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<document id="363ABDDB6D75912F19620BDD4E59C852" ID-CLB-Dataset="45364" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent &amp; Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E5CFFA5FF7FFAD2FCE1FE64" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athleta (Volutocorbis) wynnei Cox 1930" docType="treatment" docVersion="9" lastPageNumber="124" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="122" updateTime="1698333283847" updateUser="ExternalLinkService">
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<mods:title id="C4E5AD3E5DF14A6DCCA5BC70BA6EA842">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="88D1EDB7ABA64BC268B01A391669824F">Bartolini, Annachiara</mods:namePart>
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<treatment id="03A087E00E5CFFA5FF7FFAD2FCE1FE64" ID-DOI="http://doi.org/10.5281/zenodo.6137850" ID-GBIF-Taxon="119509262" ID-Zenodo-Dep="6137850" LSID="urn:lsid:plazi:treatment:03A087E00E5CFFA5FF7FFAD2FCE1FE64" httpUri="http://treatment.plazi.org/id/03A087E00E5CFFA5FF7FFAD2FCE1FE64" lastPageId="23" lastPageNumber="124" pageId="21" pageNumber="122">
<subSubSection id="C313657D0E5CFFA7FF7FFAD2FEC4FADC" pageId="21" pageNumber="122" type="nomenclature">
<paragraph id="8BB636F60E5CFFA7FF7FFAD2FD7DFAFD" blockId="21.[151,679,1340,1399]" box="[151,679,1340,1366]" pageId="21" pageNumber="122">
<heading id="D0FE819A0E5CFFA7FF7FFAD2FD7DFAFD" bold="true" box="[151,679,1340,1366]" fontSize="11" level="1" pageId="21" pageNumber="122" reason="1">
<taxonomicName id="4C094D750E5CFFA7FF7FFAD2FD7DFAFD" authority="Cox, 1930" authorityName="Cox" authorityYear="1930" box="[151,679,1340,1366]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="122" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5CFFA7FF7FFAD2FD7DFAFD" bold="true" box="[151,679,1340,1366]" pageId="21" pageNumber="122">
<emphasis id="B97DEAE40E5CFFA7FF7FFAD2FE7AFAFD" bold="true" box="[151,416,1340,1366]" italics="true" pageId="21" pageNumber="122">Athleta (Volutocorbis)</emphasis>
cf.
<emphasis id="B97DEAE40E5CFFA7FE25FAD2FDF9FAFD" bold="true" box="[461,547,1340,1366]" italics="true" pageId="21" pageNumber="122">wynnei</emphasis>
<bibRefCitation id="EF984B070E5CFFA7FDC3FAD2FD7DFAFD" author="Cox" box="[555,679,1340,1366]" pageId="21" pageNumber="122" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox, 1930</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BB636F60E5CFFA7FF7FFAB0FEC4FADC" blockId="21.[151,679,1340,1399]" box="[151,286,1374,1399]" pageId="21" pageNumber="122">
(
<figureCitation id="13322A730E5CFFA7FF77FAB0FF3BFADC" box="[159,225,1374,1399]" captionStart="FIGURE 9. A G" captionStartId="22.[151,250,1861,1883]" captionTargetBox="[242,1360,193,1849]" captionTargetId="figure@22.[242,1369,193,1851]" captionTargetPageId="22" captionText="FIGURE 9. A G. Athleta (Volutocorbis) burtoni Vredenburg, 1923, A (CPAG. RAN. I. 31, dorsal view, stn 2), B (CPAG. RAN. I. 32, ventral view, stn 3), C (CPAG. RAN. I. 33, dorsal view, stn 1), D F (MNHN. F. J 13433, Cossmann coll., ventral (D) and dorsal (E) views, enlarged view of the spire (F), Jhirak), G (MNHN. F. J 13434, Cossmann coll., dorsal view, Jhirak); H L. Athleta (Volutocorbis) cf. wynnei Cox, 1930, H I (CPAG. RAN. I. 35, dorsal view (H) and enlarged view of the spire (I), stn 4), J (CPAG. RAN. I. 36, ventral view, stn 4), K L (CPAG. RAN. I. 34, ventral (K) and dorsal (L) views). Scale bars: A E, G H, J L = 10 mm; F, I = 5 mm." httpUri="https://zenodo.org/record/228546/files/figure.png" pageId="21" pageNumber="122">Fig. 9</figureCitation>
HL)
</paragraph>
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<subSubSection id="C313657D0E5CFFA7FF7FFA48FCEAFA17" box="[151,816,1446,1469]" pageId="21" pageNumber="122" type="reference_group">
<paragraph id="8BB636F60E5CFFA7FF7FFA48FCEAFA17" blockId="21.[151,816,1446,1469]" box="[151,816,1446,1469]" pageId="21" pageNumber="122">
<treatmentCitationGroup id="AB1911D80E5CFFA7FF7FFA48FCEAFA17" box="[151,816,1446,1469]" pageId="21" pageNumber="122">
<taxonomicName id="4C094D750E5CFFA7FF7FFA48FDB8FA17" authority="Cox, 1930: 199" authorityName="Cox" authorityPageNumber="199" authorityYear="1930" box="[151,610,1446,1469]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="122" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutospina">
<emphasis id="B97DEAE40E5CFFA7FF7FFA48FE67FA16" box="[151,445,1446,1469]" italics="true" pageId="21" pageNumber="122">Athleta (Volutospina) wynnei</emphasis>
<treatmentCitation id="0AA810E70E5CFFA7FE2BFA48FDB8FA17" author="Cox" box="[451,610,1446,1468]" page="199" pageId="21" pageNumber="122" year="1930">
<bibRefCitation id="EF984B070E5CFFA7FE2BFA48FDEAFA17" author="Cox" box="[451,560,1446,1468]" pageId="21" pageNumber="122" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox, 1930</bibRefCitation>
: 199
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, pl. 31, figs 78, 11.
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</paragraph>
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<subSubSection id="C313657D0E5CFFA7FF7FFA07FF2EF825" pageId="21" pageNumber="122" type="description">
<paragraph id="8BB636F60E5CFFA7FF7FFA07FF2EF825" blockId="21.[151,1436,1513,1970]" pageId="21" pageNumber="122">
<emphasis id="B97DEAE40E5CFFA7FF7FFA07FEF3F9A9" bold="true" box="[151,297,1513,1538]" pageId="21" pageNumber="122">Description.</emphasis>
Shell biconic, ventricose, H ca.
<quantity id="4CF19B130E5CFFA7FD72FA04FD30F9AA" box="[666,746,1514,1538]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="122" unit="mm" value="40.0">40 mm</quantity>
(spire broken in adult specimen), D
<quantity id="4CF19B130E5CFFA7FB65FA07FB3AF9AA" box="[1165,1248,1513,1538]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="122" unit="mm" value="25.0">25 mm</quantity>
. Protoconch and first teleoconch whorl unknown; at least 5.5 teleoconch whorls. Spire apparently moderately high. Spire whorls convex, last whorl rather inflated. Spiral sculpture on spire of four or five rows of small nodules, subsutural row better developed than others (
<figureCitation id="13322A730E5CFFA7FE18F9BBFDECF9C5" box="[496,566,1621,1646]" captionStart="FIGURE 9. A G" captionStartId="22.[151,250,1861,1883]" captionTargetBox="[242,1360,193,1849]" captionTargetId="figure@22.[242,1369,193,1851]" captionTargetPageId="22" captionText="FIGURE 9. A G. Athleta (Volutocorbis) burtoni Vredenburg, 1923, A (CPAG. RAN. I. 31, dorsal view, stn 2), B (CPAG. RAN. I. 32, ventral view, stn 3), C (CPAG. RAN. I. 33, dorsal view, stn 1), D F (MNHN. F. J 13433, Cossmann coll., ventral (D) and dorsal (E) views, enlarged view of the spire (F), Jhirak), G (MNHN. F. J 13434, Cossmann coll., dorsal view, Jhirak); H L. Athleta (Volutocorbis) cf. wynnei Cox, 1930, H I (CPAG. RAN. I. 35, dorsal view (H) and enlarged view of the spire (I), stn 4), J (CPAG. RAN. I. 36, ventral view, stn 4), K L (CPAG. RAN. I. 34, ventral (K) and dorsal (L) views). Scale bars: A E, G H, J L = 10 mm; F, I = 5 mm." httpUri="https://zenodo.org/record/228546/files/figure.png" pageId="21" pageNumber="122">Fig. 9</figureCitation>
J). On last whorl, spiral sculpture displaying: 1) subsutural row of rounded nodules; 2) shoulder row of rounded nodules; 3) from shoulder to base, around 20 almost flat cords separated by thin spiral grooves. From second to penultimate whorl, costae interrupted by rows of nodules. On last whorl, costae obsolete between subsutural and shoulder nodules, thin but relatively low below shoulder angle, extending across base, becoming almost obsolete towards close of last whorl. Second whorl: 15 costae; third whorl: 15 costae; fourth whorl: 1617 costae; fifth and last whorl: 19 costae. Aperture ovate, rather wide. Inner lip straight, columella with two folds succeeded posteriorly by two or three smaller, weaker folds; inner lip callus narrow, apparently not spreading anteriorly. Outer lip thin, not thickened externally. Siphonal canal slightly elongate; siphonal notch shallow.
</paragraph>
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<paragraph id="8BB636F60E5CFFA7FF2FF877FAFBF819" blockId="21.[151,1436,1513,1970]" box="[199,1313,1945,1970]" pageId="21" pageNumber="122">
<emphasis id="B97DEAE40E5CFFA7FF2FF877FEECF819" bold="true" box="[199,310,1945,1970]" pageId="21" pageNumber="122">Material.</emphasis>
4 spm (stn 4: 3 spm
<collectionCode id="ED18AE330E5CFFA7FDC8F877FDB5F819" box="[544,623,1945,1970]" pageId="21" pageNumber="122">CPAG</collectionCode>
.RAN.
<date id="FFB710360E5CFFA7FD5CF877FD06F819" box="[692,732,1945,1970]" pageId="21" pageNumber="122" value="1934-01">I.34</date>
36, cast
<collectionCode id="ED18AE330E5CFFA7FCADF877FC47F819" LSID="urn:lsid:biocol.org:col:34988" box="[837,925,1945,1970]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="122">MNHN</collectionCode>
.F.A5037274 and 1 spm
<collectionCode id="ED18AE330E5CFFA7FB53F874FACCF819" LSID="urn:lsid:biocol.org:col:34988" box="[1211,1302,1946,1970]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="122">MNHN</collectionCode>
).
</paragraph>
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<caption id="DF76667E0E5FFFA4FF7FF8ABFE3DF85F" httpUri="https://zenodo.org/record/228546/files/figure.png" pageId="22" pageNumber="123" targetBox="[242,1360,193,1849]" targetPageId="22">
<paragraph id="8BB636F60E5FFFA4FF7FF8ABFE3DF85F" blockId="22.[151,1436,1861,2036]" pageId="22" pageNumber="123">
<emphasis id="B97DEAE40E5FFFA4FF7FF8ABFEBFF8F0" bold="true" box="[151,357,1861,1883]" pageId="22" pageNumber="123">FIGURE 9. AG.</emphasis>
<taxonomicName id="4C094D750E5FFFA4FE9FF8ABFC4FF8F0" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[375,917,1861,1883]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="burtoni" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5FFFA4FE9FF8ABFE1AF8F0" box="[375,448,1861,1883]" italics="true" pageId="22" pageNumber="123">Athleta</emphasis>
<emphasis id="B97DEAE40E5FFFA4FE3AF8ABFDB8F8F0" box="[466,610,1861,1883]" italics="true" pageId="22" pageNumber="123">(Volutocorbis)</emphasis>
<emphasis id="B97DEAE40E5FFFA4FD9DF8ABFD1AF8F0" box="[629,704,1861,1883]" italics="true" pageId="22" pageNumber="123">burtoni</emphasis>
Vredenburg, 1923
</taxonomicName>
, A (CPAG.RAN.I.31, dorsal view, stn 2), B (CPAG.RAN.I.32, ventral view, stn 3), C (CPAG.RAN.I.33, dorsal view, stn 1), DF (MNHN.F.
<accessionNumber id="945AAB150E5FFFA4FBB2F88AFB79F8D1" box="[1114,1187,1892,1914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J13433" pageId="22" pageNumber="123" type="EnaNcbi">J13433</accessionNumber>
, Cossmann coll., ventral (D) and dorsal (E) views, enlarged view of the spire (F), Jhirak), G (MNHN.F.
<accessionNumber id="945AAB150E5FFFA4FC2FF86CFBC8F833" box="[967,1042,1922,1944]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J13434" pageId="22" pageNumber="123" type="EnaNcbi">J13434</accessionNumber>
, Cossmann coll., dorsal view, Jhirak);
<emphasis id="B97DEAE40E5FFFA4FF7FF84FFF15F81C" bold="true" box="[151,207,1953,1975]" pageId="22" pageNumber="123">HL.</emphasis>
<taxonomicName id="4C094D750E5FFFA4FF3EF84FFD46F81C" authority="Cox, 1930" authorityName="Cox" authorityYear="1930" box="[214,668,1953,1975]" class="Gastropoda" family="Volutidae" genus="Athleta" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="123" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5FFFA4FF3EF84FFE6DF81C" box="[214,439,1953,1975]" italics="true" pageId="22" pageNumber="123">Athleta (Volutocorbis)</emphasis>
cf.
<emphasis id="B97DEAE40E5FFFA4FE08F84CFDF2F81C" box="[480,552,1954,1975]" italics="true" pageId="22" pageNumber="123">wynnei</emphasis>
Cox, 1930
</taxonomicName>
, HI (CPAG.RAN.I.35, dorsal view (H) and enlarged view of the spire (I), stn 4), J (CPAG.RAN.I.36, ventral view, stn 4), KL (CPAG.RAN.I.34, ventral (K) and dorsal (L) views). Scale bars: AE, GH, JL = 10 mm; F, I = 5 mm.
</paragraph>
</caption>
<subSubSection id="C313657D0E5EFFA5FF2FFF79FCE1FE64" pageId="23" pageNumber="124" type="discussion">
<paragraph id="8BB636F60E5EFFA5FF2FFF79FAC5FE06" blockId="23.[151,1436,151,464]" pageId="23" pageNumber="124">
<emphasis id="B97DEAE40E5EFFA5FF2FFF79FE95FF1B" bold="true" box="[199,335,151,176]" pageId="23" pageNumber="124">Comments.</emphasis>
The material collected from the Lakhra Dome includes three juvenile specimens and a broken adult. The juvenile specimens can be compared with a
<typeStatus id="54B288540E5EFFA5FCEFFF53FCBDFF7E" box="[775,871,189,213]" pageId="23" pageNumber="124" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C094D750E5EFFA5FC67FF52FA87FF7F" authority="Cox, 1930" authorityName="Cox" authorityYear="1930" box="[911,1373,188,213]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5EFFA5FC67FF52FB05FF7F" box="[911,1247,188,213]" italics="true" pageId="23" pageNumber="124">Athleta (Volutocorbis) wynnei</emphasis>
<bibRefCitation id="EF984B070E5EFFA5FB00FF52FA87FF7F" author="Cox" box="[1256,1373,188,212]" pageId="23" pageNumber="124" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox, 1930</bibRefCitation>
</taxonomicName>
(GSI no. 14,700) from the Hangu Formation, which is also a juvenile. With these young specimens of
<emphasis id="B97DEAE40E5EFFA5FB38FF0EFAA9FF53" box="[1232,1395,224,248]" italics="true" pageId="23" pageNumber="124">
<taxonomicName id="4C094D750E5EFFA5FB38FF0EFAB4FF53" box="[1232,1390,224,248]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">A. (V.) wynnei</taxonomicName>
,
</emphasis>
the Sindh specimens share close similarities, particularly the elongate shape of the spire, the sculpture on the spire (with four to five rows of small nodules) and the deep crenulations of costae on the last whorl. The adult specimens of
<taxonomicName id="4C094D750E5EFFA5FF5EFEA2FE8EFECF" box="[182,340,332,356]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5EFFA5FF5EFEA2FE8EFECF" box="[182,340,332,356]" italics="true" pageId="23" pageNumber="124">A. (V.) wynnei</emphasis>
</taxonomicName>
, as for example the
<typeStatus id="54B288540E5EFFA5FDD5FEA2FD45FECE" box="[573,671,332,357]" pageId="23" pageNumber="124" type="holotype">holotype</typeStatus>
(GSI no. 14,698), apparently differ by their more crenulated costae on the last whorl, and by their less ventricose shape. Because of these small differences, the present identification of
<taxonomicName id="4C094D750E5EFFA5FF5DFE7AFE8BFE07" box="[181,337,404,428]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="124" phylum="Mollusca" rank="species" species="wynnei" subGenus="Volutocorbis">
<emphasis id="B97DEAE40E5EFFA5FF5DFE7AFE8BFE07" box="[181,337,404,428]" italics="true" pageId="23" pageNumber="124">A. (V.) wynnei</emphasis>
</taxonomicName>
from the Lakhra Formation should be confirmed by the examination of more material.
</paragraph>
<paragraph id="8BB636F60E5EFFA5FF2FFE59FCE1FE64" blockId="23.[151,1436,151,464]" box="[199,827,439,464]" pageId="23" pageNumber="124">
<emphasis id="B97DEAE40E5EFFA5FF2FFE59FE62FE7B" bold="true" box="[199,440,439,464]" pageId="23" pageNumber="124">Stratigraphic range.</emphasis>
Lakhra Formation: Lakhra Dome.
</paragraph>
</subSubSection>
</treatment>
</document>