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<document id="BAEF96C0473E45EEC90CF07D61251E9D" ID-CLB-Dataset="72883" ID-DOI="10.11646/zootaxa.5150.3.7" ID-GBIF-Dataset="6ca82f5e-b51d-4beb-b33f-00a76c6a7bc8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6623165" ID-ZooBank="A379F26C-DCAB-43A8-B075-ED4ED10D8476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" checkinTime="1654677040107" checkinUser="plazi" docAuthor="Tang, William &amp; Dominguez, Hector Gomez" docDate="2022" docId="03A087C1FFEEFF9AFF717F33FA7352A4" docLanguage="en" docName="zootaxa.5150.3.7.pdf" docOrigin="Zootaxa 5150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila Tang, Skelley &amp; Perez-Farrera 2018" docType="key" docVersion="10" lastPageNumber="432" masterDocId="FF99FFB9FFECFF9EFFE67A00FFEE5019" masterDocTitle="Two new species of Pharaxonothinae beetles (Coleoptera: Erotylidae) inhabiting cones of the cycad Ceratozamia santillanii Pérez-Farr. &amp; Vovides (Cycadales: Zamiaceae) in Mexico" masterLastPageNumber="442" masterPageNumber="428" pageNumber="430" updateTime="1728482944120" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="2D1FF8F6EAF1FC3DF44615290C251515">Two new species of Pharaxonothinae beetles (Coleoptera: Erotylidae) inhabiting cones of the cycad Ceratozamia santillanii Pérez-Farr. &amp; Vovides (Cycadales: Zamiaceae) in Mexico</mods:title>
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<mods:namePart id="9C3B7756D02CE2F058719AD0D9311425">Tang, William</mods:namePart>
<mods:nameIdentifier id="AFCB3A882A62CB57460FCAF31A039679" type="ORCID">0000-0001-9009-3113</mods:nameIdentifier>
<mods:affiliation id="CB014995AE7C9FC9B31B0BDF2D26CE85">Florida State Collection of Arthropods, Florida Department of Agriculture - DPI, P. O. Box 147100, Gainesville, FL 32614 - 7100, USA. wlmtang 60 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9009 - 3113</mods:affiliation>
<mods:nameIdentifier id="49AE8300AC172CC1A6CDB5F60B0533FE" type="email">wlmtang60@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7802DFEDD47D12E2CFD44BA529AE49C0">Dominguez, Hector Gomez</mods:namePart>
<mods:nameIdentifier id="FD471316974135193ED034C2F8216A92" type="ORCID">0000-0001-7125-4639</mods:nameIdentifier>
<mods:affiliation id="2B5D030CA51C34F127814F3A42976485">Senda Sustentable A. C., Calle Cerro Azul, 208 B, Fraccionamiento Lomas del Pedregal, Berriozabal, CP 29130, Chiapas, Mexico hectorgomezdominguez @ hotmail. com; https: // orcid. org / 0000 - 0001 - 7125 - 4639</mods:affiliation>
<mods:nameIdentifier id="FAA17B6E21DBF21B741006246537EC93" type="email">hectorgomezdominguez@hotmail.com</mods:nameIdentifier>
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<mods:title id="7C3B98F074619220FA0C25A2CAE0C9B5">Zootaxa</mods:title>
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<mods:date id="006EE3E26707815AB5946D6AEBBE884C">2022</mods:date>
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<treatment id="03A087C1FFEEFF9AFF717F33FA7352A4" ID-DOI="http://doi.org/10.5281/zenodo.6646027" ID-GBIF-Taxon="195913643" ID-Zenodo-Dep="6646027" LSID="urn:lsid:plazi:treatment:03A087C1FFEEFF9AFF717F33FA7352A4" httpUri="http://treatment.plazi.org/id/03A087C1FFEEFF9AFF717F33FA7352A4" lastPageId="4" lastPageNumber="432" pageId="2" pageNumber="430">
<subSubSection id="C313655CFFEEFF9CFF717F33FD6F57E0" pageId="2" pageNumber="430" type="nomenclature">
<paragraph id="8BB636D7FFEEFF9CFF717F33FBDF5555" blockId="2.[151,1073,1330,1393]" box="[151,1073,1330,1357]" pageId="2" pageNumber="430">
<heading id="D0FE81BBFFEEFF9CFF717F33FBDF5555" bold="true" box="[151,1073,1330,1357]" fontSize="11" level="1" pageId="2" pageNumber="430" reason="1">
<emphasis id="B97DEAC5FFEEFF9CFF717F33FBDF5555" bold="true" box="[151,1073,1330,1357]" pageId="2" pageNumber="430">
Key to described species of
<taxonomicName id="4C094D54FFEEFF9CFE087F32FD935554" authorityName="Tang, Skelley &amp; Perez-Farrera" authorityYear="2018" box="[494,637,1330,1357]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFE087F32FD935554" bold="true" box="[494,637,1330,1357]" italics="true" pageId="2" pageNumber="430">Ceratophila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D54FFEEFF9CFD5A7F32FC845554" authorityName="Reitter" authorityYear="1875" box="[700,874,1330,1357]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFD5A7F32FC845554" bold="true" box="[700,874,1330,1357]" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
on
<taxonomicName id="4C094D54FFEEFF9CFC7E7F32FBDF5555" authorityName="Brongn" box="[920,1073,1330,1356]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFC7E7F32FBDF5555" bold="true" box="[920,1073,1330,1356]" italics="true" pageId="2" pageNumber="430">Ceratozamia</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB636D7FFEEFF9CFF717F56FDFB5568" blockId="2.[151,1073,1330,1393]" box="[151,533,1366,1393]" pageId="2" pageNumber="430">
(modified from
<bibRefCitation id="EF984B26FFEEFF9CFEAC7F57FDE35568" author="Tang, W. &amp; Perez-Farrera, M. A." box="[330,525,1366,1393]" pageId="2" pageNumber="430" pagination="151 - 178" refId="ref10958" refString="Tang, W., Skelley &amp; P. &amp; Perez-Farrera, M. A. (2018 a) Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae). Zootaxa, 4508 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 4508.2.1" type="journal article" year="2018">
Tang
<emphasis id="B97DEAC5FFEEFF9CFE6E7F57FE2E5569" box="[392,448,1366,1392]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
2018a
</bibRefCitation>
)
</paragraph>
<paragraph id="8BB636D7FFEEFF9CFF717F9EFEE55618" blockId="2.[151,1437,1438,2041]" pageId="2" pageNumber="430">
The key and species descriptions that follow rely on the morphology of spermatheca in female genitalia, particularly in
<taxonomicName id="4C094D54FFEEFF9CFF527FC2FEBA55C5" authorityName="Reitter" authorityYear="1875" box="[180,340,1474,1500]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFF527FC2FEBA55C5" box="[180,340,1474,1500]" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
(see
<figureCitation id="13322A52FFEEFF9CFE697FC2FE3E55C4" box="[399,464,1474,1501]" captionStart="FIGURE 2" captionStartId="3.[151,250,923,947]" captionTargetBox="[269,1317,181,899]" captionTargetId="figure-26@3.[269,1317,181,899]" captionTargetPageId="3" captionText="FIGURE 2. Spermatheca of Pharaxonotha from Jalpan de Cerra, Querétaro, Mexico indicating the diagnostic structures and orientations used in descriptions and key." figureDoi="http://doi.org/10.5281/zenodo.6623171" httpUri="https://zenodo.org/record/6623171/files/figure.png" pageId="2" pageNumber="430">Fig. 2</figureCitation>
for terminology and orientations used). For other diagnostic characters used in the key see
<figureCitation id="13322A52FFEEFF9CFF247FE6FEEB5618" box="[194,261,1510,1537]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="2" pageNumber="430">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="8BB636D7FFEEFF9CFF217C0AFAF65689" blockId="2.[151,1437,1438,2041]" pageId="2" pageNumber="430">
Morphological and molecular data suggest that there are more undescribed species of
<taxonomicName id="4C094D54FFEEFF9CFB637C0AFACB563D" authorityName="Reitter" authorityYear="1875" box="[1157,1317,1546,1572]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFB637C0AFACB563D" box="[1157,1317,1546,1572]" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D54FFEEFF9CFABD7C0BFF085651" authorityName="Tang, Skelley &amp; Perez-Farrera" authorityYear="2018" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFABD7C0BFF085651" italics="true" pageId="2" pageNumber="430">Ceratophila</emphasis>
</taxonomicName>
than are presented in the key below (
<bibRefCitation id="EF984B26FFEEFF9CFD617C2FFCA35650" author="Tang, W. &amp; Perez-Farrera, M. A." box="[647,845,1582,1609]" pageId="2" pageNumber="430" pagination="151 - 178" refId="ref10958" refString="Tang, W., Skelley &amp; P. &amp; Perez-Farrera, M. A. (2018 a) Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae). Zootaxa, 4508 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 4508.2.1" type="journal article" year="2018">
Tang
<emphasis id="B97DEAC5FFEEFF9CFD2E7C2FFCEE5651" box="[712,768,1582,1608]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
2018
<bibRefCitation id="EF984B26FFEEFF9CFCA77C2FFCA35650" author="Toon, A. &amp; Terry, L. I. &amp; Tang, W. &amp; Walter, G. H. &amp; Cook, L. G." box="[833,845,1583,1609]" pageId="2" pageNumber="430" pagination="1033 - 1058" refId="ref11423" refString="Toon, A., Terry, L. I., Tang, W., Walter, G. H. &amp; Cook, L. G. (2020) Insect pollination of cycads. Austral Ecology, 45, 1033 - 1058. https: // doi. org / 10.1111 / aec. 12925" type="journal article" year="2020">a</bibRefCitation>
</bibRefCitation>
, 2020; Tang unpublished data). A key to
<taxonomicName id="4C094D54FFEEFF9CFAF07C2EFF555675" authorityName="Reitter" authorityYear="1875" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFAF07C2EFF555675" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
inhabiting
<taxonomicName id="4C094D54FFEEFF9CFEDF7C53FE915675" authorityName="Lindl." box="[313,383,1619,1644]" class="Cycadopsida" family="Zamiaceae" genus="Dioon" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFEDF7C53FE915675" box="[313,383,1619,1644]" italics="true" pageId="2" pageNumber="430">Dioon</emphasis>
</taxonomicName>
was recently published by
<bibRefCitation id="EF984B26FFEEFF9CFD487C52FC7F5674" author="Skelley, P. &amp; Tang, W. &amp; Perez-Farrera, M. A." box="[686,913,1618,1645]" pageId="2" pageNumber="430" pagination="1 - 41" refId="ref10629" refString="Skelley, P., Tang, W. &amp; Perez-Farrera, M. A. (2022) Review of Pharaxonotha Reitter (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting the cycad genus Dioon Lindl. (Cycadales), with descriptions of nine new species and comments on P. kirschii Reitter. Insecta Mundi, 0917, 1 - 41." type="book chapter" year="2022">
Skelley
<emphasis id="B97DEAC5FFEEFF9CFCEE7C53FCAE5675" box="[776,832,1618,1644]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
(2022)
</bibRefCitation>
and keys for
<taxonomicName id="4C094D54FFEEFF9CFBC17C52FB295675" authorityName="Reitter" authorityYear="1875" box="[1063,1223,1618,1644]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFBC17C52FB295675" box="[1063,1223,1618,1644]" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
inhabiting
<taxonomicName id="4C094D54FFEEFF9CFAA37C53FF345689" authorityName="Brongn" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFAA37C53FF345689" italics="true" pageId="2" pageNumber="430">Ceratozamia</emphasis>
</taxonomicName>
and/or
<taxonomicName id="4C094D54FFEEFF9CFED67C77FE965689" authorityName="L." box="[304,376,1655,1680]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFED67C77FE965689" box="[304,376,1655,1680]" italics="true" pageId="2" pageNumber="430">Zamia</emphasis>
</taxonomicName>
inhabiting other regions are currently in progress and will be published elsewhere.
</paragraph>
<paragraph id="8BB636D7FFEEFF9CFF217C9BFD695738" blockId="2.[151,1437,1438,2041]" pageId="2" pageNumber="430">
Because of variable and overlapping morphological character states with some species, it is desirable, and in some cases necessary, to have a series of specimens representing color variations and both sexes to properly identify a species. In the following key, text in brackets before the species name indicate the states of
<collectingCountry id="F31E7647FFEEFF9CFB7E7CE2FB0056E5" box="[1176,1262,1762,1788]" name="Mexico" pageId="2" pageNumber="430">Mexico</collectingCountry>
where it occurs and the known host species of
<taxonomicName id="4C094D54FFEEFF9CFE097D07FD6E5738" authorityName="Brongn" box="[495,640,1799,1825]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFE097D07FD6E5738" box="[495,640,1799,1825]" italics="true" pageId="2" pageNumber="430">Ceratozamia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB636D7FFEEFF9CFF217D2BFD6F57E0" blockId="2.[151,1437,1438,2041]" pageId="2" pageNumber="430">
Concerns remain with the identity of some species described in
<bibRefCitation id="EF984B26FFEEFF9CFC467D2BFB97575C" author="Tang, W. &amp; Perez-Farrera, M. A." box="[928,1145,1834,1861]" pageId="2" pageNumber="430" pagination="151 - 178" refId="ref10958" refString="Tang, W., Skelley &amp; P. &amp; Perez-Farrera, M. A. (2018 a) Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae). Zootaxa, 4508 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 4508.2.1" type="journal article" year="2018">
Tang
<emphasis id="B97DEAC5FFEEFF9CFC067D2BFBFC575D" box="[992,1042,1834,1860]" italics="true" pageId="2" pageNumber="430">et al</emphasis>
. (2018a)
</bibRefCitation>
and those of
<bibRefCitation id="EF984B26FFEEFF9CFAF37D2AFE9E5770" author="Santiago-Jimenez, Q. J. &amp; Martinez-Dominguez, L. &amp; Nicolalde-Morejon, F." pageId="2" pageNumber="430" pagination="15 - 25" refId="ref10404" refString="Santiago-Jimenez, Q. J., Martinez-Dominguez, L. &amp; Nicolalde-Morejon, F. (2019) Two new Mexican species of Pharaxonotha Reitter, 1875 (Coleoptera: Erotylidae) from Ceratozamia tenuis (Cycadales: Zamiaceae). Dugesiana, 26 (1), 15 - 25. https: // doi. org / 10.32870 / dugesiana. v 26 i 1.7054" type="journal article" year="2019">
Santiago-Jiménez
<emphasis id="B97DEAC5FFEEFF9CFF017D4FFEF65771" box="[231,280,1870,1896]" italics="true" pageId="2" pageNumber="430">et al</emphasis>
. (2019)
</bibRefCitation>
(see
<bibRefCitation id="EF984B26FFEEFF9CFE4C7D4EFD405770" author="Skelley P. E. &amp; Tang W." box="[426,686,1870,1897]" pageId="2" pageNumber="430" pagination="1 - 11" refId="ref10580" refString="Skelley P. E. &amp; Tang W. (2020) Two new species of Pharaxonotha Reitter among the early-diverging lineages, with a key to the species of the genus (Coleoptera: Erotylidae: Pharaxonothinae). Insecta Mundi, 0837, 1 - 11." type="book chapter" year="2020">Skelley and Tang 2020</bibRefCitation>
). Based on descriptions and photographs in
<bibRefCitation id="EF984B26FFEEFF9CFB717D4EFF0A5794" author="Santiago-Jimenez, Q. J. &amp; Martinez-Dominguez, L. &amp; Nicolalde-Morejon, F." pageId="2" pageNumber="430" pagination="15 - 25" refId="ref10404" refString="Santiago-Jimenez, Q. J., Martinez-Dominguez, L. &amp; Nicolalde-Morejon, F. (2019) Two new Mexican species of Pharaxonotha Reitter, 1875 (Coleoptera: Erotylidae) from Ceratozamia tenuis (Cycadales: Zamiaceae). Dugesiana, 26 (1), 15 - 25. https: // doi. org / 10.32870 / dugesiana. v 26 i 1.7054" type="journal article" year="2019">
Santiago-Jimenez
<emphasis id="B97DEAC5FFEEFF9CFA837D4FFA735771" box="[1381,1437,1870,1896]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
(2019)
</bibRefCitation>
, compared with photographs of the
<typeStatus id="54B28875FFEEFF9CFD9C7D72FD095795" box="[634,743,1906,1932]" pageId="2" pageNumber="430" type="holotype">holotypes</typeStatus>
and additional specimens collected from the same host at the type locality, some may have been described twice. The tentative pairings are listed in the key with a question mark ‘?’. These will be more thoroughly addressed and identities resolved in a future paper covering
<taxonomicName id="4C094D54FFEEFF9CFB057DBAFA8457CD" authorityName="Tang, Skelley &amp; Perez-Farrera" authorityYear="2018" box="[1251,1386,1978,2004]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFB057DBAFA8457CD" box="[1251,1386,1978,2004]" italics="true" pageId="2" pageNumber="430">Ceratophila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D54FFEEFF9CFF717DDEFED957E1" authorityName="Reitter" authorityYear="1875" box="[151,311,2014,2040]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFF717DDEFED957E1" box="[151,311,2014,2040]" italics="true" pageId="2" pageNumber="430">Pharaxonotha</emphasis>
</taxonomicName>
on
<taxonomicName id="4C094D54FFEEFF9CFE877DDFFE1C57E0" authorityName="Brongn" box="[353,498,2015,2041]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEEFF9CFE877DDFFE1C57E0" box="[353,498,2015,2041]" italics="true" pageId="2" pageNumber="430">Ceratozamia</emphasis>
</taxonomicName>
in
<collectingRegion id="49CDF835FFEEFF9CFDF37DDFFD9257E0" box="[533,636,2015,2041]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="430">Veracruz</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="DF76665FFFEFFF9DFF71799BFDD453CE" ID-DOI="http://doi.org/10.5281/zenodo.6623171" ID-Zenodo-Dep="6623171" httpUri="https://zenodo.org/record/6623171/files/figure.png" pageId="3" pageNumber="431" startId="3.[151,250,923,947]" targetBox="[269,1317,181,899]" targetPageId="3">
<paragraph id="8BB636D7FFEFFF9DFF71799BFDD453CE" blockId="3.[151,1437,923,983]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFF71799BFEFA53AA" bold="true" box="[151,276,923,947]" pageId="3" pageNumber="431">FIGURE 2.</emphasis>
Spermatheca of
<taxonomicName id="4C094D54FFEFFF9DFE25799BFDBA53AA" authorityName="Reitter" authorityYear="1875" box="[451,596,923,947]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEFFF9DFE25799BFDBA53AA" box="[451,596,923,947]" italics="true" pageId="3" pageNumber="431">Pharaxonotha</emphasis>
</taxonomicName>
from Jalpan de Cerra, Querétaro, Mexico indicating the diagnostic structures and orientations used in descriptions and key.
</paragraph>
</caption>
<subSubSection id="C313655CFFEFFF9AFF717E03FA7352A4" lastPageId="4" lastPageNumber="432" pageId="3" pageNumber="431" type="key">
<keyStep id="30FD2E72FFEFFF9DFF717E03FA7254B3" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717E03FA72544B" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717E03FA72544B" pageId="3" pageNumber="431">
1. Lateral pronotal carinae thicker anteriorly in lateral view (
<figureCitation id="13322A52FFEFFF9DFCEA7E02FC9D5401" box="[780,883,1026,1048]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3D, E</figureCitation>
); elytra without a basal bead; head with supraocular lines, stridulatory files at base of head separated by distance ≈ ½ width of scutellar shield (
<figureCitation id="13322A52FFEFFF9DFBF47E20FBB3542F" box="[1042,1117,1056,1078]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3B</figureCitation>
); tarsomeres I–III distinctly widened, metatarsomere I width&gt; length................................................. [genus
<taxonomicName id="4C094D54FFEFFF9DFB097E3DFA8A544A" authorityName="Tang, Skelley &amp; Perez-Farrera" authorityYear="2018" box="[1263,1380,1085,1107]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEFFF9DFB097E3DFA8A544A" bold="true" box="[1263,1380,1085,1107]" italics="true" pageId="3" pageNumber="431">Ceratophila</emphasis>
</taxonomicName>
] …
<emphasis id="B97DEAC5FFEFFF9DFA767E3DFA72544B" bold="true" box="[1424,1436,1085,1106]" pageId="3" pageNumber="431">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFEFFF9DFF717E5BFA7254B3" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717E5BFA7254B3" pageId="3" pageNumber="431">
- Lateral pronotal carinae in lateral view narrow, same thickness for entire length (
<figureCitation id="13322A52FFEFFF9DFC177E5AFBD35468" box="[1009,1085,1114,1137]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3C</figureCitation>
); elytra with a basal bead (
<figureCitation id="13322A52FFEFFF9DFAA57E5AFA7F5468" box="[1347,1425,1114,1137]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3A</figureCitation>
); head without supraocular lines, stridulatory files at base of head separated by distance ≈ width of scutellar shield (
<figureCitation id="13322A52FFEFFF9DFAA47E78FA7F5497" box="[1346,1425,1144,1166]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3A</figureCitation>
); tarsomeres I–III not distinctly widened, metatarsomere I width &lt;length.................... [genus
<taxonomicName id="4C094D54FFEFFF9DFB307E95FA8B54B2" ID-CoL="6MVR" authorityName="Reitter" authorityYear="1875" box="[1238,1381,1173,1195]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEFFF9DFB307E95FA8B54B2" bold="true" box="[1238,1381,1173,1195]" italics="true" pageId="3" pageNumber="431">Pharaxonotha</emphasis>
</taxonomicName>
] …
<emphasis id="B97DEAC5FFEFFF9DFA767E95FA7254B3" bold="true" box="[1424,1436,1173,1194]" pageId="3" pageNumber="431">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFEFFF9DFF717EB3FA72558C" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717EB3FA725506" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717EB3FA725506" pageId="3" pageNumber="431">
2 (1). Pronotum in dorsal view, with anterior half of lateral carinae converging anteriorly; not explanate laterally, surface convexly curved to lateral carinae; lateral carinae in lateral view distinctly thicker anteriorly, anterior thickness 2× posterior thickness (
<figureCitation id="13322A52FFEFFF9DFF0F7EEDFEDB551A" box="[233,309,1261,1283]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3D</figureCitation>
); pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin (
<figureCitation id="13322A52FFEFFF9DFB007EEDFADF551A" box="[1254,1329,1261,1283]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3B</figureCitation>
); metatibia triangularly dilated toward apex (
<figureCitation id="13322A52FFEFFF9DFDC47F0AFD875538" box="[546,617,1290,1313]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3F</figureCitation>
)............................................ [subgenus
<taxonomicName id="4C094D54FFEFFF9DFB097F0AFA8A5539" authorityName="Tang, Skelley &amp; Perez-Farrera" authorityYear="2018" box="[1263,1380,1290,1312]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEFFF9DFB097F0AFA8A5539" bold="true" box="[1263,1380,1290,1312]" italics="true" pageId="3" pageNumber="431">Ceratophila</emphasis>
</taxonomicName>
] …
<emphasis id="B97DEAC5FFEFFF9DFA767F0AFA725506" bold="true" box="[1424,1436,1290,1311]" pageId="3" pageNumber="431">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFEFFF9DFF717F28FA72558C" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717F28FA72558C" pageId="3" pageNumber="431">
- Pronotum, in dorsal view, with central 2/3 of lateral carinae nearly parallel (
<figureCitation id="13322A52FFEFFF9DFC5D7F28FBFF5524" box="[955,1041,1320,1342]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="3" pageNumber="431">Figs. 4A</figureCitation>
,
<figureCitation id="13322A52FFEFFF9DFBFD7F28FBD55527" box="[1051,1083,1320,1342]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="3" pageNumber="431">5A</figureCitation>
), explanate laterally, surface broadly concave near lateral carinae; lateral carina weakly thickening anteriorly, anterior thickness &lt;1.5× that at base (
<figureCitation id="13322A52FFEFFF9DFAF07F45FA8C5542" box="[1302,1378,1349,1371]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3E</figureCitation>
); pronotal disc with longitudinal groove extending anteriorly from lateral basal pore in margin (
<figureCitation id="13322A52FFEFFF9DFBAF7F62FB795560" box="[1097,1175,1378,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3A</figureCitation>
); metatibia not triangularly dilated toward apex (
<figureCitation id="13322A52FFEFFF9DFE4A7F80FE19558F" box="[428,503,1408,1430]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3G</figureCitation>
)............................. [subgenus
<taxonomicName id="4C094D54FFEFFF9DFC287F7FFA8B558C" authority="Tang, Skelley &amp; Perez-Farrera" authorityName="Tang, Skelley &amp; Perez-Farrera" box="[974,1381,1407,1430]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="subGenus" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEFFF9DFC287F7FFA8B558C" bold="true" box="[974,1381,1407,1430]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFC287F7FFBCB558C" bold="true" box="[974,1061,1407,1429]" italics="true" pageId="3" pageNumber="431">Vovidesa</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</emphasis>
</taxonomicName>
] …
<emphasis id="B97DEAC5FFEFFF9DFA767F80FA72558C" bold="true" box="[1424,1436,1408,1429]" pageId="3" pageNumber="431">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFEFFF9DFF717F9DFA72567B" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717F9DFA735613" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<materialsCitation id="3B613C8AFFEFFF9DFF717F9DFA735613" ID-GBIF-Occurrence="3808181303" collectorName="Skelley &amp; Perez-Farrera" location="Veracruz" pageId="3" pageNumber="431" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<keyLead id="30F895E2FFEFFF9DFF717F9DFA735613" pageId="3" pageNumber="431">
3(2). Head with broad medial transverse ridge anterior of a distinct transverse basal groove in males (
<figureCitation id="13322A52FFEFFF9DFB707F9DFB0C55AA" box="[1174,1250,1437,1459]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3B</figureCitation>
), ridge and groove weak in females, ridge interrupting supraocular line in both male and female; head broad, width = 0.74–0.75× pronotal width; anterior clypeal margin emarginate; pronotal hypomeron densely punctate; [
<collectingRegion id="49CDF835FFEFFF9DFC227FD8FBF255F4" box="[964,1052,1496,1517]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="431">Veracruz</collectingRegion>
—
<taxonomicName id="4C094D54FFEFFF9DFBD27FD8FE8B5613" authority="Vazq. Torres, Sabato &amp; D. W. Stev." authorityName="Vazq. Torres, Sabato &amp; D. W. Stev." class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="euryphyllidia">
<emphasis id="B97DEAC5FFEFFF9DFBD27FD8FBAA55F4" box="[1076,1092,1496,1517]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.
<emphasis id="B97DEAC5FFEFFF9DFBB77FD8FB3F55F4" box="[1105,1233,1496,1517]" italics="true" pageId="3" pageNumber="431">euryphyllidia</emphasis>
Vázq.Torres, Sabato &amp; D.W. Stev.
</taxonomicName>
]................................................
<emphasis id="B97DEAC5FFEFFF9DFC4B7FF5FA735613" bold="true" box="[941,1437,1524,1547]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFC4B7FF5FC2D5613" bold="true" box="[941,963,1525,1546]" italics="true" pageId="3" pageNumber="431">C.</emphasis>
(
<emphasis id="B97DEAC5FFEFFF9DFC377FF5FC0F5613" bold="true" box="[977,993,1525,1546]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.)
<emphasis id="B97DEAC5FFEFFF9DFC137FF5FBB25612" bold="true" box="[1013,1116,1525,1547]" italics="true" pageId="3" pageNumber="431">chemnicki</emphasis>
Tang,
<collectorName id="26FC5301FFEFFF9DFB477FF5FB045612" box="[1185,1258,1525,1547]" pageId="3" pageNumber="431">Skelley</collectorName>
&amp;
<collectorName id="26FC5301FFEFFF9DFAED7FF4FA735613" box="[1291,1437,1524,1546]" pageId="3" pageNumber="431">Pérez-Farrera</collectorName>
</emphasis>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8BB636D7FFEFFF9DFF717C13FA72567B" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717C13FA72567B" pageId="3" pageNumber="431">
- Base of head at most with indistinct shallow transverse groove, supraocular line complete; head narrower, width = 0.66–0.68× pronotal width; anterior clypeal margin truncate or weakly convex; pronotal hypomeron with punctures sparse and minute or lacking.............................................................................................
<emphasis id="B97DEAC5FFEFFF9DFA767C4DFA72567B" bold="true" box="[1424,1436,1613,1634]" pageId="3" pageNumber="431">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFEFFF9DFF717C6BFA7256C1" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717C6BFA7356A3" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717C6BFA7356A3" pageId="3" pageNumber="431">
4(3). Dark elytral markings (when present) laterally covering most of disc; pronotum generally more elongate, pronotal length/pronotal width (PL/PW) = 0.83–0.90; [
<collectingRegion id="49CDF835FFEFFF9DFDD27C88FD6D5687" box="[564,643,1672,1694]" country="Mexico" name="Chiapas" pageId="3" pageNumber="431">Chiapas</collectingRegion>
—
<taxonomicName id="4C094D54FFEFFF9DFD7A7C88FBD25687" authority="Perez-Farr., Vovides &amp; Iglesias" authorityName="Perez-Farr., Vovides &amp; Iglesias" box="[668,1084,1672,1694]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="alvarezii">
<emphasis id="B97DEAC5FFEFFF9DFD7A7C88FD425684" box="[668,684,1672,1693]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.
<emphasis id="B97DEAC5FFEFFF9DFD5E7C88FCE55684" box="[696,779,1672,1693]" italics="true" pageId="3" pageNumber="431">alvarezii</emphasis>
Pérez-Farr., Vovides &amp; Iglesias
</taxonomicName>
,
<taxonomicName id="4C094D54FFEFFF9DFBA37C88FEC556A2" authority="Vovides, Perez-Farr. &amp; Iglesias" authorityName="Vovides, Perez-Farr. &amp; Iglesias" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="mirandae">
<emphasis id="B97DEAC5FFEFFF9DFBA37C88FBBB5684" box="[1093,1109,1672,1693]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.
<emphasis id="B97DEAC5FFEFFF9DFB867C88FB525684" box="[1120,1212,1672,1693]" italics="true" pageId="3" pageNumber="431">mirandae</emphasis>
Vovides, Pérez-Farr. &amp; Iglesias
</taxonomicName>
,
<taxonomicName id="4C094D54FFEFFF9DFED17CA5FDFC56A3" authority="D. W. Stev." authorityName="D. W. Stev." box="[311,530,1701,1723]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="norstogii">
<emphasis id="B97DEAC5FFEFFF9DFED17CA5FE4456A2" box="[311,426,1701,1723]" italics="true" pageId="3" pageNumber="431">C. norstogii</emphasis>
D.W.Stev.
</taxonomicName>
,
<taxonomicName id="4C094D54FFEFFF9DFDFB7CA5FC8756A3" authority="Perez-Farr. &amp; Iglesias" authorityName="Perez-Farr. &amp; Iglesias" box="[541,873,1700,1723]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="vovidesii">
<emphasis id="B97DEAC5FFEFFF9DFDFB7CA5FDC356A3" box="[541,557,1701,1722]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.
<emphasis id="B97DEAC5FFEFFF9DFDDF7CA5FD6056A3" box="[569,654,1701,1722]" italics="true" pageId="3" pageNumber="431">vovidesii</emphasis>
Pérez-Farr. &amp; Iglesias
</taxonomicName>
]....
<emphasis id="B97DEAC5FFEFFF9DFC497CA5FA7356A3" bold="true" box="[943,1437,1700,1723]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFC497CA5FC2B56A3" bold="true" box="[943,965,1701,1722]" italics="true" pageId="3" pageNumber="431">C.</emphasis>
(
<emphasis id="B97DEAC5FFEFFF9DFC357CA5FC0D56A3" bold="true" box="[979,995,1701,1722]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.)
<emphasis id="B97DEAC5FFEFFF9DFC117CA5FBB256A3" bold="true" box="[1015,1116,1701,1722]" italics="true" pageId="3" pageNumber="431">picipennis</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFEFFF9DFF717CC3FA7256C1" blockId="3.[151,1437,1026,2016]" box="[151,1436,1730,1752]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717CC3FA7256C1" box="[151,1436,1730,1752]" pageId="3" pageNumber="431">
- Dark elytral markings (when present) only along suture and laterally; pronotum generally shorter, PL/PW = 0.80–0.84.....
<emphasis id="B97DEAC5FFEFFF9DFA767CC3FA7256C1" bold="true" box="[1424,1436,1731,1752]" pageId="3" pageNumber="431">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFEFFF9DFF717CE0FA7357BC" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717CE0FA725729" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717CE0FA725729" pageId="3" pageNumber="431">
5(4). Color of pronotum and elytra similar; pronotum generally more quadrate; male genitalia with penile struts relatively long, length of penile strut relative to median lobe 2:1; setae at apex of parameres with greatest length &lt;dorsal width of parameres; [
<collectingRegion id="49CDF835FFEFFF9DFF017D1BFED15729" box="[231,319,1819,1840]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="431">Veracruz</collectingRegion>
—
<taxonomicName id="4C094D54FFEFFF9DFEB17D1BFE425729" ID-CoL="SQLK" box="[343,428,1819,1840]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="B97DEAC5FFEFFF9DFEB17D1BFE425729" box="[343,428,1819,1840]" italics="true" pageId="3" pageNumber="431">C. tenuis</emphasis>
</taxonomicName>
]..............
<emphasis id="B97DEAC5FFEFFF9DFD827D1AFBBC5729" bold="true" box="[612,1106,1818,1840]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFD827D1AFD945736" bold="true" box="[612,634,1818,1839]" italics="true" pageId="3" pageNumber="431">C.</emphasis>
(
<emphasis id="B97DEAC5FFEFFF9DFD6E7D1AFD765736" bold="true" box="[648,664,1818,1839]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.)
<emphasis id="B97DEAC5FFEFFF9DFD4A7D1AFCFF5729" bold="true" box="[684,785,1818,1840]" italics="true" pageId="3" pageNumber="431">sanchezae</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</emphasis>
(?
<taxonomicName id="4C094D54FFEFFF9DFB967D1BFA785729" ID-CoL="SQL8" authority="Santiago-Jimenez" authorityName="Santiago-Jimenez" box="[1136,1430,1818,1840]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="B97DEAC5FFEFFF9DFB967D1BFB0A5729" box="[1136,1252,1819,1840]" italics="true" pageId="3" pageNumber="431">P. mexicana</emphasis>
Santiago-Jiménez
</taxonomicName>
)
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFEFFF9DFF717D38FA7357BC" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717D38FA7357BC" pageId="3" pageNumber="431">
-
<materialsCitation id="3B613C8AFFEFFF9DFF047D38FA7357BC" ID-GBIF-Occurrence="3808181309" collectorName="Skelley &amp; Perez-Farrera" location="Oaxaca" pageId="3" pageNumber="431" specimenCount="1" stateProvince="Oaxaca">
Pronotum reddish brown contrasting with light brown of elytra; pronotum generally more trapezoidal, narrowing anteriorly; male genitalia with penile struts shorter, length of penile struts relative to median lobe 3:2; setae at apex of parameres with greatest length ≥ dorsal width of parameres; [
<collectingRegion id="49CDF835FFEFFF9DFD727D73FD335791" box="[660,733,1907,1928]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="431">Oaxaca</collectingRegion>
—
<taxonomicName id="4C094D54FFEFFF9DFD107D73FB245791" authority="Chemnick, T. J. Greg. &amp; Salas-Mor." authorityName="Chemnick, T. J. Greg. &amp; Salas-Mor." box="[758,1226,1906,1928]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="mixeorum">
<emphasis id="B97DEAC5FFEFFF9DFD107D73FCE85791" box="[758,774,1907,1928]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.
<emphasis id="B97DEAC5FFEFFF9DFCF47D73FC9C5791" box="[786,882,1907,1928]" italics="true" pageId="3" pageNumber="431">mixeorum</emphasis>
Chemnick, T.J.Greg. &amp; Salas-Mor.
</taxonomicName>
]............................................................................
<emphasis id="B97DEAC5FFEFFF9DFC247D90FA7357BC" bold="true" box="[962,1437,1935,1958]" pageId="3" pageNumber="431">
<emphasis id="B97DEAC5FFEFFF9DFC247D90FC3657BC" bold="true" box="[962,984,1936,1957]" italics="true" pageId="3" pageNumber="431">C.</emphasis>
(
<emphasis id="B97DEAC5FFEFFF9DFC007D90FC1857BC" bold="true" box="[998,1014,1936,1957]" italics="true" pageId="3" pageNumber="431">C</emphasis>
.)
<emphasis id="B97DEAC5FFEFFF9DFBEC7D90FBB357BC" bold="true" box="[1034,1117,1936,1957]" italics="true" pageId="3" pageNumber="431">gregoryi</emphasis>
Tang,
<collectorName id="26FC5301FFEFFF9DFB477D90FB0457BF" box="[1185,1258,1936,1958]" pageId="3" pageNumber="431">Skelley</collectorName>
&amp;
<collectorName id="26FC5301FFEFFF9DFAED7D8FFA7357BC" box="[1291,1437,1935,1957]" pageId="3" pageNumber="431">Pérez-Farrera</collectorName>
</emphasis>
</materialsCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFEFFF9AFF717DADFA7350FE" lastPageId="4" lastPageNumber="432" pageId="3" pageNumber="431">
<paragraph id="8BB636D7FFEFFF9DFF717DADFA7257F9" blockId="3.[151,1437,1026,2016]" pageId="3" pageNumber="431">
<keyLead id="30F895E2FFEFFF9DFF717DADFA7257F9" pageId="3" pageNumber="431">
6(2). Male meso- and metatibiae each with medial subapical emargination (
<figureCitation id="13322A52FFEFFF9DFC9C7DADFC2857DA" box="[890,966,1965,1987]" captionStart="FIGURE 3" captionStartId="6.[151,250,1783,1807]" captionTargetBox="[151,1436,560,1759]" captionTargetId="figure-200@6.[151,1436,560,1759]" captionTargetPageId="6" captionText="FIGURE 3. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) C. (Vovidesa) chiapensis; right metatibiae of male individuals, posterior view: F) C. (C.) gregoryi; G) C. (V.) mixeorum; b: basal bead of elytra; g: longitudinal grooves; s: stridulatory files t: transverse groove. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6623175" httpUri="https://zenodo.org/record/6623175/files/figure.png" pageId="3" pageNumber="431">Fig. 3G</figureCitation>
); female abdominal segments weakly alutaceous, punctures and setae visible except on lateral quarters.........................................................
<emphasis id="B97DEAC5FFEFFF9DFA767DCBFA7257F9" bold="true" box="[1424,1436,1995,2016]" pageId="3" pageNumber="431">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFE8FF9AFF717A98FA7350FE" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717A98FA7350FE" pageId="4" pageNumber="432">
- Male tibiae lacking medial subapical emargination; female abdominal segments strongly alutaceous, punctures obscured across middle, setae reduced [
<collectingRegion id="49CDF835FFE8FF9AFE587AB5FDE350D2" box="[446,525,181,203]" country="Mexico" name="Chiapas" pageId="4" pageNumber="432">Chiapas</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFDC07AB5FD7B50D3" box="[550,661,181,202]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="alvarezii">
<emphasis id="B97DEAC5FFE8FF9AFDC07AB5FDD850D3" box="[550,566,181,202]" italics="true" pageId="4" pageNumber="432">C</emphasis>
.
<emphasis id="B97DEAC5FFE8FF9AFDA47AB5FD7B50D3" box="[578,661,181,202]" italics="true" pageId="4" pageNumber="432">alvarezii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D54FFE8FF9AFD477AB5FCF750D3" box="[673,793,181,202]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="mirandae">
<emphasis id="B97DEAC5FFE8FF9AFD477AB5FCF750D3" box="[673,793,181,202]" italics="true" pageId="4" pageNumber="432">C. mirandae</emphasis>
</taxonomicName>
,
<emphasis id="B97DEAC5FFE8FF9AFCC37AB5FBE050D3" box="[805,1038,181,202]" italics="true" pageId="4" pageNumber="432">
<taxonomicName id="4C094D54FFE8FF9AFCC37AB5FC7D50D3" box="[805,915,181,202]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="norstogi">C. norstogi</taxonomicName>
,
<taxonomicName id="4C094D54FFE8FF9AFC7B7AB5FBE050D3" box="[925,1038,181,202]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="vovidesii">C. vovidesii</taxonomicName>
</emphasis>
]..........................................................................................
<emphasis id="B97DEAC5FFE8FF9AFC4A7AD2FA7350FE" bold="true" box="[940,1437,209,232]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFC4A7AD2FC2C50FE" bold="true" box="[940,962,210,231]" italics="true" pageId="4" pageNumber="432">C.</emphasis>
<taxonomicName id="4C094D54FFE8FF9AFC2F7AD2FA7350FE" authority="Tang, Skelley &amp; Perez-Farrera" authorityName="Tang, Skelley &amp; Perez-Farrera" box="[969,1437,209,232]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="chiapensis" subGenus="Vovidesa">
(
<emphasis id="B97DEAC5FFE8FF9AFC367AD2FC0E50FE" bold="true" box="[976,992,210,231]" italics="true" pageId="4" pageNumber="432">V</emphasis>
.)
<emphasis id="B97DEAC5FFE8FF9AFC127AD2FBB250F1" bold="true" box="[1012,1116,210,232]" italics="true" pageId="4" pageNumber="432">chiapensis</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFE8FF9AFF717AEFFA72518E" pageId="4" pageNumber="432">
<paragraph id="8BB636D7FFE8FF9AFF717AEFFA735126" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717AEFFA735126" pageId="4" pageNumber="432">
7(6). Submentum of male and female with sparse patch of setae-bearing punctures, male with long setae projecting anteriorly; pronotal length to width (PL/PW) = 0.65–0.73; male genitalia more dorsoventrally compressed, median lobe nearly cylindrical; [
<collectingRegion id="49CDF835FFE8FF9AFF017B2BFED15159" box="[231,319,299,320]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="432">Veracruz</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFEB17B2AFE3D5159" box="[343,467,298,320]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="mixeorum">
<emphasis id="B97DEAC5FFE8FF9AFEB17B2AFE3D5159" box="[343,467,298,320]" italics="true" pageId="4" pageNumber="432">C. mixeorum</emphasis>
</taxonomicName>
].......................................
<taxonomicName id="4C094D54FFE8FF9AFC547B2AFA735126" authority="Tang, Skelley &amp; Perez-Farrera" authorityName="Tang, Skelley &amp; Perez-Farrera" box="[946,1437,297,320]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="mixeorum" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFE8FF9AFC547B2AFA735126" bold="true" box="[946,1437,297,320]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFC547B2AFC265126" bold="true" box="[946,968,298,319]" italics="true" pageId="4" pageNumber="432">C.</emphasis>
(
<emphasis id="B97DEAC5FFE8FF9AFC307B2AFC045126" bold="true" box="[982,1002,298,319]" italics="true" pageId="4" pageNumber="432">V.</emphasis>
)
<emphasis id="B97DEAC5FFE8FF9AFC117B2AFBB25126" bold="true" box="[1015,1116,298,319]" italics="true" pageId="4" pageNumber="432">mixeorum</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFE8FF9AFF717B48FA72518E" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717B48FA72518E" pageId="4" pageNumber="432">
- Submentum of male and female with dense patch of setae-bearing punctures, male with long setae projecting laterally (
<figureCitation id="13322A52FFE8FF9AFA8A7B47FEF45162" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4E–F</figureCitation>
); PL/PW = 0.70–0.86; male genitalia laterally compressed, median lobe laterally flattened and distinctly curved in lateral view...............................................................................................
<emphasis id="B97DEAC5FFE8FF9AFA767B82FA72518E" bold="true" box="[1424,1436,386,407]" pageId="4" pageNumber="432">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFE8FF9AFF717BA0FA735214" pageId="4" pageNumber="432">
<paragraph id="8BB636D7FFE8FF9AFF717BA0FA7251CA" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717BA0FA7251CA" pageId="4" pageNumber="432">
8(7). Pronotum shorter, wider, pronotal length to width (PL/PW) of males = 0.76–0.79, of females = 0.70–0.76; [
<collectingRegion id="49CDF835FFE8FF9AFAF07BA0FA8051AC" box="[1302,1390,416,437]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="432">Veracruz</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFA607BA0FEF551CB" ID-CoL="SQLK" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="B97DEAC5FFE8FF9AFA607BA0FEF551CB" italics="true" pageId="4" pageNumber="432">C. tenuis</emphasis>
</taxonomicName>
]...............................
<taxonomicName id="4C094D54FFE8FF9AFD7D7BBDFB9A51CB" authority="Tang, Skelley &amp; Perez-Farrera" authorityName="Tang, Skelley &amp; Perez-Farrera" box="[667,1140,444,467]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="vazquezi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFE8FF9AFD7D7BBDFB9A51CB" bold="true" box="[667,1140,444,467]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFD7D7BBDFD5F51CB" bold="true" box="[667,689,445,466]" italics="true" pageId="4" pageNumber="432">C.</emphasis>
(
<emphasis id="B97DEAC5FFE8FF9AFD597BBDFD3C51CB" bold="true" box="[703,722,445,466]" italics="true" pageId="4" pageNumber="432">V.</emphasis>
)
<emphasis id="B97DEAC5FFE8FF9AFD067BBDFCDA51CB" bold="true" box="[736,820,445,466]" italics="true" pageId="4" pageNumber="432">vazquezi</emphasis>
Tang, Skelley &amp; Pérez-Farrera
</emphasis>
</taxonomicName>
(?
<taxonomicName id="4C094D54FFE8FF9AFB757BBDFA7851CA" ID-CoL="SQLK" authority="Santiago-Jimenez" authorityName="Santiago-Jimenez" box="[1171,1430,445,467]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="B97DEAC5FFE8FF9AFB757BBDFB0D51CB" box="[1171,1251,445,466]" italics="true" pageId="4" pageNumber="432">P. tenuis</emphasis>
Santiago-Jiménez
</taxonomicName>
)
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFE8FF9AFF717BDBFA735214" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717BDBFA735214" pageId="4" pageNumber="432">
- Pronotum longer, narrower, PL/PW of males = 0.81–0.86, of females = 0.78–0.82; [
<collectingRegion id="49CDF835FFE8FF9AFBE27BDAFBBD51E9" box="[1028,1107,474,496]" country="Mexico" name="Chiapas" pageId="4" pageNumber="432">Chiapas</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFB8A7BDAFB0751F6" ID-CoL="SQLH" box="[1132,1257,474,495]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFE8FF9AFB8A7BDAFB0751F6" box="[1132,1257,474,495]" italics="true" pageId="4" pageNumber="432">C. santillanii</emphasis>
</taxonomicName>
].................................................................
<emphasis id="B97DEAC5FFE8FF9AFCB37BF7FA735214" bold="true" box="[853,1437,503,525]" pageId="4" pageNumber="432">
<taxonomicName id="4C094D54FFE8FF9AFCB37BF7FAF25214" authority="Tang &amp; Gomez Dominguez" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[853,1308,503,525]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="chipjonesi" status="sp. nov." subGenus="Vovidesa">
<emphasis id="B97DEAC5FFE8FF9AFCB37BF7FC855215" bold="true" box="[853,875,503,524]" italics="true" pageId="4" pageNumber="432">C.</emphasis>
(
<emphasis id="B97DEAC5FFE8FF9AFC9F7BF8FC635214" bold="true" box="[889,909,504,525]" italics="true" pageId="4" pageNumber="432">V.</emphasis>
)
<emphasis id="B97DEAC5FFE8FF9AFC7C7BF7FC115214" bold="true" box="[922,1023,503,525]" italics="true" pageId="4" pageNumber="432">chipjonesi</emphasis>
Tang &amp; Gomez Dominguez
</taxonomicName>
,
<taxonomicNameLabel id="A24E57BEFFE8FF9AFAC07BF8FA735214" box="[1318,1437,503,525]" pageId="4" pageNumber="432" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FD2E72FFE8FF9AFF717815FA7352A4" pageId="4" pageNumber="432">
<paragraph id="8BB636D7FFE8FF9AFF717815FA73527C" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF717815FA73527C" pageId="4" pageNumber="432">
9(1). Pronotum anterior angles rounded (
<figureCitation id="13322A52FFE8FF9AFDA77815FD7E5232" box="[577,656,533,555]" captionStart="FIGURE 6" captionStartId="10.[151,250,1866,1890]" captionTargetBox="[275,1313,181,1843]" captionTargetId="figure-21@10.[275,1313,181,1843]" captionTargetPageId="10" captionText="FIGURE 6. Pharaxonotha perezi (A–F: female holotype): A–C) dorsal, lateral and ventral habitus (male habitus with no discernable difference); D–F) head, dorsal, lateral and ventral views; G) tegmen attached to median lobe (top) genital capsule (bottom), ventral views; H) tegmen attached to median lobe, lateral view; I) parameres of tegmen, (top to bottom) ventral and lateral views; J) spermatheca; K) female genitalia, ventral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G, H &amp; K; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623183" httpUri="https://zenodo.org/record/6623183/files/figure.png" pageId="4" pageNumber="432">Fig. 6A</figureCitation>
), PL/PW = 0.68–0.74; in females, spermatheca short, length/width ratio &lt;3.5, apical 1/3 wider than basal 1/3, insertion points of spermathecal and glandular ducts adjacent and in one shallow pit (
<figureCitation id="13322A52FFE8FF9AFAB77832FA7B5251" box="[1361,1429,562,584]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Fig. 4J</figureCitation>
) [
<collectingRegion id="49CDF835FFE8FF9AFF01784FFED8527C" box="[231,310,591,613]" country="Mexico" name="Chiapas" pageId="4" pageNumber="432">Chiapas</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFEA97850FE22527D" ID-CoL="SQLH" box="[335,460,591,613]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFE8FF9AFEA97850FEB1527C" box="[335,351,592,613]" italics="true" pageId="4" pageNumber="432">C</emphasis>
.
<emphasis id="B97DEAC5FFE8FF9AFE8D784FFE22527D" box="[363,460,591,612]" italics="true" pageId="4" pageNumber="432">santillanii</emphasis>
</taxonomicName>
]............................
<emphasis id="B97DEAC5FFE8FF9AFCD6784FFA73527C" bold="true" box="[816,1437,591,613]" pageId="4" pageNumber="432">
<taxonomicName id="4C094D54FFE8FF9AFCD6784FFAF4527C" authority="Tang &amp; Gomez Dominguez" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[816,1306,591,613]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="perezi" status="sp. nov.">
<emphasis id="B97DEAC5FFE8FF9AFCD6784FFBEE527D" bold="true" box="[816,1024,591,613]" italics="true" pageId="4" pageNumber="432">Pharaxonotha perezi</emphasis>
Tang &amp; Gomez Dominguez
</taxonomicName>
,
<taxonomicNameLabel id="A24E57BEFFE8FF9AFAC07850FA73527C" box="[1318,1437,591,613]" pageId="4" pageNumber="432" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB636D7FFE8FF9AFF71786DFA7352A4" blockId="4.[151,1437,151,701]" pageId="4" pageNumber="432">
<keyLead id="30F895E2FFE8FF9AFF71786DFA7352A4" pageId="4" pageNumber="432">
- Pronotum anterior angles pointed, PL/PW = 0.71–0.79; in females, spermatheca more elongated, length/width ratio&gt; 4, apical and basal 1/3 approx. equal in width; insertion points of spermathecal and glandular ducts separate and on a raised swelling, not in a pit [
<collectingRegion id="49CDF835FFE8FF9AFED678A7FE9152A4" box="[304,383,679,701]" country="Mexico" name="Chiapas" pageId="4" pageNumber="432">Chiapas</collectingRegion>
—
<taxonomicName id="4C094D54FFE8FF9AFE7E78A8FDFB52A5" ID-CoL="SQLH" box="[408,533,679,701]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFE8FF9AFE7E78A8FE4652A4" box="[408,424,680,701]" italics="true" pageId="4" pageNumber="432">C</emphasis>
.
<emphasis id="B97DEAC5FFE8FF9AFE5278A7FDFB52A5" box="[436,533,679,700]" italics="true" pageId="4" pageNumber="432">santillanii</emphasis>
</taxonomicName>
]...........................................................
<taxonomicName id="4C094D54FFE8FF9AFB0D78A7FA7352A4" box="[1259,1437,679,701]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97DEAC5FFE8FF9AFB0D78A7FA7352A4" bold="true" box="[1259,1437,679,701]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFB0D78A7FA9452A4" bold="true" box="[1259,1402,679,701]" italics="true" pageId="4" pageNumber="432">Pharaxonotha</emphasis>
sp.
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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