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<document id="59D444D6F2E367D8C2BD38A4C5815395" ID-CLB-Dataset="298364" ID-DOI="10.11646/zootaxa.5458.3.3" ID-GBIF-Dataset="b4c419df-34bc-4632-b668-4a610d362be1" ID-ISSN="1175-5326" ID-Zenodo-Dep="11547498" ID-ZooBank="55870A4F-5D5E-4C12-BAFD-F8C395D7649A" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1718021033248" checkinUser="plazi" docAuthor="Kaila, Lauri" docDate="2024" docId="03AB87D7FF8D3E0E1880FF3DFB6AFB53" docLanguage="en" docName="zootaxa.5458.3.3.pdf" docOrigin="Zootaxa 5458 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5458.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coelopoeta fissurina Kaila 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="379" masterDocId="FF92FFAFFF9F3E1C1817FFAAFFB4FFBA" masterDocTitle="A review of Coelopoetinae (Lepidoptera, Gelechioidea, Pterolonchidae), a moth subfamily confined to western North America, with descriptions of seven new species" masterLastPageNumber="384" masterPageNumber="361" pageNumber="379" updateTime="1718120867107" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3296923BF881CFFE67AD31FB0FF2D8FE">A review of Coelopoetinae (Lepidoptera, Gelechioidea, Pterolonchidae), a moth subfamily confined to western North America, with descriptions of seven new species</mods:title>
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<mods:namePart id="45E91804DEE0C82C582FAC2BDC50B384">Kaila, Lauri</mods:namePart>
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<mods:date id="183EBF58C5D4D0AC5573B4A91D7F1A8E">2024</mods:date>
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<treatment id="03AB87D7FF8D3E0E1880FF3DFB6AFB53" ID-DOI="http://doi.org/10.5281/zenodo.11547550" ID-GBIF-Taxon="232728967" ID-Zenodo-Dep="11547550" LSID="urn:lsid:plazi:treatment:03AB87D7FF8D3E0E1880FF3DFB6AFB53" httpUri="http://treatment.plazi.org/id/03AB87D7FF8D3E0E1880FF3DFB6AFB53" lastPageNumber="379" pageId="18" pageNumber="379">
<subSubSection id="C318654AFF8D3E0E1880FF3DFDFDFF0B" box="[151,585,151,178]" pageId="18" pageNumber="379" type="nomenclature">
<paragraph id="8BBD36C1FF8D3E0E1880FF3DFDFDFF0B" blockId="18.[151,585,151,213]" box="[151,585,151,178]" pageId="18" pageNumber="379">
<heading id="D0F581ADFF8D3E0E1880FF3DFDFDFF0B" bold="true" box="[151,585,151,178]" fontSize="11" level="1" pageId="18" pageNumber="379" reason="1">
<emphasis id="B976EAD3FF8D3E0E1880FF3DFDFDFF0B" bold="true" box="[151,585,151,178]" pageId="18" pageNumber="379">
<taxonomicName id="4C024D42FF8D3E0E1880FF3DFE6BFF0B" authority="Kaila" authorityName="Kaila" authorityYear="2024" box="[151,479,151,178]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="fissurina" status="sp. nov.">
<emphasis id="B976EAD3FF8D3E0E1880FF3DFE26FF0B" bold="true" box="[151,402,151,178]" italics="true" pageId="18" pageNumber="379">Coelopoeta fissurina</emphasis>
Kaila
</taxonomicName>
,
<taxonomicNameLabel id="A24557A8FF8D3E0E19FBFF3DFDFDFF0B" box="[492,585,151,177]" pageId="18" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318654AFF8D3E0E1880FF11FED9FE33" pageId="18" pageNumber="379" type="description">
<paragraph id="8BBD36C1FF8D3E0E1880FF11FEAFFF6F" blockId="18.[151,585,151,213]" box="[151,283,187,213]" pageId="18" pageNumber="379">
<figureCitation id="13392A44FF8D3E0E1880FF11FF47FF6F" box="[151,243,187,213]" captionStart="FIGURES 23–28" captionStartId="14.[151,264,1235,1259]" captionTargetBox="[294,1294,346,1212]" captionTargetId="figure-111@14.[294,1294,345,1212]" captionTargetPageId="14" captionText="FIGURES 23–28. Habitus of Coelopoeta spp.All images at the same scale as the first image. Figure 23: C. fulminea sp. nov., ♀, paratype, Canada, British Columbia, Manning Park; Figure 24: C. alboflava sp. nov., ♂, holotype; Figure 25: C. sariae sp. nov. ♂, holotype; Figure 26: C. maiadella Kaila, ♂, holotype; Figure 27: C. fissurina sp. nov., ♂, holotype; Figure 28. C. valalbui, ♂, holotype." figureDoi="http://doi.org/10.5281/zenodo.11547510" httpUri="https://zenodo.org/record/11547510/files/figure.png" pageId="18" pageNumber="379">Figs. 27</figureCitation>
,
<figureCitation id="13392A44FF8D3E0E18E8FF11FEAFFF6F" box="[255,283,187,213]" captionStart="FIGURES 29–38" captionStartId="17.[151,264,1862,1886]" captionTargetBox="[277,1382,168,1852]" captionTargetId="figure-15@17.[196,1391,166,1853]" captionTargetPageId="17" captionText="FIGURES 29–38. Male genitalia of Coelopoeta spp. Above lateral view with juxta and one valva excluded; below ventral view, with phallus excluded. Figure 29. C. glutinosi Walsingham. Figure 30: C. aprica sp. nov., holotype; Figure 31: C. phaceliae Kaila, California, Nevada Co.; Figure 32: C. aurora sp. nov., holotype; Figure 33: C. fulminea sp. nov., holotype; Figure 34: C. alboflava sp. nov., holotype; Figure 35: C. sariae sp. nov., holotype; Figure 36: C. maiadella sp. nov., holotype; Figure 37: C. fissurina sp. nov., holotype; 38: C. valalbui sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.11547518" httpUri="https://zenodo.org/record/11547518/files/figure.png" pageId="18" pageNumber="379">37</figureCitation>
</paragraph>
<paragraph id="8BBD36C1FF8D3E0E1880FEA9FED9FE33" blockId="18.[151,1437,259,1258]" pageId="18" pageNumber="379">
<materialsCitation id="3B6A3C9CFF8D3E0E1880FEA9FED9FE33" ID-GBIF-Occurrence="4876017335" collectingDate="1996-04-24" collectionCode="MZH" collectorName="L. Kaila &amp; J. Kullberg" country="United States of America" county="Cochise Co." elevation="1300" latitude="31.93" location="Portal - Paradise cyn. road, 2 mi. N. Portal" longLatPrecision="728" longitude="-109.2" municipality="Chiricahua Mts." pageId="18" pageNumber="379" specimenCode="MZH-LEO 00000294" specimenCount="1" specimenCount-male="1" stateProvince="Arizona" typeStatus="holotype">
<typeStatus id="54B98863FF8D3E0E1880FEA9FF4BFEA7" box="[151,255,259,285]" pageId="18" pageNumber="379" type="holotype">Holotype</typeStatus>
<specimenCount id="9D04FD48FF8D3E0E1910FEAEFEABFEA4" box="[263,287,260,286]" count="1" pageId="18" pageNumber="379" type="male">♂</specimenCount>
:
<collectingCountry id="F3157651FF8D3E0E193CFEA9FECEFEA7" box="[299,378,259,285]" name="United States of America" pageId="18" pageNumber="379">U.S.A.</collectingCountry>
,
<collectingRegion id="49C6F823FF8D3E0E1992FEA9FE56FEA7" box="[389,482,259,285]" country="United States of America" name="Arizona" pageId="18" pageNumber="379">Arizona</collectingRegion>
,
<geoCoordinate id="EE365006FF8D3E0E19F9FEA9FDFBFEA7" box="[494,591,259,285]" degrees="31.93" direction="north" orientation="latitude" pageId="18" pageNumber="379" precision="555" value="31.93">31.93°N</geoCoordinate>
,
<geoCoordinate id="EE365006FF8D3E0E1A4DFEA9FD79FEA7" box="[602,717,259,285]" degrees="109.20" direction="west" orientation="longitude" pageId="18" pageNumber="379" precision="555" value="-109.2">109.20°W</geoCoordinate>
,
<collectingCounty id="62DC4E4DFF8D3E0E1ACEFEA9FCD1FEA7" box="[729,869,259,285]" pageId="18" pageNumber="379">Cochise Co.</collectingCounty>
,
<collectingMunicipality id="6BD9ACBBFF8D3E0E1B66FEA9FB9CFEA4" box="[881,1064,259,286]" pageId="18" pageNumber="379">Chiricahua Mts.</collectingMunicipality>
,
<location id="8EDD601AFF8D3E0E1C23FEA9FEB6FEFB" LSID="urn:lsid:plazi:treatment:03AB87D7FF8D3E0E1880FF3DFB6AFB53:8EDD601AFF8D3E0E1C23FEA9FEB6FEFB" country="United States of America" county="Cochise Co." latitude="31.93" longLatPrecision="728" longitude="-109.2" municipality="Chiricahua Mts." name="Portal - Paradise cyn. road, 2 mi. N. Portal" pageId="18" pageNumber="379" stateProvince="Arizona">
Portal–Paradise cyn. road,
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N. Portal
</location>
,
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<elevation id="002FD1F2FF8D3E0E1918FE8DFEDDFEF8" box="[271,361,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="379" unit="m" value="1300.0">1300 m</elevation>
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, grass/
<taxonomicName id="4C024D42FF8D3E0E19AFFE82FDA0FEF8" box="[440,532,296,322]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="18" pageNumber="379" phylum="Tracheophyta" rank="genus">
<emphasis id="B976EAD3FF8D3E0E19AFFE82FDA0FEF8" box="[440,532,296,322]" italics="true" pageId="18" pageNumber="379">Opuntia</emphasis>
</taxonomicName>
,
<date id="FFBC1001FF8D3E0E1A33FE8DFD29FEFB" box="[548,669,295,321]" pageId="18" pageNumber="379" value="1996-04-24">
<collectingDate id="EFF8E9E9FF8D3E0E1A33FE8DFD29FEFB" box="[548,669,295,321]" pageId="18" pageNumber="379" value="1996-04-24">24.iv.1996</collectingDate>
</date>
, ad luc.
<collectorName id="26F75317FF8D3E0E1B16FE82FCD4FEFB" box="[769,864,295,322]" pageId="18" pageNumber="379">L. Kaila</collectorName>
&amp;
<collectorName id="26F75317FF8D3E0E1B9FFE82FBB2FEFB" box="[904,1030,295,322]" pageId="18" pageNumber="379">J. Kullberg</collectorName>
leg. L. Kaila prep. 6372; Database #
<specimenCode id="DBA49EBAFF8D3E0E18A7FEE1FE19FEDF" box="[176,429,331,357]" pageId="18" pageNumber="379">MZH-LEO 00000294</specimenCode>
;
<taxonomicName id="4C024D42FF8D3E0E19ACFEE1FDD3FEDC" box="[443,615,331,358]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B976EAD3FF8D3E0E19ACFEE1FD88FEDF" box="[443,572,331,357]" italics="true" pageId="18" pageNumber="379">Coelopoeta</emphasis>
sp.
</taxonomicName>
det. J.-F. Landry 2016; DNA-voucher FinBOL 2019; http://id.luomus. fi/KZ7707 (
<collectionCode id="ED13AE04FF8D3E0E190BFEC5FED4FE33" box="[284,352,367,393]" pageId="18" pageNumber="379">MZH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C318654AFF8D3E0E18D0FE39FC72FD87" pageId="18" pageNumber="379" type="diagnosis">
<paragraph id="8BBD36C1FF8D3E0E18D0FE39FC72FD87" blockId="18.[151,1437,259,1258]" pageId="18" pageNumber="379">
<emphasis id="B976EAD3FF8D3E0E18D0FE39FEF6FE17" bold="true" box="[199,322,403,429]" pageId="18" pageNumber="379">Diagnosis.</emphasis>
<taxonomicName id="4C024D42FF8D3E0E195EFE3EFE79FE17" authorityName="Kaila" authorityYear="2024" box="[329,461,403,430]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="fissurina">
<emphasis id="B976EAD3FF8D3E0E195EFE3EFE79FE17" box="[329,461,403,430]" italics="true" pageId="18" pageNumber="379">C. fissurina</emphasis>
</taxonomicName>
is a mottled pale grey species, with forewing colour formed of white and grey scales. The male genitalia are characterized by the bilobed uncus lobes and the basally inflated phallus. With these genital characters it is readily distinguished from all other
<taxonomicName id="4C024D42FF8D3E0E1AA8FE71FCF4FE4F" authorityName="Walsingham" authorityYear="1907" box="[703,832,475,501]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAD3FF8D3E0E1AA8FE71FCF4FE4F" box="[703,832,475,501]" italics="true" pageId="18" pageNumber="379">Coelopoeta</emphasis>
</taxonomicName>
apart from
<emphasis id="B976EAD3FF8D3E0E1BAAFE76FB8BFE4F" box="[957,1087,475,502]" italics="true" pageId="18" pageNumber="379">
<taxonomicName id="4C024D42FF8D3E0E1BAAFE76FB8EFE4F" authorityName="Kaila" authorityYear="2024" box="[957,1082,475,502]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="valalbui">C. valalbui</taxonomicName>
.
</emphasis>
These species are distinguished from each other by the shorter valva with a distally blunt apex in
<taxonomicName id="4C024D42FF8D3E0E1B79FDAAFC44FDA3" authorityName="Kaila" authorityYear="2024" box="[878,1008,511,538]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="fissurina">
<emphasis id="B976EAD3FF8D3E0E1B79FDAAFC44FDA3" box="[878,1008,511,538]" italics="true" pageId="18" pageNumber="379">C. fissurina</emphasis>
</taxonomicName>
. The uncus of
<taxonomicName id="4C024D42FF8D3E0E1C82FDAAFAA3FDA3" authorityName="Kaila" authorityYear="2024" box="[1173,1303,511,538]" class="Insecta" family="Pterolonchidae" genus="Coelopoeta" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="379" phylum="Arthropoda" rank="species" species="fissurina">
<emphasis id="B976EAD3FF8D3E0E1C82FDAAFAA3FDA3" box="[1173,1303,511,538]" italics="true" pageId="18" pageNumber="379">C. fissurina</emphasis>
</taxonomicName>
far exceeds the length of the valva in ventral view (
<figureCitation id="13392A44FF8D3E0E1A47FD89FD10FD87" box="[592,676,547,573]" captionStart="FIGURES 29–38" captionStartId="17.[151,264,1862,1886]" captionTargetBox="[277,1382,168,1852]" captionTargetId="figure-15@17.[196,1391,166,1853]" captionTargetPageId="17" captionText="FIGURES 29–38. Male genitalia of Coelopoeta spp. Above lateral view with juxta and one valva excluded; below ventral view, with phallus excluded. Figure 29. C. glutinosi Walsingham. Figure 30: C. aprica sp. nov., holotype; Figure 31: C. phaceliae Kaila, California, Nevada Co.; Figure 32: C. aurora sp. nov., holotype; Figure 33: C. fulminea sp. nov., holotype; Figure 34: C. alboflava sp. nov., holotype; Figure 35: C. sariae sp. nov., holotype; Figure 36: C. maiadella sp. nov., holotype; Figure 37: C. fissurina sp. nov., holotype; 38: C. valalbui sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.11547518" httpUri="https://zenodo.org/record/11547518/files/figure.png" pageId="18" pageNumber="379">Fig. 37</figureCitation>
). The female is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C318654AFF8D3E0E18D0FDEDFE25FBC7" pageId="18" pageNumber="379" type="description">
<paragraph id="8BBD36C1FF8D3E0E18D0FDEDFF69FD4B" blockId="18.[151,1437,259,1258]" pageId="18" pageNumber="379">
<emphasis id="B976EAD3FF8D3E0E18D0FDEDFEE5FDDB" bold="true" box="[199,337,583,609]" pageId="18" pageNumber="379">Description</emphasis>
. Forewing length
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. Labial palpus porrect, length 2/3 of diameter of head, brownish grey. Head, neck tuft, tegula, thorax, scape and pecten of antenna white. Flagellum unicolourous grey, not serrate. Legs grey. Forewing ground colour formed of unicolourous, white and pale brownish grey scales, hindwing grey with concolorous fringe. Underside of wings grey with concolorous fringe apart from forewing costa where fringe is white.
</paragraph>
<paragraph id="8BBD36C1FF8D3E0E18D0FD51FE63FBE3" blockId="18.[151,1437,259,1258]" pageId="18" pageNumber="379">Male genitalia. Uncus bilobed, lobes horn-shaped, bent inward, tapered towards curved and acute apex, as long as tegumen. Gnathos 3/4 as long as uncus, tapered in distal third to pointed apex. Tegumen triangular-shaped in lateral view, twice as long as high at its highest point; height half of width of valva in lateral view. Valva as wide as long when measured in the middle where valva is broadest, forming swelling; weakly bent inwards in distal half. Termen somewhat produced in costal part, blunt; sacculus apically separated into small, sharp-tipped lobe. Juxta distally markedly concave, laterally tapered into hook, tapered to convex anterior margin. Saccus slightly bent dorsad, evenly tapered to pointed apex, as long as its width basally. Phallus twice as long as valva, gradually widened from apex to basal 3/4; followed by abrupt narrowing and then dilating in a rounded basal part with basal opening ventrally directed; vesica with two groups of scobinations: one in the middle of phallus, another near apex; apex without carina or teeth.</paragraph>
<paragraph id="8BBD36C1FF8D3E0E18D0FBC9FE25FBC7" blockId="18.[151,1437,259,1258]" box="[199,401,1123,1149]" pageId="18" pageNumber="379">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C318654AFF8D3E0E18D0FB2DFEDEFB7C" pageId="18" pageNumber="379" type="biology_ecology">
<paragraph id="8BBD36C1FF8D3E0E18D0FB2DFEDEFB7C" blockId="18.[151,1437,259,1258]" pageId="18" pageNumber="379">
<emphasis id="B976EAD3FF8D3E0E18D0FB2DFE95FB1B" bold="true" box="[199,289,1159,1185]" pageId="18" pageNumber="379">Biology</emphasis>
. The habitat is rocky submontane prairie-like habitat with low vegetation and dense stands of
<taxonomicName id="4C024D42FF8D3E0E1D2EFB22FA21FB18" box="[1337,1429,1160,1186]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="18" pageNumber="379" phylum="Tracheophyta" rank="genus">
<emphasis id="B976EAD3FF8D3E0E1D2EFB22FA21FB18" box="[1337,1429,1160,1186]" italics="true" pageId="18" pageNumber="379">Opuntia</emphasis>
</taxonomicName>
, at altitude
<quantity id="4CFA9B24FF8D3E0E191AFB01FED1FB7C" box="[269,357,1195,1222]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="379" unit="m" value="1300.0">1300 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C318654AFF8D3E0E18D0FB65FB6AFB53" box="[199,1246,1231,1258]" pageId="18" pageNumber="379" type="distribution">
<paragraph id="8BBD36C1FF8D3E0E18D0FB65FB6AFB53" blockId="18.[151,1437,259,1258]" box="[199,1246,1231,1258]" pageId="18" pageNumber="379">
<emphasis id="B976EAD3FF8D3E0E18D0FB65FEEDFB53" bold="true" box="[199,345,1231,1257]" pageId="18" pageNumber="379">Distribution</emphasis>
. Known only from the
<typeStatus id="54B98863FF8D3E0E1A77FB7AFD24FB50" box="[608,656,1232,1258]" pageId="18" pageNumber="379">type</typeStatus>
locality in the Chiricahua Mountains in SE
<collectingRegion id="49C6F823FF8D3E0E1C6BFB65FB6EFB53" box="[1148,1242,1231,1257]" country="United States of America" name="Arizona" pageId="18" pageNumber="379">Arizona</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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