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treatments-xml/data/03/E3/88/03E38878867EFB72FEC1CD120EC26E44.xml
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<document id="4E4726E3C649A0211DA11F1E30AC55A8" ID-CLB-Dataset="27039" ID-DOI="10.5281/zenodo.2646031" ID-GBIF-Dataset="4c9ede5c-44e0-49b0-8a3c-9f176c8935bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2646031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555587772508" checkinUser="plazi" docAuthor="Paulsen, M. J." docDate="2006" docId="03E38878867EFB72FEC1CD120EC26E44" docLanguage="en" docName="zt01185p068.pdf" docOrigin="Zootaxa 1185" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Aphodius matiganae Paulsen 2006, new species" docType="treatment" docVersion="10" lastPageNumber="68" masterDocId="FFDAF000867CFB75FFC9CC4B0D276B31" masterDocTitle="A new species of Aphodius Illiger (Coleoptera: Scarabaeidae: Aphodiinae) from the central United States" masterLastPageNumber="68" masterPageNumber="61" pageNumber="63" updateTime="1698722832894" updateUser="ExternalLinkService">
<mods:mods id="71BEC7148CFAA9A0572D8C8FBACD990C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA182967B557311F6D4A19AA43B0B43D">
<mods:title id="2DFF8071806260F536392B4E8BFCD2C2">A new species of Aphodius Illiger (Coleoptera: Scarabaeidae: Aphodiinae) from the central United States</mods:title>
</mods:titleInfo>
<mods:name id="30F623884B51B5F54F05B984878348F2" type="personal">
<mods:role id="930D3507991D50EC475C39C48205AFBA">
<mods:roleTerm id="8B1885E8D8A45FB79E0D65B89ADEF9D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D891E8A2E960DAF08FBA268D8A3FB4A">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0D642EBE7793EF52529A8A2B96A46428">text</mods:typeOfResource>
<mods:relatedItem id="37EC89930C54A9648E2C61CE77CB8007" type="host">
<mods:titleInfo id="76FD3B6C8EC0A7F2835A0FB45B5923A5">
<mods:title id="59343B4B2C2E15F4F8EA81FA0970129A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7A51B27E4E53EA6FF59B4ED86E71CF20">
<mods:date id="2986D7F7A991CC5C257AC0C7459A3651">2006</mods:date>
<mods:detail id="D68EE70BF4A5C2B9512D6E2A046F8358" type="pubDate">
<mods:number id="41C24B6C5AE4227B3C09F281B1D0547E">2006-04-24</mods:number>
</mods:detail>
<mods:detail id="0693B67CE790F0504CF65D5914D91515" type="volume">
<mods:number id="DB0E746BBDF4F8691D74719B50D8BF95">1185</mods:number>
</mods:detail>
<mods:extent id="CA5801B1EF4532B8693BDDD29504438A" unit="page">
<mods:start id="C35193D5247EA653E0512C8DFFC7FB0E">61</mods:start>
<mods:end id="0FBE02911FBEE2F80C994A1036AF83F9">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ABA840CE4DC853D5D32022D27D6D010A">journal article</mods:classification>
<mods:identifier id="FFE05EFB3DC2231618BA0C9347723F9B" type="CLB-Dataset">27039</mods:identifier>
<mods:identifier id="35C8E2BFDA4D21DCB8B8620B590A7EBF" type="DOI">10.5281/zenodo.2646031</mods:identifier>
<mods:identifier id="35D01A17AC7E2C9D4BB8F76E8E079CCB" type="GBIF-Dataset">4c9ede5c-44e0-49b0-8a3c-9f176c8935bc</mods:identifier>
<mods:identifier id="6556F23F467ED00F33E6D44B456140A7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="63C9C51730AD87240332ED88425E2C7A" type="Zenodo-Dep">2646031</mods:identifier>
<mods:identifier id="5A3B81DA599FC6B4973D70808D194774" type="ZooBank">991C41B2-9DD5-4E4F-B40D-98F87A1EEC37</mods:identifier>
</mods:mods>
<treatment id="03E38878867EFB72FEC1CD120EC26E44" ID-GBIF-Taxon="156789398" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44" httpUri="http://treatment.plazi.org/id/03E38878867EFB72FEC1CD120EC26E44" lastPageId="7" lastPageNumber="68" pageId="2" pageNumber="63">
<subSubSection id="C3506AE5867EFB77FEC1CD120E2E6A42" box="[264,777,345,371]" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="8BF5396E867EFB77FEC1CD120E2E6A42" blockId="2.[264,777,283,406]" box="[264,777,345,371]" pageId="2" pageNumber="63">
<heading id="D0BD8E02867EFB77FEC1CD120E2E6A42" bold="true" box="[264,777,345,371]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
<emphasis id="B93EE57C867EFB77FEC1CD120E2E6A42" bold="true" box="[264,777,345,371]" pageId="2" pageNumber="63">
<taxonomicName id="4C4A42ED867EFB77FEC1CD120F4A6A42" authority="Paulsen" authorityName="Paulsen" authorityYear="2006" box="[264,621,345,371]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="matiganae" status="sp. nov.">
<emphasis id="B93EE57C867EFB77FEC1CD120F256A42" bold="true" box="[264,514,345,371]" italics="true" pageId="2" pageNumber="63">Aphodius matiganae</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="A20D5807867EFB77FDB0CD120E2E6A42" box="[633,777,345,371]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3506AE5867EFB77FEC1CD370CC66AA7" box="[264,481,380,406]" pageId="2" pageNumber="63" type="description">
<paragraph id="8BF5396E867EFB77FEC1CD370CC66AA7" blockId="2.[264,777,283,406]" box="[264,481,380,406]" pageId="2" pageNumber="63">
(
<figureCitation id="137125EB867EFB77FED8CD370CB86AA7" box="[273,415,380,406]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="2" pageNumber="63">Figs. 1, 4–5</figureCitation>
,
<figureCitation id="137125EB867EFB77FE62CD370CFF6AA7" box="[427,472,380,406]" captionStart-0="FIGURES 7–8" captionStart-1="FIGURE 9" captionStartId-0="5.[264,382,946,970]" captionStartId-1="6.[264,368,1865,1889]" captionTargetBox-0="[276,1308,446,934]" captionTargetId-0="figure@5.[264,1323,429,940]" captionTargetId-1="figure@6.[470,1118,846,1841]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 7–8. 7, lateral view of pronotum of female A. matiganae. 8, dorsal view of pronotum of female A. matiganae." captionText-1="FIGURE 9. Distribution (by county) of A. matiganae." httpUri-0="https://zenodo.org/record/2646035/files/figure.png" httpUri-1="https://zenodo.org/record/3387138/files/figure.png" pageId="2" pageNumber="63">7–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3506AE5867EFB77FEC1CDF20C496E2A" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="8BF5396E867EFB77FEC1CDF20C8E6AE3" blockId="2.[264,1323,441,1307]" box="[264,425,441,466]" pageId="2" pageNumber="63">
<emphasis id="B93EE57C867EFB77FEC1CDF20C8E6AE3" box="[264,425,441,466]" italics="true" pageId="2" pageNumber="63">Type material</emphasis>
</paragraph>
<paragraph id="8BF5396E867EFB77FEF1CDAA0F946FFA" blockId="2.[264,1323,441,1307]" pageId="2" pageNumber="63">
<materialsCitation id="3B223333867EFB77FEF1CDAA0E77697A" ID-GBIF-Occurrence="2236572918" collectingDate="2004-09-05" collectionCode="UNSM, NEBR" pageId="2" pageNumber="63" specimenCount="3" typeStatus="holotype">
<typeStatus id="54F187CC867EFB77FEF1CDAA0C826ACA" box="[312,421,481,507]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5BA1AB867EFB77FE3ECDAA0F686ACA" box="[503,591,481,507]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2" pageNumber="63">UNSM</collectionCode>
) labeled as: a) “
<collectionCode id="ED5BA1AB867EFB77FCD8CDAA0E416ACA" box="[785,870,481,507]" pageId="2" pageNumber="63">NEBR</collectionCode>
: ARTHUR Co. / Arapaho Prairie Nat. Con. / Reserve, in flight at dusk /
<collectingDate id="EFB0E646867EFB77FD56CE420E3D6912" box="[671,794,521,547]" pageId="2" pageNumber="63" value="2004-09-05">
<date id="FFF41FAE867EFB77FD56CE420E3D6912" box="[671,794,521,547]" pageId="2" pageNumber="63" value="2004-09-05">5­IX­2004</date>
</collectingDate>
MJ Paulsen”; b)
<typeStatus id="54F187CC867EFB77FC25CE4209756912" box="[1004,1106,521,547]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
label (red paper): “
<emphasis id="B93EE57C867EFB77FEDCCE7A0F38697B" box="[277,543,561,586]" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="4C4A42ED867EFB77FEDCCE7A0F2C697B" authorityName="Paulsen" authorityYear="2006" box="[277,523,561,586]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="matiganae">Aphodius matiganae</taxonomicName>
/
</emphasis>
Paulsen /
<typeStatus id="54F187CC867EFB77FD6CCE7A0E60697A" box="[677,839,561,587]" pageId="2" pageNumber="63" type="holotype">HOLOTYPE</typeStatus>
”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FCA2CE7A0EE369AA" ID-GBIF-Occurrence="2236572916" collectingDate="1987-08-11" collectionCode="UNSM" collectorName="M. E. Jameson" pageId="2" pageNumber="63" specimenCount="3" typeStatus="allotype">
<typeStatus id="54F187CC867EFB77FCA2CE7A0EF6697A" box="[875,977,561,587]" pageId="2" pageNumber="63" type="allotype">Allotype</typeStatus>
female (
<collectionCode id="ED5BA1AB867EFB77FB89CE7A09BF697A" box="[1088,1176,561,587]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2" pageNumber="63">UNSM</collectionCode>
) labeled: a) “NE: ARTHUR Co. / Arapaho Prairie /
<collectingDate id="EFB0E646867EFB77FD20CE120EA16942" box="[745,902,601,627]" pageId="2" pageNumber="63" value="1987-08-11">
<date id="FFF41FAE867EFB77FD20CE120EA16942" box="[745,902,601,627]" pageId="2" pageNumber="63" value="1987-08-11">VIII­11 1987</date>
</collectingDate>
/
<collectorName id="26BF5CB8867EFB77FC69CE1209776942" box="[928,1104,601,627]" pageId="2" pageNumber="63">M.E. Jameson</collectorName>
”; b)
<typeStatus id="54F187CC867EFB77FB43CE1209CF6942" box="[1162,1256,601,627]" pageId="2" pageNumber="63" type="allotype">allotype</typeStatus>
label (red paper): “
<emphasis id="B93EE57C867EFB77FE6CCECA0F8369AB" box="[421,676,641,666]" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="4C4A42ED867EFB77FE6CCECA0FB269AB" authorityName="Paulsen" authorityYear="2006" box="[421,661,641,666]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="matiganae">Aphodius matiganae</taxonomicName>
/
</emphasis>
Paulsen /
<typeStatus id="54F187CC867EFB77FCD5CECA0E9D69AA" box="[796,954,641,667]" pageId="2" pageNumber="63" type="allotype">ALLOTYPE</typeStatus>
”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FC07CECA0F00680A" ID-GBIF-Occurrence="2236572911" collectingDate="1970-08" collectingDateMax="2004-09-05" collectingDateMin="1970-08" collectionCode="NEBR" county="Custer Co." location="Same" municipality="Male" pageId="2" pageNumber="63" specimenCount="16" specimenCount-female="3" specimenCount-male="13" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FC07CECA096769AA" box="[974,1088,641,667]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D4CF2E7867EFB77FB99CECA099769AA" box="[1104,1200,641,667]" pageId="2" pageNumber="63" type="male">13 male</specimenCount>
,
<specimenCount id="9D4CF2E7867EFB77FB72CECA080569AA" box="[1211,1314,641,667]" pageId="2" pageNumber="63" type="female">3 female</specimenCount>
) labeled:
<location id="8E956FB5867EFB77FEB8CEE20C9569F2" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB5867EFB77FEB8CEE20C9569F2" box="[369,434,681,707]" county="Custer Co." municipality="Male" name="Same" pageId="2" pageNumber="63">Same</location>
data as
<typeStatus id="54F187CC867EFB77FDEBCEE20FAB69F2" box="[546,652,681,707]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
.
<location id="8E956FB5867EFB77FD63CEE20FC169F2" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB5867EFB77FD63CEE20FC169F2" box="[682,742,681,707]" county="Custer Co." municipality="Male" name="Male" pageId="2" pageNumber="63">Male</location>
<typeStatus id="54F187CC867EFB77FD3ACEE20E7F69F2" box="[755,856,681,707]" pageId="2" pageNumber="63" type="paratype">paratype</typeStatus>
labeled: “Milburn /
<collectingCounty id="629441E2867EFB77FB94CEE209C769F2" box="[1117,1248,681,707]" pageId="2" pageNumber="63">Custer Co.</collectingCounty>
/ NE BRASKA /
<collectingDate id="EFB0E646867EFB77FE6ECE9A0F0369DA" box="[423,548,721,747]" pageId="2" pageNumber="63" value="1970-08">
<date id="FFF41FAE867EFB77FE6ECE9A0F0369DA" box="[423,548,721,747]" pageId="2" pageNumber="63" value="1970-08">VIII­1970</date>
</collectingDate>
”.
<collectingMunicipality id="6B91A314867EFB77FD9BCE9A0FA969DA" box="[594,654,721,747]" pageId="2" pageNumber="63">Male</collectingMunicipality>
and disarticulated female
<typeStatus id="54F187CC867EFB77FC2CCE9A096D69DA" box="[997,1098,721,747]" pageId="2" pageNumber="63" type="paratype">paratype</typeStatus>
labeled: “
<collectionCode id="ED5BA1AB867EFB77FB18CE9A080169DA" box="[1233,1318,721,747]" pageId="2" pageNumber="63">NEBR</collectionCode>
: ARTHUR Co. / Arapaho Prairie, ex /
<taxonomicName id="4C4A42ED867EFB77FD10CEB20E106823" authorityName="Rafinesque" authorityYear="1817" box="[729,823,761,786]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="2" pageNumber="63" phylum="Chordata" rank="genus">
<emphasis id="B93EE57C867EFB77FD10CEB20E106823" box="[729,823,761,786]" italics="true" pageId="2" pageNumber="63">Geomys</emphasis>
</taxonomicName>
burrow pitfall / (became unburied); set /
<collectingDate id="EFB0E646867EFB77FEC1CF6A0CA5680A" box="[264,386,801,827]" pageId="2" pageNumber="63" value="2004-09-05">
<date id="FFF41FAE867EFB77FEC1CF6A0CA5680A" box="[264,386,801,827]" pageId="2" pageNumber="63" value="2004-09-05">5.IX.2004</date>
</collectingDate>
; MJ Paulsen”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FDF2CF6A0F5068BA" ID-GBIF-Occurrence="2236572914" collectingDate="1996-10-05" collectionCode="UV" collectorName="E. G. Riley &amp; Det. R. Gordon" county="Bastrop Co." determinerName="Det. R. Gordon" location="St. Pk." municipality="Bastrop St." pageId="2" pageNumber="63" specimenCount="19" specimenCount-female="8" specimenCount-male="11" stateProvince="Texas" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FDF2CF6A0F8A680A" box="[571,685,801,827]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D4CF2E7867EFB77FD77CF6A0E0E680A" box="[702,809,801,827]" pageId="2" pageNumber="63" type="male">11 males</specimenCount>
,
<specimenCount id="9D4CF2E7867EFB77FCFFCF6A0E8E680A" box="[822,937,801,827]" pageId="2" pageNumber="63" type="female">8 females</specimenCount>
) labeled: a) “
<collectingRegion id="498EF78C867EFB77FB8FCF6A09B6680A" box="[1094,1169,801,827]" country="United States of America" name="Texas" pageId="2" pageNumber="63">Texas</collectingRegion>
:
<collectingCounty id="629441E2867EFB77FB55CF6A080E680A" box="[1180,1321,801,827]" pageId="2" pageNumber="63">Bastrop Co.</collectingCounty>
/
<collectingMunicipality id="6B91A314867EFB77FED7CF020C816852" box="[286,422,841,867]" pageId="2" pageNumber="63">
Bastrop
<location id="8E956FB5867EFB77FE4ECF020C816852" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB5867EFB77FE4ECF020C816852" box="[391,422,841,867]" county="Bastrop Co." municipality="Bastrop St." name="St. Pk." pageId="2" pageNumber="63" stateProvince="Texas">St.</location>
</collectingMunicipality>
Pk. /
<collectingDate id="EFB0E646867EFB77FE37CF020F546852" box="[510,627,841,867]" pageId="2" pageNumber="63" value="1996-10-05">
<date id="FFF41FAE867EFB77FE37CF020F546852" box="[510,627,841,867]" pageId="2" pageNumber="63" value="1996-10-05">X­5­1996</date>
</collectingDate>
,
<collectionCode id="ED5BA1AB867EFB77FD4CCF020F976852" box="[645,688,841,867]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="2" pageNumber="63">UV</collectionCode>
light /
<collectorName id="26BF5CB8867EFB77FCDECF020EB06852" box="[791,919,841,867]" pageId="2" pageNumber="63">E.G. Riley</collectorName>
­395&quot;; b) “
<taxonomicName id="4C4A42ED867EFB77FBE5CF0209866852" authorityName="Illiger" authorityYear="1789" box="[1068,1185,841,867]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="genus">Aphodius</taxonomicName>
/
<taxonomicName id="4C4A42ED867EFB77FB0CCF020C5268BA" authority="Beauvois" authorityName="Beauvois" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="rubeolus">rubeolus Beauvois</taxonomicName>
/
<collectorName id="26BF5CB8867EFB77FE45CF3A0F6568BA" box="[396,578,881,907]" pageId="2" pageNumber="63">
<determinerName id="7E29D5AA867EFB77FE45CF3A0F6568BA" box="[396,578,881,907]" pageId="2" pageNumber="63">Det. R. Gordon</determinerName>
</collectorName>
97”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FD42CF3A0E766882" ID-GBIF-Occurrence="2236572910" collectingDate="1990-09-29" collectorName="E. G. Riley &amp; C. Wolfe" county="Hardin Co." location="Hardin Co." pageId="2" pageNumber="63" specimenCount="1" stateProvince="Texas" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FD42CF3A0FD668BA" box="[651,753,881,907]" pageId="2" pageNumber="63" type="paratype">Paratype</typeStatus>
(male) labeled: “
<collectingRegion id="498EF78C867EFB77FC0DCF3A090D68BA" box="[964,1066,881,907]" country="United States of America" name="Texas" pageId="2" pageNumber="63">TEXAS</collectingRegion>
:
<collectingCounty id="629441E2867EFB77FBFCCF3A099068BA" box="[1077,1207,881,907]" pageId="2" pageNumber="63">Hardin Co.</collectingCounty>
/ I mi. W Silsbee /
<collectingDate id="EFB0E646867EFB77FEB5CFD20F216882" box="[380,518,921,947]" pageId="2" pageNumber="63" value="1990-09-29">
<date id="FFF41FAE867EFB77FEB5CFD20F216882" box="[380,518,921,947]" pageId="2" pageNumber="63" value="1990-09-29">IX­29­1990</date>
</collectingDate>
/
<collectorName id="26BF5CB8867EFB77FDEFCFD20F836882" box="[550,676,921,947]" pageId="2" pageNumber="63">E.G. Riley</collectorName>
&amp;
<collectorName id="26BF5CB8867EFB77FD1ACFD20E6F6882" box="[723,840,921,947]" pageId="2" pageNumber="63">C. Wolfe</collectorName>
”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FCA4CFD2080068EA" ID-GBIF-Occurrence="2236572913" collectingDate="1992-10-24" collectorName="D. W. Sundberg" county="Brooks Co." location="Brooks Co." pageId="2" pageNumber="63" specimenCount="1" stateProvince="Texas" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FCA4CFD20EF46882" box="[877,979,921,947]" pageId="2" pageNumber="63" type="paratype">Paratype</typeStatus>
(female) labeled: “
<collectingRegion id="498EF78C867EFB77FB08CFD208006882" box="[1217,1319,921,947]" country="United States of America" name="Texas" pageId="2" pageNumber="63">TEXAS</collectingRegion>
:
<collectingCounty id="629441E2867EFB77FEC1CF8A0CB468EA" box="[264,403,961,987]" pageId="2" pageNumber="63">Brooks Co.</collectingCounty>
/
<locationDeviation id="9A47204C867EFB77FE7FCF8A0F8468EA" box="[438,675,961,987]" location="Falfurrias" pageId="2" pageNumber="63">
<quantity id="4CB2948B867EFB77FE7FCF8A0F2068EA" box="[438,519,961,987]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="2" pageNumber="63" unit="mi" value="10.0">10 mi.</quantity>
S Falfurrias
</locationDeviation>
/
<collectingDate id="EFB0E646867EFB77FD0CCF8A0E6268EA" box="[709,837,961,987]" pageId="2" pageNumber="63" value="1992-10-24">
<date id="FFF41FAE867EFB77FD0CCF8A0E6268EA" box="[709,837,961,987]" pageId="2" pageNumber="63" value="1992-10-24">24­X­1992</date>
</collectingDate>
/
<collectorName id="26BF5CB8867EFB77FCA1CF8A090568EA" box="[872,1058,961,987]" pageId="2" pageNumber="63">D.W. Sundberg</collectorName>
/
<collectingMethod id="520B4179867EFB77FB8DCF8A083B68EA" box="[1092,1308,961,987]" pageId="2" pageNumber="63">at blacklight trap</collectingMethod>
”
</materialsCitation>
.
<materialsCitation id="3B223333867EFB77FEC1CFA20E4A6F1A" ID-GBIF-Occurrence="2236572909" collectingDate="2004-09-26" collectorName="E. G. Riley" country="United States of America" county="Brazos Co." latitude="30.588333" location="Riley Estate" longLatPrecision="20" longitude="-96.253334" pageId="2" pageNumber="63" specimenCount="1" stateProvince="Texas" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FEC1CFA20C496F32" box="[264,366,1001,1027]" pageId="2" pageNumber="63" type="paratype">Paratype</typeStatus>
(male) labeled: “
<collectingCountry id="F35D79FE867EFB77FD99CFA20FB66F32" box="[592,657,1001,1027]" name="United States of America" pageId="2" pageNumber="63">USA</collectingCountry>
:
<collectingRegion id="498EF78C867EFB77FD6CCFA20E2F6F32" box="[677,776,1001,1027]" country="United States of America" name="Texas" pageId="2" pageNumber="63">TEXAS</collectingRegion>
:
<collectingCounty id="629441E2867EFB77FCD2CFA20E816F32" box="[795,934,1001,1027]" pageId="2" pageNumber="63">Brazos Co.</collectingCounty>
/ College Sta.,
<location id="8E956FB5867EFB77FBB2CFA208366F32" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB5867EFB77FBB2CFA208366F32" box="[1147,1297,1001,1027]" country="United States of America" county="Brazos Co." latitude="30.588333" longLatPrecision="20" longitude="-96.253334" name="Riley Estate" pageId="2" pageNumber="63" stateProvince="Texas">Riley Estate</location>
/
<geoCoordinate id="EE7E5FA9867EFB77FEC1C85A0CB06F1A" box="[264,407,1041,1067]" direction="north" orientation="latitude" pageId="2" pageNumber="63" precision="15" value="30.588333">30º35’18”N</geoCoordinate>
,
<geoCoordinate id="EE7E5FA9867EFB77FE6AC85A0F116F1A" box="[419,566,1041,1067]" direction="west" orientation="longitude" pageId="2" pageNumber="63" precision="15" value="-96.253334">96º15’12”W</geoCoordinate>
/
<collectingDate id="EFB0E646867EFB77FD85C85A0FFE6F1A" box="[588,729,1041,1067]" pageId="2" pageNumber="63" value="2004-09-26">
<date id="FFF41FAE867EFB77FD85C85A0FFE6F1A" box="[588,729,1041,1067]" pageId="2" pageNumber="63" value="2004-09-26">IX­26­2004</date>
</collectingDate>
,
<collectorName id="26BF5CB8867EFB77FD2DC85A0E456F1A" box="[740,866,1041,1067]" pageId="2" pageNumber="63">E.G. Riley</collectorName>
”
</materialsCitation>
.
<typeStatus id="54F187CC867EFB77FCB7C85A0EC36F1A" box="[894,996,1041,1067]" pageId="2" pageNumber="63" type="paratype">Paratype</typeStatus>
(female) same as preceding except date “
<date id="FFF41FAE867EFB77FE57C8720F006F62" box="[414,551,1081,1107]" pageId="2" pageNumber="63" value="2004-10-22">X­22­2004</date>
”.
<typeStatus id="54F187CC867EFB77FD8DC8720F8C6F62" box="[580,683,1081,1107]" pageId="2" pageNumber="63" type="paratype">Paratype</typeStatus>
(female) same as preceding except date “
<date id="FFF41FAE867EFB77FB58C872083D6F62" box="[1169,1306,1081,1107]" pageId="2" pageNumber="63" value="2005-10-30">X­30­2005</date>
”.
<materialsCitation id="3B223333867EFB77FEC1C82A0F886FFA" ID-GBIF-Occurrence="2236572915" collectingDate="2002-11-10" collectingDateMax="2002-11-11" collectingDateMin="2002-11-10" collectorName="D. Hildebrandt" country="United States of America" county="Brooks Co." location="Brooks Co." pageId="2" pageNumber="63" specimenCount="2" specimenCount-female="2" stateProvince="Texas" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FEC1C82A0C5D6F4A" box="[264,378,1121,1147]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D4CF2E7867EFB77FE40C82A0CDB6F4A" box="[393,508,1121,1147]" pageId="2" pageNumber="63" type="female">2 females</specimenCount>
) labeled: “TEX.,
<collectingCounty id="629441E2867EFB77FD03C82A0E686F4A" box="[714,847,1121,1147]" pageId="2" pageNumber="63">Brooks Co.</collectingCounty>
/
<locationDeviation id="9A47204C867EFB77FCACC82A091A6F4A" box="[869,1085,1121,1147]" location="Encino" pageId="2" pageNumber="63">
<quantity id="4CB2948B867EFB77FCACC82A0E8F6F4A" box="[869,936,1121,1147]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="2" pageNumber="63" unit="mi" value="10.0">10 mi</quantity>
N of Encino
</locationDeviation>
/ on Rt. 755 /
<collectingDate id="EFB0E646867EFB77FB2BC82A0CA26F92" pageId="2" pageNumber="63" value="2002-11-10" valueMax="2002-11-11" valueMin="2002-11-10">
<date id="FFF41FAE867EFB77FB2BC82A0CA26F92" pageId="2" pageNumber="63" value="2002-11-10" valueMax="2002-11-11" valueMin="2002-11-10">10–11 Nov 2002</date>
</collectingDate>
/
<collectorName id="26BF5CB8867EFB77FE64C8C20F4B6F92" box="[429,620,1161,1187]" pageId="2" pageNumber="63">D. Hildebrandt</collectorName>
”. All
<typeStatus id="54F187CC867EFB77FD06C8C20E186F92" box="[719,831,1161,1187]" pageId="2" pageNumber="63" type="paratype">paratypes</typeStatus>
labeled (on yellow paper): “
<emphasis id="B93EE57C867EFB77FB75C8C20CB46FFB" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="4C4A42ED867EFB77FB75C8C20CA46FFB" authorityName="Paulsen" authorityYear="2006" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="matiganae">Aphodius matiganae</taxonomicName>
/
</emphasis>
Paulsen /
<typeStatus id="54F187CC867EFB77FDC5C8FA0F826FFA" box="[524,677,1201,1227]" pageId="2" pageNumber="63" type="paratype">PARATYPE</typeStatus>
”
</materialsCitation>
.
</paragraph>
<paragraph id="8BF5396E867EFB77FEF1C8920C496E2A" blockId="2.[264,1323,441,1307]" pageId="2" pageNumber="63">
<materialsCitation id="3B223333867EFB77FEF1C89209D76FC2" ID-GBIF-Occurrence="2236572908" box="[312,1264,1241,1267]" collectionCode="EGRC, MJPC, PESC, PKLC, TAMU, UNSM, USNM" pageId="2" pageNumber="63" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F187CC867EFB77FEF1C8920C8D6FC2" box="[312,426,1241,1267]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
deposited in
<collectionCode id="ED5BA1AB867EFB77FD9AC8920F836FC2" box="[595,676,1241,1267]" pageId="2" pageNumber="63">EGRC</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FD7CC8920E266FC2" box="[693,769,1241,1267]" pageId="2" pageNumber="63">MJPC</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FCD8C8920E7D6FC2" box="[785,858,1241,1267]" pageId="2" pageNumber="63">PESC</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FCA2C8920E9E6FC2" box="[875,953,1241,1267]" pageId="2" pageNumber="63">PKLC</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FC00C89209076FC2" box="[969,1056,1241,1267]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="2" pageNumber="63" type="Herbarium">TAMU</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FBF9C89209AF6FC2" box="[1072,1160,1241,1267]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="2" pageNumber="63">UNSM</collectionCode>
,
<collectionCode id="ED5BA1AB867EFB77FB51C89209D76FC2" box="[1176,1264,1241,1267]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="63" type="Museum">USNM</collectionCode>
</materialsCitation>
, and WBWC.
</paragraph>
</subSubSection>
<subSubSection id="C3506AE5867EFB76FEC1C90C0F226ED0" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63" type="description">
<paragraph id="8BF5396E867EFB77FEC1C90C0CB56E51" blockId="2.[264,1323,1351,1897]" box="[264,402,1351,1376]" pageId="2" pageNumber="63">
<emphasis id="B93EE57C867EFB77FEC1C90C0CB56E51" box="[264,402,1351,1376]" italics="true" pageId="2" pageNumber="63">Description</emphasis>
</paragraph>
<paragraph id="8BF5396E867EFB76FEF1C9240F85680F" blockId="2.[264,1323,1351,1897]" lastBlockId="3.[264,1323,284,830]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
<typeStatus id="54F187CC867EFB77FEF1C9240C826EB8" box="[312,421,1391,1417]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
male.
<emphasis id="B93EE57C867EFB77FDCEC9240F7F6EB9" box="[519,600,1391,1416]" italics="true" pageId="2" pageNumber="63">Length</emphasis>
:
<quantity id="4CB2948B867EFB77FDA5C9240FEA6EB8" box="[620,717,1391,1417]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="63" unit="mm" value="5.2">5.2 mm</quantity>
.
<emphasis id="B93EE57C867EFB77FD15C9240E0F6EB9" box="[732,808,1391,1416]" italics="true" pageId="2" pageNumber="63">Width:</emphasis>
<quantity id="4CB2948B867EFB77FCFFC9240EB06EB8" box="[822,919,1391,1417]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="63" unit="mm" value="2.5">2.5 mm</quantity>
. Form robust, elongate (
<figureCitation id="137125EB867EFB77FB19C924083E6EB8" box="[1232,1305,1391,1417]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
).
<emphasis id="B93EE57C867EFB77FEC1C9DC0C716E81" box="[264,342,1431,1456]" italics="true" pageId="2" pageNumber="63">Color:</emphasis>
Everywhere yellowish red except elytral suture darker, clypeal margin and femora lighter yellowish red, abdomen and antennae lighter reddish yellow.
<emphasis id="B93EE57C867EFB77FBBAC9F4099B6EE9" box="[1139,1212,1471,1496]" italics="true" pageId="2" pageNumber="63">Head:</emphasis>
Clypeus broadly rounded on either side of median emargination, center with tumid transverse ridge (
<figureCitation id="137125EB867EFB77FED8CA440C786D18" box="[273,351,1551,1577]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="2" pageNumber="63">Fig. 4</figureCitation>
), surface punctate; punctures moderate to large. Frontoclypeal suture feebly impressed, weakly trituberculate (best seen in posterodorsal view,
<figureCitation id="137125EB867EFB77FBD3CA7C09446D60" box="[1050,1123,1591,1617]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="2" pageNumber="63">Fig. 4</figureCitation>
). Frons shining, punctate; punctures large, dense, separated by 1 diameter. Maxillary palpus with last segment longer than protibial spur.
<emphasis id="B93EE57C867EFB77FD60CACC0E016D91" box="[681,806,1671,1696]" italics="true" pageId="2" pageNumber="63">Pronotum:</emphasis>
Shape weakly convex, narrowed anteriorly, feebly rounded laterally; lateral and basal margins with bead; lateral margin in lateral view nearly straight (
<figureCitation id="137125EB867EFB77FE01CA9C0F336DC0" box="[456,532,1751,1777]" captionStart="FIGURES 7–8" captionStartId="5.[264,382,946,970]" captionTargetBox="[276,1308,446,934]" captionTargetId="figure@5.[264,1323,429,940]" captionTargetPageId="5" captionText="FIGURES 7–8. 7, lateral view of pronotum of female A. matiganae. 8, dorsal view of pronotum of female A. matiganae." httpUri="https://zenodo.org/record/2646035/files/figure.png" pageId="2" pageNumber="63">Fig. 7</figureCitation>
). Surface smooth, shining, punctate; punctures at center of disc uniform in size, size moderate, separated by 2–4 times puncture diameter (
<figureCitation id="137125EB867EFB77FB48CAB409EE6C28" box="[1153,1225,1791,1817]" captionStart="FIGURES 7–8" captionStartId="5.[264,382,946,970]" captionTargetBox="[276,1308,446,934]" captionTargetId="figure@5.[264,1323,429,940]" captionTargetPageId="5" captionText="FIGURES 7–8. 7, lateral view of pronotum of female A. matiganae. 8, dorsal view of pronotum of female A. matiganae." httpUri="https://zenodo.org/record/2646035/files/figure.png" pageId="2" pageNumber="63">Fig. 8</figureCitation>
); lateral declivity with punctures becoming less uniform, large and moderate punctures present near posterior angles and near base.
<taxonomicName id="4C4A42ED867EFB77FD7ECB040EA66C58" authority=": Shape" authorityName="Shape" box="[695,897,1871,1897]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B93EE57C867EFB77FD7ECB040E0C6C59" box="[695,811,1871,1896]" italics="true" pageId="2" pageNumber="63">Scutellum</emphasis>
: Shape
</taxonomicName>
triangular. Size small (less than 1/6 length of elytra).
<emphasis id="B93EE57C867FFB76FE15CD570F026A04" box="[476,549,284,309]" italics="true" pageId="3" pageNumber="64">Elytra</emphasis>
: Surface shining, intervals almost flat on disc, slightly convex on apical declivity, finely punctate; punctures in 2–3 irregular rows, separated by more than 5 puncture diameters; striae moderately impressed, margins more or less straight (not crenate), width uniform from base to apex, punctate; punctures moderate, separated by 1–2 puncture diameters.
<emphasis id="B93EE57C867FFB76FDC2CDF70F826AE4" box="[523,677,444,469]" italics="true" pageId="3" pageNumber="64">Metasternum</emphasis>
: Center along midline depressed; depression ovoid, longer than wide, shining, punctate; punctures large, pubescent. Anterolateral corners strongly alutaceous, deeply punctate; punctures large, pubescent. Posteromedial surface feebly alutaceous but shining, impunctate, glabrous.
<emphasis id="B93EE57C867FFB76FC4ACE7F0EE3697C" box="[899,964,564,589]" italics="true" pageId="3" pageNumber="64">Legs:</emphasis>
Protibia impunctate on dorsal surface, tridentate, serrate behind basal tooth; apical tooth long, strongly curving to outside. Protibial spur simple, triangular, laterally flattened. Mesotibia with apex fringed with unequal spinules; mesotibial spurs unmodified. Metatibia with apex fringed with more or less equal spinules; spinules extremely short, blunt (apparently worn); metatarsus with basal segment subequal in length to superior spur.
<emphasis id="B93EE57C867FFB76FC68CEB709376824" box="[929,1040,764,789]" italics="true" pageId="3" pageNumber="64">Genitalia</emphasis>
: Parameres convergent, narrowed to rounded apex (
<figureCitation id="137125EB867FFB76FD82CF6F0FB5680F" box="[587,658,804,830]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="3" pageNumber="64">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BF5396E867FFB76FEC1CF240CB568B9" blockId="3.[264,1323,879,1105]" box="[264,402,879,904]" pageId="3" pageNumber="64">
<emphasis id="B93EE57C867FFB76FEC1CF240CB568B9" box="[264,402,879,904]" italics="true" pageId="3" pageNumber="64">Description</emphasis>
</paragraph>
<paragraph id="8BF5396E867FFB76FEF1CFDC0C666F60" blockId="3.[264,1323,879,1105]" pageId="3" pageNumber="64">
<typeStatus id="54F187CC867FFB76FEF1CFDC0CB86880" box="[312,415,919,945]" pageId="3" pageNumber="64" type="allotype">Allotype</typeStatus>
female.
<emphasis id="B93EE57C867FFB76FDDECFDC0F4E6881" box="[535,617,919,944]" italics="true" pageId="3" pageNumber="64">Length</emphasis>
:
<quantity id="4CB2948B867FFB76FDB5CFDC0FFB6880" box="[636,732,919,945]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="64" unit="mm" value="4.6">4.6 mm</quantity>
.
<emphasis id="B93EE57C867FFB76FD25CFDC0E1D6881" box="[748,826,919,944]" italics="true" pageId="3" pageNumber="64">Width:</emphasis>
<quantity id="4CB2948B867FFB76FC8CCFDC0E816880" box="[837,934,919,945]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="64" unit="mm" value="2.2">2.2 mm</quantity>
. Externally similar to
<typeStatus id="54F187CC867FFB76FB0DCFDC080C6880" box="[1220,1323,919,945]" pageId="3" pageNumber="64" type="holotype">holotype</typeStatus>
male except:
<emphasis id="B93EE57C867FFB76FE6ACFF40CCB68E9" box="[419,492,959,984]" italics="true" pageId="3" pageNumber="64">Head:</emphasis>
Clypeus with transverse ridge reduced. Frontoclypeal suture distinctly impressed, trituberculate (best seen in posterodorsal view), tubercles more strongly developed. Maxillary palpus with last segment shorter, subequal in length to protibial spur.
</paragraph>
<paragraph id="8BF5396E867FFB76FEC1C8CC0C556F91" blockId="3.[264,1323,1159,1505]" box="[264,370,1159,1184]" pageId="3" pageNumber="64">
<emphasis id="B93EE57C867FFB76FEC1C8CC0C556F91" box="[264,370,1159,1184]" italics="true" pageId="3" pageNumber="64">Variation</emphasis>
</paragraph>
<paragraph id="8BF5396E867FFB76FEF1C8E40F226ED0" blockId="3.[264,1323,1159,1505]" pageId="3" pageNumber="64">
Male
<typeStatus id="54F187CC867FFB76FEB6C8E40CC86FF8" box="[383,495,1199,1225]" pageId="3" pageNumber="64" type="paratype">paratypes</typeStatus>
(n = 28).
<emphasis id="B93EE57C867FFB76FDB7C8E40FE86FF9" box="[638,719,1199,1224]" italics="true" pageId="3" pageNumber="64">Length</emphasis>
: 3.7 to 5.0 mm.
<emphasis id="B93EE57C867FFB76FC79C8E40ED46FF9" box="[944,1011,1199,1224]" italics="true" pageId="3" pageNumber="64">Width</emphasis>
:
<quantity id="4CB2948B867FFB76FBCEC8E4099D6FF8" box="[1031,1210,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="3" pageNumber="64" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7 to 2.2 mm</quantity>
. Female
<typeStatus id="54F187CC867FFB76FEC1C89C0C5F6FC0" box="[264,376,1239,1265]" pageId="3" pageNumber="64" type="paratype">paratypes</typeStatus>
(n = 17).
<emphasis id="B93EE57C867FFB76FE3BC89C0F636FC1" box="[498,580,1239,1264]" italics="true" pageId="3" pageNumber="64">Length</emphasis>
: 4.0 to
<quantity id="4CB2948B867FFB76FD52C89C0FDF6FC0" box="[667,760,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="64" unit="mm" value="4.6">4.6 mm</quantity>
.
<emphasis id="B93EE57C867FFB76FCC0C89C0E6B6FC1" box="[777,844,1239,1264]" italics="true" pageId="3" pageNumber="64">Width</emphasis>
:
<quantity id="4CB2948B867FFB76FC95C89C09256FC0" box="[860,1026,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="3" pageNumber="64" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8 to 2.2 mm</quantity>
. The tumid clypeal ridge varies from simply punctate to strongly, transversely rugose. Several males possess a more strongly impressed frontoclypeal suture than the
<typeStatus id="54F187CC867FFB76FC51C96C09256E70" box="[920,1026,1319,1345]" pageId="3" pageNumber="64" type="holotype">holotype</typeStatus>
, although it appears that the suture is most evident in females. In addition, the frontoclypeal tubercles are weakly developed in some males but are consistently well developed in the females examined. In many specimens the fringe of the metatibial apex is less worn and the unequal spinule lengths more evident.
</paragraph>
</subSubSection>
<subSubSection id="C3506AE5867FFB70FEC1CA470FA26F65" lastPageId="5" lastPageNumber="66" pageId="3" pageNumber="64" type="diagnosis">
<paragraph id="8BF5396E867FFB76FEC1CA470C586D14" blockId="3.[264,1323,1548,1894]" box="[264,383,1548,1573]" pageId="3" pageNumber="64">
<emphasis id="B93EE57C867FFB76FEC1CA470C586D14" box="[264,383,1548,1573]" italics="true" pageId="3" pageNumber="64">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF5396E867FFB71FEF1CA7F0F5B6917" blockId="3.[264,1323,1548,1894]" lastBlockId="4.[264,1323,284,550]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
This species is most similar to
<taxonomicName id="4C4A42ED867FFB76FD78CA7F0E136D7C" box="[689,820,1588,1613]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C867FFB76FD78CA7F0E136D7C" box="[689,820,1588,1613]" italics="true" pageId="3" pageNumber="64">A. rusicola</emphasis>
</taxonomicName>
(
<figureCitation id="137125EB867FFB76FC8ECA7F0EB66D7F" box="[839,913,1588,1614]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
), one of the most widespread and commonly collected species of North American
<taxonomicName id="4C4A42ED867FFB76FC87CA170E9A6D44" authorityName="Illiger" authorityYear="1789" box="[846,957,1628,1653]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B93EE57C867FFB76FC87CA170E9A6D44" box="[846,957,1628,1653]" italics="true" pageId="3" pageNumber="64">Aphodius</emphasis>
</taxonomicName>
and will key to
<taxonomicName id="4C4A42ED867FFB76FB4ECA170CED6DAF" authority="in Ratcliffe (1991)" authorityName="in Ratcliffe" authorityYear="1991" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C867FFB76FB4ECA17082D6D44" box="[1159,1290,1628,1653]" italics="true" pageId="3" pageNumber="64">A. rusicola</emphasis>
in
<bibRefCitation id="EFDB449F867FFB76FEC1CACF0CED6DAF" author="Ratcliffe, B. C." box="[264,458,1668,1694]" pageId="3" pageNumber="64" pagination="1 - 333" refId="ref2911" refString="Ratcliffe, B. C. (1991) The scarab beetles of Nebraska. Bulletin of the University of Nebraska State Museum, 12, 1 - 333." type="journal article" year="1991">Ratcliffe (1991)</bibRefCitation>
</taxonomicName>
. The yellowish red color and uniformly, moderately punctate pronotal disc characterize
<taxonomicName id="4C4A42ED867FFB76FE1ACAE70F496DF4" authorityName="Paulsen" authorityYear="2006" box="[467,622,1708,1733]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C867FFB76FE1ACAE70F496DF4" box="[467,622,1708,1733]" italics="true" pageId="3" pageNumber="64">A. matiganae</emphasis>
</taxonomicName>
. Large pronotal punctures are present in
<taxonomicName id="4C4A42ED867FFB76FB9DCAE709C86DF4" authorityName="Paulsen" authorityYear="2006" box="[1108,1263,1708,1733]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C867FFB76FB9DCAE709C86DF4" box="[1108,1263,1708,1733]" italics="true" pageId="3" pageNumber="64">A. matiganae</emphasis>
</taxonomicName>
only near the posterior angles and rarely near the basal margin. The usually dark reddish­brown to piceous
<taxonomicName id="4C4A42ED867FFB76FE43CAB70F2B6C24" authorityName="in Ratcliffe" authorityYear="1991" box="[394,524,1788,1813]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C867FFB76FE43CAB70F2B6C24" box="[394,524,1788,1813]" italics="true" pageId="3" pageNumber="64">A. rusicola</emphasis>
</taxonomicName>
frequently has large punctures mixed among the smaller punctures on the pronotal disc. Despite their overall external similarities, these two species possess distinct male genitalia; the parameres of
<taxonomicName id="4C4A42ED867FFB76FD3DCB070EB46C54" authorityName="Paulsen" authorityYear="2006" box="[756,915,1868,1893]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C867FFB76FD3DCB070EB46C54" box="[756,915,1868,1893]" italics="true" pageId="3" pageNumber="64">A. matiganae</emphasis>
</taxonomicName>
are narrow and convergent to the apex (
<figureCitation id="137125EB8678FB71FE9ACD570CB96A07" box="[339,414,284,310]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="4" pageNumber="65">Fig. 5</figureCitation>
), while those of
<taxonomicName id="4C4A42ED8678FB71FDBBCD570FD16A04" authorityName="in Ratcliffe" authorityYear="1991" box="[626,758,284,309]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C8678FB71FDBBCD570FD16A04" box="[626,758,284,309]" italics="true" pageId="4" pageNumber="65">A. rusicola</emphasis>
</taxonomicName>
are broad with sub­parallel sides (
<figureCitation id="137125EB8678FB71FB68CD5709CA6A07" box="[1185,1261,284,310]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="4" pageNumber="65">Fig. 6</figureCitation>
). In addition, the variably developed, transverse ridge of the clypeus (best seen in dorsal view) and less convex pronotum and elytra of
<taxonomicName id="4C4A42ED8678FB71FD3BCD270EB56AB4" authorityName="Paulsen" authorityYear="2006" box="[754,914,364,389]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C8678FB71FD3BCD270EB56AB4" box="[754,914,364,389]" italics="true" pageId="4" pageNumber="65">A. matiganae</emphasis>
</taxonomicName>
will separate the two species. In lateral view, the pronotal margin appears almost straight in
<emphasis id="B93EE57C8678FB71FC25CDDF09B36A9C" box="[1004,1172,404,429]" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C4A42ED8678FB71FC25CDDF09B66A9C" authorityName="Paulsen" authorityYear="2006" box="[1004,1169,404,429]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
,
</emphasis>
while in
<taxonomicName id="4C4A42ED8678FB71FAD8CDDF0C4F6AE4" authorityName="in Ratcliffe" authorityYear="1991" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C8678FB71FAD8CDDF0C4F6AE4" italics="true" pageId="4" pageNumber="65">A. rusicola</emphasis>
</taxonomicName>
the margin is curved. Due to their coloration, specimens of
<taxonomicName id="4C4A42ED8678FB71FB86CDF709C96AE4" authorityName="Paulsen" authorityYear="2006" box="[1103,1262,444,469]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C8678FB71FB86CDF709C96AE4" box="[1103,1262,444,469]" italics="true" pageId="4" pageNumber="65">A. matiganae</emphasis>
</taxonomicName>
may also be confused with
<taxonomicName id="4C4A42ED8678FB71FDEACDAF0F976ACC" box="[547,688,484,509]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rubeolus">
<emphasis id="B93EE57C8678FB71FDEACDAF0F976ACC" box="[547,688,484,509]" italics="true" pageId="4" pageNumber="65">A. rubeolus</emphasis>
</taxonomicName>
(
<figureCitation id="137125EB8678FB71FD0DCDAF0E376ACF" box="[708,784,484,510]" captionStart="FIGURES 1–6" captionStartId="4.[264,382,1722,1746]" captionTargetBox="[270,1309,593,1686]" captionTargetId="figure@4.[264,1323,590,1696]" captionTargetPageId="4" captionText="FIGURES 1–6. 1, Aphodius matiganae, n.sp., dorsal habitus. 2, A. rusicola Melsheimer, dorsal habitus. 3, A. rubeolus Palisot de Beauvois, dorsal habitus. 4, head and pronotum of A. matiganae, showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of A. matiganae, with narrowed apex. 6, male parameres of A. rusicola, with apices widened." httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
); however, the latter species has the dorsal surface of the protibia punctate.
</paragraph>
<caption id="DF3569E68678FB71FEC1CAF109976C0B" httpUri="https://zenodo.org/record/2646033/files/figure.png" pageId="4" pageNumber="65" startId="4.[264,382,1722,1746]" targetBox="[270,1309,593,1686]" targetPageId="4">
<paragraph id="8BF5396E8678FB71FEC1CAF109976C0B" blockId="4.[264,1323,1722,1850]" pageId="4" pageNumber="65">
<emphasis id="B93EE57C8678FB71FEC1CAF10C9F6DE3" bold="true" box="[264,440,1722,1746]" pageId="4" pageNumber="65">FIGURES 1–6.</emphasis>
1,
<emphasis id="B93EE57C8678FB71FE16CAF00FE36DE3" box="[479,708,1723,1746]" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C4A42ED8678FB71FE16CAF00FE66DE3" authority="Paulsen, 2006" authorityName="Paulsen" authorityYear="2006" box="[479,705,1723,1746]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae" status="sp. nov.">Aphodius matiganae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A20D58078678FB71FD06CAF00E246DE3" box="[719,771,1723,1746]" pageId="4" pageNumber="65" rank="species">n.sp.</taxonomicNameLabel>
, dorsal habitus. 2,
<taxonomicName id="4C4A42ED8678FB71FC11CAF009FB6DE3" authority="Melsheimer" authorityName="Melsheimer" box="[984,1244,1723,1746]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C8678FB71FC11CAF009776DE3" box="[984,1104,1723,1746]" italics="true" pageId="4" pageNumber="65">A. rusicola</emphasis>
Melsheimer
</taxonomicName>
, dorsal habitus. 3,
<taxonomicName id="4C4A42ED8678FB71FEB7CA960FFF6DC5" authority="Palisot de Beauvois" authorityName="Palisot de Beauvois" box="[382,728,1757,1780]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rubeolus">
<emphasis id="B93EE57C8678FB71FEB7CA960CDD6DC5" box="[382,506,1757,1780]" italics="true" pageId="4" pageNumber="65">A. rubeolus</emphasis>
Palisot de Beauvois
</taxonomicName>
, dorsal habitus. 4, head and pronotum of
<emphasis id="B93EE57C8678FB71FB5CCA96080C6DC5" box="[1173,1323,1757,1780]" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C4A42ED8678FB71FB5CCA9608016DC5" authorityName="Paulsen" authorityYear="2006" box="[1173,1318,1757,1780]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
,
</emphasis>
showing transverse clypeal ridge and uniformly punctate pronotal disc. 5, male parameres of
<emphasis id="B93EE57C8678FB71FADDCB4B0C586C0B" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C4A42ED8678FB71FADDCB4B0C5C6C0B" authorityName="Paulsen" authorityYear="2006" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
,
</emphasis>
with narrowed apex. 6, male parameres of
<emphasis id="B93EE57C8678FB71FC84CB680EEE6C0B" box="[845,969,1827,1850]" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C4A42ED8678FB71FC84CB680EE26C0B" box="[845,965,1827,1850]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="rusicola">A. rusicola</taxonomicName>
,
</emphasis>
with apices widened.
</paragraph>
</caption>
<paragraph id="8BF5396E8679FB70FEF1CD570CD06AB7" blockId="5.[264,1323,284,390]" pageId="5" pageNumber="66">
The most apparent sexually dimorphic character of
<taxonomicName id="4C4A42ED8679FB70FC58CD57090A6A04" authorityName="Paulsen" authorityYear="2006" box="[913,1069,284,309]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C8679FB70FC58CD57090A6A04" box="[913,1069,284,309]" italics="true" pageId="5" pageNumber="66">A. matiganae</emphasis>
</taxonomicName>
is the form of the last segment of the maxillary palpus: long and cylindrical in males, shorter and weakly dilated medially in females.
</paragraph>
<caption id="DF3569E68679FB70FEC1CFF90CCA68C0" httpUri="https://zenodo.org/record/2646035/files/figure.png" pageId="5" pageNumber="66" startId="5.[264,382,946,970]" targetBox="[276,1308,446,934]" targetPageId="5">
<paragraph id="8BF5396E8679FB70FEC1CFF90CCA68C0" blockId="5.[264,1322,946,1009]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FEC1CFF90C9468FB" bold="true" box="[264,435,946,970]" pageId="5" pageNumber="66">FIGURES 7–8.</emphasis>
7, lateral view of pronotum of female
<emphasis id="B93EE57C8679FB70FC9ACFF80ECF68FB" box="[851,1000,947,970]" italics="true" pageId="5" pageNumber="66">
<taxonomicName id="4C4A42ED8679FB70FC9ACFF80EC268FB" authorityName="Paulsen" authorityYear="2006" box="[851,997,947,970]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
.
</emphasis>
8, dorsal view of pronotum of female
<taxonomicName id="4C4A42ED8679FB70FE9ECF910CCE68C0" authorityName="Paulsen" authorityYear="2006" box="[343,489,986,1009]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF5396E8679FB70FEC1C8700FA26F65" blockId="5.[264,1323,1083,1309]" box="[264,645,1083,1108]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FEC1C8700FA26F65" box="[264,645,1083,1108]" italics="true" pageId="5" pageNumber="66">
Geographic distribution (
<figureCitation id="137125EB8679FB70FDFCC8700F5C6F65" box="[565,635,1083,1108]" captionStart="FIGURE 9" captionStartId="6.[264,368,1865,1889]" captionTargetId="figure@6.[470,1118,846,1841]" captionTargetPageId="6" captionText="FIGURE 9. Distribution (by county) of A. matiganae." httpUri="https://zenodo.org/record/3387138/files/figure.png" pageId="5" pageNumber="66">Fig. 9</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3506AE58679FB70FEF1C8280CAD6ED4" pageId="5" pageNumber="66" type="materials_examined">
<paragraph id="8BF5396E8679FB70FEF1C82809496E2C" blockId="5.[264,1323,1083,1309]" pageId="5" pageNumber="66">
<materialsCitation id="3B2233338679FB70FEF1C82808006F4C" ID-GBIF-Occurrence="2236572917" box="[312,1319,1123,1149]" country="United States of America" county="Arthur Co." location="Custer Co." municipality="Arapaho Prairie" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Nebraska">
<collectingCountry id="F35D79FE8679FB70FEF1C8280F3A6F4C" box="[312,541,1123,1149]" name="United States of America" pageId="5" pageNumber="66">UNITED STATES</collectingCountry>
:
<collectingRegion id="498EF78C8679FB70FDF9C8280F846F4C" box="[560,675,1123,1149]" country="United States of America" name="Nebraska" pageId="5" pageNumber="66">Nebraska</collectingRegion>
:
<collectingCounty id="629441E28679FB70FD7CC8280E1C6F4D" box="[693,827,1123,1148]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FD7CC8280E1C6F4D" box="[693,827,1123,1148]" italics="true" pageId="5" pageNumber="66">Arthur Co.</emphasis>
</collectingCounty>
:
<collectingMunicipality id="6B91A3148679FB70FC9BC82809306F4C" box="[850,1047,1123,1149]" pageId="5" pageNumber="66">Arapaho Prairie</collectingMunicipality>
;
<location id="8E956FB58679FB70FBE3C82809976F4D" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FBE3C82809976F4D" box="[1066,1200,1123,1148]" country="United States of America" county="Arthur Co." municipality="Arapaho Prairie" name="Custer Co." pageId="5" pageNumber="66" stateProvince="Nebraska">
<emphasis id="B93EE57C8679FB70FBE3C82809976F4D" box="[1066,1200,1123,1148]" italics="true" pageId="5" pageNumber="66">Custer Co.</emphasis>
</location>
:
<location id="8E956FB58679FB70FB0FC82808006F4C" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FB0FC82808006F4C" box="[1222,1319,1123,1149]" country="United States of America" county="Arthur Co." municipality="Arapaho Prairie" name="Berwyn" pageId="5" pageNumber="66" stateProvince="Nebraska">Berwyn</location>
</materialsCitation>
.
<materialsCitation id="3B2233338679FB70FEC1C8C00E226FC4" ID-GBIF-Occurrence="2236572912" country="United States of America" county="Bastrop Co." location="Brazos Co." municipality="Bastrop State Park" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Texas">
<collectingRegion id="498EF78C8679FB70FEC1C8C00C776F94" box="[264,336,1163,1189]" country="United States of America" name="Texas" pageId="5" pageNumber="66">Texas</collectingRegion>
:
<collectingCounty id="629441E28679FB70FE95C8C00CCE6F95" box="[348,489,1163,1188]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FE95C8C00CCE6F95" box="[348,489,1163,1188]" italics="true" pageId="5" pageNumber="66">Bastrop Co.</emphasis>
</collectingCounty>
:
<collectingMunicipality id="6B91A3148679FB70FE30C8C00FF16F94" box="[505,726,1163,1189]" pageId="5" pageNumber="66">Bastrop State Park</collectingMunicipality>
;
<emphasis id="B93EE57C8679FB70FD2AC8C009116F95" box="[739,1078,1163,1188]" italics="true" pageId="5" pageNumber="66">
<location id="8E956FB58679FB70FD2AC8C00E406F95" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FD2AC8C00E406F95" box="[739,871,1163,1188]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Brazos Co." pageId="5" pageNumber="66" stateProvince="Texas">Brazos Co.</location>
:
<location id="8E956FB58679FB70FCBEC8C009096F95" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FCBEC8C009096F95" box="[887,1070,1163,1188]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="College Station" pageId="5" pageNumber="66" stateProvince="Texas">College Station</location>
;
</emphasis>
<location id="8E956FB58679FB70FBF4C8C009E46F94" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FBF4C8C009E46F94" box="[1085,1219,1163,1189]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Brazos Co." pageId="5" pageNumber="66" stateProvince="Texas">Brazos Co.</location>
:
<location id="8E956FB58679FB70FB06C8C00C796FFC" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FB06C8C00C796FFC" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="College Station" pageId="5" pageNumber="66" stateProvince="Texas">College Station</location>
;
<location id="8E956FB58679FB70FEB1C8F80F2F6FFD" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FEB1C8F80F2F6FFD" box="[376,520,1203,1228]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Brooks Co." pageId="5" pageNumber="66" stateProvince="Texas">
<emphasis id="B93EE57C8679FB70FEB1C8F80F2F6FFD" box="[376,520,1203,1228]" italics="true" pageId="5" pageNumber="66">Brooks Co.</emphasis>
</location>
:
<location id="8E956FB58679FB70FDEFC8F80F5B6FFC" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FDEFC8F80F5B6FFC" box="[550,636,1203,1229]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Encino" pageId="5" pageNumber="66" stateProvince="Texas">Encino</location>
,
<location id="8E956FB58679FB70FD5CC8F80E2C6FFC" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FD5CC8F80E2C6FFC" box="[661,779,1203,1229]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Falfurrias" pageId="5" pageNumber="66" stateProvince="Texas">Falfurrias</location>
;
<location id="8E956FB58679FB70FCEDC8F80E906FFD" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FCEDC8F80E906FFD" box="[804,951,1203,1228]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Hardin Co." pageId="5" pageNumber="66" stateProvince="Texas">
<emphasis id="B93EE57C8679FB70FCEDC8F80E906FFD" box="[804,951,1203,1228]" italics="true" pageId="5" pageNumber="66">Hardin Co.</emphasis>
</location>
:
<location id="8E956FB58679FB70FC1DC8F809AA6FFC" LSID="urn:lsid:plazi:treatment:03E38878867EFB72FEC1CD120EC26E44:8E956FB58679FB70FC1DC8F809AA6FFC" box="[980,1165,1203,1229]" country="United States of America" county="Bastrop Co." municipality="Bastrop State Park" name="Silsbee. Given" pageId="5" pageNumber="66" stateProvince="Texas">Silsbee. Given</location>
the known distribution, this species probably occurs in
</materialsCitation>
<collectingRegion id="498EF78C8679FB70FCC5C8900EA16FC4" box="[780,902,1243,1269]" country="United States of America" name="Oklahoma" pageId="5" pageNumber="66">Oklahoma</collectingRegion>
and
<collectingRegion id="498EF78C8679FB70FC76C89009336FC4" box="[959,1044,1243,1269]" country="United States of America" name="Kansas" pageId="5" pageNumber="66">Kansas</collectingRegion>
as well. No specimens were located in the collection at the Snow Museum, University of
<collectingRegion id="498EF78C8679FB70FBD8C948094E6E2C" box="[1041,1129,1283,1309]" country="United States of America" name="Kansas" pageId="5" pageNumber="66">Kansas</collectingRegion>
.
</paragraph>
<paragraph id="8BF5396E8679FB70FEC1C9180F206E5D" blockId="5.[264,1323,1363,1629]" box="[264,519,1363,1388]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FEC1C9180F206E5D" box="[264,519,1363,1388]" italics="true" pageId="5" pageNumber="66">Temporal distribution</emphasis>
</paragraph>
<paragraph id="8BF5396E8679FB70FEF1C9300CAD6ED4" blockId="5.[264,1323,1363,1629]" pageId="5" pageNumber="66">
August (2), September (21), October (22), November (2). Most
<collectingRegion id="498EF78C8679FB70FBF0C93009806EA4" box="[1081,1191,1403,1429]" country="United States of America" name="Nebraska" pageId="5" pageNumber="66">Nebraska</collectingRegion>
specimens were collected in September. All but two
<collectingRegion id="498EF78C8679FB70FCD5C9E80E476E8C" box="[796,864,1443,1469]" country="United States of America" name="Texas" pageId="5" pageNumber="66">Texas</collectingRegion>
specimens were taken in October or November.
</paragraph>
</subSubSection>
<subSubSection id="C3506AE58679FB70FEC1C9B80ECD6D6C" pageId="5" pageNumber="66" type="etymology">
<paragraph id="8BF5396E8679FB70FEC1C9B80CA26D3D" blockId="5.[264,1323,1363,1629]" box="[264,389,1523,1548]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FEC1C9B80CA26D3D" box="[264,389,1523,1548]" italics="true" pageId="5" pageNumber="66">Etymology</emphasis>
</paragraph>
<paragraph id="8BF5396E8679FB70FEF1CA500ECD6D6C" blockId="5.[264,1323,1363,1629]" pageId="5" pageNumber="66">
This species is named for my niece, Matigan M. Paulsen (Lincoln,
<collectingRegion id="498EF78C8679FB70FB8ACA5009906D04" box="[1091,1207,1563,1589]" country="United States of America" name="Nebraska" pageId="5" pageNumber="66">Nebraska</collectingRegion>
), in honor of her appreciation for nature, including tiny six­legged things.
</paragraph>
</subSubSection>
<subSubSection id="C3506AE58679FB72FEC1CAD80EC26E44" lastPageId="7" lastPageNumber="68" pageId="5" pageNumber="66" type="discussion">
<paragraph id="8BF5396E8679FB70FEC1CAD80C496D9D" blockId="5.[264,1323,1683,1869]" box="[264,366,1683,1708]" pageId="5" pageNumber="66">
<emphasis id="B93EE57C8679FB70FEC1CAD80C496D9D" box="[264,366,1683,1708]" italics="true" pageId="5" pageNumber="66">Remarks</emphasis>
</paragraph>
<paragraph id="8BF5396E8679FB73FEF1CAF00FB46ACF" blockId="5.[264,1323,1683,1869]" lastBlockId="6.[264,1323,284,750]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<taxonomicName id="4C4A42ED8679FB70FEF1CAF00F0B6DE5" authorityName="Paulsen" authorityYear="2006" box="[312,556,1723,1748]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="matiganae">
<emphasis id="B93EE57C8679FB70FEF1CAF00F0B6DE5" box="[312,556,1723,1748]" italics="true" pageId="5" pageNumber="66">Aphodius matiganae</emphasis>
</taxonomicName>
is recorded from the
<collectingRegion id="498EF78C8679FB70FCFFCAF00E826DE4" box="[822,933,1723,1749]" country="United States of America" name="Nebraska" pageId="5" pageNumber="66">Nebraska</collectingRegion>
Sandhills and from
<collectingRegion id="498EF78C8679FB70FB57CAF009C16DE4" box="[1182,1254,1723,1749]" country="United States of America" name="Texas" pageId="5" pageNumber="66">Texas</collectingRegion>
. The majority of
<collectingRegion id="498EF78C8679FB70FE5BCAA80F276DCC" box="[402,512,1763,1789]" country="United States of America" name="Nebraska" pageId="5" pageNumber="66">Nebraska</collectingRegion>
specimens in the
<typeStatus id="54F187CC8679FB70FD1DCAA80E216DCC" box="[724,774,1763,1789]" pageId="5" pageNumber="66">type</typeStatus>
series were collected in flight over short­grass prairie just before sunset. Thus, the life history of this species remains unknown. Two specimens were collected concurrently with the
<typeStatus id="54F187CC8679FB70FCF0CB780E4C6C7C" box="[825,875,1843,1869]" pageId="5" pageNumber="66">type</typeStatus>
series in a dung­baited pitfall trap in a pocket gopher burrow, but I am not convinced that this species is a mammal burrow associate. Another surface species (
<taxonomicName id="4C4A42ED867AFB73FD01CD0F0E9C6A6F" authority="Muller" authorityName="Muller" box="[712,955,324,350]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B93EE57C867AFB73FD01CD0F0E7A6A6C" box="[712,861,324,349]" italics="true" pageId="6" pageNumber="67">A. distinctus</emphasis>
Müller
</taxonomicName>
) was also present in the trap, indicating that the burrow was not completely reburied or had become opened. In
<collectingRegion id="498EF78C867AFB73FEC1CDDF0C5D6A9F" box="[264,378,404,430]" country="United States of America" name="Nebraska" pageId="6" pageNumber="67">Nebraska</collectingRegion>
, no specimens were collected at blacklight traps operating at the same locality less than a half hour after the
<typeStatus id="54F187CC867AFB73FDA5CDF70FB86AE7" box="[620,671,444,470]" pageId="6" pageNumber="67">type</typeStatus>
series was collected. In contrast, the
<collectingRegion id="498EF78C867AFB73FBACCDF7098E6AE7" box="[1125,1193,444,470]" country="United States of America" name="Texas" pageId="6" pageNumber="67">Texas</collectingRegion>
specimens were collected at blacklight traps.
</paragraph>
<paragraph id="8BF5396E867AFB73FEF1CE470E5E69DF" blockId="6.[264,1323,284,750]" pageId="6" pageNumber="67">
Although the name
<taxonomicName id="4C4A42ED867AFB73FDE1CE470F8B6914" box="[552,684,524,549]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C867AFB73FDE1CE470F8B6914" box="[552,684,524,549]" italics="true" pageId="6" pageNumber="67">A. rusicola</emphasis>
</taxonomicName>
has two junior synonyms neither could pertain to
<emphasis id="B93EE57C867AFB73FAD8CE470CAC697C" italics="true" pageId="6" pageNumber="67">
<taxonomicName id="4C4A42ED867AFB73FAD8CE470CA0697C" authorityName="Paulsen" authorityYear="2006" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
.
</emphasis>
Both
<taxonomicName id="4C4A42ED867AFB73FE12CE7F0E3D697F" authority="Harold, 1863" authorityName="Harold" authorityYear="1863" box="[475,794,564,590]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="aurelianus">
<emphasis id="B93EE57C867AFB73FE12CE7F0F5F697C" box="[475,632,564,589]" italics="true" pageId="6" pageNumber="67">A. aurelianus</emphasis>
<bibRefCitation id="EFDB449F867AFB73FDB6CE7F0E3D697F" author="Harold, E. von" box="[639,794,564,590]" pageId="6" pageNumber="67" pagination="326 - 389" refId="ref2886" refString="Harold, E. von (1863) Beitrage zur Kenntnis eingier coprophagen Lamellicornien. Berliner Entomologische Zeitschrift, 7, 326 - 389." type="journal article" year="1863">Harold, 1863</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4A42ED867AFB73FC9DCE7F09AF697F" authority="Haldeman, 1848" authorityName="Haldeman" authorityYear="1848" box="[852,1160,564,590]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B93EE57C867AFB73FC9DCE7F0E99697C" box="[852,958,564,589]" italics="true" pageId="6" pageNumber="67">A. curtus</emphasis>
Haldeman, 1848
</taxonomicName>
are described as black. Furthermore, the description of
<taxonomicName id="4C4A42ED867AFB73FD35CE170EBD6944" authorityName="Harold" authorityYear="1863" box="[764,922,604,629]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="aurelianus">
<emphasis id="B93EE57C867AFB73FD35CE170EBD6944" box="[764,922,604,629]" italics="true" pageId="6" pageNumber="67">A. aurelianus</emphasis>
</taxonomicName>
mentions the lack of a transverse clypeal ridge (
<bibRefCitation id="EFDB449F867AFB73FE67CECF0F6169AF" author="Harold, E. von" box="[430,582,644,670]" pageId="6" pageNumber="67" pagination="326 - 389" refId="ref2886" refString="Harold, E. von (1863) Beitrage zur Kenntnis eingier coprophagen Lamellicornien. Berliner Entomologische Zeitschrift, 7, 326 - 389." type="journal article" year="1863">Harold 1863</bibRefCitation>
), a character that is present in
<emphasis id="B93EE57C867AFB73FC61CECF096B69AC" box="[936,1100,644,669]" italics="true" pageId="6" pageNumber="67">
<taxonomicName id="4C4A42ED867AFB73FC61CECF096069AC" authorityName="Paulsen" authorityYear="2006" box="[936,1095,644,669]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
.
</emphasis>
It should be noted that
<taxonomicName id="4C4A42ED867AFB73FEF4CEE70C9869F4" box="[317,447,684,709]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="rusicola">
<emphasis id="B93EE57C867AFB73FEF4CEE70C9869F4" box="[317,447,684,709]" italics="true" pageId="6" pageNumber="67">A. rusicola</emphasis>
</taxonomicName>
as currently delineated is an extremely variable species and may actually refer to a complex that includes more than one taxon.
</paragraph>
<caption id="DF3569E6867AFB73FEC1CB020E736C51" box="[264,852,1865,1889]" httpUri="https://zenodo.org/record/3387138/files/figure.png" pageId="6" pageNumber="67" targetBox="[470,1118,846,1841]" targetPageId="6">
<paragraph id="8BF5396E867AFB73FEC1CB020E736C51" blockId="6.[264,852,1865,1889]" box="[264,852,1865,1889]" pageId="6" pageNumber="67">
<emphasis id="B93EE57C867AFB73FEC1CB020CAC6C50" bold="true" box="[264,395,1865,1889]" pageId="6" pageNumber="67">FIGURE 9.</emphasis>
Distribution (by county) of
<taxonomicName id="4C4A42ED867AFB73FD77CB020E776C51" authorityName="Paulsen" authorityYear="2006" box="[702,848,1865,1888]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="matiganae">A. matiganae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF5396E867BFB72FEC1CD500CCA6A04" blockId="7.[264,1325,283,625]" box="[264,493,283,309]" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FEC1CD500CCA6A04" bold="true" box="[264,493,283,309]" pageId="7" pageNumber="68">Acknowledgments</emphasis>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CD2C09536940" blockId="7.[264,1325,283,625]" pageId="7" pageNumber="68">
I thank Robert Gordon (Northern Plains Entomology, Willow City, ND) and Paul Skelley (
<collectingRegion id="498EF78C867BFB72FED9CDC40C406A98" box="[272,359,399,425]" country="United States of America" name="Florida" pageId="7" pageNumber="68">Florida</collectingRegion>
State Collection of Arthropods, Gainesville, FL) for their generous assistance and Paul Skelley, Ed Riley, and Paul Lago for providing specimens for study. I also thank Paul Skelley for providing SEM images. Automontaged images were supported, in part, by NSF­DBI 0 500767. This project was supported by an NSF/PEET grant (DEB­0118669) to M. L. Jameson and B. C. Ratcliffe. Support for collecting was provided, in part, by a University of
<collectingRegion id="498EF78C867BFB72FE62CE1C0F3D6940" box="[427,538,599,625]" country="United States of America" name="Nebraska" pageId="7" pageNumber="68">Nebraska</collectingRegion>
Center for Great Plains Studies grant to the author.
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CE840CAD69D8" blockId="7.[264,394,719,745]" box="[264,394,719,745]" pageId="7" pageNumber="68">References</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CF550E076844" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1CF550E076844" pageId="7" pageNumber="68" refId="ref2829" type="journal article">
<author id="53F7D18B867BFB72FEC1CF550CBE6804" box="[264,409,798,821]" case="TC" fns="I" in="R.D." ins="D" ln="Gordon" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Gordon, R.D.</author>
&amp;
<author id="53F7D18B867BFB72FE73CF550F736804" box="[442,596,798,821]" case="TC" fns="I" in="G.A." ins="D" ln="Salsbury" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Salsbury, G.A.</author>
(
<year id="FFE2783D867BFB72FDA8CF550FBE6804" box="[609,665,798,821]" pageId="7" pageNumber="68">1999</year>
)
<title id="F94ED258867BFB72FD6FCF55095E6864" jopScore="0.6410936" pageId="7" pageNumber="68">
Studies on the genus
<emphasis id="B93EE57C867BFB72FC4ECF540ECC6807" box="[903,1003,799,822]" italics="true" pageId="7" pageNumber="68">Aphodius</emphasis>
of the United States and Canada (Coleoptera: Scarabaeidae). IX. A new species from Kansas and Texas
</title>
.
<journalOrPublisher id="518B33A2867BFB72FB46CF740FB46847" jopScore="0.7417716" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FB46CF740FB46847" italics="true" pageId="7" pageNumber="68">Journal of the New York Entomological Society</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FD68CF150FEC6844" box="[673,715,862,885]" pageId="7" pageNumber="68" type="volume">107</part>
,
<pagination id="B283C79A867BFB72FD1FCF150E3B6844" box="[726,796,862,885]" pageId="7" pageNumber="68" type="pagination">64–67</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CF350F946884" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1CF350F946884" pageId="7" pageNumber="68" refId="ref2886" type="journal article">
<author id="53F7D18B867BFB72FEC1CF350C8068A4" box="[264,423,894,917]" case="TC" fns="I" in="E." ins="D" ln="Harold" lnc="TC" npo="LnFn" pageId="7" pageNumber="68" tsw="true">Harold, E. von</author>
(
<year id="FFE2783D867BFB72FE7ECF350CC968A4" box="[439,494,894,917]" pageId="7" pageNumber="68">1863</year>
)
<title id="F94ED258867BFB72FE34CF35095C68A4" box="[509,1147,894,917]" pageId="7" pageNumber="68">Beitrage zur Kenntnis eingier coprophagen Lamellicornien</title>
.
<journalOrPublisher id="518B33A2867BFB72FB47CF340F026887" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FB47CF340F026887" italics="true" pageId="7" pageNumber="68">Berliner Entomologische Zeitschrift</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FDFACFD50F656884" box="[563,578,926,949]" pageId="7" pageNumber="68" type="volume">7</part>
,
<pagination id="B283C79A867BFB72FD87CFD50F886884" box="[590,687,926,949]" pageId="7" pageNumber="68" type="pagination">326–389</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CFF50F3668C4" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1CFF50F3668C4" pageId="7" pageNumber="68" refId="ref2911" type="journal article">
<author id="53F7D18B867BFB72FEC1CFF50C8468E4" box="[264,419,958,981]" case="TC" fns="I" in="B.C." ins="D" ln="Ratcliffe" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Ratcliffe, B.C.</author>
(
<year id="FFE2783D867BFB72FE7BCFF50CCE68E4" box="[434,489,958,981]" pageId="7" pageNumber="68">1991</year>
)
<title id="F94ED258867BFB72FE3ECFF50E6F68E4" box="[503,840,958,981]" pageId="7" pageNumber="68">The scarab beetles of Nebraska</title>
.
<journalOrPublisher id="518B33A2867BFB72FC93CFF40CB568C7" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FC93CFF40CB568C7" italics="true" pageId="7" pageNumber="68">Bulletin of the University of Nebraska State Museum</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FE69CF950C9A68C4" box="[416,445,990,1013]" pageId="7" pageNumber="68" type="volume">12</part>
,
<pagination id="B283C79A867BFB72FE01CF950F2A68C4" box="[456,525,990,1013]" pageId="7" pageNumber="68" type="pagination">1–333</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1CFB50E476F64" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1CFB50E476F64" pageId="7" pageNumber="68" refId="ref2941" type="url">
<author id="53F7D18B867BFB72FEC1CFB50C806F24" box="[264,423,1022,1045]" case="TC" fns="I" in="A." ins="D" ln="Samuelson" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Samuelson, A.</author>
,
<author id="53F7D18B867BFB72FE7BCFB50F196F24" box="[434,574,1022,1045]" case="TC" fns="I" in="N." ins="D" ln="Evenhuis" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Evenhuis, N.</author>
&amp;
<author id="53F7D18B867BFB72FDA1CFB50FC76F24" box="[616,736,1022,1045]" case="TC" fns="I" in="G." ins="D" ln="Nishida" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Nishida, G.</author>
(
<year id="FFE2783D867BFB72FD27CFB50E016F24" box="[750,806,1022,1045]" pageId="7" pageNumber="68">2001</year>
)
<title id="F94ED258867BFB72FCFDCFB40CBE6F07" jopScore="0.14863014" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FCFDCFB40CBE6F07" italics="true" pageId="7" pageNumber="68">The Insect and Spider Collections of the World Web Site</emphasis>
</title>
.
<journalOrPublisher id="518B33A2867BFB72FE71C8550E3E6F04" box="[440,793,1054,1077]" jopScore="2.722966" pageId="7" pageNumber="68">Bishop Museum, Honolulu, HI</journalOrPublisher>
. Available from:
<publicationUrl id="D3C49BA3867BFB72FC27C8550F626F64" pageId="7" pageNumber="68">http://hbs.bishopmuseum.org/ codens/codensearch.html</publicationUrl>
(accessed
<accessDate id="443C3732867BFB72FD70C8750E736F64" box="[697,852,1086,1109]" pageId="7" pageNumber="68">22 AUG 2005</accessDate>
).
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1C8150EE86FA4" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1C8150EE86FA4" pageId="7" pageNumber="68" refId="ref3000" type="journal article">
<author id="53F7D18B867BFB72FEC1C8150CA86F44" box="[264,399,1118,1141]" case="TC" fns="I" in="P.E." ins="D" ln="Skelley" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Skelley, P.E.</author>
&amp;
<author id="53F7D18B867BFB72FE7AC8150F616F44" box="[435,582,1118,1141]" case="TC" fns="I" in="R.D." ins="D" ln="Gordon" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Gordon, R.D.</author>
(
<year id="FFE2783D867BFB72FD9CC8150FAA6F44" box="[597,653,1118,1141]" pageId="7" pageNumber="68">1995</year>
)
<title id="F94ED258867BFB72FD55C8150FBE6FA4" pageId="7" pageNumber="68">
A new species of
<emphasis id="B93EE57C867BFB72FC96C8140EE36F47" box="[863,964,1119,1142]" italics="true" pageId="7" pageNumber="68">Aphodius</emphasis>
(Coleoptera: Scarabaeidae) from Alabama pocket gopher burrows
</title>
.
<journalOrPublisher id="518B33A2867BFB72FD60C8340E656FA7" box="[681,834,1151,1174]" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FD60C8340E656FA7" box="[681,834,1151,1174]" italics="true" pageId="7" pageNumber="68">Insecta Mundi</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FC86C8350E796FA4" box="[847,862,1150,1173]" pageId="7" pageNumber="68" type="volume">9</part>
,
<pagination id="B283C79A867BFB72FCA3C8350EED6FA4" box="[874,970,1150,1173]" pageId="7" pageNumber="68" type="pagination">217–219</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1C8D509636FE4" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1C8D509636FE4" pageId="7" pageNumber="68" refId="ref3041" type="journal article">
<author id="53F7D18B867BFB72FEC1C8D50CA96F84" box="[264,398,1182,1205]" case="TC" fns="I" in="P.E." ins="D" ln="Skelley" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Skelley, P.E.</author>
&amp;
<author id="53F7D18B867BFB72FE71C8D50F6D6F84" box="[440,586,1182,1205]" case="TC" fns="I" in="R.D." ins="D" ln="Gordon" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Gordon, R.D.</author>
(
<year id="FFE2783D867BFB72FD90C8D50FB66F84" box="[601,657,1182,1205]" pageId="7" pageNumber="68">2001</year>
)
<title id="F94ED258867BFB72FD69C8D50E3D6FE4" pageId="7" pageNumber="68">Scarab beetles from pocket gopher burrows in the southeastern United States (Coleoptera: Scarabaeidae)</title>
.
<journalOrPublisher id="518B33A2867BFB72FCE2C8F40EE36FE7" box="[811,964,1215,1238]" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FCE2C8F40EE36FE7" box="[811,964,1215,1238]" italics="true" pageId="7" pageNumber="68">Insecta Mundi</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FC1BC8F50EC86FE4" box="[978,1007,1214,1237]" pageId="7" pageNumber="68" type="volume">15</part>
,
<pagination id="B283C79A867BFB72FC33C8F509676FE4" box="[1018,1088,1214,1237]" pageId="7" pageNumber="68" type="pagination">77–93</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1C89509766E24" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1C89509766E24" pageId="7" pageNumber="68" refId="ref3083" type="journal article">
<author id="53F7D18B867BFB72FEC1C8950CB76FC4" box="[264,400,1246,1269]" case="TC" fns="I" in="P.E." ins="D" ln="Skelley" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Skelley, P.E.</author>
&amp;
<author id="53F7D18B867BFB72FE7FC8950F7B6FC4" box="[438,604,1246,1269]" case="TC" fns="I" in="R.E." ins="D" ln="Woodruff" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Woodruff, R.E.</author>
(
<year id="FFE2783D867BFB72FDA4C8950F836FC4" box="[621,676,1246,1269]" pageId="7" pageNumber="68">1991</year>
)
<title id="F94ED258867BFB72FD7CC8950F986E24" pageId="7" pageNumber="68">
Five new species of
<emphasis id="B93EE57C867BFB72FC50C8940ED96FC7" box="[921,1022,1247,1270]" italics="true" pageId="7" pageNumber="68">Aphodius</emphasis>
(Coleoptera: Scarabaeidae) from Florida pocket gopher burrows
</title>
.
<journalOrPublisher id="518B33A2867BFB72FD19C8B40E916E27" box="[720,950,1279,1302]" pageId="7" pageNumber="68">
<emphasis id="B93EE57C867BFB72FD19C8B40E916E27" box="[720,950,1279,1302]" italics="true" pageId="7" pageNumber="68">Florida Entomologist</emphasis>
</journalOrPublisher>
,
<part id="FFEE93F3867BFB72FC0DC8B50EC66E24" box="[964,993,1278,1301]" pageId="7" pageNumber="68" type="volume">64</part>
,
<pagination id="B283C79A867BFB72FC25C8B5096A6E24" box="[1004,1101,1278,1301]" pageId="7" pageNumber="68" type="pagination">517–536</pagination>
.
</bibRef>
</paragraph>
<paragraph id="8BF5396E867BFB72FEC1C9550EC26E44" blockId="7.[264,1325,798,1398]" pageId="7" pageNumber="68">
<bibRef id="52EAAD98867BFB72FEC1C9550EC26E44" pageId="7" pageNumber="68" refId="ref3124" type="book chapter">
<author id="53F7D18B867BFB72FEC1C9550C846E04" box="[264,419,1310,1333]" case="TC" fns="I" in="Q.D." ins="D" ln="Wheeler" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Wheeler, Q.D.</author>
&amp;
<author id="53F7D18B867BFB72FE0CC9550F736E04" box="[453,596,1310,1333]" case="TC" fns="I" in="N.I." ins="D" ln="Platnick" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Platnick, N.I.</author>
(
<year id="FFE2783D867BFB72FDA0C9550F866E04" box="[617,673,1310,1333]" pageId="7" pageNumber="68">2000</year>
)
<title id="F94ED258867BFB72FD66C9550C546E64" jopScore="0.057375986" pageId="7" pageNumber="68">The phylogenetic species concept (sensu Wheeler and Platnick)</title>
.
<emphasis id="B93EE57C867BFB72FEB7C9740CBA6E67" box="[382,413,1343,1366]" italics="true" pageId="7" pageNumber="68">In:</emphasis>
<editor id="4DD6D16D867BFB72FE6EC9750F636E64" box="[423,580,1342,1365]" case="TC" fns="I" in="Q.D." ins="D" ln="Wheeler" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Wheeler, Q.D.</editor>
&amp;
<editor id="4DD6D16D867BFB72FDBAC9750FFB6E64" box="[627,732,1342,1365]" case="TC" fns="I" in="R." ins="D" ln="Meier" lnc="TC" npo="LnFn" pageId="7" pageNumber="68">Meier, R.</editor>
, (Eds.),
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<emphasis id="B93EE57C867BFB72FCF0C9740CA26E47" italics="true" pageId="7" pageNumber="68">Species Concepts and Phylogenetic Theory, a Debate</emphasis>
</volumeTitle>
.
<journalOrPublisher id="518B33A2867BFB72FE50C9150E4E6E44" box="[409,873,1374,1397]" jopScore="1.7403507" pageId="7" pageNumber="68">Columbia University Press, New York City</journalOrPublisher>
, pp.
<pagination id="B283C79A867BFB72FC52C9150EC56E44" box="[923,994,1374,1397]" pageId="7" pageNumber="68" type="pagination">55–69</pagination>
.
</bibRef>
</paragraph>
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</document>
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