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<document id="056250E5FAFBF741DB9E488C8D5F66E4" ID-DOI="10.5281/zenodo.207679" ID-GBIF-Dataset="c8f99200-8b49-493b-8312-0f6857403b49" ID-ISSN="1175-5326" ID-Zenodo-Dep="207679" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460325380972" checkinUser="plazi" docAuthor="Janisova, Kristyna &amp; Bocakova, Milada" docDate="2011" docId="03F487ACBC71C849FF2F37EDFAF5F400" docLanguage="en" docName="zt02975p068.pdf" docOrigin="Zootaxa 2975" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hyperstoma Wittmer 1979" docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FFCDFFD4BC70C848FFB8377AFFA0F071" masterDocTitle="Review of the genus Hyperstoma (Coleoptera: Lampyridae)" masterLastPageNumber="68" masterPageNumber="64" pageNumber="65" updateTime="1698257361746" updateUser="plazi">
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<mods:title id="5FAB28B983CEEA111AC1E2205781E8D2">Review of the genus Hyperstoma (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="269416133FF1E0A26E77E1808ABA31F8">Janisova, Kristyna</mods:namePart>
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<mods:namePart id="170D9816DECF58A107D4FB4A27D920A1">Bocakova, Milada</mods:namePart>
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<mods:date id="2166D0470C936E6170846034A392E911">2011</mods:date>
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<treatment id="03F487ACBC71C849FF2F37EDFAF5F400" ID-DOI="http://doi.org/10.5281/zenodo.6194991" ID-GBIF-Taxon="119403040" ID-Zenodo-Dep="6194991" LSID="urn:lsid:plazi:treatment:03F487ACBC71C849FF2F37EDFAF5F400" httpUri="http://treatment.plazi.org/id/03F487ACBC71C849FF2F37EDFAF5F400" lastPageNumber="65" pageId="1" pageNumber="65">
<subSubSection id="C3476531BC71C849FF2F37EDFE42F0C0" box="[151,482,151,177]" pageId="1" pageNumber="65" type="nomenclature">
<paragraph id="8BE236BABC71C849FF2F37EDFE42F0C0" blockId="1.[151,482,151,177]" box="[151,482,151,177]" pageId="1" pageNumber="65">
<heading id="D0AA81D6BC71C849FF2F37EDFE42F0C0" bold="true" box="[151,482,151,177]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
<taxonomicName id="4C5D4D39BC71C849FF2F37EDFE42F0C0" ID-CoL="52GV" authority="Wittmer, 1979" authorityName="Wittmer" authorityYear="1979" box="[151,482,151,177]" class="Insecta" family="Lampyridae" genus="Hyperstoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FF2F37EDFE42F0C0" bold="true" box="[151,482,151,177]" pageId="1" pageNumber="65">
<emphasis id="B929EAA8BC71C849FF2F37EDFE89F0C0" bold="true" box="[151,297,151,177]" italics="true" pageId="1" pageNumber="65">Hyperstoma</emphasis>
<bibRefCitation id="EFCC4B4BBC71C849FE8937EDFE42F0C0" author="Wittmer" box="[305,482,151,177]" pageId="1" pageNumber="65" refString="Wittmer, W. (1979) Zur Kenntnis der Lampyridae der orientalischen Region (Coleoptera). Entomologische Arbeiten aus dem Museum G. Frey, 28, 83 - 92." type="journal article" year="1979">Wittmer, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF2F379AFE78F085" box="[151,472,222,245]" pageId="1" pageNumber="65" type="reference_group">
<paragraph id="8BE236BABC71C849FF2F379AFE78F085" blockId="1.[151,845,222,275]" box="[151,472,222,245]" pageId="1" pageNumber="65">
<treatmentCitationGroup id="AB4D1194BC71C849FF2F379AFE78F085" box="[151,472,222,245]" pageId="1" pageNumber="65">
<taxonomicName id="4C5D4D39BC71C849FF2F379AFE74F085" ID-CoL="52GV" authority="Wittmer, 1979: 84" authorityName="Wittmer" authorityPageNumber="84" authorityYear="1979" box="[151,468,222,245]" class="Insecta" family="Lampyridae" genus="Hyperstoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FF2F379AFEB3F084" box="[151,275,224,245]" italics="true" pageId="1" pageNumber="65">Hyperstoma</emphasis>
<treatmentCitation id="0AFC10ABBC71C849FEA137A4FE74F085" author="Wittmer" box="[281,468,222,244]" page="84" pageId="1" pageNumber="65" year="1979">
<bibRefCitation id="EFCC4B4BBC71C849FEA137A4FE0FF085" author="Wittmer" box="[281,431,222,244]" pageId="1" pageNumber="65" refString="Wittmer, W. (1979) Zur Kenntnis der Lampyridae der orientalischen Region (Coleoptera). Entomologische Arbeiten aus dem Museum G. Frey, 28, 83 - 92." type="journal article" year="1979">Wittmer, 1979</bibRefCitation>
: 84
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF2F3784FCEDF162" box="[151,845,253,275]" pageId="1" pageNumber="65" type="materials_examined">
<paragraph id="8BE236BABC71C849FF2F3784FCEDF162" blockId="1.[151,845,222,275]" box="[151,845,253,275]" pageId="1" pageNumber="65">
<typeStatus id="54E68818BC71C849FF2F3784FF68F162" box="[151,200,254,275]" pageId="1" pageNumber="65">Type</typeStatus>
species.
<taxonomicName id="4C5D4D39BC71C849FE9D3784FD09F163" authority="Wittmer, 1979" authorityName="Wittmer" authorityYear="1979" box="[293,681,253,275]" class="Insecta" family="Lampyridae" genus="Hyperstoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B929EAA8BC71C849FE9D3784FDB0F162" box="[293,528,254,275]" italics="true" pageId="1" pageNumber="65">Hyperstoma marginata</emphasis>
<bibRefCitation id="EFCC4B4BBC71C849FDAF3787FD09F163" author="Wittmer" box="[535,681,253,275]" pageId="1" pageNumber="65" refString="Wittmer, W. (1979) Zur Kenntnis der Lampyridae der orientalischen Region (Coleoptera). Entomologische Arbeiten aus dem Museum G. Frey, 28, 83 - 92." type="journal article" year="1979">Wittmer, 1979</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF2F3645FBE6F1EE" pageId="1" pageNumber="65" type="diagnosis">
<paragraph id="8BE236BABC71C849FF2F3645FBE6F1EE" blockId="1.[151,1437,319,1137]" pageId="1" pageNumber="65">
<emphasis id="B929EAA8BC71C849FF2F3645FEB1F129" bold="true" box="[151,273,319,344]" pageId="1" pageNumber="65">Diagnosis.</emphasis>
<taxonomicName id="4C5D4D39BC71C849FEAF363BFE3FF129" box="[279,415,321,344]" class="Insecta" family="Lampyridae" genus="Hyperstoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FEAF363BFE3FF129" box="[279,415,321,344]" italics="true" pageId="1" pageNumber="65">Hyperstoma</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="4C5D4D39BC71C849FCB53645FC2BF126" box="[781,907,319,343]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="subFamily" subFamily="Ototretinae">Ototretinae</taxonomicName>
by combination of following characters:
<emphasis id="B929EAA8BC71C849FAEE363AFAFEF129" bold="true" box="[1366,1374,320,344]" pageId="1" pageNumber="65">t</emphasis>
erminal maxillary and labial palpomere extremely long, male antennae flabellate with stout branches attached to antennomere apex, male terminal and penultimate abdominal terga fused, jointly trilobed.
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF7F36D1FCC3F475" pageId="1" pageNumber="65" type="description">
<paragraph id="8BE236BABC71C849FF7F36D1FCC3F475" blockId="1.[151,1437,319,1137]" pageId="1" pageNumber="65">
<emphasis id="B929EAA8BC71C849FF7F36D1FEF9F1B5" bold="true" box="[199,345,427,452]" pageId="1" pageNumber="65">Description.</emphasis>
Body length
<quantity id="4CA59B5FBC71C849FE4936D6FDC7F1B5" box="[497,615,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="1" pageNumber="65" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">55.5 mm</quantity>
, humeral width
<quantity id="4CA59B5FBC71C849FCA536D6FC63F1B5" box="[797,963,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.35" metricValueMin="2.05" pageId="1" pageNumber="65" unit="mm" value="2.2" valueMax="2.35" valueMin="2.05">2.052.35 mm</quantity>
, body oval, mainly yellow to light brown, distal portion of antennae and elytral margins dark brown. Head from above partly hidden by pronotum. Eyes small, distance between eyes 1.82× longer than eye diameter. Antennae 11-segmented, antennomeres 310 flabellate (
<figureCitation id="13662A3FBC71C849FF76356DFE92F25E" box="[206,306,535,559]" captionStart="FIGURES 1 4. 1 2" captionStartId="2.[151,264,615,638]" captionTargetBox="[176,1411,220,583]" captionTargetId="figure@2.[151,1436,195,591]" captionTargetPageId="2" captionText="FIGURES 1 4. 1 2 Head and pronotum, 3 4 Basal antennomeres. 1, 3 Hyperstoma marginata; 2, 4 H. wittmeri sp. nov. Scale— 0.5 mm." httpUri="https://zenodo.org/record/207680/files/figure.png" pageId="1" pageNumber="65">Figs 3, 4</figureCitation>
), lamellae strongly flattened, attached to antennomere terminal portion, branches of median antennomeres about 2.73× longer than antennomere body, branches of terminal antennomeres moderately shorter. Mandibles small, hooked, apically pointed. Maxillary palpi 4-segmented, labial palpi 3-segmented, terminal palpomeres of both palpi enlarged, flattened, oval, 2.23.3× longer than width. Pronotum (
<figureCitation id="13662A3FBC71C849FBC635FEFB45F2ED" box="[1150,1253,644,668]" captionStart="FIGURES 1 4. 1 2" captionStartId="2.[151,264,615,638]" captionTargetBox="[176,1411,220,583]" captionTargetId="figure@2.[151,1436,195,591]" captionTargetPageId="2" captionText="FIGURES 1 4. 1 2 Head and pronotum, 3 4 Basal antennomeres. 1, 3 Hyperstoma marginata; 2, 4 H. wittmeri sp. nov. Scale— 0.5 mm." httpUri="https://zenodo.org/record/207680/files/figure.png" pageId="1" pageNumber="65">Figs 1, 2</figureCitation>
) yellow to light brown, 1.8× as wide as long, anterior margin almost straight, anterior angles rounded, posterior angles triangular, projected obliquely backwards, provided with weak circular pit distally, median portion of pronotum with longitudinal groove. Scutellum triangular, yellow to light brown. Elytra elongate, each elytron about 2.73.4× longer than wide, widest medially, apex rounded. Elytra shiny, irregularly densely punctured, slightly pubescent, each elytron with 2 inconspicuous costae. Terminal abdominal sternum with oval basal portion and straight margins distally. Terminal and penultimate abdominal terga fused, jointly trilobed (
<figureCitation id="13662A3FBC71C849FC383426FC54F305" box="[896,1012,860,884]" captionStart="FIGURES 5 10. 5 8" captionStartId="2.[151,264,1124,1147]" captionTargetBox="[163,1388,722,1094]" captionTargetId="figure@2.[151,1436,717,1102]" captionTargetPageId="2" captionText="FIGURES 5 10. 5 8 Male genitalia in ventral and lateral view, 9 10 terminal and penultimate abdominal terga. 5, 6, 9 Hyperstoma marginata; 7, 8, 10 H. wittmeri sp. nov. Scale— 0.5 mm." httpUri="https://zenodo.org/record/207681/files/figure.png" pageId="1" pageNumber="65">Figs 9, 10</figureCitation>
) with straight or U-shaped basal portion. Penultimate tergum extending in 2 lateral protrusions, forming 2 longitudinal apertures along terminal tergum. Male genitalia with V-shaped phallobase (
<figureCitation id="13662A3FBC71C849FDD634DEFD73F3CD" box="[622,723,932,956]" captionStart="FIGURES 5 10. 5 8" captionStartId="2.[151,264,1124,1147]" captionTargetBox="[163,1388,722,1094]" captionTargetId="figure@2.[151,1436,717,1102]" captionTargetPageId="2" captionText="FIGURES 5 10. 5 8 Male genitalia in ventral and lateral view, 9 10 terminal and penultimate abdominal terga. 5, 6, 9 Hyperstoma marginata; 7, 8, 10 H. wittmeri sp. nov. Scale— 0.5 mm." httpUri="https://zenodo.org/record/207681/files/figure.png" pageId="1" pageNumber="65">Figs 58</figureCitation>
), paramerae robust, apex rounded or with oblique edge distally, paramerae about 1/5 longer than phallus. Phallus cochleariform, provided with short ventroapical projection and forked, angular projection basally (
<figureCitation id="13662A3FBC71C849FDA73496FD24F475" box="[543,644,1004,1028]" captionStart="FIGURES 5 10. 5 8" captionStartId="2.[151,264,1124,1147]" captionTargetBox="[163,1388,722,1094]" captionTargetId="figure@2.[151,1436,717,1102]" captionTargetPageId="2" captionText="FIGURES 5 10. 5 8 Male genitalia in ventral and lateral view, 9 10 terminal and penultimate abdominal terga. 5, 6, 9 Hyperstoma marginata; 7, 8, 10 H. wittmeri sp. nov. Scale— 0.5 mm." httpUri="https://zenodo.org/record/207681/files/figure.png" pageId="1" pageNumber="65">Figs 6, 8</figureCitation>
). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF7F3375FE7DF459" box="[199,477,1039,1064]" pageId="1" pageNumber="65" type="distribution">
<paragraph id="8BE236BABC71C849FF7F3375FE7DF459" blockId="1.[151,1437,319,1137]" box="[199,477,1039,1064]" pageId="1" pageNumber="65">
<emphasis id="B929EAA8BC71C849FF7F3375FEC0F459" bold="true" box="[199,352,1039,1064]" pageId="1" pageNumber="65">Distribution.</emphasis>
<collectingCountry id="F34A762ABC71C849FEDF336AFE79F459" box="[359,473,1040,1064]" name="Sri Lanka" pageId="1" pageNumber="65">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476531BC71C849FF7F3349FAF5F400" pageId="1" pageNumber="65" type="discussion">
<paragraph id="8BE236BABC71C849FF7F3349FAF5F400" blockId="1.[151,1437,319,1137]" pageId="1" pageNumber="65">
<emphasis id="B929EAA8BC71C849FF7F3349FEE4F43D" bold="true" box="[199,324,1075,1100]" pageId="1" pageNumber="65">Comment.</emphasis>
<taxonomicName id="4C5D4D39BC71C849FEF7334FFE78F43D" box="[335,472,1077,1100]" class="Insecta" family="Lampyridae" genus="Hyperstoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FEF7334FFE78F43D" box="[335,472,1077,1100]" italics="true" pageId="1" pageNumber="65">Hyperstoma</emphasis>
</taxonomicName>
is closely related to the genera
<taxonomicName id="4C5D4D39BC71C849FCE2334EFB02F43D" authority="Maulik, 1921" authorityName="Maulik" authorityYear="1921" box="[858,1186,1076,1100]" class="Insecta" family="Lampyridae" genus="Lamellipalpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FCE2334EFC5AF43D" box="[858,1018,1076,1100]" italics="true" pageId="1" pageNumber="65">Lamellipalpus</emphasis>
Maulik, 1921
</taxonomicName>
and
<taxonomicName id="4C5D4D39BC71C849FB59334EFE91F401" authority="Maulik, 1921" authorityName="Maulik" authorityYear="1921" class="Insecta" family="Lampyridae" genus="Lamellipalpodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B929EAA8BC71C849FB59334EFA3CF43D" box="[1249,1436,1076,1100]" italics="true" pageId="1" pageNumber="65">Lamellipalpodes</emphasis>
Maulik, 1921
</taxonomicName>
. All the genera share a unique synapomorphic character: fused terminal and penultimate terga.
</paragraph>
</subSubSection>
</treatment>
</document>