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<document id="A08CCEBBB06FABAE79AF138A69D33E1E" ID-DOI="10.11646/zootaxa.5481.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="12783020" ID-ZooBank="C9AF57F3-482F-4BCF-ABA7-C1A08E65E41D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1721383080839" checkinUser="plazi" docAuthor="Trewick, Steven A." docDate="2024" docId="03A087CE214D0B34FF18AD2586A0E24A" docLanguage="en" docName="zootaxa.5481.5.3.pdf" docOrigin="Zootaxa 5481 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crux heggi Trewick 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="542" masterDocId="FF99FFB621450B3FFF8FAB39857EE357" masterDocTitle="Two new genera of tokoriro (Orthoptera: Rhaphidophoridae: Macropathinae) from Aotearoa New Zealand" masterLastPageNumber="546" masterPageNumber="531" pageNumber="539" updateTime="1721418553730" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="C1FA11E2447976070AC212E8746DC63D">Two new genera of tokoriro (Orthoptera: Rhaphidophoridae: Macropathinae) from Aotearoa New Zealand</mods:title>
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<mods:namePart id="8C6E8CA495554108CE9CFC185E60FAA1">Trewick, Steven A.</mods:namePart>
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<mods:date id="08BCB3E5B5F18F2F2953F12824509B3F">2024</mods:date>
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<treatment id="03A087CE214D0B34FF18AD2586A0E24A" LSID="urn:lsid:plazi:treatment:03A087CE214D0B34FF18AD2586A0E24A" httpUri="http://treatment.plazi.org/id/03A087CE214D0B34FF18AD2586A0E24A" lastPageId="11" lastPageNumber="542" pageId="8" pageNumber="539">
<subSubSection id="C3136553214D0B37FF18AD25848FE561" box="[151,497,1564,1591]" pageId="8" pageNumber="539" type="nomenclature">
<paragraph id="8BB636D8214D0B37FF18AD25848FE561" blockId="8.[151,497,1564,1626]" box="[151,497,1564,1591]" pageId="8" pageNumber="539">
<heading id="D0FE81B4214D0B37FF18AD25848FE561" bold="true" box="[151,497,1564,1591]" fontSize="11" level="1" pageId="8" pageNumber="539" reason="1">
<emphasis id="B97DEACA214D0B37FF18AD25848FE561" bold="true" box="[151,497,1564,1591]" pageId="8" pageNumber="539">
<taxonomicName id="4C094D5B214D0B37FF18AD2584F3E561" authority="Trewick" authorityName="Trewick" authorityYear="2024" box="[151,397,1564,1591]" class="Holothuroidea" family="Rhaphidophoridae" genus="Crux" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="539" phylum="Echinodermata" rank="species" species="heggi" status="sp. nov.">
<emphasis id="B97DEACA214D0B37FF18AD258461E560" bold="true" box="[151,287,1564,1591]" italics="true" pageId="8" pageNumber="539">Crux heggi</emphasis>
Trewick
</taxonomicName>
<taxonomicNameLabel id="A24E57B1214D0B37FE1BAD25848FE561" box="[404,497,1564,1590]" pageId="8" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3136553214D0B37FF18AD798435E50D" box="[151,331,1600,1626]" pageId="8" pageNumber="539" type="description">
<paragraph id="8BB636D8214D0B37FF18AD798435E50D" blockId="8.[151,497,1564,1626]" box="[151,331,1600,1626]" pageId="8" pageNumber="539">
(
<figureCitation id="13322A5D214D0B37FF10AD798475E50D" box="[159,267,1600,1626]" captionStart="FIGURE 3" captionStartId="3.[151,250,1522,1546]" captionTargetBox="[151,1436,181,1489]" captionTargetId="figure-15@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURE 3. New species of Macropathinae from New Zealand. a) Crux boudica sp. nov., male, Rakiura (D. Hegg). b) Crux boudica sp. nov., female, Skippers Range (D. Hegg). c) Crux heggi sp. nov., male, Paparoa Range (D. Hegg). d) Crux heggi sp. nov., female, Saxon River (D. Hegg). e) Occultastella morgana sp. nov., male, Buller River (T. Jewell). f) Occultastella morgana sp. nov., female, Cobb Valley (A. McDonald)." figureDoi="http://doi.org/10.5281/zenodo.12783028" httpUri="https://zenodo.org/record/12783028/files/figure.png" pageId="8" pageNumber="539">Figures 3</figureCitation>
,
<figureCitation id="13322A5D214D0B37FE97AD79843DE50D" box="[280,323,1600,1626]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="9.[151,250,928,952]" captionStartId-1="9.[151,250,1891,1915]" captionStartId-2="10.[151,250,845,869]" captionTargetBox-0="[151,1436,179,902]" captionTargetBox-1="[218,1369,1165,1829]" captionTargetBox-2="[151,1436,181,822]" captionTargetId-0="figure-15@9.[151,1436,179,902]" captionTargetId-1="figure-47@9.[151,1436,1082,1851]" captionTargetId-2="figure-21@10.[151,1436,181,822]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 7. Crux heggi sp. nov. Female holotype. a) Lateral view body. bd) Detailed views of ovipositor showing position and shape of subgenital plate. Pink triangle indicates subgenital plate." captionText-1="FIGURE 8. Crux heggi sp. nov. Female holotype. Prolateral and retrolateral views of (a) fore and mid (b) legs with inset of mid femur showing apical retrolateral spine. c) Hind femur prolateral and retrolateral views. d) Hind tibia posterior view and detail of apical spine on hind femur (blue arrow). e) Hind tarsi, retrolateral view. Pink arrows indicate linear spines." captionText-2="FIGURE 9. Crux heggi sp. nov. Male Paratype. Terminalia. ac) Lateral, dorsal and ventral views of posterior end of abdomen. d) Oblique posterior view of terminalia showing subgenital plate, and styli (blue arrows) pressed against paraprocts. e) Posterior view. f) Disarticulated fused paraproct structure (MPN CW4372)." figureDoi-0="http://doi.org/10.5281/zenodo.12783038" figureDoi-1="http://doi.org/10.5281/zenodo.12783040" figureDoi-2="http://doi.org/10.5281/zenodo.12783042" httpUri-0="https://zenodo.org/record/12783038/files/figure.png" httpUri-1="https://zenodo.org/record/12783040/files/figure.png" httpUri-2="https://zenodo.org/record/12783042/files/figure.png" pageId="8" pageNumber="539">79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3136553214D0B37FF18ADB1849CE5BD" pageId="8" pageNumber="539" type="materials_examined">
<paragraph id="8BB636D8214D0B37FF18ADB1849CE5BD" blockId="8.[151,1436,1672,1951]" pageId="8" pageNumber="539">
<emphasis id="B97DEACA214D0B37FF18ADB18478E5F5" bold="true" box="[151,262,1672,1698]" pageId="8" pageNumber="539">Material.</emphasis>
<materialsCitation id="3B613C85214D0B37FE9CADB1849CE590" collectingDate="2016-04-22" collectionCode="NMNZ" collectorName="Kahurangi NP &amp; D. Hegg" country="New Zealand" location="New Zealand" pageId="8" pageNumber="539" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54B2887A214D0B37FE9CADB18405E5F5" box="[275,379,1672,1698]" pageId="8" pageNumber="539" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0FFD51214D0B37FE08ADB084E3E5F4" box="[391,413,1673,1699]" count="1" pageId="8" pageNumber="539" type="female"></specimenCount>
. Saxon hut, Heaphy Track,
<collectorName id="26FC530E214D0B37FD7FADB186E8E5F4" box="[752,918,1672,1699]" pageId="8" pageNumber="539">Kahurangi NP</collectorName>
,
<collectingCountry id="F31E7648214D0B37FC29ADB0813AE5F5" box="[934,1092,1672,1699]" name="New Zealand" pageId="8" pageNumber="539">New Zealand</collectingCountry>
,
<collectorName id="26FC530E214D0B37FBDAADB081C2E5F4" box="[1109,1212,1673,1699]" pageId="8" pageNumber="539">D. Hegg</collectorName>
,
<date id="FFB71018214D0B37FB42ADB18031E5F5" box="[1229,1359,1672,1698]" pageId="8" pageNumber="539" value="2016-04-22">
<collectingDate id="EFF3E9F0214D0B37FB42ADB18031E5F5" box="[1229,1359,1672,1698]" pageId="8" pageNumber="539" value="2016-04-22">22/04/2016</collectingDate>
</date>
, MPN CW2983,
<collectionCode id="ED18AE1D214D0B37FE83AD948420E590" box="[268,350,1709,1735]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="8" pageNumber="539" type="Museum">NMNZ</collectionCode>
AI.074305
</materialsCitation>
.
<materialsCitation id="3B613C85214D0B37FE60AD9484A1E5BD" collectingDate="2019-04-06" collectionCode="NMNZ" collectorName="D. Hegg" country="New Zealand" location="New Zealand" pageId="8" pageNumber="539" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54B2887A214D0B37FE60AD94872FE590" box="[495,593,1709,1735]" pageId="8" pageNumber="539" type="paratype">Paratype</typeStatus>
<specimenCount id="9D0FFD51214D0B37FDD5AD94870CE590" box="[602,626,1709,1735]" count="1" pageId="8" pageNumber="539" type="male"></specimenCount>
. Ces Clarke Hut, Paparoa Range,
<collectingCountry id="F31E7648214D0B37FC73AD9481E9E591" box="[1020,1175,1708,1735]" name="New Zealand" pageId="8" pageNumber="539">New Zealand</collectingCountry>
,
<collectorName id="26FC530E214D0B37FB2BAD948076E590" box="[1188,1288,1709,1735]" pageId="8" pageNumber="539">D. Hegg</collectorName>
,
<date id="FFB71018214D0B37FA99AD9580E6E591" box="[1302,1432,1708,1734]" pageId="8" pageNumber="539" value="2019-04-06">
<collectingDate id="EFF3E9F0214D0B37FA99AD9580E6E591" box="[1302,1432,1708,1734]" pageId="8" pageNumber="539" value="2019-04-06">06/04/2019</collectingDate>
</date>
, CW4443,
<collectionCode id="ED18AE1D214D0B37FE85ADE88422E5BC" box="[266,348,1745,1771]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="8" pageNumber="539" type="Museum">NMNZ</collectionCode>
AI.074306
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3136553214D0B35FF48ADCD81EAE4B1" lastPageId="10" lastPageNumber="541" pageId="8" pageNumber="539" type="description">
<paragraph id="8BB636D8214D0B37FF48ADCD8427E458" blockId="8.[151,1436,1672,1951]" box="[199,345,1780,1807]" pageId="8" pageNumber="539">
<emphasis id="B97DEACA214D0B37FF48ADCD842FE459" bold="true" box="[199,337,1780,1806]" pageId="8" pageNumber="539">Description</emphasis>
.
</paragraph>
<paragraph id="8BB636D8214D0B37FF48AC208102E4C9" blockId="8.[151,1436,1672,1951]" pageId="8" pageNumber="539">
<emphasis id="B97DEACA214D0B37FF48AC20842AE465" box="[199,340,1817,1842]" italics="true" pageId="8" pageNumber="539">Dimensions:</emphasis>
Adult female total length (from anterior of head to posterior tip of ovipositor) about
<quantity id="4CF19B3D214D0B37FAAEAC2180E6E464" box="[1313,1432,1816,1843]" metricMagnitude="-2" metricUnit="m" metricValue="2.854" pageId="8" pageNumber="539" unit="mm" value="28.54">28.54 mm</quantity>
; antenna 33.5 body length; length (anterior of head to posterior of abdomen)
<quantity id="4CF19B3D214D0B37FC7CAC058119E400" box="[1011,1127,1852,1879]" metricMagnitude="-2" metricUnit="m" metricValue="1.359" pageId="8" pageNumber="539" unit="mm" value="13.59">13.59 mm</quantity>
; pronotum length
<quantity id="4CF19B3D214D0B37FABEAC0580E6E400" box="[1329,1432,1852,1879]" metricMagnitude="-3" metricUnit="m" metricValue="4.59" pageId="8" pageNumber="539" unit="mm" value="4.59">4.59 mm</quantity>
; ovipositor
<quantity id="4CF19B3D214D0B37FE80AC59840BE42C" box="[271,373,1888,1915]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="539" unit="mm" value="14.5">14.5 mm</quantity>
; fore femur length
<quantity id="4CF19B3D214D0B37FDC9AC5987D2E42C" box="[582,684,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="8.14" pageId="8" pageNumber="539" unit="mm" value="8.14">8.14 mm</quantity>
; mid femur length
<quantity id="4CF19B3D214D0B37FCF2AC59869DE42C" box="[893,995,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="7.24" pageId="8" pageNumber="539" unit="mm" value="7.24">7.24 mm</quantity>
; hind femur length
<quantity id="4CF19B3D214D0B37FB36AC598054E42C" box="[1209,1322,1888,1915]" metricMagnitude="-2" metricUnit="m" metricValue="1.556" pageId="8" pageNumber="539" unit="mm" value="15.56">15.56 mm</quantity>
and depth
<quantity id="4CF19B3D214D0B37FF18ACBD8580E4C8" box="[151,254,1924,1951]" metricMagnitude="-3" metricUnit="m" metricValue="3.82" pageId="8" pageNumber="539" unit="mm" value="3.82">3.82 mm</quantity>
; fore tibia
<quantity id="4CF19B3D214D0B37FEF9ACBD84B1E4C8" box="[374,463,1924,1951]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="539" unit="mm" value="8.4">8.4 mm</quantity>
; mid tibia
<quantity id="4CF19B3D214D0B37FDC9ACBD87D9E4C8" box="[582,679,1924,1951]" metricMagnitude="-3" metricUnit="m" metricValue="7.56" pageId="8" pageNumber="539" unit="mm" value="7.56">7.56mm</quantity>
; hind tibia length
<quantity id="4CF19B3D214D0B37FCFFACBD869CE4C8" box="[880,994,1924,1951]" metricMagnitude="-2" metricUnit="m" metricValue="1.571" pageId="8" pageNumber="539" unit="mm" value="15.71">15.71 mm</quantity>
(
<figureCitation id="13322A5D214D0B37FC7FACBD812FE4C9" box="[1008,1105,1924,1950]" captionStart="FIGURE 3" captionStartId="3.[151,250,1522,1546]" captionTargetBox="[151,1436,181,1489]" captionTargetId="figure-15@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURE 3. New species of Macropathinae from New Zealand. a) Crux boudica sp. nov., male, Rakiura (D. Hegg). b) Crux boudica sp. nov., female, Skippers Range (D. Hegg). c) Crux heggi sp. nov., male, Paparoa Range (D. Hegg). d) Crux heggi sp. nov., female, Saxon River (D. Hegg). e) Occultastella morgana sp. nov., male, Buller River (T. Jewell). f) Occultastella morgana sp. nov., female, Cobb Valley (A. McDonald)." figureDoi="http://doi.org/10.5281/zenodo.12783028" httpUri="https://zenodo.org/record/12783028/files/figure.png" pageId="8" pageNumber="539">Figure 3</figureCitation>
,
<figureCitation id="13322A5D214D0B37FBD1ACBD8110E4C9" box="[1118,1134,1924,1950]" captionStart="FIGURE 7" captionStartId="9.[151,250,928,952]" captionTargetBox="[151,1436,179,902]" captionTargetId="figure-15@9.[151,1436,179,902]" captionTargetPageId="9" captionText="FIGURE 7. Crux heggi sp. nov. Female holotype. a) Lateral view body. bd) Detailed views of ovipositor showing position and shape of subgenital plate. Pink triangle indicates subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.12783038" httpUri="https://zenodo.org/record/12783038/files/figure.png" pageId="8" pageNumber="539">7</figureCitation>
).
</paragraph>
<caption id="DF766650214C0B36FF18A8998651E08B" ID-DOI="http://doi.org/10.5281/zenodo.12783038" ID-Zenodo-Dep="12783038" httpUri="https://zenodo.org/record/12783038/files/figure.png" pageId="9" pageNumber="540" startId="9.[151,250,928,952]" targetBox="[151,1436,179,902]" targetPageId="9" targetType="figure">
<paragraph id="8BB636D8214C0B36FF18A8998651E08B" blockId="9.[151,1437,928,988]" pageId="9" pageNumber="540">
<emphasis id="B97DEACA214C0B36FF18A899846CE0EF" bold="true" box="[151,274,928,952]" pageId="9" pageNumber="540">FIGURE 7.</emphasis>
<taxonomicName id="4C094D5B214C0B36FE97A89984F6E0EF" authorityName="Trewick" authorityYear="2024" box="[280,392,928,952]" class="Holothuroidea" genus="Crux" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="540" phylum="Echinodermata" rank="species" species="heggi" status="sp. nov.">
<emphasis id="B97DEACA214C0B36FE97A89984F6E0EF" box="[280,392,928,952]" italics="true" pageId="9" pageNumber="540">Crux heggi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B1214C0B36FE01A89984A2E0E0" box="[398,476,928,951]" pageId="9" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
Female holotype. a) Lateral view body. bd) Detailed views of ovipositor showing position and shape of subgenital plate. Pink triangle indicates subgenital plate.
</paragraph>
</caption>
<caption id="DF766650214C0B36FF18AC5A8192E493" ID-DOI="http://doi.org/10.5281/zenodo.12783040" ID-Zenodo-Dep="12783040" httpUri="https://zenodo.org/record/12783040/files/figure.png" pageId="9" pageNumber="540" startId="9.[151,250,1891,1915]" targetBox="[218,1369,1165,1829]" targetPageId="9" targetType="figure">
<paragraph id="8BB636D8214C0B36FF18AC5A8192E493" blockId="9.[151,1436,1891,1988]" pageId="9" pageNumber="540">
<emphasis id="B97DEACA214C0B36FF18AC5A846CE42C" bold="true" box="[151,274,1891,1915]" pageId="9" pageNumber="540">FIGURE 8.</emphasis>
<taxonomicName id="4C094D5B214C0B36FE97AC5D84F6E42B" authorityName="Trewick" authorityYear="2024" box="[280,392,1892,1916]" class="Holothuroidea" genus="Crux" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="540" phylum="Echinodermata" rank="species" species="heggi" status="sp. nov.">
<emphasis id="B97DEACA214C0B36FE97AC5D84F6E42B" box="[280,392,1892,1916]" italics="true" pageId="9" pageNumber="540">Crux heggi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B1214C0B36FE01AC5D84A2E42C" box="[398,476,1892,1915]" pageId="9" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
Female holotype. Prolateral and retrolateral views of (a) fore and mid (b) legs with inset of mid femur showing apical retrolateral spine. c) Hind femur prolateral and retrolateral views. d) Hind tibia posterior view and detail of apical spine on hind femur (blue arrow). e) Hind tarsi, retrolateral view. Pink arrows indicate linear spines.
</paragraph>
</caption>
<caption id="DF766650214F0B35FF18A874864CE0F9" ID-DOI="http://doi.org/10.5281/zenodo.12783042" ID-Zenodo-Dep="12783042" httpUri="https://zenodo.org/record/12783042/files/figure.png" pageId="10" pageNumber="541" startId="10.[151,250,845,869]" targetBox="[151,1436,181,822]" targetPageId="10" targetType="figure">
<paragraph id="8BB636D8214F0B35FF18A874864CE0F9" blockId="10.[151,1437,845,942]" pageId="10" pageNumber="541">
<emphasis id="B97DEACA214F0B35FF18A874846CE031" bold="true" box="[151,274,845,870]" pageId="10" pageNumber="541">FIGURE 9.</emphasis>
<taxonomicName id="4C094D5B214F0B35FE97A87784F7E031" authorityName="Trewick" authorityYear="2024" box="[280,393,846,870]" class="Holothuroidea" genus="Crux" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="541" phylum="Echinodermata" rank="species" species="heggi" status="sp. nov.">
<emphasis id="B97DEACA214F0B35FE97A87784F7E031" box="[280,393,846,870]" italics="true" pageId="10" pageNumber="541">Crux heggi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57B1214F0B35FE00A87784A3E032" box="[399,477,846,869]" pageId="10" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
Male Paratype. Terminalia. ac) Lateral, dorsal and ventral views of posterior end of abdomen. d) Oblique posterior view of terminalia showing subgenital plate, and styli (blue arrows) pressed against paraprocts. e) Posterior view. f) Disarticulated fused paraproct structure (MPN CW4372).
</paragraph>
</caption>
<paragraph id="8BB636D8214F0B35FF48AF388128E69D" blockId="10.[151,1437,1024,2022]" pageId="10" pageNumber="541">
<emphasis id="B97DEACA214F0B35FF48AF388582E74D" box="[199,252,1025,1050]" italics="true" pageId="10" pageNumber="541">Legs</emphasis>
: Femora, fore tibiae and mid tibiae paler than dorsal abdomen, but with some darker markings. Legs are robust and glabrous except for inferior surfaces, which are clothed in fine gold-brown hairs. Most spines are dark and stout. Hind tibiae and femora similar in length, approximately equal to the body length. Fore femora with no apical spines. Mid femora with one retrolateral apical spine. Hind femora short, stout, with stout, fixed, spines towards the posterior end of the prominent inferior carinae; 2 retrolateral and 1 main prolateral towards distal end plus several small spines. Hind femora bear a short, fixed, retrolateral apical spine on knee. Fore tibiae with three inferior linear spines, 2 prolateral and 1retrolateral. Fore tibiae with 2 superior apical spines (1 prolateral and 1 retrolateral), and 2 inferior apical spines spines (1 prolateral and 1 retrolateral). Mid tibiae with same spines as fore tibiae. Hind tibiae naked above, fine gold hairs beneath and with two rows of prominent, stout, sharp, fixed spines; ~7 along superior prolateral margin and ~8 along superior retrolateral margin. Hind tibiae with a pair of small articulated, inferior apical spines, a pair of articulated, lateral apical spines, and a pair of longer, articulated superior apical spines. Hind tarsi with 1
<superScript id="7C7C9B90214F0B35FE9BAEB58461E6CD" attach="left" box="[276,287,1420,1434]" fontSize="6" pageId="10" pageNumber="541">st</superScript>
and 4
<superScript id="7C7C9B90214F0B35FEEEAEB58410E6CD" attach="left" box="[353,366,1420,1434]" fontSize="6" pageId="10" pageNumber="541">th</superScript>
segments longer than the 2
<superScript id="7C7C9B90214F0B35FD12AEB587D3E6CD" attach="left" box="[669,685,1420,1434]" fontSize="6" pageId="10" pageNumber="541">nd</superScript>
and 3
<superScript id="7C7C9B90214F0B35FD7FAEB58780E6CD" attach="none" box="[752,766,1420,1434]" fontSize="6" pageId="10" pageNumber="541">rd</superScript>
; unarmed except 1
<superScript id="7C7C9B90214F0B35FC5EAEB586A2E6CD" attach="left" box="[977,988,1420,1434]" fontSize="6" pageId="10" pageNumber="541">st</superScript>
and 2
<superScript id="7C7C9B90214F0B35FB90AEB58151E6CD" attach="left" box="[1055,1071,1420,1434]" fontSize="6" pageId="10" pageNumber="541">nd</superScript>
segments which each bear a pair of backward leaning, stout, fixed spines near distal end of superior surface (
<figureCitation id="13322A5D214F0B35FC69AE898136E69D" box="[998,1096,1456,1482]" captionStart="FIGURE 8" captionStartId="9.[151,250,1891,1915]" captionTargetBox="[218,1369,1165,1829]" captionTargetId="figure-47@9.[151,1436,1082,1851]" captionTargetPageId="9" captionText="FIGURE 8. Crux heggi sp. nov. Female holotype. Prolateral and retrolateral views of (a) fore and mid (b) legs with inset of mid femur showing apical retrolateral spine. c) Hind femur prolateral and retrolateral views. d) Hind tibia posterior view and detail of apical spine on hind femur (blue arrow). e) Hind tarsi, retrolateral view. Pink arrows indicate linear spines." figureDoi="http://doi.org/10.5281/zenodo.12783040" httpUri="https://zenodo.org/record/12783040/files/figure.png" pageId="10" pageNumber="541">Figure 8</figureCitation>
).
</paragraph>
<paragraph id="8BB636D8214F0B35FF48AEED85CBE529" blockId="10.[151,1437,1024,2022]" pageId="10" pageNumber="541">
<emphasis id="B97DEACA214F0B35FF48AEED8465E6B9" box="[199,283,1492,1518]" italics="true" pageId="10" pageNumber="541">Female</emphasis>
. Ventral surface of sternites hirsute. A small bifid subgenital plate with few hairs. Ovipositor long and almost straight (notably along the ventral margin), curving only slightly upwards towards the posterior end. Very pale cream at base especially on underside, rich amber brown in posterior half. Valves mostly smooth, but ventral valves with undulations at the tip. Cerci stout, pale, with fine amber hairs, and a dark ring below a cream tip (
<figureCitation id="13322A5D214F0B35FADEAD7985D9E529" captionStart="FIGURE 7" captionStartId="9.[151,250,928,952]" captionTargetBox="[151,1436,179,902]" captionTargetId="figure-15@9.[151,1436,179,902]" captionTargetPageId="9" captionText="FIGURE 7. Crux heggi sp. nov. Female holotype. a) Lateral view body. bd) Detailed views of ovipositor showing position and shape of subgenital plate. Pink triangle indicates subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.12783038" httpUri="https://zenodo.org/record/12783038/files/figure.png" pageId="10" pageNumber="541">Figure 7</figureCitation>
).
</paragraph>
<paragraph id="8BB636D8214F0B35FF48ADB184E6E591" blockId="10.[151,1437,1024,2022]" pageId="10" pageNumber="541">
<emphasis id="B97DEACA214F0B35FF48ADB1842EE5F5" box="[199,336,1672,1698]" italics="true" pageId="10" pageNumber="541">Colouration</emphasis>
: Predominantly cream/beige with dark brown markings primarily on dorsal surface of thorax and abdomen (
<figureCitation id="13322A5D214F0B35FE83AD958413E591" box="[268,365,1708,1734]" captionStart="FIGURE 3" captionStartId="3.[151,250,1522,1546]" captionTargetBox="[151,1436,181,1489]" captionTargetId="figure-15@3.[151,1436,181,1489]" captionTargetPageId="3" captionText="FIGURE 3. New species of Macropathinae from New Zealand. a) Crux boudica sp. nov., male, Rakiura (D. Hegg). b) Crux boudica sp. nov., female, Skippers Range (D. Hegg). c) Crux heggi sp. nov., male, Paparoa Range (D. Hegg). d) Crux heggi sp. nov., female, Saxon River (D. Hegg). e) Occultastella morgana sp. nov., male, Buller River (T. Jewell). f) Occultastella morgana sp. nov., female, Cobb Valley (A. McDonald)." figureDoi="http://doi.org/10.5281/zenodo.12783028" httpUri="https://zenodo.org/record/12783028/files/figure.png" pageId="10" pageNumber="541">Figure 3</figureCitation>
,
<figureCitation id="13322A5D214F0B35FEF5AD9584F4E591" box="[378,394,1708,1734]" captionStart="FIGURE 7" captionStartId="9.[151,250,928,952]" captionTargetBox="[151,1436,179,902]" captionTargetId="figure-15@9.[151,1436,179,902]" captionTargetPageId="9" captionText="FIGURE 7. Crux heggi sp. nov. Female holotype. a) Lateral view body. bd) Detailed views of ovipositor showing position and shape of subgenital plate. Pink triangle indicates subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.12783038" httpUri="https://zenodo.org/record/12783038/files/figure.png" pageId="10" pageNumber="541">7</figureCitation>
).
</paragraph>
<paragraph id="8BB636D8214F0B35FF48ADE981EAE4B1" blockId="10.[151,1437,1024,2022]" pageId="10" pageNumber="541">
<emphasis id="B97DEACA214F0B35FF48ADE9847FE5BD" box="[199,257,1744,1770]" italics="true" pageId="10" pageNumber="541">Male</emphasis>
: Similar to female in colour and leg spine combinations. Length
<quantity id="4CF19B3D214F0B35FC70ADE98134E5BD" box="[1023,1098,1744,1770]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="541" unit="mm" value="17.0">17mm</quantity>
; fore femur length
<quantity id="4CF19B3D214F0B35FAA1ADE980E6E5BD" box="[1326,1432,1744,1770]" metricMagnitude="-3" metricUnit="m" metricValue="9.16" pageId="10" pageNumber="541" unit="mm" value="9.16">9.16 mm</quantity>
; pronotum length 4.74; mid femur length
<quantity id="4CF19B3D214F0B35FDEFADCD87B9E459" box="[608,711,1780,1806]" metricMagnitude="-3" metricUnit="m" metricValue="8.27" pageId="10" pageNumber="541" unit="mm" value="8.27">8.27 mm</quantity>
; hind femur length
<quantity id="4CF19B3D214F0B35FC2EADCD816EE458" box="[929,1040,1780,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.558" pageId="10" pageNumber="541" unit="mm" value="15.58">15.58mm</quantity>
; fore tibia 9.0 mm; mid tibia length
<quantity id="4CF19B3D214F0B35FF18AC21847EE465" box="[151,256,1816,1842]" metricMagnitude="-3" metricUnit="m" metricValue="8.86" pageId="10" pageNumber="541" unit="mm" value="8.86">8.86 mm</quantity>
; hind tibia length
<quantity id="4CF19B3D214F0B35FE5DAC218737E465" box="[466,585,1816,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.6949999999999998" pageId="10" pageNumber="541" unit="mm" value="16.95">16.95 mm</quantity>
. Tergite IX simple. Subgenital plate broad, simple with straight or slightly concave posterior margin. Paraprocts extend beyond subgenital plate and styli to about middle of cerci. Paraprocts large, sturdy, and in close contact with each other forming a median carina on a sturdy, almost angular, hook that turns up and then forward to the tip of the abdomen under tergite IX. Styli short and pressed against the paraprocts and so difficult to see. Cerci gently tapering towards tip and extending well beyond the paraprocts and subgenital plate, diffuse darker band prior to pale tip, abundant long and short golden hairs (
<figureCitation id="13322A5D214F0B35FBABACF581F8E4B1" box="[1060,1158,1996,2022]" captionStart="FIGURE 9" captionStartId="10.[151,250,845,869]" captionTargetBox="[151,1436,181,822]" captionTargetId="figure-21@10.[151,1436,181,822]" captionTargetPageId="10" captionText="FIGURE 9. Crux heggi sp. nov. Male Paratype. Terminalia. ac) Lateral, dorsal and ventral views of posterior end of abdomen. d) Oblique posterior view of terminalia showing subgenital plate, and styli (blue arrows) pressed against paraprocts. e) Posterior view. f) Disarticulated fused paraproct structure (MPN CW4372)." figureDoi="http://doi.org/10.5281/zenodo.12783042" httpUri="https://zenodo.org/record/12783042/files/figure.png" pageId="10" pageNumber="541">Figure 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3136553214E0B34FF48ABAF8439E382" pageId="11" pageNumber="542" type="distribution">
<paragraph id="8BB636D8214E0B34FF48ABAF8439E382" blockId="11.[151,1436,150,285]" pageId="11" pageNumber="542">
<emphasis id="B97DEACA214E0B34FF48ABAF841DE3E7" bold="true" box="[199,355,150,176]" pageId="11" pageNumber="542">Distribution:</emphasis>
<collectingRegion id="49CDF83A214E0B34FEE5ABAE8497E3E6" box="[362,489,151,177]" country="New Zealand" name="West Coast" pageId="11" pageNumber="542">West Coast</collectingRegion>
and
<collectingRegion id="49CDF83A214E0B34FDAEABAE8705E3E6" box="[545,635,151,177]" country="New Zealand" name="Tasman" pageId="11" pageNumber="542">Tasman</collectingRegion>
, South Island,
<collectingCountry id="F31E7648214E0B34FCACABAE86C0E3E6" box="[803,958,151,177]" name="New Zealand" pageId="11" pageNumber="542">New Zealand</collectingCountry>
. Entomological regions Northwest
<collectingRegion id="49CDF83A214E0B34FAC2ABAE80E2E3E6" box="[1357,1436,151,177]" country="New Zealand" name="Nelson" pageId="11" pageNumber="542">Nelson</collectingRegion>
NN, Buller BR.
</paragraph>
</subSubSection>
<subSubSection id="C3136553214E0B34FF48ABE686A0E24A" pageId="11" pageNumber="542" type="etymology">
<paragraph id="8BB636D8214E0B34FF48ABE686A0E24A" blockId="11.[151,1436,150,285]" pageId="11" pageNumber="542">
<emphasis id="B97DEACA214E0B34FF48ABE68436E3AE" bold="true" box="[199,328,223,249]" pageId="11" pageNumber="542">Etymology</emphasis>
: Named for the orthopterist Danilo Hegg who collected specimens of this taxon and who has contributed extensively to revision of the
<collectingCountry id="F31E7648214E0B34FDE5AA3A867EE24A" box="[618,768,259,285]" name="New Zealand" pageId="11" pageNumber="542">New Zealand</collectingCountry>
<taxonomicName id="4C094D5B214E0B34FC88AA3A86A4E24A" authorityName="Walker" authorityYear="1869" box="[775,986,259,285]" class="Insecta" family="Rhaphidophoridae" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="542" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>