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<document id="4DB6E6F0B4CA457C4FE0DB1F4505BD11" ID-DOI="10.11646/zootaxa.3838.5.5" ID-GBIF-Dataset="030bf385-7938-4b8a-b7b7-ef7a3663285d" ID-ISSN="1175-5326" ID-Zenodo-Dep="231513" ID-ZooBank="36769F07-1455-4569-83D4-9BC0E94C30EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460460580464" checkinUser="plazi" docAuthor="Kondorosy, Előd, Rédei, Dávid &amp; Mejlon, Hans" docDate="2014" docId="03C38782F614FFC4FF5208B7FD90FC27" docLanguage="en" docName="zt03838p574.pdf" docOrigin="Zootaxa 3838 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lethaeus ater Thunberg 1822, new combination" docType="treatment" docVersion="8" lastPageNumber="569" masterDocId="FFFAFFFAF615FFC6FFC50A74FF9DFF82" masterDocTitle="Taxonomic corrections to species of Rhyparochromidae (Hemiptera: Heteroptera) described by Carl Peter Thunberg" masterLastPageNumber="574" masterPageNumber="567" pageNumber="568" updateTime="1698303729205" updateUser="plazi">
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<mods:title id="227951C5FA8DDC05C412904D91791728">Taxonomic corrections to species of Rhyparochromidae (Hemiptera: Heteroptera) described by Carl Peter Thunberg</mods:title>
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<mods:namePart id="D4C316FF354E34FDF331433226E24E6F">Kondorosy, Előd</mods:namePart>
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<mods:namePart id="A232DDEB7AD473EE420028D32479B3D6">Rédei, Dávid</mods:namePart>
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<mods:namePart id="313F2060D4364094ABC511BABCC51E63">Mejlon, Hans</mods:namePart>
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<mods:typeOfResource id="E3BCC56B210ECF7F7228D39208D7C829">text</mods:typeOfResource>
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<mods:title id="E41AF115D56602AB15B1EB7BB57B15C0">Zootaxa</mods:title>
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<mods:date id="517EEB49EA39E1C72D102B61E74A870F">2014</mods:date>
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<treatment id="03C38782F614FFC4FF5208B7FD90FC27" ID-DOI="http://doi.org/10.5281/zenodo.6136039" ID-GBIF-Taxon="119512346" ID-Zenodo-Dep="6136039" LSID="urn:lsid:plazi:treatment:03C38782F614FFC4FF5208B7FD90FC27" httpUri="http://treatment.plazi.org/id/03C38782F614FFC4FF5208B7FD90FC27" lastPageId="2" lastPageNumber="569" pageId="1" pageNumber="568">
<subSubSection id="C370651FF614FFC7FF5208B7FE8CFC82" pageId="1" pageNumber="568" type="nomenclature">
<paragraph id="8BD53694F614FFC7FF5208B7FD7BFD5F" blockId="1.[151,742,707,768]" box="[151,742,707,733]" pageId="1" pageNumber="568">
<heading id="D09D81F8F614FFC7FF5208B7FD7BFD5F" bold="true" box="[151,742,707,733]" fontSize="10" level="2" pageId="1" pageNumber="568" reason="2">
<emphasis id="B91EEA86F614FFC7FF5208B7FD7BFD5F" bold="true" box="[151,742,707,733]" pageId="1" pageNumber="568">
<taxonomicName id="4C6A4D17F614FFC7FF5208B7FD90FD5F" authority="Thunberg, 1822" authorityName="Thunberg" authorityYear="1822" box="[151,525,707,733]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="ater" status="comb. nov.">
<emphasis id="B91EEA86F614FFC7FF5208B7FEA8FD59" bold="true" box="[151,309,707,732]" italics="true" pageId="1" pageNumber="568">Lethaeus ater</emphasis>
(Thunberg, 1822)
</taxonomicName>
,
<taxonomicNameLabel id="A22D57FDF614FFC7FDDF08B0FD7BFD5F" box="[538,742,708,733]" pageId="1" pageNumber="568">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD53694F614FFC7FF520893FE8CFC82" blockId="1.[151,742,707,768]" box="[151,273,743,768]" pageId="1" pageNumber="568">
(
<figureCitation id="13512A11F614FFC7FF5A0893FE94FC82" box="[159,265,743,768]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="1" pageNumber="568">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C370651FF614FFC7FF520945FBD7FC09" pageId="1" pageNumber="568" type="reference_group">
<paragraph id="8BD53694F614FFC7FF520945FBFDFCC4" blockId="1.[151,1120,816,838]" box="[151,1120,816,838]" pageId="1" pageNumber="568">
<treatmentCitationGroup id="AB7A11BAF614FFC7FF520945FBFDFCC4" box="[151,1120,816,838]" pageId="1" pageNumber="568">
<taxonomicName id="4C6A4D17F614FFC7FF520945FE77FCC4" authority="Thunberg, 1822b: 3" authorityName="Thunberg" authorityPageNumber="3" authorityYear="1822" box="[151,490,816,838]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91EEA86F614FFC7FF520945FE87FCC4" box="[151,282,817,838]" italics="true" pageId="1" pageNumber="568">Lygaeus ater</emphasis>
<treatmentCitation id="0ACB1085F614FFC7FEDA0944FE77FCC4" author="Thunberg" box="[287,490,816,838]" page="3" pageId="1" pageNumber="568" year="1822">
<bibRefCitation id="EFFB4B65F614FFC7FEDA0944FE4CFCC4" author="Thunberg" box="[287,465,816,838]" pageId="1" pageNumber="568" refString="Thunberg, C. P. (1822 b) Dissertatio entomologica de hemipteris rostratis capensibus, 4, 1 - 6. Upsaliae." type="journal article" year="1822" yearSuffix="b">Thunberg, 1822b</bibRefCitation>
: 3
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54D18836F614FFC7FE310944FDD3FCC4" box="[500,590,816,838]" pageId="1" pageNumber="568" type="syntype">Syntype</typeStatus>
(s): Capensis [=
<collectingCountry id="F37D7604F614FFC7FD2B0944FCEBFCC4" box="[750,886,816,838]" name="South Africa" pageId="1" pageNumber="568">South Africa</collectingCountry>
]; depository not stated.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD53694F614FFC7FF520906FBD7FC09" blockId="1.[151,1437,882,2023]" box="[151,1098,882,907]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF520906FEBDFC09" bold="true" box="[151,288,882,907]" pageId="1" pageNumber="568">References.</emphasis>
<bibRefCitation id="EFFB4B65F614FFC7FEE30906FE35FC09" author="Stal" box="[294,424,882,907]" pageId="1" pageNumber="568" refString="Stal, C. (1855) Om Thunbergska Hemipterarter. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 12 (7), 345 - 347." type="journal article" year="1855">Stål (1855)</bibRefCitation>
: 347 (synonymy);
<bibRefCitation id="EFFB4B65F614FFC7FDBC0906FC8CFC09" author="Slater" box="[633,785,882,907]" pageId="1" pageNumber="568" refString="Slater, J. A. (1964) A catalogue of the Lygaeidae of the world. Vols. I. - II. University of Connecticut, Storrs, xviii + 1668 pp. [I: i - xviii, 1 - 778, II: 779 - 1668]" type="book chapter" year="1964">Slater (1964)</bibRefCitation>
: 1192 (identity, synonymy).
</paragraph>
</subSubSection>
<subSubSection id="C370651FF614FFC7FF0209E2FEF2FBE1" pageId="1" pageNumber="568" type="materials_examined">
<paragraph id="8BD53694F614FFC7FF0209E2FEF2FBE1" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF0209E2FDE2FC2D" bold="true" box="[199,639,918,943]" pageId="1" pageNumber="568">
<typeStatus id="54D18836F614FFC7FF0209E2FE9FFC2D" box="[199,258,918,943]" pageId="1" pageNumber="568">Type</typeStatus>
material examined.
<typeStatus id="54D18836F614FFC7FDCC09E2FDE2FC2D" box="[521,639,918,943]" pageId="1" pageNumber="568" type="lectotype">Lectotype</typeStatus>
</emphasis>
(present designation) ♂ (
<figureCitation id="13512A11F614FFC7FC7309E2FBB4FC2D" box="[950,1065,918,943]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="1" pageNumber="568">Figs. 12</figureCitation>
): “Uppsala Univ. Zool.
<taxonomicName id="4C6A4D17F614FFC7FA8B09E3FA1EFC2C" box="[1358,1411,919,942]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="1" pageNumber="571" phylum="Chordata" rank="genus">Mus</taxonomicName>
. / Thunbergsaml. nr. 18550 /
<taxonomicName id="4C6A4D17F614FFC7FE0909C8FDC2FC51" box="[460,607,956,979]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91EEA86F614FFC7FE0909C8FDC2FC51" box="[460,607,956,979]" italics="true" pageId="1" pageNumber="568">Lygaeus ater</emphasis>
</taxonomicName>
/
<emphasis id="B91EEA86F614FFC7FDBD09CFFD3AFC50" box="[632,679,955,978]" italics="true" pageId="1" pageNumber="568">Cap</emphasis>
. TYP” [red, printed]; “
<typeStatus id="54D18836F614FFC7FC7409CFFBFBFC51" box="[945,1126,955,979]" pageId="1" pageNumber="568" type="lectotype">LECTOTYPUS</typeStatus>
/
<taxonomicName id="4C6A4D17F614FFC7FBBA09CFFA92FC50" box="[1151,1295,955,979]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="ater">Lygaeus ater</taxonomicName>
/ Thunberg, 1822 / desig. Kondorosy, 2013” [with red frame, printed]; “
<taxonomicName id="4C6A4D17F614FFC7FCEA09ABFCC0FC75" box="[815,861,991,1015]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="ater">ater</taxonomicName>
. / Cap. 19” [handwritten, pinned in the drawer next to the specimen (
<figureCitation id="13512A11F614FFC7FEA50E76FE35FB99" box="[352,424,1026,1051]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="1" pageNumber="568">Fig. 3</figureCitation>
)]; deposited in
<collectionCode id="ED7BAE51F614FFC7FDA50E77FD35FB99" box="[608,680,1027,1051]" pageId="1" pageNumber="568">UPSZ</collectionCode>
. Pinned; left fore wing, left antenna and segments IIIV of right antenna, and all legs distad of coxae with the exception of right fore leg, missing; body broken between pro- and mesothorax by pin.
</paragraph>
</subSubSection>
<subSubSection id="C370651FF614FFC4FF020E1AFEA8FF7A" lastPageId="2" lastPageNumber="569" pageId="1" pageNumber="568" type="description">
<paragraph id="8BD53694F614FFC7FF020E1AFAE7FB71" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF020E1AFEF0FB05" bold="true" box="[199,365,1134,1159]" pageId="1" pageNumber="568">Redescription</emphasis>
.
<emphasis id="B91EEA86F614FFC7FEB80E1AFE56FB05" box="[381,459,1134,1159]" italics="true" pageId="1" pageNumber="568">Colour</emphasis>
. Ground colour of body and appendages fuscous brown (antennal segments IIIV as well as mid and hind legs missing in the single examined specimen); head, anterior part of pronotum, scutellum and posterior half of abdominal tergum (segments VVII) almost black except apical projection of tergite VII; membrane, labial segments IIIII, fore tibia and metepimeron, brown; veins of corium with tiny yellow streaks.
</paragraph>
<paragraph id="8BD53694F614FFC7FF020E8BFB16FAB8" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF020E8BFEB1FA94" box="[199,300,1279,1302]" italics="true" pageId="1" pageNumber="568">Structure</emphasis>
.
<emphasis id="B91EEA86F614FFC7FEF90E8AFEEBFA95" box="[316,374,1278,1303]" italics="true" pageId="1" pageNumber="568">Body</emphasis>
glabrous except abdomen and extremities; slightly shining, almost opaque. Head and anterior half of pronotum with fine punctation, remaining parts of dorsum with stronger punctures.
</paragraph>
<paragraph id="8BD53694F614FFC7FF020F32FDAAFA49" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF020F32FE99FADD" box="[199,260,1350,1375]" italics="true" pageId="1" pageNumber="568">Head</emphasis>
pentagonal, antennae inserted far from eyes, lateral outlines of antenniferous tubercles parallel in dorsal view. Ocelli situated relatively close to eyes, eyes reaching anterior margin of pronotum. Antennal segment I surpassing apex of clypeus by about half of its length. Apex of labium reaching about base of mid coxae (surpassing middle of mesosternum).
</paragraph>
<paragraph id="8BD53694F614FFC7FF020FA2FD91F8D5" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF020FA2FE82FA6D" box="[199,287,1494,1519]" italics="true" pageId="1" pageNumber="568">Thorax.</emphasis>
Pronotum convex, without any trace of transverse furrow or median carina, anterior margin concave, posterior margin almost straight, laterally convex; lateral margin moderately broadly laminate. Scutellum triangular, flat, without carinae. Hemelytra abbreviated (
<figureCitation id="13512A11F614FFC7FCEC0C6AFCF2F9B5" box="[809,879,1566,1591]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="1" pageNumber="568">Fig. 1</figureCitation>
), reaching only base of tergite VII; clavus and corium immovably fused (claval furrow weakly developed), apex of corium broadly rounded, membranal margin of corium straight, oblique; clavus with four irregular rows of punctures; membrane reduced to a narrow stripe. Metathoracic scent gland ostiole close to coxae, slit-like, provided with short, auriculate peritreme curved posteriorly; evaporatorium relatively small, occupying only about medial half of metapleuron, extending to the posterior supracoxal lobe of the mesopleuron. Fore femur slightly thickened, more slender than width of vertex, ventral surface with some long setae and four or five very short spines apically; fore tibia straight, unarmed; first segment of fore tarsus about as long as combined lengths of the remaining two segments; mid and hind legs absent in the single examined specimen.
</paragraph>
<paragraph id="8BD53694F614FFC7FF020D16FBCAF865" blockId="1.[151,1437,882,2023]" pageId="1" pageNumber="568">
<emphasis id="B91EEA86F614FFC7FF020D16FEACF8F9" box="[199,305,1890,1915]" italics="true" pageId="1" pageNumber="568">Abdomen</emphasis>
with suture between sternites IV and V curved anteriorly and ending in a trichobothrial furrow as typical in
<taxonomicName id="4C6A4D17F614FFC7FEC80DF2FE7AF81D" box="[269,487,1926,1951]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="568" phylum="Arthropoda" rank="family">Rhyparochromidae</taxonomicName>
. Trichobothria of sternum V rectilinear, anterior and middle trichobothria close to each other on anterior part of sterna, posterior one situated posteriad of spiracle. All spiracles ventral. Connexivum upturned, posterolateral angle of connexival segments VII broadly, obtusely angulate.
</paragraph>
<paragraph id="8BD53694F617FFC4FF020AEDFEA8FF7A" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="569">
<emphasis id="B91EEA86F617FFC4FF020AEDFEF4FF32" box="[199,361,153,176]" italics="true" pageId="2" pageNumber="569">Measurements</emphasis>
(in mm): body length 8.6, length of head 1.15, greatest width across eyes 1.55, width immediately frontad of eyes 1.15, interocular distance 1.0, length of pronotum 2.0, humeral width 3.05, length of scutellum 1.6.
</paragraph>
</subSubSection>
<subSubSection id="C370651FF617FFC4FF020B70FC5EFC96" pageId="2" pageNumber="569" type="discussion">
<paragraph id="8BD53694F617FFC4FF020B70FB31FE0A" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="569">
<emphasis id="B91EEA86F617FFC4FF020B70FED1FE9F" bold="true" box="[199,332,260,285]" pageId="2" pageNumber="569">Discussion.</emphasis>
A single male (
<figureCitation id="13512A11F617FFC4FDD90B70FD1FFE9E" box="[540,642,260,285]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="2" pageNumber="569">Fig. 12</figureCitation>
) is present in Thunbergs collection in UPSZ. The brief original description of
<taxonomicName id="4C6A4D17F617FFC4FEFC0B5DFE57FEC2" box="[313,458,297,320]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91EEA86F617FFC4FEFC0B5DFE57FEC2" box="[313,458,297,320]" italics="true" pageId="2" pageNumber="569">Lygaeus ater</emphasis>
</taxonomicName>
fits well with this specimen, except the fore wings which are not completely devoid of spots (“immaculatus hemelytris opacis”). Based on the ancient handwritten label pinned next to the specimen (
<figureCitation id="13512A11F617FFC4FAAA0B38FF38FE0A" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1943,1965]" captionTargetBox="[150,1435,988,1915]" captionTargetId="figure@2.[150,1436,988,1922]" captionTargetPageId="2" captionText="FIGURES 1 7. Lectotypes of three of Thunbergs species and selected relevant labels. 1 3, Lygaeus ater Thunberg, 1822; 4 5, Lygaeus biguttatus Thunberg, 1822; 6 7, Cimex caffer Thunberg, 1784. Scales in mm; labels not to scale." httpUri="https://zenodo.org/record/231514/files/figure.png" pageId="2" pageNumber="569">Fig. 3</figureCitation>
) it is considered as part of the original
<typeStatus id="54D18836F617FFC4FD990B04FD11FE0A" box="[604,652,368,392]" pageId="2" pageNumber="569">type</typeStatus>
series and it is hereby designated as a
<typeStatus id="54D18836F617FFC4FBFB0B1BFB34FE0A" box="[1086,1193,367,392]" pageId="2" pageNumber="569" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BD53694F617FFC4FF020BE0FE94FDBF" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="569">
<bibRefCitation id="EFFB4B65F617FFC4FF020BE0FED8FE2F" author="Stal" box="[199,325,404,429]" pageId="2" pageNumber="569" refString="Stal, C. (1855) Om Thunbergska Hemipterarter. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 12 (7), 345 - 347." type="journal article" year="1855">Stål (1855)</bibRefCitation>
synonymized
<taxonomicName id="4C6A4D17F617FFC4FE290BE1FDE3FE2E" box="[492,638,405,428]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91EEA86F617FFC4FE290BE1FDE3FE2E" box="[492,638,405,428]" italics="true" pageId="2" pageNumber="569">Lygaeus ater</emphasis>
</taxonomicName>
with
<taxonomicName id="4C6A4D17F617FFC4FD040BE0FBF2FE2E" authority="Panzer, 1799" authorityName="Panzer" authorityYear="1799" box="[705,1135,404,429]" class="Insecta" family="Rhyparochromidae" genus="Rhyparochromus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="echii">
<emphasis id="B91EEA86F617FFC4FD040BE0FC23FE2F" box="[705,958,404,429]" italics="true" pageId="2" pageNumber="569">Rhyparochromus echii</emphasis>
(Panzer, 1799)
</taxonomicName>
, now a junior synonym of
<taxonomicName id="4C6A4D17F617FFC4FF520BC3FD80FE52" authority="Goeze, 1778" authorityName="Goeze" authorityYear="1778" box="[151,541,439,464]" class="Insecta" family="Lygaeidae" genus="Aellopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="B91EEA86F617FFC4FF520BC3FEC6FE52" box="[151,347,439,464]" italics="true" pageId="2" pageNumber="569">Aellopus atratus</emphasis>
(
<bibRefCitation id="EFFB4B65F617FFC4FEB30BCCFD89FE52" author="Goeze" box="[374,532,440,464]" pageId="2" pageNumber="569" refString="Goeze, J. A. E. (1778) Entomologische Beytrage zu des Ritter Linne zwolften Ausgabe des Natursystems. Zweyter Theil. Weidmanns Erben &amp; Reich, Leipzig, lxxii + 352 pp." type="book" year="1778">Goeze, 1778</bibRefCitation>
)
</taxonomicName>
; the same synonymy was cited by
<bibRefCitation id="EFFB4B65F617FFC4FC310BC3FB0FFE52" author="Slater" box="[1012,1170,439,464]" pageId="2" pageNumber="569" refString="Slater, J. A. (1964) A catalogue of the Lygaeidae of the world. Vols. I. - II. University of Connecticut, Storrs, xviii + 1668 pp. [I: i - xviii, 1 - 778, II: 779 - 1668]" type="book chapter" year="1964">Slater (1964)</bibRefCitation>
and
<bibRefCitation id="EFFB4B65F617FFC4FB250BC3FA05FE52" author="Pericart" box="[1248,1432,439,464]" pageId="2" pageNumber="569" refString="Pericart, J. (2001) Family Lygaeidae Schilling, 1829 - Seed-bugs. In: Aukema, B. &amp; Rieger, Chr. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. The Netherlands Entomological Society, Amsterdam, pp. 35 - 220." type="book" year="2001">Péricart (2001)</bibRefCitation>
. Reexamination of the
<typeStatus id="54D18836F617FFC4FE640BA9FE4CFE77" box="[417,465,477,501]" pageId="2" pageNumber="569">type</typeStatus>
material revealed that the act of
<bibRefCitation id="EFFB4B65F617FFC4FCAE0BA8FC70FE77" author="Stal" box="[875,1005,476,501]" pageId="2" pageNumber="569" refString="Stal, C. (1855) Om Thunbergska Hemipterarter. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 12 (7), 345 - 347." type="journal article" year="1855">Stål (1855)</bibRefCitation>
was erroneous,
<taxonomicName id="4C6A4D17F617FFC4FB700BA9FAD6FE76" box="[1205,1355,477,500]" class="Insecta" family="Lygaeidae" genus="Lygaeus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="ater">
<emphasis id="B91EEA86F617FFC4FB700BA9FAD6FE76" box="[1205,1355,477,500]" italics="true" pageId="2" pageNumber="569">Lygaeus ater</emphasis>
</taxonomicName>
is not conspecific with
<taxonomicName id="4C6A4D17F617FFC4FEA40874FE48FD9A" box="[353,469,512,536]" class="Insecta" family="Lygaeidae" genus="Aellopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="B91EEA86F617FFC4FEA40874FE48FD9A" box="[353,469,512,536]" italics="true" pageId="2" pageNumber="569">A. atratus</emphasis>
</taxonomicName>
. The spiracles and trichobothria undoubtedly indicate that it belongs to the tribe Lethaeini.
</paragraph>
<paragraph id="8BD53694F617FFC4FF020833FC5EFC96" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="569">
Since all members of
<taxonomicName id="4C6A4D17F617FFC4FE010833FD5CFDE2" authority="Dallas, 1852" authorityName="Dallas" authorityYear="1852" box="[452,705,583,608]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86F617FFC4FE010833FDB4FDE2" box="[452,553,583,608]" italics="true" pageId="2" pageNumber="569">Lethaeus</emphasis>
<bibRefCitation id="EFFB4B65F617FFC4FDF70833FD5CFDE2" author="Dallas" box="[562,705,583,608]" pageId="2" pageNumber="569" refString="Dallas, W. S. (1852) List of the specimens of hemipterous insects in the collection of the British Museum. Part II. Trustees of the British Museum, London, pp. 369 - 592, plates xii - xv." type="book chapter" year="1852">Dallas, 1852</bibRefCitation>
</taxonomicName>
and related genera are known from macropterous morphs only, brachyptery is an unusual character of this species. Several characters, particularly the shape of the head, the absence of a pronotal collar, the absence of anterolateral setae of the pronotum, as well as the hemelytral colour, and the armature of the fore femora, still suggest that in spite of the reduced wings it belongs to
<taxonomicName id="4C6A4D17F617FFC4FB3708C0FACAFD4F" box="[1266,1367,692,717]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86F617FFC4FB3708C0FACAFD4F" box="[1266,1367,692,717]" italics="true" pageId="2" pageNumber="569">Lethaeus</emphasis>
</taxonomicName>
. As a consequence, the new combination
<taxonomicName id="4C6A4D17F617FFC4FDE608A3FC15FD72" authority="Thunberg, 1822" authorityName="Thunberg" authorityYear="1822" box="[547,904,727,752]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="ater" status="comb. nov.">
<emphasis id="B91EEA86F617FFC4FDE608A3FD26FD72" box="[547,699,727,752]" italics="true" pageId="2" pageNumber="569">Lethaeus ater</emphasis>
(Thunberg, 1822)
</taxonomicName>
,
<taxonomicNameLabel id="A22D57FDF617FFC4FC5108A3FB8EFD72" box="[916,1043,727,752]" pageId="2" pageNumber="569">
<emphasis id="B91EEA86F617FFC4FC5108A3FB8EFD72" bold="true" box="[916,1043,727,752]" pageId="2" pageNumber="569">comb. nov.</emphasis>
</taxonomicNameLabel>
, is hereby proposed. The species is apparently not conspecific with any other described species of
<taxonomicName id="4C6A4D17F617FFC4FC930888FC26FC97" box="[854,955,764,789]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86F617FFC4FC930888FC26FC97" box="[854,955,764,789]" italics="true" pageId="2" pageNumber="569">Lethaeus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C370651FF617FFC4FF02096BFD90FC27" pageId="2" pageNumber="569" type="reference_group">
<paragraph id="8BD53694F617FFC4FF02096BFD90FC27" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="569">
<taxonomicName id="4C6A4D17F617FFC4FF02096BFD82FCBA" authority="Slater, 1964" authorityName="Slater" authorityYear="1964" box="[199,543,799,824]" class="Insecta" family="Lygaeidae" genus="Lethaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="barberi">
<emphasis id="B91EEA86F617FFC4FF02096BFE17FCBA" box="[199,394,799,824]" italics="true" pageId="2" pageNumber="569">Lethaeus barberi</emphasis>
<bibRefCitation id="EFFB4B65F617FFC4FE51096BFD82FCBA" author="Slater" box="[404,543,799,824]" pageId="2" pageNumber="569" refString="Slater, J. A. (1964) A catalogue of the Lygaeidae of the world. Vols. I. - II. University of Connecticut, Storrs, xviii + 1668 pp. [I: i - xviii, 1 - 778, II: 779 - 1668]" type="book chapter" year="1964">Slater, 1964</bibRefCitation>
</taxonomicName>
, another brachypterous species of Lethaeini from
<collectingCountry id="F37D7604F617FFC4FBA3096BFB66FCBA" box="[1126,1275,799,824]" name="South Africa" pageId="2" pageNumber="569">South Africa</collectingCountry>
, based on the reexamination of its
<typeStatus id="54D18836F617FFC4FE480930FE72FCDF" box="[397,495,836,861]" pageId="2" pageNumber="569" type="holotype">holotype</typeStatus>
deposited in MZLU certainly does not belong to this genus as its pronotum is provided with a rather narrow collar and anterolateral setae. Currently it is not possible to place this species with confidence in any existing genus.
</paragraph>
</subSubSection>
</treatment>
</document>