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<document id="B15903E197C75DFEB1837423C940A15E" ID-DOI="10.11646/zootaxa.5541.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14241202" ID-ZooBank="26AD8B52-4B4A-4152-8350-C2DF28E288AD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732836169173" checkinUser="plazi" docAuthor="Obrial, Graden G., Agbas, Daven Jayson D., Medina, Milton Norman D. &amp; Cabras, Analyn A." docDate="2024" docId="03A2985E4F29760EFF0AFA3D3A32C3E7" docLanguage="en" docName="zootaxa.5541.4.2.pdf" docOrigin="Zootaxa 5541 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metapocyrtus inangsabong Cabras, Obrial &amp; Agbas 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="441" masterDocId="FF9BE0264F28760DFF9DFFF63E53C45E" masterDocTitle="Three New Mimetic Weevils (Coleoptera, Curculionidae, Entiminae) from Mt. Candalaga, Davao de Oro, Mindanao Island, Philippines" masterLastPageNumber="454" masterPageNumber="438" pageNumber="439" updateTime="1732920951879" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5D4C8487AB72AAE91E51AAAED7184C9B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1B9DE3ED9C8AD3B29C239330049A8F1B">
<mods:title id="9297F544E5B2130DBE9CF994FA9C13EB">Three New Mimetic Weevils (Coleoptera, Curculionidae, Entiminae) from Mt. Candalaga, Davao de Oro, Mindanao Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="AFCB371BF4ED5965185AE3A623A88136" type="personal">
<mods:role id="16234CAE40453E7E755C9F71662D6C3D">
<mods:roleTerm id="EEFD7FB4D43E9F18395B53E4624D2031">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="930580BB26F2215A998967CE91B30DDC">Obrial, Graden G.</mods:namePart>
<mods:nameIdentifier id="8EE69A49BBF6A32B2C62B5A33D39C656" type="ORCID">0009-0004-4163-6942</mods:nameIdentifier>
<mods:nameIdentifier id="3F0D1142CE5E25B6356F2CA03952B728" type="email">graden.obrial@dorsu.edu.ph</mods:nameIdentifier>
</mods:name>
<mods:name id="9D1C7B63C77C21BC26313236BF6D7A9A" type="personal">
<mods:role id="E0C26ACBC228507D088FD0D987DB731B">
<mods:roleTerm id="4513EBCDBBC7DD4BEA8383A37CD894F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A9DF89C5D47A41AEDE23853FCD0B5C1">Agbas, Daven Jayson D.</mods:namePart>
<mods:affiliation id="4DCE348777CC99A3526FF3ABA7017545">Faculty of Computing, Data Sciences, Engineering, and Technology, Information Technology Program, Davao Oriental State University, Dahican, Mati City 8200 Philippines</mods:affiliation>
</mods:name>
<mods:name id="3070F6D169CAB9E1AB9F5E2B1AFED7BE" type="personal">
<mods:role id="A444EB5CA2C4E20876C361AFE8CEF5B3">
<mods:roleTerm id="BC0855AE64E7CB3C716BEA742CA1B9D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8776511CB543F3A4634FD07CF652819D">Medina, Milton Norman D.</mods:namePart>
<mods:affiliation id="05FB3F052459A3A18A00E85CAA2E997B">Tropical Genomics Laboratory, URESCOM, Davao Oriental State University, Dahican, City of Mati, 8200, Philippines</mods:affiliation>
</mods:name>
<mods:name id="2B3DF27BBA2A6541782C3CF8FDC1FA05" type="personal">
<mods:role id="F7B10EC8DA1E3445054A7DA510A05F56">
<mods:roleTerm id="E6CC7C94645F653CDDE8BEB96665D080">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB9ECFBCB37AC2995FF4C48F3461F0E1">Cabras, Analyn A.</mods:namePart>
<mods:nameIdentifier id="43DFF609566AA6C5B1B833778AB401AC" type="ORCID">0000-0002-0980-1651</mods:nameIdentifier>
<mods:nameIdentifier id="60B18D54905DEA886D6E29205DC94DFB" type="email">ann.cabras24@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="83ABCDD6B8DA575ECE92F66C5A74D82E">text</mods:typeOfResource>
<mods:relatedItem id="95F40BAA9012ED790963EACEBC288385" type="host">
<mods:titleInfo id="6E7270B97C83B5A0583C648E11F8A126">
<mods:title id="5363B6A07044071755A5A397FC70BC01">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="82DA5644E58D22456D562EB7736ABB3D">
<mods:date id="C4F9D924FB5EC0D8898AF56E106AF735">2024</mods:date>
<mods:detail id="D9EDB6983C5BC473A550BAA92F2FA91E" type="pubDate">
<mods:number id="0CEC6D9903357D5097C2BF1A738031D6">2024-11-28</mods:number>
</mods:detail>
<mods:detail id="54D459B9C1421A4B3F5613FB685C108A" type="volume">
<mods:number id="321E1862A50FB4402F9E79717A1F062D">5541</mods:number>
</mods:detail>
<mods:detail id="E942319040190EC127382DE5823FCEE0" type="issue">
<mods:number id="CFC71D91D3F69872DEC99DF878AC0ECE">4</mods:number>
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<mods:start id="3B4B62C06664204A587B37E9CF099446">438</mods:start>
<mods:end id="25A806E61081CF434DD2FFAB9F5075C8">454</mods:end>
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<mods:location id="154DCE012444AA2F2717009E25B24ED4">
<mods:url id="E161CC90A4C8537F4687B4E8272E0967">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
<mods:classification id="7AEC93029242B5A74D4AA85A696F4527">journal article</mods:classification>
<mods:identifier id="1DE5A5C08E4CC73CB47D6F5983C519BB" type="DOI">10.11646/zootaxa.5541.4.2</mods:identifier>
<mods:identifier id="4C3F3F4877726A3F9B8E2C7C5291111A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A290F61FBFA00B450A2FD8B330537596" type="Zenodo-Dep">14241202</mods:identifier>
<mods:identifier id="3CFC96955A4EE536BF305C1884D5137B" type="ZooBank">26AD8B52-4B4A-4152-8350-C2DF28E288AD</mods:identifier>
</mods:mods>
<treatment id="03A2985E4F29760EFF0AFA3D3A32C3E7" ID-DOI="http://doi.org/10.5281/zenodo.14248471" ID-Zenodo-Dep="14248471" LSID="urn:lsid:plazi:treatment:03A2985E4F29760EFF0AFA3D3A32C3E7" httpUri="http://treatment.plazi.org/id/03A2985E4F29760EFF0AFA3D3A32C3E7" lastPageId="3" lastPageNumber="441" pageId="1" pageNumber="439">
<subSubSection id="C3117AC34F29760CFF0AFA3D3D83C257" pageId="1" pageNumber="439" type="nomenclature">
<paragraph id="8BB429484F29760CFF0AFA3D3D2EC1BB" blockId="1.[151,976,1482,1581]" box="[151,893,1482,1509]" pageId="1" pageNumber="439">
<heading id="D0FC9E244F29760CFF0AFA3D3D2EC1BB" bold="true" box="[151,893,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="439" reason="1">
<emphasis id="B97FF55A4F29760CFF0AFA3D3D2EC1BB" bold="true" box="[151,893,1482,1509]" pageId="1" pageNumber="439">
<taxonomicName id="4C0B52CB4F29760CFF0AFA3D3D44C1BA" authority="Cabras, Obrial &amp; Agbas" authorityName="Cabras, Obrial &amp; Agbas" authorityYear="2024" box="[151,791,1482,1509]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="inangsabong" status="sp. nov.">
<emphasis id="B97FF55A4F29760CFF0AFA3D3F8CC1BB" bold="true" box="[151,479,1482,1509]" italics="true" pageId="1" pageNumber="439">Metapocyrtus inangsabong</emphasis>
Cabras, Obrial &amp; Agbas
</taxonomicName>
<taxonomicNameLabel id="A24C48214F29760CFC82FA3D3D2EC1BB" box="[799,893,1483,1509]" pageId="1" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB429484F29760CFF0AFA183D83C257" blockId="1.[151,976,1482,1581]" box="[151,976,1518,1545]" pageId="1" pageNumber="439">
<uri id="FF9A254A4F29760CFF0AFA183D83C257" box="[151,976,1518,1545]" pageId="1" pageNumber="439">
urn:lsid:zoobank.org:act:
<uuid id="FFAD139D4F29760CFE37FA183D83C257" box="[426,976,1518,1545]" pageId="1" pageNumber="439">582E0F1C-EC8B-410E-9C6E-B92B97ABC65F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3117AC34F29760CFF0AF9E53F61C273" box="[151,306,1555,1581]" pageId="1" pageNumber="439" type="description">
<paragraph id="8BB429484F29760CFF0AF9E53F61C273" blockId="1.[151,976,1482,1581]" box="[151,306,1555,1581]" pageId="1" pageNumber="439">
<figureCitation id="133035CD4F29760CFF0AF9E53F61C273" box="[151,306,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-22@2.[151,1436,181,1755]" captionTargetPageId="2" captionText="FIGURE 1. Metapocyrtus inangsabong A—♂ holotype, dorsal view, C­—idem, lateral view; B—♀ paratype, dorsal view, D—idem, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14241204" httpUri="https://zenodo.org/record/14241204/files/figure.png" pageId="1" pageNumber="439">Figure 1 A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3117AC34F29760CFF0AF9AD3A8FC313" pageId="1" pageNumber="439" type="materials_examined">
<paragraph id="8BB429484F29760CFF0AF9AD3A8FC313" blockId="1.[151,1436,1627,2013]" pageId="1" pageNumber="439">
<materialsCitation id="3B6323154F29760CFF0AF9AD3A8FC313" collectingDate="2024-01" collectingDateMax="2024-06" collectingDateMin="2024-01" collectionCode="TIRL, PNM, SMTD, CMNC" collectorName="E. Avergonzado &amp; G. Obrial &amp; D. Agbas &amp; Y. Japitan" country="Philippines" county="Presently" location="Mindanao Island" municipality="Mindanao Island" pageId="1" pageNumber="439" specimenCount="26" specimenCount-female="18" specimenCount-male="8" stateProvince="Davao" typeStatus="holotype">
<emphasis id="B97FF55A4F29760CFF0AF9AD3F57C22B" bold="true" box="[151,260,1627,1653]" pageId="1" pageNumber="439">
<typeStatus id="54B097EA4F29760CFF0AF9AD3F57C22B" box="[151,260,1627,1653]" pageId="1" pageNumber="439" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="133035CD4F29760CFE88F9AD3FC4C22B" box="[277,407,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-22@2.[151,1436,181,1755]" captionTargetPageId="2" captionText="FIGURE 1. Metapocyrtus inangsabong A—♂ holotype, dorsal view, C­—idem, lateral view; B—♀ paratype, dorsal view, D—idem, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14241204" httpUri="https://zenodo.org/record/14241204/files/figure.png" pageId="1" pageNumber="439">Fig. 1 A, C</figureCitation>
), male: “
<collectingCountry id="F31C69D84F29760CFE63F9AD3C2EC22B" box="[510,637,1627,1653]" name="Philippines" pageId="1" pageNumber="439">Philippines</collectingCountry>
—Mindanao Island,
<collectingRegion id="49CFE7AA4F29760CFCF5F9AD3DE2C22B" box="[872,945,1627,1653]" country="Philippines" name="Davao" pageId="1" pageNumber="439">Davao</collectingRegion>
de Oro, Maragusan/
<date id="FFB50F884F29760CFB3EF9AD3B69C22B" box="[1187,1338,1627,1653]" pageId="1" pageNumber="439" value="2024-01">
<collectingDate id="EFF1F6604F29760CFB3EF9AD3B69C22B" box="[1187,1338,1627,1653]" pageId="1" pageNumber="439" value="2024-01">January 2024</collectingDate>
</date>
/ leg.
<collectorName id="26FE4C9E4F29760CFA19F9AD3F78C2C7" pageId="1" pageNumber="439">E. Avergonzado</collectorName>
(typed on white card) //
<typeStatus id="54B097EA4F29760CFDCEF9893CB4C2C7" box="[595,743,1663,1689]" pageId="1" pageNumber="439" type="holotype">HOLOTYPE</typeStatus>
male /
<taxonomicName id="4C0B52CB4F29760CFCDEF9893A2BC2C7" authorityName="Cabras, Obrial &amp; Agbas" authorityYear="2024" box="[835,1144,1663,1689]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="inangsabong">
<emphasis id="B97FF55A4F29760CFCDEF9893A2BC2C7" box="[835,1144,1663,1689]" italics="true" pageId="1" pageNumber="439">Metapocyrtus inangsabong</emphasis>
</taxonomicName>
/ CABRAS, OBRIAL &amp; AGBAS, 2024 (typed on red card)” (
<collectingCounty id="62D551C44F29760CFDABF9553CF3C2E3" box="[566,672,1699,1725]" pageId="1" pageNumber="439">Presently</collectingCounty>
at
<collectionCode id="ED1AB18D4F29760CFD5FF9553D53C2E3" box="[706,768,1699,1725]" pageId="1" pageNumber="439">TIRL</collectionCode>
, to be deposited at
<collectionCode id="ED1AB18D4F29760CFC41F9553A4FC2E3" box="[988,1052,1699,1725]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="1" pageNumber="439">PNM</collectionCode>
.
<emphasis id="B97FF55A4F29760CFBBAF9553AC7C2E3" bold="true" box="[1063,1172,1699,1725]" pageId="1" pageNumber="439">
<typeStatus id="54B097EA4F29760CFBBAF9553AC7C2E3" box="[1063,1172,1699,1725]" pageId="1" pageNumber="439" type="paratype">Paratype</typeStatus>
</emphasis>
(
<specimenCount id="9D0DE2C14F29760CFB3EF9553AADC2E3" box="[1187,1278,1699,1725]" count="4" pageId="1" pageNumber="439" type="male">4 males</specimenCount>
,
<specimenCount id="9D0DE2C14F29760CFA94F9553B28C2E3" box="[1289,1403,1699,1725]" count="9" pageId="1" pageNumber="439" type="female">9 females</specimenCount>
);
<specimenCount id="9D0DE2C14F29760CFA13F9553EB5C2BF" count="1" pageId="1" pageNumber="439" type="female">1 female</specimenCount>
, same data as the holotype;
<specimenCount id="9D0DE2C14F29760CFDB8F9313C27C2BF" box="[549,628,1735,1761]" count="1" pageId="1" pageNumber="439" type="male">1 male</specimenCount>
,
<specimenCount id="9D0DE2C14F29760CFD1DF9313CA3C2BF" box="[640,752,1735,1761]" count="3" pageId="1" pageNumber="439" type="female">3 females</specimenCount>
,
<collectingCountry id="F31C69D84F29760CFD61F9313D29C2BF" box="[764,890,1735,1761]" name="Philippines" pageId="1" pageNumber="439">Philippines</collectingCountry>
—
<collectingMunicipality id="6BD0B3324F29760CFC0BF9313A05C2BF" box="[918,1110,1735,1761]" pageId="1" pageNumber="439">Mindanao Island</collectingMunicipality>
,
<collectingRegion id="49CFE7AA4F29760CFBFEF9313AFFC2BF" box="[1123,1196,1735,1761]" country="Philippines" name="Davao" pageId="1" pageNumber="439">Davao</collectingRegion>
de Oro, Maragusan /
<date id="FFB50F884F29760CFF0AF91D3F6BC35B" box="[151,312,1771,1797]" pageId="1" pageNumber="439" value="2024-02">
<collectingDate id="EFF1F6604F29760CFF0AF91D3F6BC35B" box="[151,312,1771,1797]" pageId="1" pageNumber="439" value="2024-02">February 2024</collectingDate>
</date>
/leg.
<collectorName id="26FE4C9E4F29760CFEEEF91D3F88C35B" box="[371,475,1771,1797]" pageId="1" pageNumber="439">G.Obrial</collectorName>
,
<collectorName id="26FE4C9E4F29760CFE7EF91D3C18C35B" box="[483,587,1771,1797]" pageId="1" pageNumber="439">D.Agbas</collectorName>
;
<specimenCount id="9D0DE2C14F29760CFDCEF91D3CFBC35B" box="[595,680,1771,1797]" count="3" pageId="1" pageNumber="439" type="male">3males</specimenCount>
,
<specimenCount id="9D0DE2C14F29760CFD32F91D3D48C35B" box="[687,795,1771,1797]" count="3" pageId="1" pageNumber="439" type="female">3 females</specimenCount>
,
<collectingCountry id="F31C69D84F29760CFCBFF91D3DF3C35B" box="[802,928,1771,1797]" name="Philippines" pageId="1" pageNumber="439">Philippines</collectingCountry>
—
<location id="8ED47F934F29760CFC20F91D3A2BC35B" LSID="urn:lsid:plazi:treatment:03A2985E4F29760EFF0AFA3D3A32C3E7:8ED47F934F29760CFC20F91D3A2BC35B" box="[957,1144,1771,1797]" country="Philippines" county="Presently" municipality="Mindanao Island" name="Mindanao Island" pageId="1" pageNumber="439" stateProvince="Davao">Mindanao Island</location>
,
<collectingRegion id="49CFE7AA4F29760CFB1DF91D3A9AC35B" box="[1152,1225,1771,1797]" country="Philippines" name="Davao" pageId="1" pageNumber="439">Davao</collectingRegion>
de Oro, Maragusan /
<date id="FFB50F884F29760CFF3BF8F93F4BC377" box="[166,280,1807,1833]" pageId="1" pageNumber="439" value="2024-06">
<collectingDate id="EFF1F6604F29760CFF3BF8F93F4BC377" box="[166,280,1807,1833]" pageId="1" pageNumber="439" value="2024-06">June 2024</collectingDate>
</date>
/ leg.
<collectorName id="26FE4C9E4F29760CFEC3F8F93F99C377" box="[350,458,1807,1833]" pageId="1" pageNumber="439">G. Obrial</collectorName>
,
<collectorName id="26FE4C9E4F29760CFE4BF8F93C12C377" box="[470,577,1807,1833]" pageId="1" pageNumber="439">D. Agbas</collectorName>
;
<specimenCount id="9D0DE2C14F29760CFDD0F8F93CEFC377" box="[589,700,1807,1833]" count="2" pageId="1" pageNumber="439" type="female">2 females</specimenCount>
,
<collectingCountry id="F31C69D84F29760CFD5AF8F93D16C377" box="[711,837,1807,1833]" name="Philippines" pageId="1" pageNumber="439">Philippines</collectingCountry>
—
<location id="8ED47F934F29760CFCFCF8F93A73C377" LSID="urn:lsid:plazi:treatment:03A2985E4F29760EFF0AFA3D3A32C3E7:8ED47F934F29760CFCFCF8F93A73C377" box="[865,1056,1807,1833]" country="Philippines" county="Presently" municipality="Mindanao Island" name="Mindanao Island" pageId="1" pageNumber="439" stateProvince="Davao">Mindanao Island</location>
,
<collectingRegion id="49CFE7AA4F29760CFBB1F8F93A26C377" box="[1068,1141,1807,1833]" country="Philippines" name="Davao" pageId="1" pageNumber="439">Davao</collectingRegion>
de Oro, Maragusan /
<date id="FFB50F884F29760CFAF4F8F93E9CC313" pageId="1" pageNumber="439" value="2024-06">
<collectingDate id="EFF1F6604F29760CFAF4F8F93E9CC313" pageId="1" pageNumber="439" value="2024-06">June 2024</collectingDate>
</date>
/ leg.
<collectorName id="26FE4C9E4F29760CFE88F8C53F99C313" box="[277,458,1843,1869]" pageId="1" pageNumber="439">E. Avergonzado</collectorName>
,
<collectorName id="26FE4C9E4F29760CFE49F8C53C11C313" box="[468,578,1843,1869]" pageId="1" pageNumber="439">Y. Japitan</collectorName>
(all in
<collectionCode id="ED1AB18D4F29760CFD0CF8C53C9CC313" box="[657,719,1843,1869]" pageId="1" pageNumber="439">TIRL</collectionCode>
, to be deposited at
<collectionCode id="ED1AB18D4F29760CFC34F8C53DBAC313" box="[937,1001,1843,1869]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="1" pageNumber="439">PNM</collectionCode>
,
<collectionCode id="ED1AB18D4F29760CFC69F8C53A11C313" box="[1012,1090,1843,1869]" pageId="1" pageNumber="439">SMTD</collectionCode>
and
<collectionCode id="ED1AB18D4F29760CFBE5F8C53A83C313" box="[1144,1232,1843,1869]" pageId="1" pageNumber="439">CMNC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3117AC34F29760CFF5AF8A03A83C383" pageId="1" pageNumber="439" type="diagnosis">
<paragraph id="8BB429484F29760CFF5AF8A03A83C383" blockId="1.[151,1436,1627,2013]" pageId="1" pageNumber="439">
<emphasis id="B97FF55A4F29760CFF5AF8A03F11C32E" bold="true" box="[199,322,1878,1904]" pageId="1" pageNumber="439">Diagnosis.</emphasis>
<taxonomicName id="4C0B52CB4F29760CFED7F8A13C2DC32F" authorityName="Cabras, Obrial &amp; Agbas" authorityYear="2024" box="[330,638,1879,1905]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="439" phylum="Arthropoda" rank="species" species="inangsabong">
<emphasis id="B97FF55A4F29760CFED7F8A13C2DC32F" box="[330,638,1879,1905]" italics="true" pageId="1" pageNumber="439">Metapocyrtus inangsabong</emphasis>
</taxonomicName>
is unique from its congeners for its combination of characters having shiny dark crimson integument, subglabrous head, pronotum, and elytra, head and rostrum almost continuous, separated only by short transverse groove not reaching lateral margin, and apex of elytra in females with two, minute spines. So far, it is only found in the higher elevation of Mt. Candalaga in
<collectingRegion id="49CFE7AA4F29760CFBB6F8353A27C383" box="[1067,1140,1987,2013]" country="Philippines" name="Davao" pageId="1" pageNumber="439">Davao</collectingRegion>
de Oro.
</paragraph>
</subSubSection>
<caption id="DF7479C04F2A760FFF0AF9053F2CC371" ID-DOI="http://doi.org/10.5281/zenodo.14241204" ID-Zenodo-Dep="14241204" httpUri="https://zenodo.org/record/14241204/files/figure.png" pageId="2" pageNumber="440" startId="2.[151,250,1779,1803]" targetBox="[151,1436,181,1755]" targetPageId="2" targetType="figure">
<paragraph id="8BB429484F2A760FFF0AF9053F2CC371" blockId="2.[151,1436,1779,1839]" pageId="2" pageNumber="440">
<emphasis id="B97FF55A4F2A760FFF0AF9053F45C355" bold="true" box="[151,278,1779,1803]" pageId="2" pageNumber="440">FIGURE 1.</emphasis>
<taxonomicName id="4C0B52CB4F2A760FFEBDF9023C6BC355" authorityName="Cabras, Obrial &amp; Agbas" authorityYear="2024" box="[288,568,1779,1803]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="440" phylum="Arthropoda" rank="species" species="inangsabong">
<emphasis id="B97FF55A4F2A760FFEBDF9023C6BC355" box="[288,568,1779,1803]" italics="true" pageId="2" pageNumber="440">Metapocyrtus inangsabong</emphasis>
</taxonomicName>
A—♂ holotype, dorsal view, C­—idem, lateral view; B—♀ paratype, dorsal view, D—idem, lateral view.
</paragraph>
</caption>
<subSubSection id="C3117AC34F2B760EFF5AFF603AC2C377" pageId="3" pageNumber="441" type="description">
<paragraph id="8BB429484F2B760EFF5AFF603FF0C4EF" blockId="3.[151,1437,150,1977]" box="[199,419,150,177]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFF603FF0C4EF" bold="true" box="[199,419,150,177]" pageId="3" pageNumber="441">Description. Male.</emphasis>
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFF4D3D0FC543" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
Dimensions: LB:
<quantity id="4CF384AD4F2B760EFE11FF4C3C58C48B" box="[396,523,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.5" metricValueMin="7.1" pageId="3" pageNumber="441" unit="mm" value="7.3" valueMax="7.5" valueMin="7.1">7.1–7.5mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFD85FF4D3C2FC48B" box="[536,636,187,213]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFD1FFF4C3C86C48B" box="[642,725,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="441" unit="mm" value="7.5">7.5mm</quantity>
). LR: 1.0–
<quantity id="4CF384AD4F2B760EFCD5FF4D3DC5C48B" box="[840,918,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFC39FF4D3A5BC48B" box="[932,1032,187,213]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFB90FF4D3A33C48B" box="[1037,1120,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3mm</quantity>
). WR: 1.0–
<quantity id="4CF384AD4F2B760EFB40FF4D3B78C48B" box="[1245,1323,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFAA5FF4D3BCFC48B" box="[1336,1436,187,213]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFF0AFF293EB9C4A7" box="[151,234,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3mm</quantity>
). LP:
<quantity id="4CF384AD4F2B760EFEB7FF283FF9C4A7" box="[298,426,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="3" pageNumber="441" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1–2.5mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFE2AFF293C48C4A7" box="[439,539,223,249]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFDBDFF283C27C4A7" box="[544,628,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5mm</quantity>
). WP:
<quantity id="4CF384AD4F2B760EFD21FF283D6FC4A7" box="[700,828,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="3" pageNumber="441" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFCD4FF293DFEC4A7" box="[841,941,223,249]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFC2EFF283A55C4A7" box="[947,1030,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5mm</quantity>
). LE: 4.5–5.0mm (
<typeStatus id="54B097EA4F2B760EFB49FF293B6BC4A7" box="[1236,1336,223,249]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
5.0mm). WE:
<quantity id="4CF384AD4F2B760EFF4FFEF43F02C543" box="[210,337,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="3" pageNumber="441" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5mm</quantity>
(
<typeStatus id="54B097EA4F2B760EFEFDFEF53F97C543" box="[352,452,259,285]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
<quantity id="4CF384AD4F2B760EFE57FEF43C4DC543" box="[458,542,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="441" unit="mm" value="3.5">3.5mm</quantity>
). N=3 for all measurements.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFED13F1CC5D7" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFED13F02C51F" bold="true" box="[199,337,295,321]" pageId="3" pageNumber="441">Integument</emphasis>
of antenna, apex of femora and tibiae and tarsi black; head, rostrum, pronotum, and elytra, dark crimson while legs are lighter crimson. Body and pronotal surface, rostrum, head, and underside lustrous; legs weakly lustrous.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFE653A88C5AB" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFE653F54C5F3" bold="true" box="[199,263,403,429]" pageId="3" pageNumber="441">Head</emphasis>
dorsal surface punctate with sparse and appressed piliform bluish scales; lateral side near the eye with weak rugae, and below the eye with sparse and appressed, piliform, bluish and turquoise scales; forehead between eyes very weakly depressed with distinct median furrow; eyes medium-sized and weakly convex.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFE093E86C7DF" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFE093F62C647" bold="true" box="[199,305,511,537]" pageId="3" pageNumber="441">Rostrum</emphasis>
almost continuous with head, separated only by a short but deep transverse groove not reaching the dorsolateral margin, weakly punctate on basal half with appressed and sparse, piliform, bluish scales and finely punctate on apical half with fewer setae, as long as wide (LR
<quantity id="4CF384AD4F2B760EFCC8FDB13DFCC63F" box="[853,943,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3 mm</quantity>
; WR
<quantity id="4CF384AD4F2B760EFC6DFDB13A18C63F" box="[1008,1099,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="441" unit="mm" value="1.3">1.3 mm</quantity>
), lateral sides with appressed, piliform, bluish scales interspersed with long, suberect, white setae, especially towards anterior margin; dorsum with distinct median furrow reaching the middle; dorsal contour in lateral view almost flat then subapically gradually declined towards apex, lateral contour in dorsal view anteriorly widened; dorsolateral margin weakly rounded, not angled. Antennal scape and funicle subequal in length. Scape covered with appressed, thin, light-colored setae, funicle with suberect, light-colored setae. Antennae with segments 1 and 2 subequal in length, three times longer than wide; segments 3 nearly a third shorter than segment 2 but slightly longer than segment 4; segments 4 to 6 nearly as long as wide; segment 7 slightly longer than 4 to 6; club subelliptical, nearly three times longer than wide.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFC7D3B6EC797" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFC7D3F12C7FB" bold="true" box="[199,321,907,933]" pageId="3" pageNumber="441">Prothorax</emphasis>
subcylindrical; finely punctate and sparsely covered with appressed setae; as long as wide (LP
<quantity id="4CF384AD4F2B760EFAE4FC7C3E95C797" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5 mm</quantity>
; WP
<quantity id="4CF384AD4F2B760EFE9CFC583F08C797" box="[257,347,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5 mm</quantity>
), dorsal contour in lateral view nearly flat, lateral contour in dorsal view weakly convex.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFC253B65C153" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFC253F40C7B3" bold="true" box="[199,275,979,1005]" pageId="3" pageNumber="441">Elytra</emphasis>
ovate, longer than wide (LE 5.0 mm; WE
<quantity id="4CF384AD4F2B760EFC8EFC243D3CC7B3" box="[787,879,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="441" unit="mm" value="3.5">3.5 mm</quantity>
), slightly wider and twice longer than prothorax (WE
<quantity id="4CF384AD4F2B760EFF49FC003F7EC04F" box="[212,301,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="441" unit="mm" value="3.5">3.5 mm</quantity>
; WP
<quantity id="4CF384AD4F2B760EFEF4FC003F90C04F" box="[361,451,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5 mm</quantity>
; LE 5.0 mm; LP
<quantity id="4CF384AD4F2B760EFD1BFC003CB2C04F" box="[646,737,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5 mm</quantity>
), body surface sparsely covered with minute setae and minute piliform, bluish scales along distal margins; dorsal contour in lateral view uniformly convex; lateral contour in dorsal view uniformly convex, slightly narrow at base then gradually widens towards middle then uniformly tapers towards a subobtuse apex.
<emphasis id="B97FF55A4F2B760EFE56FB953C50C023" bold="true" box="[459,515,1123,1149]" pageId="3" pageNumber="441">Legs</emphasis>
with moderately clavate femorae covered in white and bluish setae and piliform scales. Tibiae covered with subadpressed and suberect white setae, weakly serrate along inner edge. Protibiae and mesotibiae mucronate at apex. Tarsi moderately long and pubscent. Tarsomere 1 elongate, and subtriangular, longer than tarsomere 2; tarsomere 2 short, triangular, and simple without sharp projections at apical corners; tarsomere 3 bilobed, slightly shorter than tarsomere 2; tarsomere 1 and 5 slightly subequal in length; tarsal claws free.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFAE03CDEC1C3" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AFAE03F41C16E" bold="true" box="[199,274,1302,1328]" pageId="3" pageNumber="441">Coxae</emphasis>
covered with colored piliform scales and setae. Mesoventrite covered with setae; metaventrite covered with setae. Ventrite 1 weakly depressed in the middle with some punctures and covered with dense setae. Ventrite 2 with some punctures and covered with dense setae. Ventrite 3–4 sparsely covered with white setae. Ventrite 5 flattened, and rugosely punctured with setae.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AFA513D61C19F" blockId="3.[151,1437,150,1977]" box="[199,818,1447,1473]" pageId="3" pageNumber="441">Male genitalia and sternite 9 as shown in figure 4 A–C.</paragraph>
<paragraph id="8BB429484F2B760EFF5AFA3D3D4BC257" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
Female Dimensions: LB: 9.0–
<quantity id="4CF384AD4F2B760EFDBEFA3D3C20C1BB" box="[547,627,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="441" unit="mm" value="9.1">9.1mm</quantity>
. LR:
<quantity id="4CF384AD4F2B760EFD2BFA3D3D6AC1BB" box="[694,825,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="441" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2mm</quantity>
. WR:
<quantity id="4CF384AD4F2B760EFC18FA3C3A5BC1BB" box="[901,1032,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="3" pageNumber="441" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5mm</quantity>
. LP:
<quantity id="4CF384AD4F2B760EFBDAFA3C3AC9C1BB" box="[1095,1178,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5mm</quantity>
. WP:
<quantity id="4CF384AD4F2B760EFB7FFA3C3B36C1BB" box="[1250,1381,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="3" pageNumber="441" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6mm</quantity>
. LE:
<quantity id="4CF384AD4F2B760EFF0AFA183F48C257" box="[151,283,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.5" metricValueMin="6.3" pageId="3" pageNumber="441" unit="mm" value="6.4" valueMax="6.5" valueMin="6.3">6.3–6.5mm</quantity>
. WE:
<quantity id="4CF384AD4F2B760EFEC2FA183FB2C257" box="[351,481,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="3" pageNumber="441" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5mm</quantity>
. N=6 for all measurements.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AF9E53C01C273" blockId="3.[151,1437,150,1977]" box="[199,594,1555,1581]" pageId="3" pageNumber="441">
Habitus as shown in
<figureCitation id="133035CD4F2B760EFE2CF9E53C1EC273" box="[433,589,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-22@2.[151,1436,181,1755]" captionTargetPageId="2" captionText="FIGURE 1. Metapocyrtus inangsabong A—♂ holotype, dorsal view, C­—idem, lateral view; B—♀ paratype, dorsal view, D—idem, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14241204" httpUri="https://zenodo.org/record/14241204/files/figure.png" pageId="3" pageNumber="441">Figure 1 B, D</figureCitation>
.
</paragraph>
<paragraph id="8BB429484F2B760EFF5AF9C13AC2C377" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
Female (
<figureCitation id="133035CD4F2B760EFEB4F9C13FF9C20F" box="[297,426,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-22@2.[151,1436,181,1755]" captionTargetPageId="2" captionText="FIGURE 1. Metapocyrtus inangsabong A—♂ holotype, dorsal view, C­—idem, lateral view; B—♀ paratype, dorsal view, D—idem, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14241204" httpUri="https://zenodo.org/record/14241204/files/figure.png" pageId="3" pageNumber="441">Fig. 1 B, D</figureCitation>
) differ from males in the following characters: a) rostrum weakly depressed at middle, b) integuments of head and rostrum black, c) pronotum more cylindrical with nearly straight dorsal contour in lateral view, d) elytra longer and wider (LE/WE
<quantity id="4CF384AD4F2B760EFD15F9883C84C2C7" box="[648,727,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="441" unit="mm" value="6.5">6.5mm</quantity>
/
<quantity id="4CF384AD4F2B760EFD7DF9883D7CC2C7" box="[736,815,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="441" unit="mm" value="4.5">4.5mm</quantity>
, WE/WP
<quantity id="4CF384AD4F2B760EFC3BF9883DA5C2C7" box="[934,1014,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="441" unit="mm" value="4.5">4.5mm</quantity>
/
<quantity id="4CF384AD4F2B760EFC63F9883A1DC2C7" box="[1022,1102,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="441" unit="mm" value="2.6">2.6mm</quantity>
, LE/LP
<quantity id="4CF384AD4F2B760EFB2FF9883B52C2C7" box="[1202,1281,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="441" unit="mm" value="6.5">6.5mm</quantity>
/
<quantity id="4CF384AD4F2B760EFA97F9883B0AC2C7" box="[1290,1369,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="441" unit="mm" value="2.5">2.5mm</quantity>
) than male, e) lateral contour of elytra in dorsal view subapically interrupted, f) apical declivity gradual but interrupted subapically then abruptly declines towards apex; g) apex of elytra with two minute but acute spines; h) lateral margin of elytra, coxae and mesoventrite and metaventrite with white elliptical scales, i) with Ventrite 1 slightly convex in the middle; j) Ventrite 5 deeply concave. Otherwise, females similar to the male.
</paragraph>
</subSubSection>
<subSubSection id="C3117AC34F2B760EFF5AF8C53FC2C3CB" pageId="3" pageNumber="441" type="etymology">
<paragraph id="8BB429484F2B760EFF5AF8C53FC2C3CB" blockId="3.[151,1437,150,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AF8C53F1DC313" bold="true" box="[199,334,1843,1869]" pageId="3" pageNumber="441">Etymology.</emphasis>
The species name
<taxonomicName id="4C0B52CB4F2B760EFD83F8C53CE8C313" authorityName="Cabras, Obrial &amp; Agbas" authorityYear="2024" box="[542,699,1843,1869]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="441" phylum="Arthropoda" rank="species" species="inangsabong">
“
<emphasis id="B97FF55A4F2B760EFDB7F8C53CE8C313" box="[554,699,1843,1869]" italics="true" pageId="3" pageNumber="441">inangsabong</emphasis>
</taxonomicName>
” refers to the legendary first Mansaka man to settle in
<collectingRegion id="49CFE7AA4F2B760EFA8BF8C53EB2C32F" country="Philippines" name="Compostela" pageId="3" pageNumber="441">Compostela Valley</collectingRegion>
. According to stories, Inangsabong had seven wives, who later established various Mansaka communities throughout the region.
</paragraph>
</subSubSection>
<subSubSection id="C3117AC34F2B760EFF5AF8683A32C3E7" box="[199,1121,1950,1977]" pageId="3" pageNumber="441" type="distribution">
<paragraph id="8BB429484F2B760EFF5AF8683A32C3E7" blockId="3.[151,1437,150,1977]" box="[199,1121,1950,1977]" pageId="3" pageNumber="441">
<emphasis id="B97FF55A4F2B760EFF5AF8683F32C3E6" bold="true" box="[199,353,1950,1976]" pageId="3" pageNumber="441">Distribution.</emphasis>
<collectingCountry id="F31C69D84F2B760EFEFAF8693FB6C3E7" box="[359,485,1951,1977]" name="Philippines" pageId="3" pageNumber="441">Philippines</collectingCountry>
(Mindanao:
<collectingRegion id="49CFE7AA4F2B760EFDEBF8693CECC3E7" box="[630,703,1951,1977]" country="Philippines" name="Davao" pageId="3" pageNumber="441">Davao</collectingRegion>
de Oro, Maragusan, Mt. Candalaga).
</paragraph>
</subSubSection>
</treatment>
</document>
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