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<document id="81FBB182CB2B051D09D6053769801747" ID-DOI="10.11646/phytotaxa.323.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13696684" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725566521507" checkinUser="felipe" docAuthor="Henderson, Andrew &amp; Dung, Nguyen Quoc" docDate="2017" docId="03C38781FF8CFFF7FF776F83FD6CFB95" docLanguage="en" docName="phytotaxa.323.2.4.pdf" docOrigin="Phytotaxa 323 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.323.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lanonia verrucosa Henderson &amp; Nguyen Quoc Dung 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="163" masterDocId="FFFAFFF9FF88FFF3FFFF6F1DFFD3FFA9" masterDocTitle="New species of Lanonia, Licuala, and Pinanga (Arecaceae) from Vietnam" masterLastPageNumber="172" masterPageNumber="159" pageNumber="163" updateTime="1725567521063" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A6CF9E927C9EDFA36C4E5C2AF1E9B88C">New species of Lanonia, Licuala, and Pinanga (Arecaceae) from Vietnam</mods:title>
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<mods:name id="B0CBBDDA1DAD643A5C8527532B36F3C5" type="personal">
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<mods:namePart id="EEACABD5D47BF795B341A4E6FE7C784F">Henderson, Andrew</mods:namePart>
<mods:affiliation id="5ABB3A2C06E517F58DAC4C4697F1030E">Institute of Systematic Botany, The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
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<mods:namePart id="0A292B0860D59737403AA9438B5C8112">Dung, Nguyen Quoc</mods:namePart>
<mods:affiliation id="8326FAF472200A055776948529BCF40B">Forest Inventory and Planning Institute, Thanh Tri, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="F4F5D99E155BDC3EDB4F45DE76C8E759">Phytotaxa</mods:title>
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<mods:date id="D6DCA67138A0B491A5F817446680E6F4">2017</mods:date>
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<mods:number id="7B1443F39267D21B1DDC2C48C437A5D4">2017-09-29</mods:number>
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<mods:identifier id="655D9EB323C9E55151F849F54C999BA4" type="DOI">10.11646/phytotaxa.323.2.4</mods:identifier>
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<treatment id="03C38781FF8CFFF7FF776F83FD6CFB95" ID-DOI="http://doi.org/10.5281/zenodo.13696700" ID-Zenodo-Dep="13696700" LSID="urn:lsid:plazi:treatment:03C38781FF8CFFF7FF776F83FD6CFB95" httpUri="http://treatment.plazi.org/id/03C38781FF8CFFF7FF776F83FD6CFB95" lastPageNumber="163" pageId="4" pageNumber="163">
<subSubSection id="C370651CFF8CFFF7FF776F83FC4DFF11" box="[136,926,158,184]" pageId="4" pageNumber="163" type="nomenclature">
<paragraph id="8BD53697FF8CFFF7FF776F83FC4DFF11" blockId="4.[136,926,158,184]" box="[136,926,158,184]" pageId="4" pageNumber="163">
<heading id="D09D81FBFF8CFFF7FF776F83FC4DFF11" box="[136,926,158,184]" fontSize="11" level="2" pageId="4" pageNumber="163" reason="3">
<taxonomicName id="4C6A4D14FF8CFFF7FF776F83FD3BFF11" ID-CoL="8WWY5" authority="Henderson &amp; Nguyen Quoc Dung" authorityName="Henderson &amp; Nguyen Quoc Dung" authorityYear="2017" box="[136,744,158,184]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="verrucosa" status="sp. nov.">
<emphasis id="B91EEA85FF8CFFF7FF776F83FEB0FF11" bold="true" box="[136,355,158,184]" italics="true" pageId="4" pageNumber="163">Lanonia verrucosa</emphasis>
Henderson &amp; Nguyen Quoc Dung
</taxonomicName>
<emphasis id="B91EEA85FF8CFFF7FD106F82FC97FF11" box="[751,836,159,184]" italics="true" pageId="4" pageNumber="163">
<taxonomicNameLabel id="A22D57FEFF8CFFF7FD106F82FC97FF11" box="[751,836,159,184]" pageId="4" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13512A12FF8CFFF7FCAC6F82FC45FF11" box="[851,918,159,184]" captionStart="FIGURE 4" captionStartId="4.[136,229,2007,2028]" captionTargetBox="[154,1435,1112,1983]" captionTargetId="figure-447@4.[151,1436,1109,1983]" captionTargetPageId="4" captionText="FIGURE 4. Lanonia verrucosa. Left, leaf blade with wide central segment lobes. Right, infructescence with warty fruits (from Henderson &amp; Nguyen Quoc Dung 4079)." figureDoi="http://doi.org/10.5281/zenodo.13696698" httpUri="https://zenodo.org/record/13696698/files/figure.png" pageId="4" pageNumber="163">Fig. 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C370651CFF8CFFF7FF776FDBFA76FCE9" pageId="4" pageNumber="163" type="materials_examined">
<paragraph id="8BD53697FF8CFFF7FF776FDBFBF9FF56" blockId="4.[136,1452,198,255]" pageId="4" pageNumber="163">
Type:—
<collectingCountry id="F37D7607FF8CFFF7FF296FDBFE9BFF72" box="[214,328,198,219]" name="Vietnam" pageId="4" pageNumber="163">VIETNAM</collectingCountry>
.
<collectingRegion id="49AEF875FF8CFFF7FEAF6FDBFDCFFF72" box="[336,540,198,219]" country="Vietnam" name="Khanh Hoa" pageId="4" pageNumber="163">Khanh Hoa Province</collectingRegion>
: Ninh Hoa District, Ninh Tay Commune, road to hydroelectric dam,
<geoCoordinate id="EE5E5050FF8CFFF7FB566FDBFB2FFF72" box="[1193,1276,198,219]" degrees="12" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="163" precision="925" value="12.583333">12°35N</geoCoordinate>
<geoCoordinate id="EE5E5050FF8CFFF7FAFD6FDBFAB3FF72" box="[1282,1376,198,219]" degrees="108" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="163" precision="925" value="108.96667">108°58E</geoCoordinate>
,
<quantity id="4C929B72FF8CFFF7FA966FDBFA7BFF72" box="[1385,1448,198,219]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="4" pageNumber="163" unit="m" value="433.0">433 m</quantity>
,
<date id="FFD41057FF8CFFF7FF426FF7FE86FF56" box="[189,341,234,255]" pageId="4" pageNumber="163" value="2016-10-07">7 October 2016</date>
,
<emphasis id="B91EEA85FF8CFFF7FEA06FF7FD25FF56" box="[351,758,234,255]" italics="true" pageId="4" pageNumber="163">
<collectionCode id="ED7BAE52FF8CFFF7FEA06FF7FEA2FF56" box="[351,369,234,255]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="163" type="Herbarium">A</collectionCode>
. Henderson &amp; Nguyen Quoc Dung 4079
</emphasis>
(
<typeStatus id="54D18835FF8CFFF7FCFD6FF7FC8BFF56" box="[770,856,234,255]" pageId="4" pageNumber="163" type="holotype">holotype</typeStatus>
<collectionCode id="ED7BAE52FF8CFFF7FCA16FF7FC4AFF56" box="[862,921,234,255]" pageId="4" pageNumber="163">VFM</collectionCode>
!,
<typeStatus id="54D18835FF8CFFF7FC596FF7FC38FF56" box="[934,1003,234,255]" pageId="4" pageNumber="163" type="isotype">isotype</typeStatus>
<collectionCode id="ED7BAE52FF8CFFF7FC0E6FF7FBC8FF56" box="[1009,1051,234,255]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="4" pageNumber="163" type="Herbarium">NY</collectionCode>
!).
</paragraph>
<paragraph id="8BD53697FF8CFFF7FF776E32FA43FEC5" blockId="4.[136,1452,303,1084]" pageId="4" pageNumber="163">
It differs from similar species with few (313) segments and the central segment much wider than the others (
<emphasis id="B91EEA85FF8CFFF7FA6A6E32FDE7FEC5" italics="true" pageId="4" pageNumber="163">
<taxonomicName id="4C6A4D14FF8CFFF7FA6A6E32FF28FEC5" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="calciphila">L. calciphila</taxonomicName>
,
<taxonomicName id="4C6A4D14FF8CFFF7FEF96E4EFE49FEC5" box="[262,410,339,364]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="dasyantha">L. dasyantha</taxonomicName>
,
<taxonomicName id="4C6A4D14FF8CFFF7FE5A6E4EFDE7FEC5" box="[421,564,339,364]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="magalonii">L. magalonii</taxonomicName>
</emphasis>
) in its exceptionally wide central segment and distantly spaced, warty fruits.
</paragraph>
<paragraph id="8BD53697FF8CFFF7FF426E6AFA76FCE9" blockId="4.[136,1452,303,1084]" pageId="4" pageNumber="163">
Stems solitary,
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tall, 6.0 cm diameter. Leaf number not recorded; sheaths not recorded; ligules not recorded; petioles 130.0 cm long, thorny along the margins; blades to 80.0 cm across, split into 68, unequal segments; costa
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long; central segment split into 2 lobes, these 45.0 cm long, 18.0 cm wide at the apex; lateral segment 36.0 cm long,
<quantity id="4C929B72FF8CFFF7FF076EFEFE87FE55" box="[248,340,483,508]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="163" unit="cm" value="11.5">11.5 cm</quantity>
wide at apex; segments briefly indented at the apex. Inflorescences erect amongst the petioles; staminate inflorescences branched to 1 order; prophylls not recorded; peduncles to 10.0 cm long; partial inflorescence 1; rachillae to 12.0 cm long, minutely hairy; staminate flowers 3.0 mm long, solitary or in clusters of 23; calyx 2.0 mm long, tubular at the base for ca. 1.0 mm, cupular and shallowly lobed above; corolla 3.0 mm long, split for ca. half its length into 3, valvate petals; stamens 6; filaments with scarcely developed staminal ring, unequal with 2 longer than the other 4; pistillode 3-parted; pistillate inflorescences larger than the staminate, branched to 2 orders; prophylls 30.0 cm long; peduncles to 60.0 cm long with 1, narrow, tubular, glabrous peduncular bract; partial inflorescence 1; rachillae
<quantity id="4C929B72FF8CFFF7FF106DC2FEAAFD51" box="[239,377,735,760]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.65" metricValueMin="0.95" pageId="4" pageNumber="163" unit="cm" value="13.0" valueMax="16.5" valueMin="9.5">9.516.5 cm</quantity>
long, minutely hairy; pistillate flowers not seen at anthesis, solitary; calyx tubular at the base, 3- lobed above, each lobe splitting again as the fruits develop to give a 6-lobed calyx; corolla with 3 petals, these reflexed as fruits develop; fruits ellipsoid,
<quantity id="4C929B72FF8CFFF7FDFE6C3AFD9EFCE9" box="[513,589,807,832]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="163" unit="cm" value="1.1">1.1 cm</quantity>
long,
<quantity id="4C929B72FF8CFFF7FD6B6C3AFD33FCE9" box="[660,736,807,832]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="163" unit="cm" value="0.7">0.7 cm</quantity>
diameter, orange, the surface warty; endosperm homogeneous.
</paragraph>
</subSubSection>
<subSubSection id="C370651CFF8CFFF7FF426C56FD6CFB95" pageId="4" pageNumber="163" type="distribution">
<paragraph id="8BD53697FF8CFFF7FF426C56FE8CFC21" blockId="4.[136,1452,303,1084]" pageId="4" pageNumber="163">
<emphasis id="B91EEA85FF8CFFF7FF426C56FE32FCCD" bold="true" box="[189,481,843,868]" pageId="4" pageNumber="163">Distribution and habitat</emphasis>
:—Southern
<collectingCountry id="F37D7607FF8CFFF7FD906C56FD1DFCCD" box="[623,718,843,868]" name="Vietnam" pageId="4" pageNumber="163">Vietnam</collectingCountry>
in
<collectingRegion id="49AEF875FF8CFFF7FD0D6C56FC08FCCD" box="[754,987,843,868]" country="Vietnam" name="Khanh Hoa" pageId="4" pageNumber="163">Khanh Hoa province</collectingRegion>
in disturbed forest on steep, rocky slopes at
<quantity id="4C929B72FF8CFFF7FF5C6C72FF39FC21" box="[163,234,879,904]" metricMagnitude="2" metricUnit="m" metricValue="4.33" pageId="4" pageNumber="163" unit="m" value="433.0">433 m</quantity>
elevation.
</paragraph>
<paragraph id="8BD53697FF8CFFF7FF426C8EFD6CFB95" blockId="4.[136,1452,303,1084]" pageId="4" pageNumber="163">
<emphasis id="B91EEA85FF8CFFF7FF426C8EFE7DFC05" bold="true" box="[189,430,915,940]" pageId="4" pageNumber="163">Taxonomic notes:—</emphasis>
<taxonomicName id="4C6A4D14FF8CFFF7FE516C8EFD56FC05" authorityName="Henderson &amp; Nguyen Quoc Dung" authorityYear="2017" box="[430,645,915,940]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="verrucosa">
<emphasis id="B91EEA85FF8CFFF7FE516C8EFD56FC05" box="[430,645,915,940]" italics="true" pageId="4" pageNumber="163">Lanonia verrucosa</emphasis>
</taxonomicName>
most resembles
<taxonomicName id="4C6A4D14FF8CFFF7FCB56C8EFC0EFC05" box="[842,989,915,940]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="dasyantha">
<emphasis id="B91EEA85FF8CFFF7FCB56C8EFC0EFC05" box="[842,989,915,940]" italics="true" pageId="4" pageNumber="163">L. dasyantha</emphasis>
</taxonomicName>
, although this occurs almost
<quantity id="4C929B72FF8CFFF7FACB6C8EFA5FFC05" box="[1332,1420,915,940]" metricMagnitude="5" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="163" unit="km" value="900.0">900 km</quantity>
to the north. It differs from
<taxonomicName id="4C6A4D14FF8CFFF7FE5F6CAAFDE2FC79" box="[416,561,951,976]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="dasyantha">
<emphasis id="B91EEA85FF8CFFF7FE5F6CAAFDE2FC79" box="[416,561,951,976]" italics="true" pageId="4" pageNumber="163">L. dasyantha</emphasis>
</taxonomicName>
in its non-mottled leaves (
<emphasis id="B91EEA85FF8CFFF7FCA26CAAFC76FC79" box="[861,933,951,976]" italics="true" pageId="4" pageNumber="163">versus</emphasis>
mottled), erect inflorescences (
<emphasis id="B91EEA85FF8CFFF7FAF86CAAFA9CFC79" box="[1287,1359,951,976]" italics="true" pageId="4" pageNumber="163">versus</emphasis>
curving down below the leaves), and wiry flowering braches (
<emphasis id="B91EEA85FF8CFFF7FD1B6CC6FCFFFC5D" box="[740,812,987,1012]" italics="true" pageId="4" pageNumber="163">versus</emphasis>
thick and fleshy). Flowers are distantly spaced along the flowering branches, as opposed to the arrangement in
<taxonomicName id="4C6A4D14FF8CFFF7FD1A6CE2FCA5FBB1" box="[741,886,1023,1048]" class="Liliopsida" family="Arecaceae" genus="Lanonia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="163" phylum="Tracheophyta" rank="species" species="dasyantha">
<emphasis id="B91EEA85FF8CFFF7FD1A6CE2FCA5FBB1" box="[741,886,1023,1048]" italics="true" pageId="4" pageNumber="163">L. dasyantha</emphasis>
</taxonomicName>
in which they are closely spaced and the fruits are consequently closely packed in the infructescence.
</paragraph>
</subSubSection>
</treatment>
</document>