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<mods:title id="185DB5CC459ABAAE27B27EC6C50C67B9">A new species of Opilioacarus With, 1902 (Acari: Opilioacaridae) from Italy, and a new diagnosis of the genus</mods:title>
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<mods:namePart id="C585868311996C6AF7FF0A332481F82B">Araújo, Marcel Santos De</mods:namePart>
<mods:affiliation id="4CA4672FE26D4FE542C6980514F252E9">Laboratório de Estudos Subterrâneos, Departamento de Ecologia e Biologia Evolutiva, Universidade Federal de São Carlos, Via Washington Luís, km 235, Caixa Postal 676, CEP 13565 - 905, São Carlos, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="EE515CCD839FFDAD5A705225A312DDF2">Palma, Antonella Di</mods:namePart>
<mods:affiliation id="8A6AC0C9BC313183631A4039EBFF6D60">Department of the Sciences of Agriculture, Food and Environment, University of Foggia, 71100, Foggia, Italy.</mods:affiliation>
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<mods:namePart id="56AA011647BC584F79126CEB51827A6D">Feres, Reinaldo José Fazzio</mods:namePart>
<mods:affiliation id="499C500F358FCE806DE96EED76BD9514">Departamento de Zoologia e Botânica, IBILCE, Universidade Estadual Paulista, Rua Cristóvão Colombo 2265, CEP 15054 - 000, São José do Rio Preto, SP, Brazil.</mods:affiliation>
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<paragraph id="8BD536D4142A94590EA5FF6670F9FF2F" blockId="2.[151,451,186,212]" box="[151,451,186,212]" pageId="2" pageNumber="137">
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<taxonomicName id="4C6A4D57142A94590EA5FF6670F9FF2F" ID-CoL="693X" authority="With, 1902" authorityName="With" authorityYear="1902" box="[151,451,186,212]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142A94590EA5FF6670F9FF2F" bold="true" box="[151,451,186,212]" pageId="2" pageNumber="137">
<emphasis id="B91EEAC6142A94590EA5FF667009FF2F" bold="true" box="[151,307,186,212]" italics="true" pageId="2" pageNumber="137">Opilioacarus</emphasis>
<bibRefCitation id="EFFB4B25142A94590F08FF6670F9FF2F" author="With, C. J." box="[314,451,186,212]" pageId="2" pageNumber="137" pagination="4 - 5" refId="ref6779" refString="With, C. J. (1902) A new acaride Opilioacarus segmentatus. Forhandlingar vid Nordiska Naturforskareog Lakaremotet i Helsingfors den 7 till 12 Juli 1902 (Comptes Rendus du Congres des Naturalistes et Medecins du Nord tenu a Helsingfors), Sektionen for Zoologi, 20, 4 - 5." type="journal article" year="1902">With, 1902</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C370655F142A94580EA5FEDD705BFD9B" lastPageId="3" lastPageNumber="138" pageId="2" pageNumber="137" type="diagnosis">
<paragraph id="8BD536D4142A94590EA5FEDD71DAFE7D" blockId="2.[151,1437,257,390]" pageId="2" pageNumber="137">
<emphasis id="B91EEAC6142A94590EA5FEDD7029FEE1" bold="true" box="[151,275,257,282]" pageId="2" pageNumber="137">Diagnosis:</emphasis>
45 foliate
<emphasis id="B91EEAC6142A94590FACFEDD7096FEE2" box="[414,428,257,281]" italics="true" pageId="2" pageNumber="137">d</emphasis>
setae (
<emphasis id="B91EEAC6142A94590FCCFEDD7321FEE2" box="[510,539,257,281]" italics="true" pageId="2" pageNumber="137">d1</emphasis>
) and one slightly pectinate
<emphasis id="B91EEAC6142A94590D6AFEDD725CFEE2" box="[856,870,257,281]" italics="true" pageId="2" pageNumber="137">d</emphasis>
seta (
<emphasis id="B91EEAC6142A94590D9EFEDD72FDFEE2" box="[940,967,257,281]" italics="true" pageId="2" pageNumber="137">d2</emphasis>
). Withs organ membranous and discoid. Two pairs of dorsolateral eyes, preanal region with 36 dorsal setae and 2 lateroventral setae. Dorsal segments between prodorsal shield and preanal region without setae. Leg I crown-like eupathidion (
<emphasis id="B91EEAC6142A94590AB1FE9575A7FE9A" box="[1155,1181,329,353]" italics="true" pageId="2" pageNumber="137">ζ1</emphasis>
) close to main sensilla group.
</paragraph>
<caption id="DF15665C142A94590EA5F87D74A6F827" ID-DOI="http://doi.org/10.5281/zenodo.5297426" ID-Zenodo-Dep="5297426" httpUri="https://zenodo.org/record/5297426/files/figure.png" pageId="2" pageNumber="137" startId="2.[151,250,1953,1975]" targetBox="[243,1385,531,1832]" targetPageId="2">
<paragraph id="8BD536D4142A94590EA5F87D74A6F827" blockId="2.[151,1436,1953,2012]" pageId="2" pageNumber="137">
<emphasis id="B91EEAC6142A94590EA5F87D7029F84D" bold="true" box="[151,275,1953,1975]" pageId="2" pageNumber="137">FIGURE 1.</emphasis>
<taxonomicName id="4C6A4D57142A94590F2BF87D70C4F84C" authority="Araújo &amp; Palma &amp; Feres, 2018" authorityName="Araújo &amp; Palma &amp; Feres" authorityYear="2018" box="[281,510,1953,1975]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="brignolii" status="sp. nov.">
<emphasis id="B91EEAC6142A94590F2BF87D70C4F84C" box="[281,510,1953,1975]" italics="true" pageId="2" pageNumber="137">Opilioacarus brignolii</emphasis>
</taxonomicName>
<emphasis id="B91EEAC6142A94590C36F87D7307F84D" bold="true" box="[516,573,1953,1974]" pageId="2" pageNumber="137">
<taxonomicNameLabel id="A22D57BD142A94590C36F87D7307F84D" box="[516,573,1953,1974]" pageId="2" pageNumber="137" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
collection sites in Italy, and original collection site of
<taxonomicName id="4C6A4D57142A94590A51F87D7400F84C" baseAuthorityName="With" baseAuthorityYear="1904" box="[1123,1338,1953,1975]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B91EEAC6142A94590A51F87D7400F84C" box="[1123,1338,1953,1975]" italics="true" pageId="2" pageNumber="137">Opilioacarus italicus</emphasis>
</taxonomicName>
.
<emphasis id="B91EEAC6142A94590EF5F81F7007F827" bold="true" box="[199,317,1987,2012]" pageId="2" pageNumber="137">Remarks:</emphasis>
The genus
<taxonomicName id="4C6A4D57142A94590FFEF818735AF827" authorityName="With" authorityYear="1902" box="[460,608,1988,2012]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142A94590FFEF818735AF827" box="[460,608,1988,2012]" italics="true" pageId="2" pageNumber="137">Opilioacarus</emphasis>
</taxonomicName>
presents similar foliate
<emphasis id="B91EEAC6142A94590D4CF81872B6F827" box="[894,908,1988,2012]" italics="true" pageId="2" pageNumber="137">d</emphasis>
type setae (
<emphasis id="B91EEAC6142A94590A2DF8187506F827" box="[1055,1084,1988,2012]" italics="true" pageId="2" pageNumber="137">d1</emphasis>
) to
<emphasis id="B91EEAC6142A94590A43F8187452F827" box="[1137,1384,1988,2012]" italics="true" pageId="2" pageNumber="137">
<taxonomicName id="4C6A4D57142A94590A43F81875E5F820" authorityName="Chamberlin &amp; Mulaik" authorityYear="1942" box="[1137,1247,1988,2011]" class="Arachnida" family="Opilioacaridae" genus="Neocarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Neocarus</taxonomicName>
,
<taxonomicName id="4C6A4D57142A94590ADDF8187452F827" authorityName="Das &amp; Bastawade" authorityYear="2007" box="[1263,1384,1988,2012]" class="Arachnida" family="Opilioacaridae" genus="Indiacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Indiacarus</taxonomicName>
</emphasis>
and
</paragraph>
</caption>
<paragraph id="8BD536D4142B94580EA5FF44737EFE9E" blockId="3.[151,1436,151,357]" pageId="3" pageNumber="138">
<taxonomicName id="4C6A4D57142B94580EA5FF447022FF4B" authorityName="Naudo" authorityYear="1963" box="[151,280,152,176]" class="Arachnida" family="Opilioacaridae" genus="Panchaetes" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580EA5FF447022FF4B" box="[151,280,152,176]" italics="true" pageId="3" pageNumber="138">Panchaetes</emphasis>
</taxonomicName>
, but differs from those genera by the presence of the single pectinate
<emphasis id="B91EEAC6142B94580A1EFF447500FF4B" box="[1068,1082,152,176]" italics="true" pageId="3" pageNumber="138">d</emphasis>
<typeStatus id="54D18876142B94580A73FF44754BFF4B" box="[1089,1137,152,176]" pageId="3" pageNumber="138">type</typeStatus>
seta (
<emphasis id="B91EEAC6142B94580A87FF4475EBFF4B" box="[1205,1233,152,176]" italics="true" pageId="3" pageNumber="138">d2</emphasis>
), similar in shape to the
<emphasis id="B91EEAC6142B94580EEDFF6071D7FF2F" box="[223,237,188,212]" italics="true" pageId="3" pageNumber="138">d</emphasis>
<typeStatus id="54D18876142B94580EC7FF60701FFF2F" box="[245,293,188,212]" pageId="3" pageNumber="138">type</typeStatus>
setae found in
<taxonomicName id="4C6A4D57142B94580FE7FF607250FF2E" authority="Vazquez et al. 2014" authorityName="Vazquez" authorityYear="2014" box="[469,874,188,213]" class="Arachnida" family="Opilioacaridae" genus="Amazonacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580FE7FF607341FF28" box="[469,635,188,211]" italics="true" pageId="3" pageNumber="138">Amazonacarus</emphasis>
<bibRefCitation id="EFFB4B25142B94580CB6FF607250FF2E" author="Vazquez, M. M. &amp; De Araujo, M. S. &amp; Feres, R. J. F." box="[644,874,188,213]" pageId="3" pageNumber="138" pagination="151 - 176" refId="ref6342" refString="Vazquez, M. M., De Araujo, M. S. &amp; Feres, R. J. F. (2014) A new genus and two new species of Opilioacaridae (Acari: Parasitiformes) from Amazonia, Brazil with a key to world genera. Zootaxa, 3814 (2), 151 - 176. https: // doi. org / 10.11646 / zootaxa. 3814.2.1" type="journal article" year="2014">
Vázquez
<emphasis id="B91EEAC6142B94580CDCFF61721AFF2F" box="[750,800,188,212]" italics="true" pageId="3" pageNumber="138">et al</emphasis>
. 2014
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A4D57142B94580D44FF6075C9FF2E" authority="Vazquez et al. 2015" authorityName="Vazquez" authorityYear="2015" box="[886,1267,188,213]" class="Arachnida" family="Opilioacaridae" genus="Brasilacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580D44FF60753DFF2F" box="[886,1031,188,212]" italics="true" pageId="3" pageNumber="138">Brasilacarus</emphasis>
Vázquez
<emphasis id="B91EEAC6142B94580A4BFF617596FF2F" box="[1145,1196,188,212]" italics="true" pageId="3" pageNumber="138">et al</emphasis>
. 2015
</taxonomicName>
,
<taxonomicName id="4C6A4D57142B94580B30FF6070FDFF03" authority="Vazquez &amp; Klompen 2009" authorityName="Vazquez &amp; Klompen" authorityYear="2009" class="Arachnida" family="Opilioacaridae" genus="Caribeacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580B30FF6074A6FF2F" box="[1282,1436,188,212]" italics="true" pageId="3" pageNumber="138">Caribeacarus</emphasis>
<bibRefCitation id="EFFB4B25142B94580EA5FF0370FDFF03" author="Vazquez, M. M. &amp; Klompen, H." box="[151,455,223,248]" pageId="3" pageNumber="138" pagination="23 - 44" refId="ref6581" refString="Vazquez, M. M. &amp; Klompen, H. (2009) New species of New World Opilioacaridae (Acari: Parasitiformes) with the description of a new genus from the Caribbean region. Zootaxa, 2061, 23 - 44." type="journal article" year="2009">Vázquez &amp; Klompen 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6A4D57142B94580FCDFF3C7219FF03" authority="Leclerc 1989" authorityName="Leclerc" authorityYear="1989" box="[511,803,223,248]" class="Arachnida" family="Opilioacaridae" genus="Siamacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580FCDFF3C73B9FF0C" box="[511,643,224,247]" italics="true" pageId="3" pageNumber="138">Siamacarus</emphasis>
Leclerc 1989
</taxonomicName>
. Also, the presence of more than a single dorsal seta on the preanal segment (XVII segment) is shared with
<taxonomicName id="4C6A4D57142B94580CEDFED875FCFEE6" authority="Coineau &amp; Hammen 1979" authorityName="Coineau &amp; Hammen" authorityYear="1979" box="[735,1222,260,285]" class="Arachnida" family="Opilioacaridae" genus="Phalangiacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580CEDFED872A8FEE7" box="[735,914,260,284]" italics="true" pageId="3" pageNumber="138">Phalangiacarus</emphasis>
<bibRefCitation id="EFFB4B25142B94580DA8FED875FCFEE6" author="Coineau, Y. &amp; Hammen, L. van der" box="[922,1222,260,285]" pageId="3" pageNumber="138" pagination="437 - 441" refId="ref5654" refString="Coineau, Y. &amp; Hammen, L. van der (1979) The postembryonic development of Opilioacarida, with notes on new taxa and on a general model for the evolution. In: E. Pfiffl (Ed), Proceedings of the 4 th International Congress of Acarology, Saalfelden 1974. Akademiai Kiado, Budapest, pp. 437 - 441." type="journal article" year="1979">Coineau &amp; Hammen 1979</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91EEAC6142B94580AE0FED8747AFEE7" box="[1234,1344,260,284]" italics="true" pageId="3" pageNumber="138">Indiacaru</emphasis>
s Das &amp; Bastawade (genera with no setae on VIIXVI segments),
<taxonomicName id="4C6A4D57142B94580D00FEF47571FEBB" authority="Naudo 1963" authorityName="Naudo" authorityYear="1963" box="[818,1099,295,320]" class="Arachnida" family="Opilioacaridae" genus="Panchaetes" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580D00FEF47289FEBB" box="[818,947,296,320]" italics="true" pageId="3" pageNumber="138">Panchaetes</emphasis>
Naudo 1963
</taxonomicName>
, and
<taxonomicName id="4C6A4D57142B94580AB9FEF47439FEBB" authorityName="Hammen" authorityYear="1977" box="[1163,1283,296,320]" class="Arachnida" family="Opilioacaridae" genus="Salfacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAC6142B94580AB9FEF47439FEBB" box="[1163,1283,296,320]" italics="true" pageId="3" pageNumber="138">Salfacarus</emphasis>
</taxonomicName>
(genera with segments VIIXVII with many setae).
</paragraph>
<paragraph id="8BD536D4142B94580EA5FE487066FE56" blockId="3.[151,348,404,429]" box="[151,348,404,429]" pageId="3" pageNumber="138">Species included:</paragraph>
<paragraph id="8BD536D4142B94580EA5FE0075BAFE0E" blockId="3.[151,1152,476,608]" box="[151,1152,476,501]" pageId="3" pageNumber="138">
<taxonomicName id="4C6A4D57142B94580EA5FE007379FE0E" authority="With, 1902" authorityName="With" authorityYear="1902" box="[151,579,476,501]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="segmentatus">
<emphasis id="B91EEAC6142B94580EA5FE007086FE0F" box="[151,444,476,500]" italics="true" pageId="3" pageNumber="138">Opilioacarus segmentatus</emphasis>
<bibRefCitation id="EFFB4B25142B94580FF1FE007379FE0E" author="With, C. J." box="[451,579,476,501]" pageId="3" pageNumber="138" pagination="4 - 5" refId="ref6779" refString="With, C. J. (1902) A new acaride Opilioacarus segmentatus. Forhandlingar vid Nordiska Naturforskareog Lakaremotet i Helsingfors den 7 till 12 Juli 1902 (Comptes Rendus du Congres des Naturalistes et Medecins du Nord tenu a Helsingfors), Sektionen for Zoologi, 20, 4 - 5." type="journal article" year="1902">With, 1902</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54D18876142B94580C7CFE0073BCFE0F" box="[590,646,476,500]" pageId="3" pageNumber="138">Type</typeStatus>
species (=
<taxonomicName id="4C6A4D57142B94580D36FE017542FE0E" authority="With, 1904" authorityName="With" authorityYear="1904" box="[772,1144,476,501]" class="Arachnida" family="Opilioacaridae" genus="Eucarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="segmentatus">
<emphasis id="B91EEAC6142B94580D36FE0172C9FE0F" box="[772,1011,477,500]" italics="true" pageId="3" pageNumber="138">Eucarus segmentatus</emphasis>
<bibRefCitation id="EFFB4B25142B94580DC8FE007542FE0E" author="With, C. J." box="[1018,1144,476,501]" pageId="3" pageNumber="138" pagination="137 - 191" refId="ref6833" refString="With, C. J. (1904) The Notostigmata, a new suborder of Acari. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KJObenhavn, 137 - 191." type="book chapter" year="1904">With, 1904</bibRefCitation>
</taxonomicName>
)
</paragraph>
<paragraph id="8BD536D4142B94580EA5FDDC72BEFDE3" blockId="3.[151,1152,476,608]" box="[151,900,511,536]" pageId="3" pageNumber="138">
<taxonomicName id="4C6A4D57142B94580EA5FDDC70A1FDE3" authority="(With 1904)" baseAuthorityName="With" baseAuthorityYear="1904" box="[151,411,511,536]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B91EEAC6142B94580EA5FDDC7036FDE3" box="[151,268,512,536]" italics="true" pageId="3" pageNumber="138">O. italicus</emphasis>
(
<bibRefCitation id="EFFB4B25142B94580F28FE2370A9FDE3" author="With, C. J." box="[282,403,511,536]" pageId="3" pageNumber="138" pagination="137 - 191" refId="ref6833" refString="With, C. J. (1904) The Notostigmata, a new suborder of Acari. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KJObenhavn, 137 - 191." type="book chapter" year="1904">With 1904</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="B91EEAC6142B94580F99FDDD736BFDE3" box="[427,593,512,536]" italics="true" pageId="3" pageNumber="138">nomen dubium</emphasis>
, this work, see Discussion)
</paragraph>
<paragraph id="8BD536D4142B94580EA5FDF873C8FDC6" blockId="3.[151,1152,476,608]" box="[151,754,548,573]" pageId="3" pageNumber="138">
<taxonomicName id="4C6A4D57142B94580E97FDF8737EFDC6" authority="Dunlop &amp; Bernardi, 2014" authorityName="Dunlop &amp; Bernardi" authorityYear="2014" box="[165,580,548,573]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="B91EEAC6142B94580E97FDF87027FDC7" box="[165,285,548,572]" italics="true" pageId="3" pageNumber="138">O. groehni</emphasis>
Dunlop &amp;
<bibRefCitation id="EFFB4B25142B94580FAEFDF8737EFDC6" author="Bernardi, L. F. O. &amp; Klompen, H. &amp; Ferreira, R. L." box="[412,580,548,573]" pageId="3" pageNumber="138" pagination="47 - 56" refId="ref5528" refString="Bernardi, L. F. O., Klompen, H. &amp; Ferreira, R. L. (2014) Neocarus caipora, a new mite species (Parasitiformes: Opilioacarida: Opilioacaridae) from Brazilian amazon caves. Acarologia, 54, 47 - 56. https: // doi. org / 10.1051 / acarologia / 20142113" type="journal article" year="2014">Bernardi, 2014</bibRefCitation>
</taxonomicName>
(fossil species)
</paragraph>
<paragraph id="8BD536D4142B94580EA5FD94705BFD9B" blockId="3.[151,1152,476,608]" box="[151,353,583,608]" pageId="3" pageNumber="138">
<taxonomicName id="4C6A4D57142B94580EA5FD947021FD9B" authority="Araújo &amp; Palma &amp; Feres, 2018" authorityName="Araújo &amp; Palma &amp; Feres" authorityYear="2018" box="[151,283,584,608]" class="Arachnida" family="Opilioacaridae" genus="Opilioacarus" kingdom="Animalia" order="Opilioacarida" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="brignolii" status="sp. nov.">
<emphasis id="B91EEAC6142B94580EA5FD947021FD9B" box="[151,283,584,608]" italics="true" pageId="3" pageNumber="138">O. brignolii</emphasis>
</taxonomicName>
<emphasis id="B91EEAC6142B94580F10FD9B705BFD9B" bold="true" box="[290,353,583,608]" pageId="3" pageNumber="138">
<taxonomicNameLabel id="A22D57BD142B94580F10FD9B705BFD9B" box="[290,353,583,608]" pageId="3" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>