu. Kustov (
<specimenCount id="9D5BFD4BFF891A71FDE0FEA4BFA3FED1" box="[627,674,260,284]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
); same data,
<date id="FFE31002FF891A71FCD8FEA4BECCFED0" box="[843,973,260,285]" pageId="14" pageNumber="365" value="2014-07-17">17.vii.2014</date>
, Е. Pavlova, (1 ♀); env. Guamka (3),
<date id="FFE31002FF891A71FF04FE87BC19FE8D" box="[151,280,295,320]" pageId="14" pageNumber="365" value="2015-04-30">30.04.2015</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FE42FE88BF02FEF2" box="[465,515,296,320]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
, 1 ♀).
<collectingCountry id="F34A7652FF891A71FDF3FE87BFE6FE8D" box="[608,743,295,320]" name="Georgia" pageId="14" pageNumber="365">
<emphasis id="B929EAD0FF891A71FDF3FE87BFE6FE8D" bold="true" box="[608,743,295,320]" pageId="14" pageNumber="365">GEORGIA</emphasis>
</collectingCountry>
: Samtskhe-Javakheti (29),
<date id="FFE31002FF891A71FBBAFE87B9A0FE8D" box="[1065,1185,295,320]" pageId="14" pageNumber="365" value="2014-04-20">20.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FACEFE88B88CFE8D" box="[1373,1421,296,320]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
); Shavsheti ridge (24),
<date id="FFE31002FF891A71FE1AFEECBCF2FEA8" box="[393,499,332,357]" pageId="14" pageNumber="365" value="2015-04-04">4.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FD25FEECBFE5FEA9" box="[694,740,332,356]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
); env. Hihadziri (33),
<date id="FFE31002FF891A71FC4DFEECB957FEA8" box="[990,1110,332,357]" pageId="14" pageNumber="365" value="2015-04-06">06.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FA89FEECB845FEA9" box="[1306,1348,332,357]" pageId="14" pageNumber="365" type="generic">5 ♂</specimenCount>
, 10 ♀); env. Chinkadziebi,
<quantity id="4CA59B27FF891A71FEE4FECFBCC2FE4A" box="[375,451,367,392]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="365" unit="m" value="360.0">360 m</quantity>
a.s.l.,
<date id="FFE31002FF891A71FD87FECFBF8CFE45" box="[532,653,367,392]" pageId="14" pageNumber="365" value="2015-04-06">06.iv.2015</date>
, A.V. Zherebilo (1 ♀); env. Zedachukhuneti (27),
<date id="FFE31002FF891A71FB72FECFB85BFE45" box="[1249,1370,367,392]" pageId="14" pageNumber="365" value="2015-04-07">07.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FE8AFE34BC48FE61" box="[281,329,404,428]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
, 1 ♀); env. Chaisubani (26),
<date id="FFE31002FF891A71FD21FE34BE2AFE60" box="[690,811,404,429]" pageId="14" pageNumber="365" value="2015-04-08">08.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FC6DFE34B92CFE61" box="[1022,1069,404,428]" pageId="14" pageNumber="365" type="generic">3 ♂</specimenCount>
, 5 ♀); env. Zedakinchha (34),
<date id="FFE31002FF891A71FF04FE17BC11FE1D" box="[151,272,439,464]" pageId="14" pageNumber="365" value="2015-04-09">09.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FE4DFE18BF0BFE02" box="[478,522,440,464]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
, 1 ♀); env. Opitara (35),
<date id="FFE31002FF891A71FCA8FE17BEB5FE1D" box="[827,948,439,464]" pageId="14" pageNumber="365" value="2015-04-10">10.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FB11FE18B9B1FE1D" box="[1154,1200,440,464]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
); env. Chkumi (36),
<date id="FFE31002FF891A71FF04FE7CBC0FFE38" box="[151,270,476,501]" pageId="14" pageNumber="365" value="2015-04-11">11.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FE43FE7CBCFCFE39" box="[464,509,476,500]" pageId="14" pageNumber="365" type="generic">2 ♂</specimenCount>
, 1 ♀).
<emphasis id="B929EAD0FF891A71FDD9FE7CBFE3FE38" bold="true" box="[586,738,476,501]" pageId="14" pageNumber="365">ABKHAZIA</emphasis>
: River Kelasuri (21),
<date id="FFE31002FF891A71FC4AFE7CB950FE38" box="[985,1105,476,501]" pageId="14" pageNumber="365" value="2014-04-12">12.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF891A71FB6DFE7CB828FE39" box="[1278,1321,476,500]" pageId="14" pageNumber="365" type="generic">1 ♂</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF891A71FF54FE5FB970FD49" pageId="14" pageNumber="365" type="diagnosis">
<paragraph id="8BE236C2FF891A71FF54FE5FB970FD49" blockId="14.[151,1437,151,861]" pageId="14" pageNumber="365">
<emphasis id="B929EAD0FF891A71FF54FE5FBC3BFDD5" bold="true" box="[199,314,511,536]" pageId="14" pageNumber="365">Diagnosis</emphasis>
. This species is distinguished by the completely dark brown scutum, acrostichals divergent around the prescutellar depression, 56 pairs of dorsocentrals with few intermediate setae, scutellum with several marginal setulae, fore femur apically without strong anterior setae; clasping cercus with anterior lobe short, pointed, bearing cluster of apical setae; dorsal process subtriangular, inner margin clothed in dense setae.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF891A71FF54FD2FBC73FD00" pageId="14" pageNumber="365" type="distribution">
<paragraph id="8BE236C2FF891A71FF54FD2FBC73FD00" blockId="14.[151,1437,151,861]" pageId="14" pageNumber="365">
<emphasis id="B929EAD0FF891A71FF54FD2FBC58FD65" bold="true" box="[199,345,655,680]" pageId="14" pageNumber="365">Distribution</emphasis>
. PALAEARCTIC: Caucasus:
<collectingCountry id="F34A7652FF891A71FD2CFD2FBE0BFD65" box="[703,778,655,680]" name="Russia" pageId="14" pageNumber="365">Russia</collectingCountry>
(Krasnodarsky Territory, Karachay-Cherkess Republic),
<collectingCountry id="F34A7652FF891A71FF04FD14BDF5FD00" box="[151,244,692,717]" name="Georgia" pageId="14" pageNumber="365">Georgia</collectingCountry>
, Abkhazia.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF891A71FF54FD77B863FC90" pageId="14" pageNumber="365" type="discussion">
<paragraph id="8BE236C2FF891A71FF54FD77B863FC90" blockId="14.[151,1437,151,861]" pageId="14" pageNumber="365">
<emphasis id="B929EAD0FF891A71FF54FD77BC35FD3D" bold="true" box="[199,308,727,752]" pageId="14" pageNumber="365">Remarks</emphasis>
.
<bibRefCitation id="EFCC4B33FF891A71FED7FD78BCD0FD3D" author="Joost" box="[324,465,728,752]" pageId="14" pageNumber="365" refString="Joost, W. (1981) Beitrag zur Kenntnis der Hemerodromiiniae des Kaukasus (I) (Diptera, Empididae). Reichenbachia, 19 (31), 183 - 191." type="journal article" year="1981">Joost (1981)</bibRefCitation>
reported this species from the territory of Russian Caucasus—Dombai and Teberda, but
<bibRefCitation id="EFCC4B33FF891A71FF52FD5CBCDBFCD9" author="Chvala" box="[193,474,764,789]" pageId="14" pageNumber="365" refString="Chvala, M. &amp; Wagner, R. (1989) Hybotidae. In: Catalogue of Palaearctic Diptera. Volume 6. Therevidae-Empididae. Elsevier Science Publishing, pp. 228 - 336." type="book" year="1989">Chvála &amp; Wagner (1989)</bibRefCitation>
in the Catalogue of Palaearctic
<taxonomicName id="4C5D4D41FF891A71FCAFFD5CBE91FCD8" box="[828,912,764,789]" kingdom="Plantae" order="Diptera" pageId="14" pageNumber="365" rank="order">Diptera</taxonomicName>
mistakenly indicated the species only from the Georgian territory. Therefore the species is noted for
<collectingCountry id="F34A7652FF891A71FC9FFCBFBE67FCF5" box="[780,870,799,824]" name="Georgia" pageId="14" pageNumber="365">Georgia</collectingCountry>
and Abkhazia for the first time. This species is commonly known from mountain rivers at elevations from
<quantity id="4CA59B27FF891A71FCBCFCE4BEBDFC91" box="[815,956,836,861]" metricMagnitude="2" metricUnit="m" metricValue="8.94" metricValueMax="15.0" metricValueMin="2.88" pageId="14" pageNumber="365" unit="m" value="894.0" valueMax="1500.0" valueMin="288.0">2881500 m</quantity>
a.s.l. and is active from April to July.
</paragraph>
</subSubSection>
</treatment>
</document>