treatments-xml/data/03/D3/FD/03D3FD33FFA0FFE84E8F745BF394F96A.xml

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<mods:title id="593B58DC98CC23FBB3CC6F03DB3603AC">Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, “ Protoneuridae ”), with emphasis on its Brazilian species</mods:title>
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<paragraph id="8BC54C25FFA0FFE94E8F745BF40BFCB3" blockId="2.[151,516,865,891]" box="[151,516,865,891]" pageId="2" pageNumber="203">
<heading id="D08DFB49FFA0FFE94E8F745BF40BFCB3" bold="true" box="[151,516,865,891]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
<emphasis id="B90E9037FFA0FFE94E8F745BF40BFCB3" bold="true" box="[151,516,865,891]" pageId="2" pageNumber="203">
Key to males of 
<taxonomicName id="4C7A37A6FFA0FFE94F47745BF40BFCB3" ID-CoL="6349D" box="[351,516,865,891]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B90E9037FFA0FFE94F47745BF40BFCB3" bold="true" box="[351,516,865,891]" italics="true" pageId="2" pageNumber="203">Epipleoneura</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F7492F36FFBC0" blockId="2.[151,1436,935,1032]" pageId="2" pageNumber="203">A key to females is not included since only half of the females have been described for the known species. Caution should be used in using this key. Identification relies on subtle differences of the epiproct, cerci, and apex of the genital ligula some of which should be examined at high (ca 100x) power and proper lighting for best results.</paragraph>
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<paragraph id="8BC54C25FFA0FFE94E8F730DF394FBA2" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
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1. Cercus in posterior view with an inner-basal branch, lobe or tooth visible on posterior or latero-posterior view (e.g. 
<figureCitation id="134150A0FFA0FFE94B2E730DF37CFB84" box="[1334,1395,1079,1100]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs. 1</figureCitation>
b-c, 3, 7b-c, CT, CIB)..................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F7348F394FB4F" blockId="2.[151,1437,1079,2010]" box="[151,1435,1138,1159]" pageId="2" pageNumber="203">
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1’. Cercus in posterior view without an inner-basal branch or tooth (e. g. 
<figureCitation id="134150A0FFA0FFE94D647348F5B1FB4F" box="[892,958,1138,1159]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Figs. 8</figureCitation>
b, 17bc, 19b, 24c)........................ 3
</keyLead>
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<keyStep id="308E5480FFA0FFE94E8F73B5F394FB34" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F73B5F394FB0A" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F73B5F394FB0A" pageId="2" pageNumber="203">
2. Epiproct in posterior view with a well-developed quadrangular or convexly curved medial lobe on its distal margin (ML, 
<figureCitation id="134150A0FFA0FFE94B7473B5F6E2FB0A" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Figs. 4</figureCitation>
b and 5b), with its apex surpassing the level of poorly developed lateral branches (LB)............................. 4
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F73F0F394FB34" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
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2’. Epiproct in posterior view without a medial lobe on its distal margin (
<figureCitation id="134150A0FFA0FFE94D8E73F0F5D9FB17" box="[918,982,1226,1247]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs. 1</figureCitation>
b–3b, 6b–9b), if present (in some males of 
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<emphasis id="B90E9037FFA0FFE94B9F73F0F734FB34" italics="true" pageId="2" pageNumber="203">E. metallica</emphasis>
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and 
<taxonomicName id="4C7A37A6FFA0FFE94F7173DDF7E7FB34" ID-CoL="6G6VC" box="[361,488,1255,1276]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="B90E9037FFA0FFE94F7173DDF7E7FB34" box="[361,488,1255,1276]" italics="true" pageId="2" pageNumber="203">E. fernandezi</emphasis>
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, figs 6b, 13b), its apex never surpassing the level of lateral branches...................... 5
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<paragraph id="8BC54C25FFA0FFE94E8F723FF394FAD2" blockId="2.[151,1437,1079,2010]" box="[151,1435,1285,1306]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F723FF394FAD2" box="[151,1435,1285,1306]" pageId="2" pageNumber="203">
3. Cercus dorsal branch with an apical hook (
<figureCitation id="134150A0FFA0FFE94C63723FF4B1FAD2" box="[635,702,1285,1306]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Figs. 8</figureCitation>
, 
<figureCitation id="134150A0FFA0FFE94CD0723FF4EEFAD2" box="[712,737,1285,1306]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="2" pageNumber="203">18</figureCitation>
c, 19).................................................. 21
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F7218F393FAFF" blockId="2.[151,1437,1079,2010]" box="[151,1436,1314,1335]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7218F393FAFF" box="[151,1436,1314,1335]" pageId="2" pageNumber="203">
3’. Cercus dorsal branch without an apical hook (
<figureCitation id="134150A0FFA0FFE94C817218F4E6FAFF" box="[665,745,1314,1335]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="2" pageNumber="203">Figs. 17</figureCitation>
, 
<figureCitation id="134150A0FFA0FFE94CEB7218F53EFAFF" box="[755,817,1314,1335]" captionStart-0="FIGURES 22 – 24" captionStart-1="FIGURES 25 – 27" captionStartId-0="14.[151,264,1502,1524]" captionStartId-1="15.[151,264,1645,1667]" captionTargetBox-0="[161,1426,198,1476]" captionTargetBox-1="[158,1436,205,1614]" captionTargetId-0="figure@14.[151,1436,195,1480]" captionTargetId-1="figure@15.[151,1436,195,1623]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." captionText-1="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri-0="https://zenodo.org/record/230982/files/figure.png" httpUri-1="https://zenodo.org/record/230983/files/figure.png" pageId="2" pageNumber="203">22–27</figureCitation>
)................................................ 22
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<keyStep id="308E5480FFA0FFE94E8F7205F394FABA" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F7205F394FA9C" blockId="2.[151,1437,1079,2010]" box="[151,1435,1343,1364]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7205F394FA9C" box="[151,1435,1343,1364]" pageId="2" pageNumber="203">
4. Medial lobe of epiproct in posterior view quadrangular (
<figureCitation id="134150A0FFA0FFE94CE47205F53AFA9C" box="[764,821,1343,1364]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Fig. 5</figureCitation>
b).................................... 
<taxonomicName id="4C7A37A6FFA0FFE94B1E7205F394FA9C" ID-CoL="6G77N" box="[1286,1435,1343,1364]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="capilliformis">
<emphasis id="B90E9037FFA0FFE94B1E7205F394FA9C" box="[1286,1435,1343,1364]" italics="true" pageId="2" pageNumber="203">E. capilliformis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F7267F394FABA" blockId="2.[151,1437,1079,2010]" box="[151,1435,1373,1394]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7267F394FABA" box="[151,1435,1373,1394]" pageId="2" pageNumber="203">
4’. Medial lobe of epiproct in posterior view convexly curved (
<figureCitation id="134150A0FFA0FFE94D057267F559FABA" box="[797,854,1373,1394]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Fig. 4</figureCitation>
b).................................. 
<taxonomicName id="4C7A37A6FFA0FFE94B097267F394FABA" ID-CoL="6FV7C" box="[1297,1435,1373,1394]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="B90E9037FFA0FFE94B097267F394FABA" box="[1297,1435,1373,1394]" italics="true" pageId="2" pageNumber="203">E. demarmelsi</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="308E5480FFA0FFE94E8F7240F394FA64" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F7240F394FA47" blockId="2.[151,1437,1079,2010]" box="[151,1435,1402,1423]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7240F394FA47" box="[151,1435,1402,1423]" pageId="2" pageNumber="203">
5. Epiproct in posterior view with an additional lateral lobe, apical branch short and rounded (
<figureCitation id="134150A0FFA0FFE94A547240F29EFA47" box="[1100,1169,1402,1423]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="2" pageNumber="203">Fig. 12</figureCitation>
b)....... 
<taxonomicName id="4C7A37A6FFA0FFE94AE67240F394FA47" ID-CoL="6FV6X" box="[1278,1435,1402,1423]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B90E9037FFA0FFE94AE67240F394FA47" box="[1278,1435,1402,1423]" italics="true" pageId="2" pageNumber="203">E. venezuelensis</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F72ADF394FA64" blockId="2.[151,1437,1079,2010]" box="[151,1435,1431,1452]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F72ADF394FA64" box="[151,1435,1431,1452]" pageId="2" pageNumber="203">5’. Epiproct in posterior view lacking an additional lateral lobe, apical branch variable................................ 6</keyLead>
</paragraph>
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<keyStep id="308E5480FFA0FFE94E8F728FF394F9CC" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F728FF393FA02" blockId="2.[151,1437,1079,2010]" box="[151,1436,1461,1482]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F728FF393FA02" box="[151,1436,1461,1482]" pageId="2" pageNumber="203">
6. Epiproct in posterior view with a wide (
<figureCitation id="134150A0FFA0FFE94C47728FF4AEFA02" box="[607,673,1461,1482]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Figs. 7</figureCitation>
b–9b, 13b, 21b) or narrow (
<figureCitation id="134150A0FFA0FFE94DB2728FF5EAFA02" box="[938,997,1461,1482]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs.1</figureCitation>
and 
<figureCitation id="134150A0FFA0FFE94A0C728FF223FA02" box="[1044,1068,1461,1482]" captionStart="FIGURES 67 – 69" captionStartId="20.[151,264,856,878]" captionText="FIGURES 67 – 69. Male epiproct, posterior view, morphological variation." pageId="2" pageNumber="203">67</figureCitation>
c–f) basal gap between branches.. 7
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F72E8F394F9CC" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F72E8F394F9CC" pageId="2" pageNumber="203">
6’. Epiproct in posterior view with branches fused basally (
<figureCitation id="134150A0FFA0FFE94D1F72E8F544FA2F" box="[775,843,1490,1511]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs. 2</figureCitation>
b, 3b, 67 a–b) from about half (
<figureCitation id="134150A0FFA0FFE94A9C72E8F2CEFA2F" box="[1156,1217,1490,1511]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Figs.6</figureCitation>
b, 20b) to almost their entire length (
<figureCitation id="134150A0FFA0FFE94F7E72D5F7BAF9CC" box="[358,437,1519,1540]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="2" pageNumber="203">Figs. 10</figureCitation>
b, 11b, 14b, 15b)................................................................... 14
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</paragraph>
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<keyStep id="308E5480FFA0FFE94E8F7137F394F9B2" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F7137F394F9F7" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
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7. Cercus ventral branch small and approximately rounded, with two posterior small teeth visible in inner view, dorsal branch with a big inner apical spine or small hook anteriorly directed (
<figureCitation id="134150A0FFA0FFE94D257110F58CF9F7" box="[829,899,1578,1599]" captionStart="FIGURES 25 – 27" captionStartId="15.[151,264,1645,1667]" captionTargetBox="[158,1436,205,1614]" captionTargetId="figure@15.[151,1436,195,1623]" captionTargetPageId="15" captionText="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri="https://zenodo.org/record/230983/files/figure.png" pageId="2" pageNumber="203">Fig. 25</figureCitation>
a–c,e).......................... 
<taxonomicName id="4C7A37A6FFA0FFE94B197110F394F9F7" ID-CoL="6FV7N" box="[1281,1435,1578,1599]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="albuquerquei">
<emphasis id="B90E9037FFA0FFE94B197110F394F9F7" box="[1281,1435,1578,1599]" italics="true" pageId="2" pageNumber="203">E. albuquerquei</emphasis>
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F717DF394F9B2" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
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7’. Cercus ventral branch comparatively larger, with a clearly visible inner-basal lobe or tooth (i.e. 
<figureCitation id="134150A0FFA0FFE94A95717DF2DEF994" box="[1165,1233,1607,1628]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Figs. 6</figureCitation>
b, 9b), dorsal branch without or with a ventral apical hook ventrally or postero-ventrally directed (i.e. 
<figureCitation id="134150A0FFA0FFE94DFE715FF210F9B2" box="[998,1055,1637,1658]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Fig. 2</figureCitation>
a, 9a)......................... 8
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<keyStep id="308E5480FFA0FFE94E8F71B8F394F97C" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F71B8F394F95F" blockId="2.[151,1437,1079,2010]" box="[151,1435,1666,1687]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F71B8F394F95F" box="[151,1435,1666,1687]" pageId="2" pageNumber="203">
8. Cercus without apical hook (
<figureCitation id="134150A0FFA0FFE94FED71B8F436F95F" box="[501,569,1666,1687]" captionStart="FIGURES 19 – 21" captionStartId="12.[151,264,1717,1739]" captionTargetBox="[158,1429,198,1690]" captionTargetId="figure@12.[151,1436,193,1695]" captionTargetPageId="12" captionText="FIGURES 19 – 21. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 19 and 21 modified from Rácenis (1960) and Machado (1985) respectively." httpUri="https://zenodo.org/record/230981/files/figure.png" pageId="2" pageNumber="203">Fig. 21</figureCitation>
a)........................................................... 
<taxonomicName id="4C7A37A6FFA0FFE94B0F71B8F394F95F" ID-CoL="6FV79" box="[1303,1435,1666,1687]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="waiwaiana">
<emphasis id="B90E9037FFA0FFE94B0F71B8F394F95F" box="[1303,1435,1666,1687]" italics="true" pageId="2" pageNumber="203">E. waiwaiana</emphasis>
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F71A5F394F97C" blockId="2.[151,1437,1079,2010]" box="[151,1435,1695,1716]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F71A5F394F97C" box="[151,1435,1695,1716]" pageId="2" pageNumber="203">
8’. Cercus with apical hook (
<figureCitation id="134150A0FFA0FFE94FCE71A5F43EF97C" box="[470,561,1695,1716]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs. 1–3</figureCitation>
, 
<figureCitation id="134150A0FFA0FFE94C2471A5F445F97C" box="[572,586,1695,1716]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">7</figureCitation>
, 
<figureCitation id="134150A0FFA0FFE94C4C71A5F462F97C" box="[596,621,1695,1716]" captionStart="FIGURES 13 – 15" captionStartId="10.[151,264,1588,1610]" captionTargetBox="[160,1430,205,1536]" captionTargetId="figure@10.[151,1436,195,1546]" captionTargetPageId="10" captionText="FIGURES 13 – 15. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230979/files/figure.png" pageId="2" pageNumber="203">13</figureCitation>
)................................................................. 9
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<paragraph id="8BC54C25FFA0FFE94E8F7187F394F927" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7187F394F927" pageId="2" pageNumber="203">
9. Epiproct in posterior view with branches widely separated at their bases, distance between them greater than half length of epiproct branch (
<figureCitation id="134150A0FFA0FFE94F9C71E0F7B2F927" box="[388,445,1754,1775]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Fig. 6</figureCitation>
b, 7b, 9b, 13b, 68 a–f).............................................................. 10
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F71CDF394F8E2" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F71CDF394F8E2" pageId="2" pageNumber="203">
9’. Epiproct in posterior view with branches slightly separated at their bases, distance between them considerably less than half length of epiproct branch (
<figureCitation id="134150A0FFA0FFE94FC7702FF410F8E2" box="[479,543,1813,1834]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="2" pageNumber="203">Figs. 1</figureCitation>
b, 67c–f)................................................................ 13
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</paragraph>
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<keyStep id="308E5480FFA0FFE94E8F7008F394F8AC" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F7008F394F88F" blockId="2.[151,1437,1079,2010]" box="[151,1435,1842,1863]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7008F394F88F" box="[151,1435,1842,1863]" pageId="2" pageNumber="203">
10. Epiproct branches short and triangular (
<figureCitation id="134150A0FFA0FFE94C457008F498F88F" box="[605,663,1842,1863]" captionStart="FIGURES 4 – 6" captionStartId="7.[151,264,1680,1702]" captionTargetBox="[161,1423,198,1649]" captionTargetId="figure@7.[151,1436,193,1659]" captionTargetPageId="7" captionText="FIGURES 4 – 6. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 4 modified from Garrison et al. (2010). CIB: cercus inner-basal branch. LB: epiproct lateral branches. ML: epiproct medial lobe." httpUri="https://zenodo.org/record/230976/files/figure.png" pageId="2" pageNumber="203">Fig. 6</figureCitation>
b).................................................... 
<taxonomicName id="4C7A37A6FFA0FFE94B047008F394F88F" ID-CoL="6G6VC" box="[1308,1435,1842,1863]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="B90E9037FFA0FFE94B047008F394F88F" box="[1308,1435,1842,1863]" italics="true" pageId="2" pageNumber="203">E. fernandezi</emphasis>
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F7075F394F8AC" blockId="2.[151,1437,1079,2010]" box="[151,1435,1871,1892]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7075F394F8AC" box="[151,1435,1871,1892]" pageId="2" pageNumber="203">
10’ Epiproct branches long and narrow (
<figureCitation id="134150A0FFA0FFE94C277075F48EF8AC" box="[575,641,1871,1892]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Figs. 7</figureCitation>
b, 9b, 13b)...................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA0FFE94E8F7057F394F874" pageId="2" pageNumber="203">
<paragraph id="8BC54C25FFA0FFE94E8F7057F394F857" blockId="2.[151,1437,1079,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F7057F394F857" pageId="2" pageNumber="203">
11. Epiproct branches in posterior view divergent, their apices straight (
<figureCitation id="134150A0FFA0FFE94D7F7057F5B8F84A" box="[871,951,1901,1922]" captionStart="FIGURES 13 – 15" captionStartId="10.[151,264,1588,1610]" captionTargetBox="[160,1430,205,1536]" captionTargetId="figure@10.[151,1436,195,1546]" captionTargetPageId="10" captionText="FIGURES 13 – 15. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230979/files/figure.png" pageId="2" pageNumber="203">Figs. 13</figureCitation>
), bent medially, or rarely convergent (
<figureCitation id="134150A0FFA0FFE94B3A7057F37FF84A" box="[1314,1392,1901,1922]" captionStart="FIGURES 67 – 69" captionStartId="20.[151,264,856,878]" captionText="FIGURES 67 – 69. Male epiproct, posterior view, morphological variation." pageId="2" pageNumber="203">Figs. 68</figureCitation>
e–f)........................................................................................... 
<taxonomicName id="4C7A37A6FFA0FFE94B3F70B0F394F857" ID-CoL="6G77G" box="[1319,1435,1930,1951]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="B90E9037FFA0FFE94B3F70B0F394F857" box="[1319,1435,1930,1951]" italics="true" pageId="2" pageNumber="203">E. metallica</emphasis>
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</paragraph>
<paragraph id="8BC54C25FFA0FFE94E8F709DF394F874" blockId="2.[151,1437,1079,2010]" box="[151,1435,1959,1980]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F709DF394F874" box="[151,1435,1959,1980]" pageId="2" pageNumber="203">
11’. Epiproct branches in posterior view parallel, their tips slightly convergent apically (
<figureCitation id="134150A0FFA0FFE94A1A709DF24BF874" box="[1026,1092,1959,1980]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Figs. 7</figureCitation>
b, 9b)..................... 12
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</paragraph>
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<paragraph id="8BC54C25FFA0FFE94E8F70FFF397F812" blockId="2.[151,1437,1079,2010]" box="[151,1432,1989,2010]" pageId="2" pageNumber="203">
<keyLead id="308BEF10FFA0FFE94E8F70FFF397F812" box="[151,1432,1989,2010]" pageId="2" pageNumber="203">
12. Epiproct branches comparatively long, in posterior view reaching or surpassing cercus ventral margin (
<figureCitation id="134150A0FFA0FFE94AA470FFF2FAF812" box="[1212,1269,1989,2010]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="2" pageNumber="203">Fig. 7</figureCitation>
b).. 
<taxonomicName id="4C7A37A6FFA0FFE94B2D70FFF397F812" ID-CoL="6G77H" box="[1333,1432,1989,2010]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B90E9037FFA0FFE94B2D70FFF397F812" box="[1333,1432,1989,2010]" italics="true" pageId="2" pageNumber="203">E.pereirai</emphasis>
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</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F77ADF394FF64" blockId="3.[151,1437,151,1698]" box="[151,1435,151,172]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F77ADF394FF64" box="[151,1435,151,172]" pageId="3" pageNumber="204">
12’. Epiproct branches comparatively short, in posterior view not reaching cercus ventral margin (
<figureCitation id="134150A0FFA1FFE84A7977ADF295FF64" box="[1121,1178,151,172]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="3" pageNumber="204">Fig. 9</figureCitation>
b).......... 
<taxonomicName id="4C7A37A6FFA1FFE84B2C77ADF394FF64" box="[1332,1435,151,172]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B90E9037FFA1FFE84B2C77ADF394FF64" box="[1332,1435,151,172]" italics="true" pageId="3" pageNumber="204">E. susanae</emphasis>
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<paragraph id="8BC54C25FFA1FFE84E8F778FF394FF2F" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F778FF394FF2F" pageId="3" pageNumber="204">
13. Epiproct branches in posterior view separated by a triangular structure at their bases (
<figureCitation id="134150A0FFA1FFE84A16778FF247FF02" box="[1038,1096,181,202]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Fig. 2</figureCitation>
b, TS), inner-basal branch of cercus triangular, visible in lateral view (
<figureCitation id="134150A0FFA1FFE84C3D77E8F451FF2F" box="[549,606,210,231]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Fig. 2</figureCitation>
a).................................................. 
<taxonomicName id="4C7A37A6FFA1FFE84AD677E8F348FF2F" ID-CoL="6FV7G" box="[1230,1351,210,231]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="B90E9037FFA1FFE84AD677E8F348FF2F" box="[1230,1351,210,231]" italics="true" pageId="3" pageNumber="204">E. machadoi</emphasis>
</taxonomicName>
(in part)
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F77D5F394FEEA" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F77D5F394FEEA" pageId="3" pageNumber="204">
13’. Epiproct branches in posterior view not separated by a triangular structure at their bases (
<figureCitation id="134150A0FFA1FFE84A2D77D5F264FECC" box="[1077,1131,239,260]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Fig. 1</figureCitation>
b), inner-basal branch of cercus not triangular, medially directed and with its apex directed posteriorly, not visible in lateral view (
<figureCitation id="134150A0FFA1FFE84A927637F2B0FEEA" box="[1162,1215,269,290]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Fig. 1</figureCitation>
a)...... 
<taxonomicName id="4C7A37A6FFA1FFE84B057637F394FEEA" ID-CoL="6G6VB" box="[1309,1435,269,290]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="kaxuriana">
<emphasis id="B90E9037FFA1FFE84B057637F394FEEA" box="[1309,1435,269,290]" italics="true" pageId="3" pageNumber="204">E. kaxuriana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7610F393FE94" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7610F393FEF7" blockId="3.[151,1437,151,1698]" box="[151,1436,298,319]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7610F393FEF7" box="[151,1436,298,319]" pageId="3" pageNumber="204">
14. Epiproct branches in posterior view fused along their entire length (
<figureCitation id="134150A0FFA1FFE84D717610F5B7FEF7" box="[873,952,298,319]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="3" pageNumber="204">Figs. 10</figureCitation>
b, 11b, 14b, 15b)....................... 15
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F767DF393FE94" blockId="3.[151,1437,151,1698]" box="[151,1436,327,348]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F767DF393FE94" box="[151,1436,327,348]" pageId="3" pageNumber="204">
14’. Epiproct branches in posterior view fused at their bases or along basal half (
<figureCitation id="134150A0FFA1FFE84DA0767DF5F3FE94" box="[952,1020,327,348]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Figs. 2</figureCitation>
b, 67 a–b)....................... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F765FF394FE5F" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F765FF393FEB2" blockId="3.[151,1437,151,1698]" box="[151,1436,357,378]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F765FF393FEB2" box="[151,1436,357,378]" pageId="3" pageNumber="204">
15. Epiproct in posterior view distinctly longer than wide, apex with two narrow lobes (
<figureCitation id="134150A0FFA1FFE84A1D765FF25BFEB2" box="[1029,1108,357,378]" captionStart="FIGURES 13 – 15" captionStartId="10.[151,264,1588,1610]" captionTargetBox="[160,1430,205,1536]" captionTargetId="figure@10.[151,1436,195,1546]" captionTargetPageId="10" captionText="FIGURES 13 – 15. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230979/files/figure.png" pageId="3" pageNumber="204">Figs. 14</figureCitation>
b, 15b).................. 16
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F76B8F394FE5F" blockId="3.[151,1437,151,1698]" box="[151,1435,386,407]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F76B8F394FE5F" box="[151,1435,386,407]" pageId="3" pageNumber="204">
15’. Epiproct in posterior view about as long as wide, apex with two short, wide rounded lobes (
<figureCitation id="134150A0FFA1FFE84A5676B8F292FE5F" box="[1102,1181,386,407]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="3" pageNumber="204">Figs. 10</figureCitation>
b, 11b)............. 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F76A5F394FDE2" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F76A5F394FE27" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F76A5F394FE27" pageId="3" pageNumber="204">
16. Cercus in posterior view with a narrow mesally directed inner-basal process, its apex directed posteriorly and reaching midline of S10 medially (
<figureCitation id="134150A0FFA1FFE84F9D7687F7C4FE1A" box="[389,459,445,466]" captionStart="FIGURES 13 – 15" captionStartId="10.[151,264,1588,1610]" captionTargetBox="[160,1430,205,1536]" captionTargetId="figure@10.[151,1436,195,1546]" captionTargetPageId="10" captionText="FIGURES 13 – 15. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230979/files/figure.png" pageId="3" pageNumber="204">Fig. 15</figureCitation>
b). Apex of genital ligula with two long and slender apical lobes and a V cleft on its distal margin (
<figureCitation id="134150A0FFA1FFE84B6D7687F6F5FE27" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Fig. 30</figureCitation>
)......................................................................................... 
<taxonomicName id="4C7A37A6FFA1FFE84B3176E0F394FE27" ID-CoL="6G76Z" box="[1321,1435,474,495]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="B90E9037FFA1FFE84B3176E0F394FE27" box="[1321,1435,474,495]" italics="true" pageId="3" pageNumber="204">E. uncinata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F76CDF394FDE2" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F76CDF394FDE2" pageId="3" pageNumber="204">
16’ Cercus in posterior view with a well-developed posteriorly directed inner-basal tooth (
<figureCitation id="134150A0FFA1FFE84A0376CDF26FFDC4" box="[1051,1120,503,524]" captionStart="FIGURES 13 – 15" captionStartId="10.[151,264,1588,1610]" captionTargetBox="[160,1430,205,1536]" captionTargetId="figure@10.[151,1436,195,1546]" captionTargetPageId="10" captionText="FIGURES 13 – 15. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230979/files/figure.png" pageId="3" pageNumber="204">Fig. 14</figureCitation>
b). Genital ligula with two apical quadrangular lobes and a wide medial U cleft on its distal margin (
<figureCitation id="134150A0FFA1FFE84D47752FF5AAFDE2" box="[863,933,533,554]" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Fig. 37</figureCitation>
)................................ 
<taxonomicName id="4C7A37A6FFA1FFE84B23752FF394FDE2" ID-CoL="6FV7B" box="[1339,1435,533,554]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="janirae">
<emphasis id="B90E9037FFA1FFE84B23752FF394FDE2" box="[1339,1435,533,554]" italics="true" pageId="3" pageNumber="204">E. janirae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7508F393FDAC" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7508F393FD8F" blockId="3.[151,1437,151,1698]" box="[151,1436,562,583]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7508F393FD8F" box="[151,1436,562,583]" pageId="3" pageNumber="204">
17. Cercus dorsal branch parallel to main body axis in lateral view, with apex blunt with a small inner spine (
<figureCitation id="134150A0FFA1FFE84AC87508F31AFD8F" box="[1232,1301,562,583]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="3" pageNumber="204">Fig. 10</figureCitation>
a) 
<taxonomicName id="4C7A37A6FFA1FFE84B227508F393FD8F" ID-CoL="6FV6Z" box="[1338,1436,562,583]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="tariana">
<emphasis id="B90E9037FFA1FFE84B227508F393FD8F" box="[1338,1436,562,583]" italics="true" pageId="3" pageNumber="204">E. tariana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F7575F393FDAC" blockId="3.[151,1437,151,1698]" box="[151,1436,591,612]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7575F393FDAC" box="[151,1436,591,612]" pageId="3" pageNumber="204">
17’. Cercus dorsal branch directed upward in lateral view, with an apical hook directed meso-ventrally (
<figureCitation id="134150A0FFA1FFE84A807575F2D3FDAC" box="[1176,1244,591,612]" captionStart="FIGURES 10 – 12" captionStartId="9.[151,264,1621,1643]" captionTargetBox="[160,1426,202,1590]" captionTargetId="figure@9.[151,1436,193,1600]" captionTargetPageId="9" captionText="FIGURES 10 – 12. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 10 modified from Machado (1985)." httpUri="https://zenodo.org/record/230978/files/figure.png" pageId="3" pageNumber="204">Fig. 11</figureCitation>
a)...... 
<taxonomicName id="4C7A37A6FFA1FFE84B267575F393FDAC" ID-CoL="6G77B" box="[1342,1436,591,612]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="lamina">
<emphasis id="B90E9037FFA1FFE84B267575F393FDAC" box="[1342,1436,591,612]" italics="true" pageId="3" pageNumber="204">E. lamina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7557F393FD57" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7557F394FD4A" blockId="3.[151,1437,151,1698]" box="[151,1435,621,642]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7557F394FD4A" box="[151,1435,621,642]" pageId="3" pageNumber="204">
18. Epiproct branches in posterior view fused along ca basal half (
<figureCitation id="134150A0FFA1FFE84D2F7557F589FD4A" box="[823,902,621,642]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="3" pageNumber="204">Figs. 16</figureCitation>
b, 20b)..................................... 19
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F75B0F393FD57" blockId="3.[151,1437,151,1698]" box="[151,1436,650,671]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F75B0F393FD57" box="[151,1436,650,671]" pageId="3" pageNumber="204">
18’. Epiproct branches in posterior view fused at their bases (
<figureCitation id="134150A0FFA1FFE84D1975B0F54BFD57" box="[769,836,650,671]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Figs. 2</figureCitation>
b, 3b).......................................... 20
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F759DF394FC87" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F759DF394FD3F" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F759DF394FD3F" pageId="3" pageNumber="204">
19. Gap between epiproct branches at their apices wider than epiproct base, branches comparatively thick (
<figureCitation id="134150A0FFA1FFE84ADB759DF303FD74" box="[1219,1292,679,700]" captionStart="FIGURES 19 – 21" captionStartId="12.[151,264,1717,1739]" captionTargetBox="[158,1429,198,1690]" captionTargetId="figure@12.[151,1436,193,1695]" captionTargetPageId="12" captionText="FIGURES 19 – 21. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 19 and 21 modified from Rácenis (1960) and Machado (1985) respectively." httpUri="https://zenodo.org/record/230981/files/figure.png" pageId="3" pageNumber="204">Fig. 20</figureCitation>
b). Peduncle of posterior processes of genital ligula long and clearly visible, apex of genital ligula triangular, not bent upward and lacking spines (
<figureCitation id="134150A0FFA1FFE84F3275D8F77FFD3F" box="[298,368,738,759]" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Fig. 36</figureCitation>
).............................................................................. 
<taxonomicName id="4C7A37A6FFA1FFE84B0575D8F394FD3F" ID-CoL="6G6VD" box="[1309,1435,738,759]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B90E9037FFA1FFE84B0575D8F394FD3F" box="[1309,1435,738,759]" italics="true" pageId="3" pageNumber="204">E. humeralis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F75C5F394FC87" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F75C5F394FC87" pageId="3" pageNumber="204">
19’. Gap between epiproct branches at their apices narrower than epiproct base, branches comparatively thin (
<figureCitation id="134150A0FFA1FFE84AC575C5F32BFCDC" box="[1245,1316,767,788]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="3" pageNumber="204">Fig. 16</figureCitation>
b). Peduncle posterior processes of genital ligula short, barely visible. Apex of genital ligula folded upwards, with a lateral spine on each side, posterior process widely rounded in lateral view, (
<figureCitation id="134150A0FFA1FFE84CEB7400F536FC87" box="[755,825,826,847]" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Fig. 43</figureCitation>
)......................................... 
<taxonomicName id="4C7A37A6FFA1FFE84B377400F394FC87" ID-CoL="6G6V9" box="[1327,1435,826,847]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="westfalli">
<emphasis id="B90E9037FFA1FFE84B377400F394FC87" box="[1327,1435,826,847]" italics="true" pageId="3" pageNumber="204">E. westfalli</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F746DF393FC2A" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F746DF394FC6F" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F746DF394FC6F" pageId="3" pageNumber="204">
20. Inner basal margin of cercus with a well-developed triangular tooth clearly visible in lateral view, cercus in lateral view approximately L-shaped. Branches of epiproct comparatively longer, reaching ventral level of cercus inner basal tooth (
<figureCitation id="134150A0FFA1FFE84B73744FF6E1FC6F" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Figs. 2</figureCitation>
a, b) (rarely separated at their bases, also keyed under couplet 11 due to intraspecific variation)...... 
<taxonomicName id="4C7A37A6FFA1FFE84AD774A8F347FC6F" ID-CoL="6FV7G" box="[1231,1352,914,935]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="machadoi">
<emphasis id="B90E9037FFA1FFE84AD774A8F347FC6F" box="[1231,1352,914,935]" italics="true" pageId="3" pageNumber="204">E. machadoi</emphasis>
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(in part)
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</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F7495F393FC2A" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7495F393FC2A" pageId="3" pageNumber="204">
20’. Inner basal margin of cercus with a small rounded tooth, barely visible in lateral view; cercus in lateral view approximately C shaped. Branches of epiproct comparatively shorter, not reaching ventral level of cercus inner basal tooth (
<figureCitation id="134150A0FFA1FFE84AC574F7F319FC2A" box="[1245,1302,973,994]" captionStart="FIGURES 1 – 3" captionStartId="5.[151,264,1656,1678]" captionTargetBox="[154,1428,189,1639]" captionTargetId="figure@5.[143,1429,189,1639]" captionTargetPageId="5" captionText="FIGURES 1 – 3. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. AH: apical hook. DB: cercus dorsal branch. VB: cercus ventral branch. CIB: cercus inner-basal branch. CT: cercus inner-basal tooth. TS: triangular swollen structure." httpUri="https://zenodo.org/record/230975/files/figure.png" pageId="3" pageNumber="204">Fig. 3</figureCitation>
a, b) 
<taxonomicName id="4C7A37A6FFA1FFE84B4D74F7F393FC2A" box="[1365,1436,973,994]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="ottoi">
<emphasis id="B90E9037FFA1FFE84B4D74F7F393FC2A" box="[1365,1436,973,994]" italics="true" pageId="3" pageNumber="204">E. ottoi</emphasis>
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</paragraph>
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<keyStep id="308E5480FFA1FFE84E8F74D0F394FB9F" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F74D0F394FBD4" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F74D0F394FBD4" pageId="3" pageNumber="204">
21. Cercus dorsal branch parallel to main body axis, epiproct branches in posterior view long and slender, widely separated at their bases, divergent, with apex closely appressed against medial surface of cercus (
<figureCitation id="134150A0FFA1FFE84DC1733DF211FBD4" box="[985,1054,1031,1052]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="3" pageNumber="204">Fig. 18</figureCitation>
a)................... 
<taxonomicName id="4C7A37A6FFA1FFE84B05733DF394FBD4" ID-CoL="6G77C" box="[1309,1435,1031,1052]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="fuscaenea">
<emphasis id="B90E9037FFA1FFE84B05733DF394FBD4" box="[1309,1435,1031,1052]" italics="true" pageId="3" pageNumber="204">E. fuscaenea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F731FF394FB9F" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F731FF394FB9F" pageId="3" pageNumber="204">
21’ Cercus dorsal branch directed postero-dorsally, epiproct branches in posterior view relatively short and not widely separated at their bases, if divergent, then branches short and with apex bluntly pointed (
<figureCitation id="134150A0FFA1FFE84DA17378F5F4FB9F" box="[953,1019,1090,1111]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="3" pageNumber="204">Figs. 8</figureCitation>
b, 19b).......................... 23
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7365F394FB5A" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7365F394FBBC" blockId="3.[151,1437,151,1698]" box="[151,1435,1119,1140]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7365F394FBBC" box="[151,1435,1119,1140]" pageId="3" pageNumber="204">
22. Epiproct branches in posterior view adjacent or close together at their bases (
<figureCitation id="134150A0FFA1FFE84DDA7365F21EFBBC" box="[962,1041,1119,1140]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="3" pageNumber="204">Figs. 17</figureCitation>
b, 27 b)....................... 24
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F7347F394FB5A" blockId="3.[151,1437,151,1698]" box="[151,1435,1149,1170]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7347F394FB5A" box="[151,1435,1149,1170]" pageId="3" pageNumber="204">
22’. Epiproct branches in posterior view widely separated at their bases (
<figureCitation id="134150A0FFA1FFE84D767347F5EDFB5A" box="[878,994,1149,1170]" captionStart-0="FIGURES 22 – 24" captionStart-1="FIGURES 25 – 27" captionStartId-0="14.[151,264,1502,1524]" captionStartId-1="15.[151,264,1645,1667]" captionTargetBox-0="[161,1426,198,1476]" captionTargetBox-1="[158,1436,205,1614]" captionTargetId-0="figure@14.[151,1436,195,1480]" captionTargetId-1="figure@15.[151,1436,195,1623]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." captionText-1="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri-0="https://zenodo.org/record/230982/files/figure.png" httpUri-1="https://zenodo.org/record/230983/files/figure.png" pageId="3" pageNumber="204">Figs. 22–26</figureCitation>
)................................. 25
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F73A0F394FACF" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F73A0F394FB04" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F73A0F394FB04" pageId="3" pageNumber="204">
23. Branches of epiproct in posterior view divergent, apex of branches rounded, cleft between them inverted V shaped (
<figureCitation id="134150A0FFA1FFE84B2F73A0F373FB67" box="[1335,1404,1178,1199]" captionStart="FIGURES 19 – 21" captionStartId="12.[151,264,1717,1739]" captionTargetBox="[158,1429,198,1690]" captionTargetId="figure@12.[151,1436,193,1695]" captionTargetPageId="12" captionText="FIGURES 19 – 21. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 19 and 21 modified from Rácenis (1960) and Machado (1985) respectively." httpUri="https://zenodo.org/record/230981/files/figure.png" pageId="3" pageNumber="204">Fig. 19</figureCitation>
b)............................................................................................. 
<taxonomicName id="4C7A37A6FFA1FFE84B27738DF394FB04" ID-CoL="6G6VH" box="[1343,1435,1207,1228]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="pallida">E. pallida</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F73EFF394FACF" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F73EFF394FACF" pageId="3" pageNumber="204">
23’. Branches of epiproct in posterior view parallel or convergent, apex of branches acute, cleft between them inverted U shaped (
<figureCitation id="134150A0FFA1FFE84EF073C8F72FFACF" box="[232,288,1266,1287]" captionStart="FIGURES 7 – 9" captionStartId="8.[151,264,1939,1961]" captionTargetBox="[164,1425,352,1913]" captionTargetId="figure@8.[151,1436,339,1918]" captionTargetPageId="8" captionText="FIGURES 7 – 9. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figure 8 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230977/files/figure.png" pageId="3" pageNumber="204">Fig. 8</figureCitation>
b)..................................................................................... 
<taxonomicName id="4C7A37A6FFA1FFE84B2573C8F394FACF" ID-CoL="6FV7H" box="[1341,1435,1266,1287]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="ocuene">
<emphasis id="B90E9037FFA1FFE84B2573C8F394FACF" box="[1341,1435,1266,1287]" italics="true" pageId="3" pageNumber="204">E. ocuene</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7235F394FA8A" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7235F394FAEC" blockId="3.[151,1437,151,1698]" box="[151,1435,1295,1316]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7235F394FAEC" box="[151,1435,1295,1316]" pageId="3" pageNumber="204">
24. Cercus longer than S10, dorsal branch parallel to main body axis (
<figureCitation id="134150A0FFA1FFE84D437235F590FAEC" box="[859,927,1295,1316]" captionStart="FIGURES 25 – 27" captionStartId="15.[151,264,1645,1667]" captionTargetBox="[158,1436,205,1614]" captionTargetId="figure@15.[151,1436,195,1623]" captionTargetPageId="15" captionText="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri="https://zenodo.org/record/230983/files/figure.png" pageId="3" pageNumber="204">Fig. 27</figureCitation>
a)........................... 
<taxonomicName id="4C7A37A6FFA1FFE84B177235F394FAEC" ID-CoL="6G6VG" box="[1295,1435,1295,1316]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="manauensis">
<emphasis id="B90E9037FFA1FFE84B177235F394FAEC" box="[1295,1435,1295,1316]" italics="true" pageId="3" pageNumber="204">E. manauensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F7217F394FA8A" blockId="3.[151,1437,151,1698]" box="[151,1435,1325,1346]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7217F394FA8A" box="[151,1435,1325,1346]" pageId="3" pageNumber="204">
24’. Cercus shorter than S10, dorsal branch directed postero-dorsally (
<figureCitation id="134150A0FFA1FFE84D4E7217F594FA8A" box="[854,923,1325,1346]" captionStart="FIGURES 16 – 18" captionStartId="11.[151,264,1828,1850]" captionTargetBox="[158,1430,349,1786]" captionTargetId="figure@11.[151,1436,339,1791]" captionTargetPageId="11" captionText="FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth." httpUri="https://zenodo.org/record/230980/files/figure.png" pageId="3" pageNumber="204">Fig. 17</figureCitation>
a)............................... 
<taxonomicName id="4C7A37A6FFA1FFE84B377217F394FA8A" ID-CoL="6G6VF" box="[1327,1435,1325,1346]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="B90E9037FFA1FFE84B377217F394FA8A" box="[1327,1435,1325,1346]" italics="true" pageId="3" pageNumber="204">E. solitaria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7270F394FA7F" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7270F394FAB4" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7270F394FAB4" pageId="3" pageNumber="204">
25. Apical margin of genital ligula segment 
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ectal view very wide and with two triangular lobes directed postero-laterally (
<figureCitation id="134150A0FFA1FFE84B6D7270F6F5FAB4" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Fig. 47</figureCitation>
)......................................................................................... 
<taxonomicName id="4C7A37A6FFA1FFE84B37725DF394FAB4" ID-CoL="6FV7D" box="[1327,1435,1383,1404]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="haroldoi">
<emphasis id="B90E9037FFA1FFE84B37725DF394FAB4" box="[1327,1435,1383,1404]" italics="true" pageId="3" pageNumber="204">E. haroldoi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F72BFF394FA7F" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F72BFF394FA7F" pageId="3" pageNumber="204">
25’. Apical margin of genital ligula segment 
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ectal view comparatively much narrower, without lobes directed postero-laterally (
<figureCitation id="134150A0FFA1FFE84EF07298F753FA7F" box="[232,348,1442,1463]" captionStart="FIGURES 28 – 47" captionStartId="17.[151,264,1969,1991]" captionTargetBox="[233,1339,231,1909]" captionTargetId="figure@17.[212,1378,204,1943]" captionTargetPageId="17" captionText="FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection." httpUri="https://zenodo.org/record/230984/files/figure.png" pageId="3" pageNumber="204">Figs. 45, 46</figureCitation>
, 
<figureCitation id="134150A0FFA1FFE84F7E7298F770FA7F" box="[358,383,1442,1463]" captionStart="FIGURES 48 – 52" captionStartId="18.[151,264,1135,1157]" captionTargetBox="[359,1226,206,1082]" captionTargetId="figure@18.[339,1246,193,1114]" captionTargetPageId="18" captionText="FIGURES 48 – 52. Genital ligula, a: lateral view; b: ectal view. Figures 48 and 51 modified from De Marmels (1989)." httpUri="https://zenodo.org/record/230985/files/figure.png" pageId="3" pageNumber="204">52</figureCitation>
).................................................................................... 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F7285F394F9E4" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F7285F394FA3A" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F7285F394FA3A" pageId="3" pageNumber="204">
26. Cercus ventral branch in lateral view separated by a narrow excavation from dorsal branch and almost parallel to it. Apex of paraproct not reaching level of ventral branch apex (
<figureCitation id="134150A0FFA1FFE84CCF72E7F513FA3A" box="[727,796,1501,1522]" captionStart="FIGURES 22 – 24" captionStartId="14.[151,264,1502,1524]" captionTargetBox="[161,1426,198,1476]" captionTargetId="figure@14.[151,1436,195,1480]" captionTargetPageId="14" captionText="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." httpUri="https://zenodo.org/record/230982/files/figure.png" pageId="3" pageNumber="204">Fig. 23</figureCitation>
b)........................................... 
<taxonomicName id="4C7A37A6FFA1FFE84B2D72E7F394FA3A" ID-CoL="6G6VN" box="[1333,1435,1501,1522]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="angeloi">
<emphasis id="B90E9037FFA1FFE84B2D72E7F394FA3A" box="[1333,1435,1501,1522]" italics="true" pageId="3" pageNumber="204">E. angeloi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F72C0F394F9E4" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F72C0F394F9E4" pageId="3" pageNumber="204">
26’. Cercus ventral branch in lateral view separated by a wide excavation from dorsal branch. Apex of paraproct surpassing level of ventral branch apex (
<figureCitation id="134150A0FFA1FFE84FB3712DF7F4F9E4" box="[427,507,1559,1580]" captionStart="FIGURES 22 – 24" captionStartId="14.[151,264,1502,1524]" captionTargetBox="[161,1426,198,1476]" captionTargetId="figure@14.[151,1436,195,1480]" captionTargetPageId="14" captionText="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." httpUri="https://zenodo.org/record/230982/files/figure.png" pageId="3" pageNumber="204">Figs. 24</figureCitation>
b, 26b)..................................................................... 27
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E5480FFA1FFE84E8F710FF394F96A" pageId="3" pageNumber="204">
<paragraph id="8BC54C25FFA1FFE84E8F710FF394F9AF" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
<keyLead id="308BEF10FFA1FFE84E8F710FF394F9AF" pageId="3" pageNumber="204">
27. Cercus ventral branch forming a ca 45° angle with dorsal branch in lateral view (
<figureCitation id="134150A0FFA1FFE84A3B710FF262F982" box="[1059,1133,1589,1610]" captionStart="FIGURES 22 – 24" captionStartId="14.[151,264,1502,1524]" captionTargetBox="[161,1426,198,1476]" captionTargetId="figure@14.[151,1436,195,1480]" captionTargetPageId="14" captionText="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." httpUri="https://zenodo.org/record/230982/files/figure.png" pageId="3" pageNumber="204">Fig. 24</figureCitation>
b), concave medially; epiproct branches separated at their bases by a distance longer than length of each branch (
<figureCitation id="134150A0FFA1FFE84DE97168F238F9AF" box="[1009,1079,1618,1639]" captionStart="FIGURES 22 – 24" captionStartId="14.[151,264,1502,1524]" captionTargetBox="[161,1426,198,1476]" captionTargetId="figure@14.[151,1436,195,1480]" captionTargetPageId="14" captionText="FIGURES 22 – 24. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. SS: subapical spine." httpUri="https://zenodo.org/record/230982/files/figure.png" pageId="3" pageNumber="204">Fig. 24</figureCitation>
a)................ 
<taxonomicName id="4C7A37A6FFA1FFE84B097168F394F9AF" ID-CoL="6G76X" box="[1297,1435,1618,1639]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="williamsoni">
<emphasis id="B90E9037FFA1FFE84B097168F394F9AF" box="[1297,1435,1618,1639]" italics="true" pageId="3" pageNumber="204">E. williamsoni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC54C25FFA1FFE84E8F7155F394F96A" blockId="3.[151,1437,151,1698]" pageId="3" pageNumber="204">
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27’. Cercus ventral branch directed posteriorly in lateral view (
<figureCitation id="134150A0FFA1FFE84D007155F56FF94C" box="[792,864,1647,1668]" captionStart="FIGURES 25 – 27" captionStartId="15.[151,264,1645,1667]" captionTargetBox="[158,1436,205,1614]" captionTargetId="figure@15.[151,1436,195,1623]" captionTargetPageId="15" captionText="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri="https://zenodo.org/record/230983/files/figure.png" pageId="3" pageNumber="204">Fig. 26</figureCitation>
a), flattened dorso-ventrally, epiproct branches separated at their bases by a distance shorter than length of each branch (
<figureCitation id="134150A0FFA1FFE84D3B71B7F566F96A" box="[803,873,1677,1698]" captionStart="FIGURES 25 – 27" captionStartId="15.[151,264,1645,1667]" captionTargetBox="[158,1436,205,1614]" captionTargetId="figure@15.[151,1436,195,1623]" captionTargetPageId="15" captionText="FIGURES 25 – 27. Male caudal appendages, 25 a: cercus dorsal view, 25 b: cercus inner view, 25 c: cercus lateral view, 25 d: epiproct posterior view, 25 e: caudal appendages latero-posterior view, 26 – 27 a: lateral view; b: posterior view, c: lateroposterior view. Figure 25 e original drawings by R. W. Garrison, modified by the author." httpUri="https://zenodo.org/record/230983/files/figure.png" pageId="3" pageNumber="204">Fig. 26</figureCitation>
b).................................. 
<taxonomicName id="4C7A37A6FFA1FFE84B3F71B7F394F96A" ID-CoL="6G77F" box="[1319,1435,1677,1698]" class="Insecta" family="Protoneuridae" genus="Epipleoneura" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="B90E9037FFA1FFE84B3F71B7F394F96A" box="[1319,1435,1677,1698]" italics="true" pageId="3" pageNumber="204">E. spatulata</emphasis>
</taxonomicName>
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