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<document id="CA25D7E29E46196F9822125D96113E56" ID-DOI="10.5281/zenodo.185534" ID-GBIF-Dataset="2dbe7cbb-40a2-4b2d-9281-84318b5700bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="185534" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459989612214" checkinUser="plazi" docAuthor="Lehr, Edgar, Catenazzi, Alessandro &amp; Rodríguez, Daniel" docDate="2009" docId="03A087A7FFB2EC4E74B3FA19FF19FE37" docLanguage="en" docName="zt01990p040.pdf" docOrigin="Zootaxa 1990" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pristimantis orcus Lehr, Catenazzi &amp; Rodríguez, 2009, new species" docType="treatment" docVersion="10" lastPageNumber="39" masterDocId="FF99FFDFFFB3EC477424FFAFFFE3FFF7" masterDocTitle="A new species of Pristimantis (Anura: Strabomantidae) from the Amazonian lowlands of northern Peru (Region Loreto and San Martín)" masterLastPageNumber="40" masterPageNumber="30" pageNumber="31" updateTime="1698232561086" updateUser="plazi">
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<mods:title id="1AA02D56FE81B48E1FE1E34FB2D150F0">A new species of Pristimantis (Anura: Strabomantidae) from the Amazonian lowlands of northern Peru (Region Loreto and San Martín)</mods:title>
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<mods:namePart id="9379E472E8630D8A497FCECE5FD2016A">Lehr, Edgar</mods:namePart>
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<mods:namePart id="7573C00C8C45DA2DA9BFC2C469CC689F">Catenazzi, Alessandro</mods:namePart>
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<mods:namePart id="39EDDB856EEC4A88D13EB488947B5A63">Rodríguez, Daniel</mods:namePart>
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<mods:date id="D9DE4D17149104FBC57993B79C1561C0">2009</mods:date>
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<treatment id="03A087A7FFB2EC4E74B3FA19FF19FE37" ID-DOI="http://doi.org/10.5281/zenodo.5625489" ID-GBIF-Taxon="119379837" ID-Zenodo-Dep="5625489" LSID="urn:lsid:plazi:treatment:03A087A7FFB2EC4E74B3FA19FF19FE37" httpUri="http://treatment.plazi.org/id/03A087A7FFB2EC4E74B3FA19FF19FE37" lastPageId="9" lastPageNumber="39" pageId="1" pageNumber="31">
<subSubSection id="C313653AFFB2EC4674B3FA19FB07F993" pageId="1" pageNumber="31" type="nomenclature">
<paragraph id="8BB636B1FFB2EC4674B3FA19FDE9FA27" blockId="1.[151,522,1462,1488]" box="[151,522,1462,1488]" pageId="1" pageNumber="31">
<heading id="D0FE81DDFFB2EC4674B3FA19FDE9FA27" bold="true" box="[151,522,1462,1488]" fontSize="11" level="1" pageId="1" pageNumber="31" reason="1">
<emphasis id="B97DEAA3FFB2EC4674B3FA19FDE9FA27" bold="true" box="[151,522,1462,1488]" pageId="1" pageNumber="31">
<taxonomicName id="4C094D32FFB2EC4674B3FA19FE97FA27" box="[151,372,1462,1488]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="31" phylum="Chordata" rank="species" species="orcus" status="sp. nov.">
<emphasis id="B97DEAA3FFB2EC4674B3FA19FE97FA27" bold="true" box="[151,372,1462,1488]" italics="true" pageId="1" pageNumber="31">Pristimantis orcus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57D8FFB2EC46755FFA19FDE9FA27" box="[379,522,1462,1488]" pageId="1" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB636B1FFB2EC4674B3F9AFFB07F993" blockId="1.[151,1437,1536,2018]" pageId="1" pageNumber="31">
<taxonomicName id="4C094D32FFB2EC4674B3F9AFFE8CF9EE" box="[151,367,1536,1561]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB2EC4674B3F9AFFE8CF9EE" box="[151,367,1536,1561]" italics="true" pageId="1" pageNumber="31">Eleutherodactylus</emphasis>
</taxonomicName>
sp. 5, Fig. 7A -
<bibRefCitation id="EF984B40FFB2EC467612F9AFFC29F9ED" author="Catenazzi" box="[566,970,1536,1562]" pageId="1" pageNumber="39" refString="Catenazzi, A. &amp; Bustamante, M. (2007) Amphibians and reptiles; Appendix 4; In: Vriesendorp, C., Alvarez, J. A., Barbagelata, N., Anderson, W. S. &amp; Moskovits, D. K. (Eds.). Nanay-Mazan-Arabela. Rapid Biological Inventories 18. The Field Museum, Chicago, pp. 130 - 134, 206 - 211." type="journal article" year="2007">Catenazzi and Bustamante (2007)</bibRefCitation>
Amphibians and reptiles; Appendix 4. pp. 130134, 206211. In: Vriesendorp, C., J. A. Álvarez, N. Barbagelata, W. S. Anderson &amp; D. K. Moskovits (Eds.). Nanay-Mazán-Arabela. Rapid Biological Inventories 18. The Field Museum, Chicago.
</paragraph>
</subSubSection>
<subSubSection id="C313653AFFB2EC4574E1F9DFFCE3FED1" lastPageId="2" lastPageNumber="32" pageId="1" pageNumber="31" type="materials_examined">
<paragraph id="8BB636B1FFB2EC4674E1F9DFFD8DF923" blockId="1.[151,1437,1536,2018]" pageId="1" pageNumber="31">
<typeStatus id="54B28813FFB2EC4674E1F9DFFEDBF97D" box="[197,312,1648,1674]" pageId="1" pageNumber="31" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED18AE74FFB2EC467567F9DFFE7EF97D" box="[323,413,1648,1674]" pageId="1" pageNumber="31">MUSM</collectionCode>
27435 (
<figureCitation id="13322A34FFB2EC467624F9DFFDAEF97D" box="[512,589,1648,1674]" captionStart="FIGURE 1" captionStartId="2.[151,258,1596,1620]" captionTargetBox="[151,1434,351,1570]" captionTargetId="figure@2.[151,1436,351,1571]" captionTargetPageId="2" captionText="FIGURE 1. Left column: Female Pristimantis orcus (MUSM 27435, holotype, SVL 36.5 mm) in dorsolateral (A), lateral (B), and ventral (C) views; right column: male Pristimantis orcus (MUSM 25938, paratype, SVL 22.4 mm) in dorsolateral (D), lateral (E), and ventral (F) views. Photos A C by D. Rodríguez, photos D F by A. Catenazzi." httpUri="https://zenodo.org/record/185535/files/figure.png" pageId="1" pageNumber="31">Figs. 1</figureCitation>
AC; 23), an adult female collected at Comunidad de Nueva Vida (
<geoCoordinate id="EE3D5076FFB2EC4671A7F9DFFEF4F958" direction="south" orientation="latitude" pageId="1" pageNumber="31" precision="1" value="-3.6325557">3º 37' 57.2'' S</geoCoordinate>
,
<geoCoordinate id="EE3D5076FFB2EC467507F93AFE38F958" box="[291,475,1685,1711]" direction="west" orientation="longitude" pageId="1" pageNumber="31" precision="1" value="-75.34692">75º 20' 48.9'' W</geoCoordinate>
) at
<quantity id="4CF19B54FFB2EC467622F93AFDB3F958" box="[518,592,1685,1711]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="31" unit="m" value="158.0">158 m</quantity>
elevation, Distrito de Trompeteros, Provincia de Loreto, Región Loreto,
<collectingCountry id="F31E7621FFB2EC4674B3F915FF33F923" box="[151,208,1722,1748]" name="Peru" pageId="1" pageNumber="31">Peru</collectingCountry>
, on
<date id="FFB71071FFB2EC467524F915FE7FF923" box="[256,412,1722,1748]" pageId="1" pageNumber="31" value="2008-06-18">18 June 2008</date>
by D. Rodríguez.
</paragraph>
<paragraph id="8BB636B1FFB2EC4574E1F94FFCE3FED1" blockId="1.[151,1437,1536,2018]" lastBlockId="2.[151,1436,152,294]" lastPageId="2" lastPageNumber="32" pageId="1" pageNumber="31">
<typeStatus id="54B28813FFB2EC4674E1F94FFEA1F90D" box="[197,322,1760,1786]" pageId="1" pageNumber="31" type="paratype">Paratypes</typeStatus>
: Six: four males (
<collectionCode id="ED18AE74FFB2EC467639F94FFD9FF90D" box="[541,636,1760,1786]" pageId="1" pageNumber="31">MUSM</collectionCode>
22437, 25938, 27436, 27392), and two females (
<collectionCode id="ED18AE74FFB2EC4670C5F94FFAA3F90D" box="[1249,1344,1760,1786]" pageId="1" pageNumber="31">MUSM</collectionCode>
22444, 27456). One male (
<collectionCode id="ED18AE74FFB2EC46755DF8A9FE36F8D7" box="[377,469,1798,1824]" pageId="1" pageNumber="31">MUSM</collectionCode>
27436) collected in axillar amplexus with the
<typeStatus id="54B28813FFB2EC4677D2F8A9FB83F8D7" box="[1014,1120,1798,1824]" pageId="1" pageNumber="31" type="holotype">holotype</typeStatus>
; one male (
<collectionCode id="ED18AE74FFB2EC4670C2F8A9FAA1F8D7" box="[1254,1346,1798,1824]" pageId="1" pageNumber="31">MUSM</collectionCode>
27392) collected at
<geoCoordinate id="EE3D5076FFB2EC467501F882FE24F8B0" box="[293,455,1837,1863]" direction="south" orientation="latitude" pageId="1" pageNumber="31" precision="1" value="-3.6800835">3º 40' 48.3'' S</geoCoordinate>
,
<geoCoordinate id="EE3D5076FFB2EC4675F1F882FD73F8B0" box="[469,656,1837,1863]" direction="west" orientation="longitude" pageId="1" pageNumber="31" precision="1" value="-75.58295">75º 34' 58.6'' W</geoCoordinate>
at
<quantity id="4CF19B54FFB2EC467691F882FD1CF8B0" box="[693,767,1837,1863]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="1" pageNumber="31" unit="m" value="163.0">163 m</quantity>
elevation, Distrito de Trompeteros, Provincia de Loreto, on 0
<date id="FFB71071FFB2EC4674E9F8FBFE84F899" box="[205,359,1876,1902]" pageId="1" pageNumber="31" value="2008-04-03">3 April 2008</date>
by D. Rodríguez; one female (
<collectionCode id="ED18AE74FFB2EC4676C4F8FBFCDEF899" box="[736,829,1876,1902]" pageId="1" pageNumber="31">MUSM</collectionCode>
27456) collected at Comunidad de Santa Rosa de Siamba (
<geoCoordinate id="EE3D5076FFB2EC4674DBF8D5FE41F863" box="[255,418,1914,1940]" direction="south" orientation="latitude" pageId="1" pageNumber="31" precision="1" value="-4.1048055">4º 06' 17.3'' S</geoCoordinate>
,
<geoCoordinate id="EE3D5076FFB2EC467594F8D5FD8FF863" box="[432,620,1914,1940]" direction="west" orientation="longitude" pageId="1" pageNumber="31" precision="1" value="-75.53345">75º 32' 00.4'' W</geoCoordinate>
) at
<quantity id="4CF19B54FFB2EC4676BDF8D5FD07F863" box="[665,740,1914,1940]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="31" unit="m" value="133.0">133 m</quantity>
elevation, Distrito de Urarinas, Provincia de Loreto, on
<date id="FFB71071FFB2EC46715BF8D5FEC5F84C" pageId="1" pageNumber="31" value="2008-03-20">20 March 2008</date>
by J. Ramírez; one male (
<collectionCode id="ED18AE74FFB2EC46767EF80EFD55F84C" box="[602,694,1953,1979]" pageId="1" pageNumber="31">MUSM</collectionCode>
25938) collected at the upper Panguana watershed (an affluent of the Río Napo;
<geoCoordinate id="EE3D5076FFB2EC467542F867FE12F815" box="[358,497,1992,2018]" direction="south" orientation="latitude" pageId="1" pageNumber="31" precision="15" value="-2.136389">2º 08' 11'' S</geoCoordinate>
,
<geoCoordinate id="EE3D5076FFB2EC4675DBF867FD46F815" box="[511,677,1992,2018]" direction="west" orientation="longitude" pageId="1" pageNumber="31" precision="15" value="-75.14916">75º 08' 57'' W</geoCoordinate>
) at
<quantity id="4CF19B54FFB2EC4676F7F867FCFDF815" box="[723,798,1992,2018]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="31" unit="m" value="160.0">160 m</quantity>
elevation, Distrito de Napo, Provincia de Maynas, on
<date id="FFB71071FFB1EC4574B3FF37FEBFFF45" box="[151,348,152,178]" pageId="2" pageNumber="32" value="2006-08-29">29 August 2006</date>
by A. Catenazzi and M. Bustamante. All from Región Loreto,
<collectingCountry id="F31E7621FFB1EC45704AFF37FB49FF45" box="[1134,1194,152,178]" name="Peru" pageId="2" pageNumber="32">Peru</collectingCountry>
. One male (
<collectionCode id="ED18AE74FFB1EC45711AFF37FA7FFF45" box="[1342,1436,152,178]" pageId="2" pageNumber="32">MUSM</collectionCode>
22437) and one female (
<collectionCode id="ED18AE74FFB1EC457596FF11FDEDFF2F" box="[434,526,190,216]" pageId="2" pageNumber="32">MUSM</collectionCode>
22444) collected at km 34 on road from Tarapoto to Yurimaguas (
<geoCoordinate id="EE3D5076FFB1EC457131FF11FF4AFF08" direction="south" orientation="latitude" pageId="2" pageNumber="32" precision="1" value="-6.4273887">6º 25' 38.6'' S</geoCoordinate>
,
<geoCoordinate id="EE3D5076FFB1EC457493FF4AFE96FF08" box="[183,373,229,255]" direction="west" orientation="longitude" pageId="2" pageNumber="32" precision="1" value="-76.30019">76º 18' 00.7'' W</geoCoordinate>
), Valle de Cainarachi, Distrito de La Banda de Shilcayo, Provincia de San Martín, Región de San Martín,
<collectingCountry id="F31E7621FFB1EC457568FEA3FE67FED1" box="[332,388,268,294]" name="Peru" pageId="2" pageNumber="32">Peru</collectingCountry>
, on
<date id="FFB71071FFB1EC457590FEA3FD83FED1" box="[436,608,268,294]" pageId="2" pageNumber="32" value="2005-02">February 2005</date>
by C. Torres.
</paragraph>
</subSubSection>
<caption id="DF766639FFB1EC4574B3F993FAD7F96E" httpUri="https://zenodo.org/record/185535/files/figure.png" pageId="2" pageNumber="32" targetBox="[151,1434,351,1570]" targetPageId="2">
<paragraph id="8BB636B1FFB1EC4574B3F993FAD7F96E" blockId="2.[151,1436,1596,1689]" pageId="2" pageNumber="32">
<emphasis id="B97DEAA3FFB1EC4574B3F993FEFCF9A3" bold="true" box="[151,287,1596,1620]" pageId="2" pageNumber="32">FIGURE 1.</emphasis>
Left column: Female
<taxonomicName id="4C094D32FFB1EC45763CF992FD06F9A3" box="[536,741,1597,1620]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="32" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB1EC45763CF992FD06F9A3" box="[536,741,1597,1620]" italics="true" pageId="2" pageNumber="32">Pristimantis orcus</emphasis>
</taxonomicName>
(MUSM 27435, holotype, SVL 36.5 mm) in dorsolateral (A), lateral (B), and ventral (C) views; right column: male
<taxonomicName id="4C094D32FFB1EC4576CEF9CFFC51F980" box="[746,946,1632,1655]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="32" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB1EC4576CEF9CFFC51F980" box="[746,946,1632,1655]" italics="true" pageId="2" pageNumber="32">Pristimantis orcus</emphasis>
</taxonomicName>
(MUSM 25938, paratype, SVL 22.4 mm) in dorsolateral (D), lateral (E), and ventral (F) views. Photos AC by D. Rodríguez, photos DF by A. Catenazzi.
</paragraph>
</caption>
<subSubSection id="C313653AFFB1EC4274E1F964FD90FF22" lastPageId="5" lastPageNumber="35" pageId="2" pageNumber="32" type="diagnosis">
<paragraph id="8BB636B1FFB1EC4474E1F964FC5DFE84" blockId="2.[151,1437,1739,2036]" lastBlockId="3.[151,1436,152,371]" lastPageId="3" lastPageNumber="33" pageId="2" pageNumber="32">
<emphasis id="B97DEAA3FFB1EC4574E1F964FEA7F912" bold="true" box="[197,324,1739,1765]" pageId="2" pageNumber="32">Diagnosis.</emphasis>
A member of
<taxonomicName id="4C094D32FFB1EC4575CAF963FD9EF912" box="[494,637,1740,1765]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB1EC4575CAF963FD9EF912" box="[494,637,1740,1765]" italics="true" pageId="2" pageNumber="32">Pristimantis</emphasis>
</taxonomicName>
based on the diagnostic set of characters as defined by
<bibRefCitation id="EF984B40FFB1EC457120F964FF0FF8FB" pageId="2" pageNumber="39" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article">Hedges et al. (2008)</bibRefCitation>
assigned to the
<taxonomicName id="4C094D32FFB1EC4575E6F95DFC20F8FC" box="[450,963,1778,1803]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="32" phylum="Chordata" rank="species" species="unistrigatus" subGenus="Pristimantis">
<emphasis id="B97DEAA3FFB1EC4575E6F95DFC20F8FC" box="[450,963,1778,1803]" italics="true" pageId="2" pageNumber="32">Pristimantis (Pristimantis) unistrigatus</emphasis>
</taxonomicName>
species Group having the following combination of characters: (1) Skin on dorsum finely shagreen with low scattered tubercles, skin on venter weakly areolate; discoidal fold present; dorsolateral fold absent; (2) tympanic membrane and tympanic annulus present; (3) snout acutely rounded in dorsal view, rounded in lateral view; (4) upper eyelid with minute tubercles; width of upper eyelid narrower than IOD; cranial crests absent; (5) dentigerous processes of vomers prominent, oblique; (6) males without vocal slits, nuptial pads present on dorsal and medial surfaces of thumb; (7) Finger I shorter than Finger II; discs of digits broadly expanded, truncate; (8) fingers with broad lateral fringes; (9) ulnar and tarsal tubercles present; (10) heel with small tubercles; inner tarsal fold present; (11) inner metatarsal tubercle ovoid, about three times as large as outer; outer metatarsal tubercle rounded; supernumerary plantar tubercles present; (12) toes with broad lateral fringes; webbing between Toes IIV; Toe V much longer than Toe III; discs of digits broadly expanded, truncate, about as large as those on fingers; (13) in ethanol, dorsum grayish brown with tan blotches, venter grayish brown and tan mottled; (14) SVL in adult females
<quantity id="4CF19B54FFB0EC4474DDFEF6FE47FE84" box="[249,420,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="3.4549999999999996" metricValueMax="3.65" metricValueMin="3.26" pageId="3" pageNumber="33" unit="mm" value="34.55" valueMax="36.5" valueMin="32.6">32.636.5 mm</quantity>
(
<emphasis id="B97DEAA3FFB0EC447590FEF6FE20FE85" box="[436,451,345,370]" italics="true" pageId="3" pageNumber="33">n</emphasis>
= 3), in adult males 20.0
<quantity id="4CF19B54FFB0EC4476DCFEF6FCBCFE84" box="[760,863,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="3" pageNumber="33" unit="mm" value="25.1">25.1 mm</quantity>
(
<emphasis id="B97DEAA3FFB0EC447754FEF6FC9CFE85" box="[880,895,345,370]" italics="true" pageId="3" pageNumber="33">n</emphasis>
= 4).
</paragraph>
<caption id="DF766639FFB0EC4474B3FB6DFAFDFB2E" box="[151,1310,1218,1242]" httpUri="https://zenodo.org/record/185536/files/figure.png" pageId="3" pageNumber="33" targetBox="[157,1425,430,1194]" targetPageId="3">
<paragraph id="8BB636B1FFB0EC4474B3FB6DFAFDFB2E" blockId="3.[151,1310,1218,1242]" box="[151,1310,1218,1242]" pageId="3" pageNumber="33">
<emphasis id="B97DEAA3FFB0EC4474B3FB6DFEFAFB2D" bold="true" box="[151,281,1218,1242]" pageId="3" pageNumber="33">FIGURE 2.</emphasis>
Preserved
<taxonomicName id="4C094D32FFB0EC4475ABFB6DFDB0FB2E" box="[399,595,1218,1241]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC4475ABFB6DFDB0FB2E" box="[399,595,1218,1241]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
(holotype) in dorsal (A), and ventral (B) views. Photos by E. Lehr.
</paragraph>
</caption>
<paragraph id="8BB636B1FFB0EC4474E1FAA4FB0CF8B4" blockId="3.[151,1437,1291,2013]" pageId="3" pageNumber="33">
<taxonomicName id="4C094D32FFB0EC4474E1FAA4FE49FAD3" box="[197,426,1291,1316]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC4474E1FAA4FE49FAD3" box="[197,426,1291,1316]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
is most similar to following five medium-sized species of
<taxonomicName id="4C094D32FFB0EC4470B4FAA4FACAFAD3" box="[1168,1321,1291,1316]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB0EC4470B4FAA4FACAFAD3" box="[1168,1321,1291,1316]" italics="true" pageId="3" pageNumber="33">Pristimantis</emphasis>
</taxonomicName>
from the Amazonian lowlands and lower eastern Andes (
<emphasis id="B97DEAA3FFB0EC4476E8FA9DFA78FABC" box="[716,1435,1330,1355]" italics="true" pageId="3" pageNumber="33">
<taxonomicName id="4C094D32FFB0EC4476E8FA9DFC9BFABC" box="[716,888,1330,1355]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="altamazonicus">altamazonicus</taxonomicName>
,
<taxonomicName id="4C094D32FFB0EC4477A1FA9DFBECFABC" box="[901,1039,1330,1355]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="diadematus">diadematus</taxonomicName>
,
<taxonomicName id="4C094D32FFB0EC447038FA9DFB56FABC" box="[1052,1205,1330,1355]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="eurydactylus">eurydactylus</taxonomicName>
,
<taxonomicName id="4C094D32FFB0EC4470E5FA9DFA7BFABC" box="[1217,1432,1330,1355]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="ventrimarmoratus">ventrimarmoratus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C094D32FFB0EC4474EEFAF6FEBFFA85" box="[202,348,1369,1394]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB0EC4474EEFAF6FEBFFA85" box="[202,348,1369,1394]" italics="true" pageId="3" pageNumber="33">Pristimantis</emphasis>
</taxonomicName>
sp.) in having Finger I &lt;II, discs on fingers and toes broadly extended, dorsolateral folds absent, dentigerous processes of vomers, males without vocal slits, a pale venter with darker blotches, and the groin with a contrasting pattern consisting of white, blueish white, red or yellow with dark brown or black (
<tableCitation id="C68B030AFFB0EC4474BAFA62FF14FA10" box="[158,247,1485,1511]" captionStart="TABLE 1" captionStartId="5.[151,240,151,175]" captionTargetBox="[151,1436,241,1557]" captionTargetPageId="5" captionText="TABLE 1. Selected characters (+ = character present; — = character absent) and character conditions among selected" httpUri="http://table.plazi.org/id/DF766639FFB6EC4274B3FF38FA7FFF58" pageId="3" pageNumber="33" tableUuid="DF766639FFB6EC4274B3FF38FA7FFF58">Table 1</tableCitation>
).
<taxonomicName id="4C094D32FFB0EC44752BFA62FE05FA11" box="[271,486,1485,1510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC44752BFA62FE05FA11" box="[271,486,1485,1510]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
differs from all by its unique coloration pattern consisting of black with white or whitish blue blotches in axilla, groin, anterior and posterior surfaces of the thighs and concealed surfaces of the tibia and by its broad lateral fringes on fingers and toes. Furthermore, the new species differs from
<taxonomicName id="4C094D32FFB0EC4471A2F9B5FEA5F9AD" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="altamazonicus">
<emphasis id="B97DEAA3FFB0EC4471A2F9B5FEA5F9AD" italics="true" pageId="3" pageNumber="33">P. altamazonicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D32FFB0EC4475A7F9EEFD9EF9AD" box="[387,637,1601,1626]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="ventrimarmoratus">
<emphasis id="B97DEAA3FFB0EC4475A7F9EEFD9EF9AD" box="[387,637,1601,1626]" italics="true" pageId="3" pageNumber="33">P. ventrimarmoratus</emphasis>
</taxonomicName>
by having a distinct tympanic annulus and membrane, which are absent in the latter two species.
<taxonomicName id="4C094D32FFB0EC44762BF9C7FD05F976" box="[527,742,1640,1665]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC44762BF9C7FD05F976" box="[527,742,1640,1665]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C094D32FFB0EC4477A6F9C7FBF2F976" box="[898,1041,1640,1665]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB0EC4477A6F9C7FBF2F976" box="[898,1041,1640,1665]" italics="true" pageId="3" pageNumber="33">Pristimantis</emphasis>
</taxonomicName>
sp. (currently being described by Lehr &amp; von May, in press) in having a tympanic membrane and males with nuptial pads, both characters are absent in
<taxonomicName id="4C094D32FFB0EC44752DF91AFE7AF939" box="[265,409,1717,1742]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB0EC44752DF91AFE7AF939" box="[265,409,1717,1742]" italics="true" pageId="3" pageNumber="33">Pristimantis</emphasis>
</taxonomicName>
sp.
<taxonomicName id="4C094D32FFB0EC4475EEF91AFD41F939" box="[458,674,1717,1742]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC4475EEF91AFD41F939" box="[458,674,1717,1742]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C094D32FFB0EC447764F91AFC06F939" box="[832,997,1717,1742]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="diadematus">
<emphasis id="B97DEAA3FFB0EC447764F91AFC06F939" box="[832,997,1717,1742]" italics="true" pageId="3" pageNumber="33">P. diadematus</emphasis>
</taxonomicName>
in having the venter weakly areolate (smooth in
<taxonomicName id="4C094D32FFB0EC44753BF973FE26F902" box="[287,453,1756,1781]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="diadematus">
<emphasis id="B97DEAA3FFB0EC44753BF973FE26F902" box="[287,453,1756,1781]" italics="true" pageId="3" pageNumber="33">P. diadematus</emphasis>
</taxonomicName>
) and by being smaller.
<taxonomicName id="4C094D32FFB0EC4476C4F973FC59F902" box="[736,954,1756,1781]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC4476C4F973FC59F902" box="[736,954,1756,1781]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C094D32FFB0EC44707DF973FAEDF902" box="[1113,1294,1756,1781]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="eurydactylus">
<emphasis id="B97DEAA3FFB0EC44707DF973FAEDF902" box="[1113,1294,1756,1781]" italics="true" pageId="3" pageNumber="33">P. eurydactylus</emphasis>
</taxonomicName>
in having a shagreen dorsum with minute scattered tubercles (tuberculate in
<taxonomicName id="4C094D32FFB0EC4477B5F8ADFBA6F8EC" box="[913,1093,1794,1819]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="eurydactylus">
<emphasis id="B97DEAA3FFB0EC4477B5F8ADFBA6F8EC" box="[913,1093,1794,1819]" italics="true" pageId="3" pageNumber="33">P. eurydactylus</emphasis>
</taxonomicName>
) and by being smaller.
<tableCitation id="C68B030AFFB0EC44717FF8ADFF45F8B4" captionStart="TABLE 1" captionStartId="5.[151,240,151,175]" captionTargetBox="[151,1436,241,1557]" captionTargetPageId="5" captionText="TABLE 1. Selected characters (+ = character present; — = character absent) and character conditions among selected" httpUri="http://table.plazi.org/id/DF766639FFB6EC4274B3FF38FA7FFF58" pageId="3" pageNumber="33" tableUuid="DF766639FFB6EC4274B3FF38FA7FFF58">Table 1</tableCitation>
presents a detailed comparison between
<taxonomicName id="4C094D32FFB0EC4476A1F886FCBFF8B5" box="[645,860,1833,1858]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB0EC4476A1F886FCBFF8B5" box="[645,860,1833,1858]" italics="true" pageId="3" pageNumber="33">Pristimantis orcus</emphasis>
</taxonomicName>
and the above-mentioned species.
</paragraph>
<paragraph id="8BB636B1FFB0EC4374E1F8E0FB41FCEB" blockId="3.[151,1437,1291,2013]" lastBlockId="4.[151,1437,152,796]" lastPageId="4" lastPageNumber="34" pageId="3" pageNumber="33">
Four other species of
<taxonomicName id="4C094D32FFB0EC4475FAF8FFFD98F89E" box="[478,635,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB0EC4475FAF8FFFD98F89E" box="[478,635,1872,1897]" italics="true" pageId="3" pageNumber="33">Pristimantis</emphasis>
</taxonomicName>
(
<emphasis id="B97DEAA3FFB0EC4476AAF8FFFBB4F89E" box="[654,1111,1872,1897]" italics="true" pageId="3" pageNumber="33">
<taxonomicName id="4C094D32FFB0EC4476AAF8FFFCEFF89E" box="[654,780,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="carvalhoi">carvalhoi</taxonomicName>
,
<taxonomicName id="4C094D32FFB0EC44773EF8FFFC39F89E" box="[794,986,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="croceoinguinis">croceoinguinis</taxonomicName>
,
<taxonomicName id="4C094D32FFB0EC4477CCF8FFFBB4F89E" box="[1000,1111,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="imitatrix">imitatrix</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C094D32FFB0EC4470BBF8FFFAE1F89E" box="[1183,1282,1872,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="martiae">
<emphasis id="B97DEAA3FFB0EC4470BBF8FFFAE1F89E" box="[1183,1282,1872,1897]" italics="true" pageId="3" pageNumber="33">martiae</emphasis>
</taxonomicName>
) are widely distributed in the Amazonian lowlands, but can be easily distinguished from
<emphasis id="B97DEAA3FFB0EC447015F8D9FB6FF878" box="[1073,1164,1910,1935]" italics="true" pageId="3" pageNumber="33">P. o rc u s</emphasis>
in lacking a tympanic annulus and tympanic membrane (present in
<emphasis id="B97DEAA3FFB0EC447688F832FCE5F841" box="[684,774,1949,1974]" italics="true" pageId="3" pageNumber="33">P. o rc u s</emphasis>
) and males lacking nuptial pads (present). Furthermore
<taxonomicName id="4C094D32FFB0EC4474B3F86BFE7DF82A" box="[151,414,1988,2013]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="33" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B97DEAA3FFB0EC4474B3F86BFE7DF82A" box="[151,414,1988,2013]" italics="true" pageId="3" pageNumber="33">Pristimantis carvalhoi</emphasis>
</taxonomicName>
has the dorsum with conical tubercles (absent), fingers and toes without lateral fringes (present), and in life the groin yellow (black with white blueish blotches), and the venter creamy tan (dark brown and tan mottled).
<taxonomicName id="4C094D32FFB7EC4375E6FF10FCF3FF2F" box="[450,784,191,216]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="croceoinguinis">
<emphasis id="B97DEAA3FFB7EC4375E6FF10FCF3FF2F" box="[450,784,191,216]" italics="true" pageId="4" pageNumber="34">Pristimantis croceoinguinis</emphasis>
</taxonomicName>
lacks lateral fringes on fingers and toes (present in
<taxonomicName id="4C094D32FFB7EC4371A2FF10FF34FF09" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB7EC4371A2FF10FF34FF09" italics="true" pageId="4" pageNumber="34">P. orcus</emphasis>
</taxonomicName>
), and has in life a yellow to orange spot in the groin (black with white blueish blotches), and the venter gray with white flecks (dark brown and tan mottled).
<taxonomicName id="4C094D32FFB7EC437705FEA3FBC7FED2" box="[801,1060,268,293]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="imitatrix">
<emphasis id="B97DEAA3FFB7EC437705FEA3FBC7FED2" box="[801,1060,268,293]" italics="true" pageId="4" pageNumber="34">Pristimantis imitatrix</emphasis>
</taxonomicName>
lacks dentigerous processes of vomers (present), has in life the groin mottled black and white (black with white blueish blotches), and the venter pale blueish gray with dark brown or black flecks or reticulations (dark brown and tan mottled), and is much smaller (SVL to
<quantity id="4CF19B54FFB7EC4375E5FE2FFDD1FE6D" box="[449,562,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="4" pageNumber="34" unit="mm" value="20.2">20.2 mm</quantity>
in
<taxonomicName id="4C094D32FFB7EC43767AFE2FFD0CFE6E" box="[606,751,384,409]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="imitatrix">
<emphasis id="B97DEAA3FFB7EC43767AFE2FFD0CFE6E" box="[606,751,384,409]" italics="true" pageId="4" pageNumber="34">P. imitatrix</emphasis>
</taxonomicName>
[
<bibRefCitation id="EF984B40FFB7EC437724FE2FFC3DFE6D" author="Duellman" box="[768,990,384,410]" pageId="4" pageNumber="39" refString="Duellman, W. E. (1978) Three new species of Eleutherodactylus from Amazonian Peru (Amphibia: Anura: Leptodactylidae). Herpetologica, 34, 264 - 270." type="journal article" year="1978" yearSuffix="b">Duellman 1978b</bibRefCitation>
] vs. SVL to
<quantity id="4CF19B54FFB7EC4370A0FE2FFB15FE6D" box="[1156,1270,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="4" pageNumber="34" unit="mm" value="36.5">36.5 mm</quantity>
in
<emphasis id="B97DEAA3FFB7EC437106FE2FFA65FE6E" box="[1314,1414,384,409]" italics="true" pageId="4" pageNumber="34">P. o rc u s</emphasis>
).
<taxonomicName id="4C094D32FFB7EC4374B3FE08FE6CFE37" box="[151,399,423,448]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="martiae">
<emphasis id="B97DEAA3FFB7EC4374B3FE08FE6CFE37" box="[151,399,423,448]" italics="true" pageId="4" pageNumber="34">Pristimantis martiae</emphasis>
</taxonomicName>
has fingers and toes with narrow lateral fringes (broad), and in life the venter grayish brown with small white to pale orange flecks (black with white blueish blotches). Three other species of
<taxonomicName id="4C094D32FFB7EC4374B3FE5BFEC5FDFA" box="[151,294,500,525]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB7EC4374B3FE5BFEC5FDFA" box="[151,294,500,525]" italics="true" pageId="4" pageNumber="34">Pristimantis</emphasis>
</taxonomicName>
(
<emphasis id="B97DEAA3FFB7EC43751CFE5BFDE4FDFA" box="[312,519,500,525]" italics="true" pageId="4" pageNumber="34">
<taxonomicName id="4C094D32FFB7EC43751CFE5BFE51FDFA" box="[312,434,500,525]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="ockendeni">ockendeni</taxonomicName>
,
<taxonomicName id="4C094D32FFB7EC43759AFE5BFDE7FDFA" box="[446,516,500,525]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="toftae">toftae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C094D32FFB7EC437666FE5BFD51FDFA" box="[578,690,500,525]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B97DEAA3FFB7EC437666FE5BFD51FDFA" box="[578,690,500,525]" italics="true" pageId="4" pageNumber="34">variabilis</emphasis>
</taxonomicName>
) from the Amazonian lowlands share with
<emphasis id="B97DEAA3FFB7EC437090FE5BFAEDFDFA" box="[1204,1294,500,525]" italics="true" pageId="4" pageNumber="34">P. o rc u s</emphasis>
a tympanic annulus and tympanic membrane, but males of these species lack nuptial pads (present). Furthermore
<taxonomicName id="4C094D32FFB7EC4374B3FDEEFE4EFDAD" box="[151,429,577,602]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="ockendeni">
<emphasis id="B97DEAA3FFB7EC4374B3FDEEFE4EFDAD" box="[151,429,577,602]" italics="true" pageId="4" pageNumber="34">Pristimantis ockendeni</emphasis>
</taxonomicName>
usually has dorsolateral folds (absent), fingers and toes with narrow lateral fringes (broad), and the scapular with a H- or W-shaped fold (absent).
<taxonomicName id="4C094D32FFB7EC4377B1FDC7FB96FD76" box="[917,1141,616,641]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="toftae">
<emphasis id="B97DEAA3FFB7EC4377B1FDC7FB96FD76" box="[917,1141,616,641]" italics="true" pageId="4" pageNumber="34">Pristimantis toftae</emphasis>
</taxonomicName>
has the upper lip with a white margin (absent), in life the groin bears a yellow to orange spot (black with white blueish blotches), and the venter is pale cream (dark brown and tan mottled).
<taxonomicName id="4C094D32FFB7EC437700FD1AFBCEFD39" box="[804,1069,693,718]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B97DEAA3FFB7EC437700FD1AFBCEFD39" box="[804,1069,693,718]" italics="true" pageId="4" pageNumber="34">Pristimantis variabilis</emphasis>
</taxonomicName>
lacks lateral fringes on fingers and toes (present), has in life the groin with a yellow fleck bordered with black (black with white blueish blotches), and the venter cream with brown or gray flecks (dark brown and tan mottled).
</paragraph>
<caption id="DF766639FFB7EC4374B3F8D0FCE3F84E" httpUri="https://zenodo.org/record/185537/files/figure.png" pageId="4" pageNumber="34" targetBox="[181,1373,882,1880]" targetPageId="4">
<paragraph id="8BB636B1FFB7EC4374B3F8D0FCE3F84E" blockId="4.[151,1436,1919,1977]" pageId="4" pageNumber="34">
<emphasis id="B97DEAA3FFB7EC4374B3F8D0FEF9F860" bold="true" box="[151,282,1919,1943]" pageId="4" pageNumber="34">FIGURE 3.</emphasis>
Dorsal (A) and lateral (B) views of head and ventral views of hand (C) and foot (D) of
<taxonomicName id="4C094D32FFB7EC4370F2F82FFA7FF860" box="[1238,1436,1920,1943]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="34" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB7EC4370F2F82FFA7FF860" box="[1238,1436,1920,1943]" italics="true" pageId="4" pageNumber="34">Pristimantis orcus</emphasis>
</taxonomicName>
(MUSM 27435). Scale bar = 5 mm. Drawings by E. Lehr.
</paragraph>
</caption>
<caption id="DF766639FFB6EC4274B3FF38FA7FFF58" ID-Table-UUID="DF766639FFB6EC4274B3FF38FA7FFF58" box="[151,1436,151,175]" httpUri="http://table.plazi.org/id/DF766639FFB6EC4274B3FF38FA7FFF58" pageId="5" pageNumber="35" targetBox="[151,1436,241,1557]" targetIsTable="true" targetPageId="5">
<paragraph id="8BB636B1FFB6EC4274B3FF38FA7FFF58" blockId="5.[151,1436,151,213]" box="[151,1436,151,175]" pageId="5" pageNumber="35">
<emphasis id="B97DEAA3FFB6EC4274B3FF38FEE8FF58" bold="true" box="[151,267,151,175]" pageId="5" pageNumber="35">TABLE 1.</emphasis>
Selected characters (+ = character present; — = character absent) and character conditions among selected
</paragraph>
</caption>
<paragraph id="8BB636B1FFB6EC4274B3FF11FD90FF22" blockId="5.[151,1436,151,213]" box="[151,627,189,213]" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC4274B3FF11FEFAFF22" box="[151,281,190,213]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFB6EC4274B3FF11FEFAFF22" box="[151,281,190,213]" italics="true" pageId="5" pageNumber="35">Pristimantis</emphasis>
</taxonomicName>
from the Amazonian Lowlands.
</paragraph>
</subSubSection>
<paragraph id="8BB636B1FFB6EC4274BBFF5EFAD9FAB2" pageId="5" pageNumber="35">
<table id="F909C411FFB613B874B3FF5EFA7FF9E2" box="[151,1436,241,1557]" gridcols="7" gridrows="11" pageId="5" pageNumber="35">
<tr id="353934F3FFB613B874B3FF5EFA7FFE64" box="[151,1436,241,403]" gridrow="0" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FF5EFEDDFE64" box="[151,318,241,403]" gridcol="0" gridrow="0" pageId="5" pageNumber="35">Characters and source</th>
<th id="76E85D8FFFB613B8757EFF5EFE07FE64" box="[346,484,241,403]" gridcol="1" gridrow="0" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC42757EFF5EFE6AFEDC" authority="This" authorityName="This" box="[346,404,241,299]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB6EC42757EFF5EFE77FEFF" box="[346,404,241,264]" italics="true" pageId="5" pageNumber="35">orcus</emphasis>
This
</taxonomicName>
paper
</th>
<th id="76E85D8FFFB613B875DEFF5EFD4DFE64" box="[506,686,241,403]" gridcol="2" gridrow="0" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC4275DEFF5EFD69FEDC" authority="Lynch 1980" authorityName="Lynch" authorityYear="1980" box="[506,660,241,299]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="altamazonicus">
<emphasis id="B97DEAA3FFB6EC4275DEFF5EFD77FEFF" box="[506,660,241,264]" italics="true" pageId="5" pageNumber="35">altamazonicus</emphasis>
(Lynch 1980)
</taxonomicName>
</th>
<th id="76E85D8FFFB613B876E7FF5EFCAEFE64" box="[707,845,241,403]" gridcol="3" gridrow="0" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC4276E7FF5EFCE2FEBA" authority="Lynch 1980" authorityName="Lynch" authorityYear="1980" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="diadematus">
<emphasis id="B97DEAA3FFB6EC4276E7FF5EFCDDFEFF" box="[707,830,241,264]" italics="true" pageId="5" pageNumber="35">diadematus</emphasis>
(Lynch 1980)
</taxonomicName>
</th>
<th id="76E85D8FFFB613B87744FF5EFBE4FE64" box="[864,1031,241,403]" gridcol="4" gridrow="0" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC427744FF5EFC0BFEFF" box="[864,1000,241,264]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="eurydactylus">
<emphasis id="B97DEAA3FFB6EC427744FF5EFC0BFEFF" box="[864,1000,241,264]" italics="true" pageId="5" pageNumber="35">eurydactylus</emphasis>
</taxonomicName>
(Hedges &amp; Schlüter 1992; Lehr and von May, in press)
</th>
<th id="76E85D8FFFB613B8700CFF5EFB09FE64" box="[1064,1258,241,403]" gridcol="5" gridrow="0" pageId="5" pageNumber="35">
<taxonomicName id="4C094D32FFB6EC42700CFF5EFB36FEBA" authority="Lynch &amp; Duellman 1980" authorityName="Lynch &amp; Duellman" authorityYear="1980" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="ventrimarmoratus">
<emphasis id="B97DEAA3FFB6EC42700CFF5EFB0BFEFF" box="[1064,1256,241,264]" italics="true" pageId="5" pageNumber="35">ventrimarmoratus</emphasis>
(Lynch &amp; Duellman 1980)
</taxonomicName>
</th>
<th id="76E85D8FFFB613B87124FF5EFA7FFE64" box="[1280,1436,241,403]" gridcol="6" gridrow="0" pageId="5" pageNumber="35">species (Lehr &amp; von May, in press)</th>
</tr>
<tr id="353934F3FFB613B874B3FE09FA7FFE4A" box="[151,1436,422,445]" gridrow="1" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FE09FEDDFE4A" box="[151,318,422,445]" gridcol="0" gridrow="1" pageId="5" pageNumber="35">SVL (mm)</th>
<td id="76E85D8FFFB613B8757EFE09FE07FE4A" box="[346,484,422,445]" gridcol="1" gridrow="1" pageId="5" pageNumber="35">20.025.1</td>
<td id="76E85D8FFFB613B875DEFE09FD4DFE4A" box="[506,686,422,445]" gridcol="2" gridrow="1" pageId="5" pageNumber="35">14.423.1</td>
<td id="76E85D8FFFB613B876E7FE09FCAEFE4A" box="[707,845,422,445]" gridcol="3" gridrow="1" pageId="5" pageNumber="35">20.027.4</td>
<td id="76E85D8FFFB613B87744FE09FBE4FE4A" box="[864,1031,422,445]" gridcol="4" gridrow="1" pageId="5" pageNumber="35">18.231.8</td>
<td id="76E85D8FFFB613B8700CFE09FB09FE4A" box="[1064,1258,422,445]" gridcol="5" gridrow="1" pageId="5" pageNumber="35">17.825.5</td>
<td id="76E85D8FFFB613B87124FE09FA7FFE4A" box="[1280,1436,422,445]" gridcol="6" gridrow="1" pageId="5" pageNumber="35">22.824.4</td>
</tr>
<tr id="353934F3FFB613B874B3FE7EFA7FFD62" box="[151,1436,465,661]" gridrow="2" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FE7EFEDDFD62" box="[151,318,465,661]" gridcol="0" gridrow="2" pageId="5" pageNumber="35">Dorsal skin</th>
<td id="76E85D8FFFB613B8757EFE7EFE07FD62" box="[346,484,465,661]" gridcol="1" gridrow="2" pageId="5" pageNumber="35">finely shagreen with scattered minute tubercles</td>
<td id="76E85D8FFFB613B875DEFE7EFD4DFD62" box="[506,686,465,661]" gridcol="2" gridrow="2" pageId="5" pageNumber="35">shagreen</td>
<td id="76E85D8FFFB613B876E7FE7EFCAEFD62" box="[707,845,465,661]" gridcol="3" gridrow="2" pageId="5" pageNumber="35">shagreen</td>
<td id="76E85D8FFFB613B87744FE7EFBE4FD62" box="[864,1031,465,661]" gridcol="4" gridrow="2" pageId="5" pageNumber="35">tuberculate</td>
<td id="76E85D8FFFB613B8700CFE7EFB09FD62" box="[1064,1258,465,661]" gridcol="5" gridrow="2" pageId="5" pageNumber="35">shagreen</td>
<td id="76E85D8FFFB613B87124FE7EFA7FFD62" box="[1280,1436,465,661]" gridcol="6" gridrow="2" pageId="5" pageNumber="35">shagreen with scattered minute tubercles</td>
</tr>
<tr id="353934F3FFB613B874B3FD06FA7FFD14" box="[151,1436,681,739]" gridrow="3" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FD06FEDDFD14" box="[151,318,681,739]" gridcol="0" gridrow="3" pageId="5" pageNumber="35">Ventral skin</th>
<td id="76E85D8FFFB613B8757EFD06FE07FD14" box="[346,484,681,739]" gridcol="1" gridrow="3" pageId="5" pageNumber="35">weakly areolate</td>
<td id="76E85D8FFFB613B875DEFD06FD4DFD14" box="[506,686,681,739]" gridcol="2" gridrow="3" pageId="5" pageNumber="35">coarsely areolate</td>
<td id="76E85D8FFFB613B876E7FD06FCAEFD14" box="[707,845,681,739]" gridcol="3" gridrow="3" pageId="5" pageNumber="35">smooth</td>
<td id="76E85D8FFFB613B87744FD06FBE4FD14" box="[864,1031,681,739]" gridcol="4" gridrow="3" pageId="5" pageNumber="35">weakly areolate</td>
<td id="76E85D8FFFB613B8700CFD06FB09FD14" box="[1064,1258,681,739]" gridcol="5" gridrow="3" pageId="5" pageNumber="35">coarsely areolate</td>
<td id="76E85D8FFFB613B87124FD06FA7FFD14" box="[1280,1436,681,739]" gridcol="6" gridrow="3" pageId="5" pageNumber="35">areolate</td>
</tr>
<tr id="353934F3FFB613B874B3FD59FA7FFCC7" box="[151,1436,758,816]" gridrow="4" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FD59FEDDFCC7" box="[151,318,758,816]" gridcol="0" gridrow="4" pageId="5" pageNumber="35">Tympanic annulus</th>
<td id="76E85D8FFFB613B8757EFD59FE07FCC7" box="[346,484,758,816]" gridcol="1" gridrow="4" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B875DEFD59FD4DFCC7" box="[506,686,758,816]" gridcol="2" gridrow="4" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B876E7FD59FCAEFCC7" box="[707,845,758,816]" gridcol="3" gridrow="4" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B87744FD59FBE4FCC7" box="[864,1031,758,816]" gridcol="4" gridrow="4" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B8700CFD59FB09FCC7" box="[1064,1258,758,816]" gridcol="5" gridrow="4" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B87124FD59FA7FFCC7" box="[1280,1436,758,816]" gridcol="6" gridrow="4" pageId="5" pageNumber="35">+</td>
</tr>
<tr id="353934F3FFB613B874B3FCEBFA7FFC8A" box="[151,1436,836,893]" gridrow="5" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FCEBFEDDFC8A" box="[151,318,836,893]" gridcol="0" gridrow="5" pageId="5" pageNumber="35">Tympanic membrane</th>
<td id="76E85D8FFFB613B8757EFCEBFE07FC8A" box="[346,484,836,893]" gridcol="1" gridrow="5" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B875DEFCEBFD4DFC8A" box="[506,686,836,893]" gridcol="2" gridrow="5" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B876E7FCEBFCAEFC8A" box="[707,845,836,893]" gridcol="3" gridrow="5" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B87744FCEBFBE4FC8A" box="[864,1031,836,893]" gridcol="4" gridrow="5" pageId="5" pageNumber="35">+, prominent</td>
<td id="76E85D8FFFB613B8700CFCEBFB09FC8A" box="[1064,1258,836,893]" gridcol="5" gridrow="5" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B87124FCEBFA7FFC8A" box="[1280,1436,836,893]" gridcol="6" gridrow="5" pageId="5" pageNumber="35"></td>
</tr>
<tr id="353934F3FFB613B874B3FC3EFA7FFC5F" box="[151,1436,913,936]" gridrow="6" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FC3EFEDDFC5F" box="[151,318,913,936]" gridcol="0" gridrow="6" pageId="5" pageNumber="35">Nuptial pads</th>
<td id="76E85D8FFFB613B8757EFC3EFE07FC5F" box="[346,484,913,936]" gridcol="1" gridrow="6" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B875DEFC3EFD4DFC5F" box="[506,686,913,936]" gridcol="2" gridrow="6" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B876E7FC3EFCAEFC5F" box="[707,845,913,936]" gridcol="3" gridrow="6" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B87744FC3EFBE4FC5F" box="[864,1031,913,936]" gridcol="4" gridrow="6" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B8700CFC3EFB09FC5F" box="[1064,1258,913,936]" gridcol="5" gridrow="6" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B87124FC3EFA7FFC5F" box="[1280,1436,913,936]" gridcol="6" gridrow="6" pageId="5" pageNumber="35"></td>
</tr>
<tr id="353934F3FFB613B874B3FC13FA7FFBEF" box="[151,1436,956,1048]" gridrow="7" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FC13FEDDFBEF" box="[151,318,956,1048]" gridcol="0" gridrow="7" pageId="5" pageNumber="35">Lateral fringes on toes and fingers</th>
<td id="76E85D8FFFB613B8757EFC13FE07FBEF" box="[346,484,956,1048]" gridcol="1" gridrow="7" pageId="5" pageNumber="35">+, broad</td>
<td id="76E85D8FFFB613B875DEFC13FD4DFBEF" box="[506,686,956,1048]" gridcol="2" gridrow="7" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B876E7FC13FCAEFBEF" box="[707,845,956,1048]" gridcol="3" gridrow="7" pageId="5" pageNumber="35">+, narrow</td>
<td id="76E85D8FFFB613B87744FC13FBE4FBEF" box="[864,1031,956,1048]" gridcol="4" gridrow="7" pageId="5" pageNumber="35">+, narrow</td>
<td id="76E85D8FFFB613B8700CFC13FB09FBEF" box="[1064,1258,956,1048]" gridcol="5" gridrow="7" pageId="5" pageNumber="35">+, narrow</td>
<td id="76E85D8FFFB613B87124FC13FA7FFBEF" box="[1280,1436,956,1048]" gridcol="6" gridrow="7" pageId="5" pageNumber="35">+, narrow</td>
</tr>
<tr id="353934F3FFB613B874B3FB83FA7FFBB4" box="[151,1436,1068,1091]" gridrow="8" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FB83FEDDFBB4" box="[151,318,1068,1091]" gridcol="0" gridrow="8" pageId="5" pageNumber="35">Toe webbing</th>
<td id="76E85D8FFFB613B8757EFB83FE07FBB4" box="[346,484,1068,1091]" gridcol="1" gridrow="8" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B875DEFB83FD4DFBB4" box="[506,686,1068,1091]" gridcol="2" gridrow="8" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B876E7FB83FCAEFBB4" box="[707,845,1068,1091]" gridcol="3" gridrow="8" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B87744FB83FBE4FBB4" box="[864,1031,1068,1091]" gridcol="4" gridrow="8" pageId="5" pageNumber="35"></td>
<td id="76E85D8FFFB613B8700CFB83FB09FBB4" box="[1064,1258,1068,1091]" gridcol="5" gridrow="8" pageId="5" pageNumber="35">+</td>
<td id="76E85D8FFFB613B87124FB83FA7FFBB4" box="[1280,1436,1068,1091]" gridcol="6" gridrow="8" pageId="5" pageNumber="35">+</td>
</tr>
<tr id="353934F3FFB613B874B3FBF9FA7FFB0F" box="[151,1436,1110,1272]" gridrow="9" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FBF9FEDDFB0F" box="[151,318,1110,1272]" gridcol="0" gridrow="9" pageId="5" pageNumber="35">Life coloration on chest and belly</th>
<td id="76E85D8FFFB613B8757EFBF9FE07FB0F" box="[346,484,1110,1272]" gridcol="1" gridrow="9" pageId="5" pageNumber="35">dark brown and tan mottled</td>
<td id="76E85D8FFFB613B875DEFBF9FD4DFB0F" box="[506,686,1110,1272]" gridcol="2" gridrow="9" pageId="5" pageNumber="35">gray or grayish brown, with or without small white flecks</td>
<td id="76E85D8FFFB613B876E7FBF9FCAEFB0F" box="[707,845,1110,1272]" gridcol="3" gridrow="9" pageId="5" pageNumber="35">creamy white with a greenish tint posteriorly</td>
<td id="76E85D8FFFB613B87744FBF9FBE4FB0F" box="[864,1031,1110,1272]" gridcol="4" gridrow="9" pageId="5" pageNumber="35">in ethanol: venter is white with or without brown flecks or small spots</td>
<td id="76E85D8FFFB613B8700CFBF9FB09FB0F" box="[1064,1258,1110,1272]" gridcol="5" gridrow="9" pageId="5" pageNumber="35">white with bold black mottling</td>
<td id="76E85D8FFFB613B87124FBF9FA7FFB0F" box="[1280,1436,1110,1272]" gridcol="6" gridrow="9" pageId="5" pageNumber="35">dark gray with white mottling</td>
</tr>
<tr id="353934F3FFB613B874B3FAA3FA7FF9E2" box="[151,1436,1292,1557]" gridrow="10" pageId="5" pageNumber="35">
<th id="76E85D8FFFB613B874B3FAA3FEDDF9E2" box="[151,318,1292,1557]" gridcol="0" gridrow="10" pageId="5" pageNumber="35">Groin coloration</th>
<td id="76E85D8FFFB613B8757EFAA3FE07F9E2" box="[346,484,1292,1557]" gridcol="1" gridrow="10" pageId="5" pageNumber="35">white or whitish blue and black</td>
<td id="76E85D8FFFB613B875DEFAA3FD4DF9E2" box="[506,686,1292,1557]" gridcol="2" gridrow="10" pageId="5" pageNumber="35">red or salmon and black</td>
<td id="76E85D8FFFB613B876E7FAA3FCAEF9E2" box="[707,845,1292,1557]" gridcol="3" gridrow="10" pageId="5" pageNumber="35">bluish white, yellowish tan, pink, or pale green with diagonal dark brown bars</td>
<td id="76E85D8FFFB613B87744FAA3FBE4F9E2" box="[864,1031,1292,1557]" gridcol="4" gridrow="10" pageId="5" pageNumber="35">in ethanol: tan and brown</td>
<td id="76E85D8FFFB613B8700CFAA3FB09F9E2" box="[1064,1258,1292,1557]" gridcol="5" gridrow="10" pageId="5" pageNumber="35">creamy white with brown marks</td>
<td id="76E85D8FFFB613B87124FAA3FA7FF9E2" box="[1280,1436,1292,1557]" gridcol="6" gridrow="10" pageId="5" pageNumber="35">yellow and black</td>
</tr>
</table>
</paragraph>
<subSubSection id="C313653AFFB6EC4174E1F9CFFAB0F97C" lastPageId="6" lastPageNumber="36" pageId="5" pageNumber="35" type="description">
<paragraph id="8BB636B1FFB6EC4274E1F9CFFE41F820" blockId="5.[151,1437,1632,2007]" pageId="5" pageNumber="35">
<emphasis id="B97DEAA3FFB6EC4274E1F9CFFDC0F98D" bold="true" box="[197,547,1632,1658]" pageId="5" pageNumber="35">
Description of the
<typeStatus id="54B28813FFB6EC42758BF9CFFDFDF98D" box="[431,542,1632,1658]" pageId="5" pageNumber="35" type="holotype">holotype</typeStatus>
.
</emphasis>
Head narrower than body, slightly longer than wide; head width 37.0% of SVL; head length 38.6% of SVL; snout moderate, acutely rounded in dorsal view, rounded in lateral view, (
<figureCitation id="13322A34FFB6EC427484F901FF17F93F" box="[160,244,1710,1736]" captionStart="FIGURE 3" captionStartId="4.[151,255,1919,1943]" captionTargetBox="[181,1373,882,1880]" captionTargetId="figure@4.[151,1436,852,1896]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and lateral (B) views of head and ventral views of hand (C) and foot (D) of Pristimantis orcus (MUSM 27435). Scale bar = 5 mm. Drawings by E. Lehr." httpUri="https://zenodo.org/record/185537/files/figure.png" pageId="5" pageNumber="35">Figs. 3</figureCitation>
A, B), eyenostril distance larger than eye diameter; nostrils protuberant, directed dorsolaterally; canthus rostralis slightly concave in dorsal view, rounded in profile; loreal region slightly concave; lips flared; upper eyelid with minute tubercles; width of upper eyelid narrower than IOD (upper eyelid width 86.8% of IOD); supratympanic fold short, broad, concealing posterior margin of tympanum; tympanic membrane and tympanic annulus present, prominent, about half the size of ED (54.3%), separated from eye by half its diameter; postrictal tubercles absent. Choanae large, ovoid, not concealed by palatal shelf of maxilla; dentigerous processes of vomers prominent, oblique; tongue two times as long as wide, posteriorly notched, posterior one half free.
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FF37FAA6FE40" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
Skin on dorsum shagreen with scattered tubercles more dense on head, dorsolateral folds absent; skin on flanks areolate; skin on throat, chest, and belly weakly areolate; weak discoidal fold present, more prominent posteriorly; cloacal sheath short; large tubercles absent in cloacal region. Outer surface of ulnar with minute tubercles; palmar tubercles low, outer palmar tubercle bifid, ovoid, approximately three times the size of elongate, inner palmar tubercle; one low supernumerary tubercle at base of each finger, about half the size of subarticular tubercles; subarticular tubercles well defined, ovoid in dorsal view, conical in lateral view; fingers with lateral fringes broadest on base of fingers; Finger I shorter than Finger II; discs on digits of outer fingers broadly expanded, about twice the size of digit proximal to it on Fingers III and IV, truncate (
<figureCitation id="13322A34FFB5EC4170FFFE32FAC2FE40" box="[1243,1313,413,439]" captionStart="FIGURE 3" captionStartId="4.[151,255,1919,1943]" captionTargetBox="[181,1373,882,1880]" captionTargetId="figure@4.[151,1436,852,1896]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and lateral (B) views of head and ventral views of hand (C) and foot (D) of Pristimantis orcus (MUSM 27435). Scale bar = 5 mm. Drawings by E. Lehr." httpUri="https://zenodo.org/record/185537/files/figure.png" pageId="6" pageNumber="36">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FE6DFEE8FCA8" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
<collectingCountry id="F31E7621FFB5EC4174E1FE6DFEE3FE2B" box="[197,256,450,476]" name="India" pageId="6" pageNumber="36">Hind</collectingCountry>
limbs relatively short, tibia length 44.1% of SVL; foot length 41.4% of SVL; upper surface of hind limbs shagreen with small, scattered tubercles; posterior and ventral surfaces of thighs weakly areolate; heel with minute tubercles and one slightly larger conical tubercle in its middle; outer surface of tarsus with minute tubercles; tarsal fold long, distinct, not contacting inner metatarsal tubercle, metatarsal tubercles low, inner metatarsal tubercle ovoid, about four times as large as elongate outer metatarsal tubercle; one plantar supernumerary tubercle on basis of each toe, about one third the size of subarticular tubercles; subarticular tubercles well defined, less prominent than those on fingers, ovoid in dorsal view, slightly conical in profile; toes with broad and thin lateral fringes, concealing sides of toes, basal webbing present between Toes IIV; discs on digits of toes broadly expanded, about twice the size of digit proximal to it on Toes IV and V, about as large as those on fingers, truncate; relative lengths of toes: 1 &lt;2 &lt;3 &lt;5 &lt;4; Toe V much longer than Toe III (
<figureCitation id="13322A34FFB5EC4174BBFCEAFF07FCA8" box="[159,228,837,863]" captionStart="FIGURE 3" captionStartId="4.[151,255,1919,1943]" captionTargetBox="[181,1373,882,1880]" captionTargetId="figure@4.[151,1436,852,1896]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and lateral (B) views of head and ventral views of hand (C) and foot (D) of Pristimantis orcus (MUSM 27435). Scale bar = 5 mm. Drawings by E. Lehr." httpUri="https://zenodo.org/record/185537/files/figure.png" pageId="6" pageNumber="36">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FCC3FD2CFC24" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
Measurements (in mm) of
<typeStatus id="54B28813FFB5EC4175DBFCC3FD89FC71" box="[511,618,876,902]" pageId="6" pageNumber="36" type="holotype">holotype</typeStatus>
: SVL 36.5; tibia length 16.1; foot length 15.1; head length 14.1; head width 13.5; eye diameter 3.5; tympanum diameter 1.9; interorbital distance 3.8; upper eyelid width 3.3; internarial distance 3.0; eyenostril distance 4.7.
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FC4FFE02FAFF" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
Coloration of
<typeStatus id="54B28813FFB5EC417552FC4FFE00FC0D" box="[374,483,992,1018]" pageId="6" pageNumber="36" type="holotype">holotype</typeStatus>
in preservative (
<figureCitation id="13322A34FFB5EC41769DFC4FFCE4FC0D" box="[697,775,992,1018]" captionStart="FIGURE 2" captionStartId="3.[151,255,1218,1242]" captionTargetBox="[157,1425,430,1194]" captionTargetId="figure@3.[151,1436,426,1200]" captionTargetPageId="3" captionText="FIGURE 2. Preserved Pristimantis orcus (holotype) in dorsal (A), and ventral (B) views. Photos by E. Lehr." httpUri="https://zenodo.org/record/185536/files/figure.png" pageId="6" pageNumber="36">Fig. 2</figureCitation>
): Dorsum grayish brown with irregularly shaped tan blotches on region between interorbital area to cloaca; each lower arm with one dark brown blotch; hind limbs grayish brown without bars, heels tan; Fingers I and II gray, Fingers III and IV grayish brown, toes grayish brown; head laterally grayish brown without canthal and sypratympanic stripes and without blotches on upper lip; upper half of tympanum grayish brown, lower half tan; flanks paler than dorsum, tan and brown mottled; axilla, groin, and anterior and posterior surfaces of thighs and concealed surfaces of tibia black with white blotches; throat, chest, and belly dark brown and tan mottled; thighs grayish brown with few tan flecks; hands and feet gray; iris dark gray.
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FABAFCCCFAA1" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
Coloration of
<typeStatus id="54B28813FFB5EC417548FABAFE30FAD8" box="[364,467,1301,1327]" pageId="6" pageNumber="36" type="holotype">holotype</typeStatus>
in life (
<figureCitation id="13322A34FFB5EC417616FABAFD9CFAD8" box="[562,639,1301,1327]" captionStart="FIGURE 1" captionStartId="2.[151,258,1596,1620]" captionTargetBox="[151,1434,351,1570]" captionTargetId="figure@2.[151,1436,351,1571]" captionTargetPageId="2" captionText="FIGURE 1. Left column: Female Pristimantis orcus (MUSM 27435, holotype, SVL 36.5 mm) in dorsolateral (A), lateral (B), and ventral (C) views; right column: male Pristimantis orcus (MUSM 25938, paratype, SVL 22.4 mm) in dorsolateral (D), lateral (E), and ventral (F) views. Photos A C by D. Rodríguez, photos D F by A. Catenazzi." httpUri="https://zenodo.org/record/185535/files/figure.png" pageId="6" pageNumber="36">Figs. 1</figureCitation>
AC): As described above except for white being whitish blue and for iris gold with cupper tint and fine black reticulations.
</paragraph>
<paragraph id="8BB636B1FFB5EC4174E1FACDFAB0F97C" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
<emphasis id="B97DEAA3FFB5EC4174E1FACDFEA5FA8B" bold="true" box="[197,326,1378,1404]" pageId="6" pageNumber="36">Variation.</emphasis>
In respect to morphology and coloration the
<typeStatus id="54B28813FFB5EC4177A6FACDFC1AFA8B" box="[898,1017,1378,1404]" pageId="6" pageNumber="36" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="54B28813FFB5EC4170C4FACDFAB2FA8B" box="[1248,1361,1378,1404]" pageId="6" pageNumber="36" type="holotype">holotype</typeStatus>
. Male specimens are smaller than females, lack vocal slits, and have tan nuptial pads on dorsal and medial surfaces of thumb. The ventral coloration tends to be paler in males than in females. In life one male (MUSM 25938,
<figureCitation id="13322A34FFB5EC4174B3FA79FF05FA07" box="[151,230,1494,1520]" captionStart="FIGURE 1" captionStartId="2.[151,258,1596,1620]" captionTargetBox="[151,1434,351,1570]" captionTargetId="figure@2.[151,1436,351,1571]" captionTargetPageId="2" captionText="FIGURE 1. Left column: Female Pristimantis orcus (MUSM 27435, holotype, SVL 36.5 mm) in dorsolateral (A), lateral (B), and ventral (C) views; right column: male Pristimantis orcus (MUSM 25938, paratype, SVL 22.4 mm) in dorsolateral (D), lateral (E), and ventral (F) views. Photos A C by D. Rodríguez, photos D F by A. Catenazzi." httpUri="https://zenodo.org/record/185535/files/figure.png" pageId="6" pageNumber="36">Figs. 1</figureCitation>
DF) is dorsally brown with tan and olive green blotches, axilla, groin, anterior surface of thighs, concealed surfaces of tibia and tarsus are black with whitish blue blotches, the iris is pale brown with dark brown reticulations and a dark brown vertical streak from pupil to lower half of iris. In life, the iris of one male (MUSM 27436) is pale brown with fine black reticulations with a black vertical streak through the pupil giving a cross-like appearance. See Tables 2 and 3 for measurements and proportions of the
<typeStatus id="54B28813FFB5EC4170EBF9DEFAE2F97C" box="[1231,1281,1649,1675]" pageId="6" pageNumber="36">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C313653AFFB5EC4174E2F938FA6EF8D1" pageId="6" pageNumber="36" type="distribution">
<paragraph id="8BB636B1FFB5EC4174E2F938FA6EF8D1" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
<emphasis id="B97DEAA3FFB5EC4174E2F938FDE0F946" bold="true" box="[198,515,1687,1713]" pageId="6" pageNumber="36">Distribution and ecology.</emphasis>
<taxonomicName id="4C094D32FFB5EC417628F937FD06F946" box="[524,741,1688,1713]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="36" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB5EC417628F937FD06F946" box="[524,741,1688,1713]" italics="true" pageId="6" pageNumber="36">Pristimantis orcus</emphasis>
</taxonomicName>
is known from three localities in Región Loreto and from one locality in San Martín (
<figureCitation id="13322A34FFB5EC4175CDF911FDD5F92F" box="[489,566,1726,1752]" captionStart="FIGURE 4" captionStartId="7.[151,255,1964,1988]" captionTargetBox="[176,1419,194,1938]" captionTargetId="figure@7.[176,1420,194,1939]" captionTargetPageId="7" captionText="FIGURE 4. Map illustrating collecting sites (indicated by circles) and type locality (indicated by a star) of the type series of Pristimantis orcus in northern Peru." httpUri="https://zenodo.org/record/185538/files/figure.png" pageId="6" pageNumber="36">Fig. 4</figureCitation>
). All specimens were found at night on the ground in primary (MUSM 27456) and secondary forests (MUSM 2743536, 27392). For description of the collecting site of one specimen (MUSM 25938) from near the Panguana stream in northern Loreto see Vriesdendorp
<emphasis id="B97DEAA3FFB5EC4170D1F8A3FAD3F8D2" box="[1269,1328,1804,1829]" italics="true" pageId="6" pageNumber="36">et al.</emphasis>
(2007).
</paragraph>
</subSubSection>
<subSubSection id="C313653AFFB5EC4F74E2F89DFDBDFC34" lastPageId="8" lastPageNumber="38" pageId="6" pageNumber="36" type="etymology">
<paragraph id="8BB636B1FFB5EC4174E2F89DFE6EF837" blockId="6.[151,1437,152,1984]" pageId="6" pageNumber="36">
<emphasis id="B97DEAA3FFB5EC4174E2F89DFEB0F8BB" bold="true" box="[198,339,1842,1868]" pageId="6" pageNumber="36">Etymology.</emphasis>
The specific name
<taxonomicName id="4C094D32FFB5EC417666F89CFD61F8BB" box="[578,642,1843,1868]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="36" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB5EC417666F89CFD61F8BB" box="[578,642,1843,1868]" italics="true" pageId="6" pageNumber="36">orcus</emphasis>
</taxonomicName>
is the Latin noun for underworld. The specific name is a noun in apposition and refers to the contrasting pattern of black and white in axilla, groin, anterior and posterior surfaces of thighs and concealed surfaces of tibia of preserved specimens, which remind us of the pattern in Orcas (
<taxonomicName id="4C094D32FFB5EC4174CFF809FE61F837" box="[235,386,1958,1984]" class="Mammalia" family="Delphinidae" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="36" phylum="Chordata" rank="family">Delphinidae</taxonomicName>
).
</paragraph>
<caption id="DF766639FFB4EC4074B3F803FDD1F81E" httpUri="https://zenodo.org/record/185538/files/figure.png" pageId="7" pageNumber="37" targetBox="[176,1419,194,1938]" targetPageId="7">
<paragraph id="8BB636B1FFB4EC4074B3F803FDD1F81E" blockId="7.[151,1436,1964,2025]" pageId="7" pageNumber="37">
<emphasis id="B97DEAA3FFB4EC4074B3F803FEFBF833" bold="true" box="[151,280,1964,1988]" pageId="7" pageNumber="37">FIGURE 4.</emphasis>
Map illustrating collecting sites (indicated by circles) and type locality (indicated by a star) of the type series of
<taxonomicName id="4C094D32FFB4EC407490F87DFE94F81E" box="[180,375,2002,2025]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="37" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFB4EC407490F87DFE94F81E" box="[180,375,2002,2025]" italics="true" pageId="7" pageNumber="37">Pristimantis orcus</emphasis>
</taxonomicName>
in northern Peru.
</paragraph>
</caption>
<paragraph id="8BB636B1FFBBEC4F74B3FF36FACBFCF0" pageId="8" pageNumber="38">
<table id="F909C411FFBB13B874B3FF36FABBFCF0" box="[151,1368,153,775]" gridcols="5" gridrows="14" pageId="8" pageNumber="38">
<tr id="353934F3FFBB13B874B3FF36FABBFF46" box="[151,1368,153,177]" gridrow="0" pageId="8" pageNumber="38" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76E85D8FFFBB13B874B3FF36FD15FF46" box="[151,758,153,177]" gridcol="0" gridrow="0" pageId="8" pageNumber="38">
<emphasis id="B97DEAA3FFBBEC4F74B3FF36FEEAFF46" bold="true" box="[151,265,153,177]" pageId="8" pageNumber="38">TABLE 2.</emphasis>
Measurements (in mm) of
<taxonomicName id="4C094D32FFBBEC4F7608FF35FD0CFF46" box="[556,751,154,177]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="38" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFBBEC4F7608FF35FD0CFF46" box="[556,751,154,177]" italics="true" pageId="8" pageNumber="38">Pristimantis orcus</emphasis>
</taxonomicName>
.
</th>
</tr>
<tr id="353934F3FFBB13B874B3FF62FABBFEF0" box="[151,1368,205,263]" gridrow="1" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FF62FD15FEF0" box="[151,758,205,263]" gridcol="0" gridrow="1" pageId="8" pageNumber="38">MUSM MUSM MUSM Character 27435 27456 22444</th>
<td id="76E85D8FFFBB13B87716FF62FC67FEF0" box="[818,900,205,263]" gridcol="1" gridrow="1" pageId="8" pageNumber="38">MUSM 27392</td>
<td id="76E85D8FFFBB13B877EAFF62FBFCFEF0" box="[974,1055,205,263]" gridcol="2" gridrow="1" pageId="8" pageNumber="38">MUSM 27436</td>
<td id="76E85D8FFFBB13B8704FFF62FB5FFEF0" box="[1131,1212,205,263]" gridcol="3" gridrow="1" pageId="8" pageNumber="38">MUSM 25938</td>
<td id="76E85D8FFFBB13B87123FF62FABBFEF0" box="[1287,1368,205,263]" gridcol="4" gridrow="1" pageId="8" pageNumber="38">MUSM 22437</td>
</tr>
<tr id="353934F3FFBB13B874B3FEB5FABBFEC6" box="[151,1368,282,305]" gridrow="2" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FEB5FD15FEC6" box="[151,758,282,305]" gridcol="0" gridrow="2" pageId="8" pageNumber="38">sex female female female</th>
<td id="76E85D8FFFBB13B87716FEB5FC67FEC6" box="[818,900,282,305]" gridcol="1" gridrow="2" pageId="8" pageNumber="38">male</td>
<td id="76E85D8FFFBB13B877EAFEB5FBFCFEC6" box="[974,1055,282,305]" gridcol="2" gridrow="2" pageId="8" pageNumber="38">male</td>
<td id="76E85D8FFFBB13B8704FFEB5FB5FFEC6" box="[1131,1212,282,305]" gridcol="3" gridrow="2" pageId="8" pageNumber="38">male</td>
<td id="76E85D8FFFBB13B87123FEB5FABBFEC6" box="[1287,1368,282,305]" gridcol="4" gridrow="2" pageId="8" pageNumber="38">male</td>
</tr>
<tr id="353934F3FFBB13B874B3FEEAFABBFEAB" box="[151,1368,325,348]" gridrow="3" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FEEAFD15FEAB" box="[151,758,325,348]" gridcol="0" gridrow="3" pageId="8" pageNumber="38">SVL 36.5 33.0 32.6</th>
<td id="76E85D8FFFBB13B87716FEEAFC67FEAB" box="[818,900,325,348]" gridcol="1" gridrow="3" pageId="8" pageNumber="38">25.1</td>
<td id="76E85D8FFFBB13B877EAFEEAFBFCFEAB" box="[974,1055,325,348]" gridcol="2" gridrow="3" pageId="8" pageNumber="38">24.0</td>
<td id="76E85D8FFFBB13B8704FFEEAFB5FFEAB" box="[1131,1212,325,348]" gridcol="3" gridrow="3" pageId="8" pageNumber="38">22.4</td>
<td id="76E85D8FFFBB13B87123FEEAFABBFEAB" box="[1287,1368,325,348]" gridcol="4" gridrow="3" pageId="8" pageNumber="38">20.0</td>
</tr>
<tr id="353934F3FFBB13B874B3FEDFFABBFE70" box="[151,1368,368,391]" gridrow="4" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FEDFFD15FE70" box="[151,758,368,391]" gridcol="0" gridrow="4" pageId="8" pageNumber="38">TL 16.1 15.9 15.0</th>
<td id="76E85D8FFFBB13B87716FEDFFC67FE70" box="[818,900,368,391]" gridcol="1" gridrow="4" pageId="8" pageNumber="38">12.6</td>
<td id="76E85D8FFFBB13B877EAFEDFFBFCFE70" box="[974,1055,368,391]" gridcol="2" gridrow="4" pageId="8" pageNumber="38">11.3</td>
<td id="76E85D8FFFBB13B8704FFEDFFB5FFE70" box="[1131,1212,368,391]" gridcol="3" gridrow="4" pageId="8" pageNumber="38">11.5</td>
<td id="76E85D8FFFBB13B87123FEDFFABBFE70" box="[1287,1368,368,391]" gridcol="4" gridrow="4" pageId="8" pageNumber="38">9.9</td>
</tr>
<tr id="353934F3FFBB13B874B3FE35FABBFE46" box="[151,1368,410,433]" gridrow="5" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FE35FD15FE46" box="[151,758,410,433]" gridcol="0" gridrow="5" pageId="8" pageNumber="38">FL 15.1 14.6 13.3</th>
<td id="76E85D8FFFBB13B87716FE35FC67FE46" box="[818,900,410,433]" gridcol="1" gridrow="5" pageId="8" pageNumber="38">10.6</td>
<td id="76E85D8FFFBB13B877EAFE35FBFCFE46" box="[974,1055,410,433]" gridcol="2" gridrow="5" pageId="8" pageNumber="38">10.1</td>
<td id="76E85D8FFFBB13B8704FFE35FB5FFE46" box="[1131,1212,410,433]" gridcol="3" gridrow="5" pageId="8" pageNumber="38">9.5</td>
<td id="76E85D8FFFBB13B87123FE35FABBFE46" box="[1287,1368,410,433]" gridcol="4" gridrow="5" pageId="8" pageNumber="38">8.1</td>
</tr>
<tr id="353934F3FFBB13B874B3FE6AFABBFE2B" box="[151,1368,453,476]" gridrow="6" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FE6AFD15FE2B" box="[151,758,453,476]" gridcol="0" gridrow="6" pageId="8" pageNumber="38">HL 14.1 13.2 12.7</th>
<td id="76E85D8FFFBB13B87716FE6AFC67FE2B" box="[818,900,453,476]" gridcol="1" gridrow="6" pageId="8" pageNumber="38">9.3</td>
<td id="76E85D8FFFBB13B877EAFE6AFBFCFE2B" box="[974,1055,453,476]" gridcol="2" gridrow="6" pageId="8" pageNumber="38">10.1</td>
<td id="76E85D8FFFBB13B8704FFE6AFB5FFE2B" box="[1131,1212,453,476]" gridcol="3" gridrow="6" pageId="8" pageNumber="38">8.8</td>
<td id="76E85D8FFFBB13B87123FE6AFABBFE2B" box="[1287,1368,453,476]" gridcol="4" gridrow="6" pageId="8" pageNumber="38">8.3</td>
</tr>
<tr id="353934F3FFBB13B874B3FE5FFABBFDF0" box="[151,1368,496,519]" gridrow="7" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FE5FFD15FDF0" box="[151,758,496,519]" gridcol="0" gridrow="7" pageId="8" pageNumber="38">HW 13.5 12.5 13.0</th>
<td id="76E85D8FFFBB13B87716FE5FFC67FDF0" box="[818,900,496,519]" gridcol="1" gridrow="7" pageId="8" pageNumber="38">9.2</td>
<td id="76E85D8FFFBB13B877EAFE5FFBFCFDF0" box="[974,1055,496,519]" gridcol="2" gridrow="7" pageId="8" pageNumber="38">8.6</td>
<td id="76E85D8FFFBB13B8704FFE5FFB5FFDF0" box="[1131,1212,496,519]" gridcol="3" gridrow="7" pageId="8" pageNumber="38">8.5</td>
<td id="76E85D8FFFBB13B87123FE5FFABBFDF0" box="[1287,1368,496,519]" gridcol="4" gridrow="7" pageId="8" pageNumber="38">7.8</td>
</tr>
<tr id="353934F3FFBB13B874B3FDB5FABBFDC6" box="[151,1368,538,561]" gridrow="8" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FDB5FD15FDC6" box="[151,758,538,561]" gridcol="0" gridrow="8" pageId="8" pageNumber="38">ED 3.5 3.6 3.4</th>
<td id="76E85D8FFFBB13B87716FDB5FC67FDC6" box="[818,900,538,561]" gridcol="1" gridrow="8" pageId="8" pageNumber="38">3.7</td>
<td id="76E85D8FFFBB13B877EAFDB5FBFCFDC6" box="[974,1055,538,561]" gridcol="2" gridrow="8" pageId="8" pageNumber="38">3.3</td>
<td id="76E85D8FFFBB13B8704FFDB5FB5FFDC6" box="[1131,1212,538,561]" gridcol="3" gridrow="8" pageId="8" pageNumber="38">3.0</td>
<td id="76E85D8FFFBB13B87123FDB5FABBFDC6" box="[1287,1368,538,561]" gridcol="4" gridrow="8" pageId="8" pageNumber="38">2.4</td>
</tr>
<tr id="353934F3FFBB13B874B3FDEAFABBFDAB" box="[151,1368,581,604]" gridrow="9" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FDEAFD15FDAB" box="[151,758,581,604]" gridcol="0" gridrow="9" pageId="8" pageNumber="38">TY 1.9 1.5 1.9</th>
<td id="76E85D8FFFBB13B87716FDEAFC67FDAB" box="[818,900,581,604]" gridcol="1" gridrow="9" pageId="8" pageNumber="38">1.5</td>
<td id="76E85D8FFFBB13B877EAFDEAFBFCFDAB" box="[974,1055,581,604]" gridcol="2" gridrow="9" pageId="8" pageNumber="38">1.3</td>
<td id="76E85D8FFFBB13B8704FFDEAFB5FFDAB" box="[1131,1212,581,604]" gridcol="3" gridrow="9" pageId="8" pageNumber="38">1.3</td>
<td id="76E85D8FFFBB13B87123FDEAFABBFDAB" box="[1287,1368,581,604]" gridcol="4" gridrow="9" pageId="8" pageNumber="38">1.1</td>
</tr>
<tr id="353934F3FFBB13B874B3FDDFFABBFD70" box="[151,1368,624,647]" gridrow="10" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FDDFFD15FD70" box="[151,758,624,647]" gridcol="0" gridrow="10" pageId="8" pageNumber="38">IOD 3.8 3.4 3.8</th>
<td id="76E85D8FFFBB13B87716FDDFFC67FD70" box="[818,900,624,647]" gridcol="1" gridrow="10" pageId="8" pageNumber="38">2.3</td>
<td id="76E85D8FFFBB13B877EAFDDFFBFCFD70" box="[974,1055,624,647]" gridcol="2" gridrow="10" pageId="8" pageNumber="38">3.1</td>
<td id="76E85D8FFFBB13B8704FFDDFFB5FFD70" box="[1131,1212,624,647]" gridcol="3" gridrow="10" pageId="8" pageNumber="38">2.5</td>
<td id="76E85D8FFFBB13B87123FDDFFABBFD70" box="[1287,1368,624,647]" gridcol="4" gridrow="10" pageId="8" pageNumber="38">2.4</td>
</tr>
<tr id="353934F3FFBB13B874B3FD35FABBFD46" box="[151,1368,666,689]" gridrow="11" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FD35FD15FD46" box="[151,758,666,689]" gridcol="0" gridrow="11" pageId="8" pageNumber="38">EW 3.3 3.1 3.3</th>
<td id="76E85D8FFFBB13B87716FD35FC67FD46" box="[818,900,666,689]" gridcol="1" gridrow="11" pageId="8" pageNumber="38">2.8</td>
<td id="76E85D8FFFBB13B877EAFD35FBFCFD46" box="[974,1055,666,689]" gridcol="2" gridrow="11" pageId="8" pageNumber="38">2.4</td>
<td id="76E85D8FFFBB13B8704FFD35FB5FFD46" box="[1131,1212,666,689]" gridcol="3" gridrow="11" pageId="8" pageNumber="38">1.9</td>
<td id="76E85D8FFFBB13B87123FD35FABBFD46" box="[1287,1368,666,689]" gridcol="4" gridrow="11" pageId="8" pageNumber="38">1.8</td>
</tr>
<tr id="353934F3FFBB13B874B3FD6AFABBFD2B" box="[151,1368,709,732]" gridrow="12" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FD6AFD15FD2B" box="[151,758,709,732]" gridcol="0" gridrow="12" pageId="8" pageNumber="38">IND 3.0 2.7 2.6</th>
<td id="76E85D8FFFBB13B87716FD6AFC67FD2B" box="[818,900,709,732]" gridcol="1" gridrow="12" pageId="8" pageNumber="38">2.2</td>
<td id="76E85D8FFFBB13B877EAFD6AFBFCFD2B" box="[974,1055,709,732]" gridcol="2" gridrow="12" pageId="8" pageNumber="38">2.1</td>
<td id="76E85D8FFFBB13B8704FFD6AFB5FFD2B" box="[1131,1212,709,732]" gridcol="3" gridrow="12" pageId="8" pageNumber="38">2.2</td>
<td id="76E85D8FFFBB13B87123FD6AFABBFD2B" box="[1287,1368,709,732]" gridcol="4" gridrow="12" pageId="8" pageNumber="38">1.8</td>
</tr>
<tr id="353934F3FFBB13B874B3FD5FFABBFCF0" box="[151,1368,752,775]" gridrow="13" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FD5FFD15FCF0" box="[151,758,752,775]" gridcol="0" gridrow="13" pageId="8" pageNumber="38">EN 4.7 4.2 3.8</th>
<td id="76E85D8FFFBB13B87716FD5FFC67FCF0" box="[818,900,752,775]" gridcol="1" gridrow="13" pageId="8" pageNumber="38">3.0</td>
<td id="76E85D8FFFBB13B877EAFD5FFBFCFCF0" box="[974,1055,752,775]" gridcol="2" gridrow="13" pageId="8" pageNumber="38">2.9</td>
<td id="76E85D8FFFBB13B8704FFD5FFB5FFCF0" box="[1131,1212,752,775]" gridcol="3" gridrow="13" pageId="8" pageNumber="38">2.7</td>
<td id="76E85D8FFFBB13B87123FD5FFABBFCF0" box="[1287,1368,752,775]" gridcol="4" gridrow="13" pageId="8" pageNumber="38">2.8</td>
</tr>
</table>
</paragraph>
<caption id="DF766639FFBBEC4F74B3FCFDFDCDFC78" ID-Table-UUID="DF766639FFBBEC4F74B3FCFDFDCDFC78" httpUri="http://table.plazi.org/id/DF766639FFBBEC4F74B3FCFDFDCDFC78" pageId="8" pageNumber="38" targetBox="[151,1436,982,1816]" targetIsTable="true" targetPageId="8">
<paragraph id="8BB636B1FFBBEC4F74B3FCFDFDCDFC78" blockId="8.[151,1436,850,963]" pageId="8" pageNumber="38">
<emphasis id="B97DEAA3FFBBEC4F74B3FCFDFEEFFC9D" bold="true" box="[151,268,850,874]" pageId="8" pageNumber="38">TABLE 3.</emphasis>
Measurements (in mm) and proportions of type series of
<emphasis id="B97DEAA3FFBBEC4F7759FCFDFBA8FC9E" box="[893,1099,850,873]" italics="true" pageId="8" pageNumber="38">
<taxonomicName id="4C094D32FFBBEC4F7759FCFDFBA6FC9E" box="[893,1093,850,873]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="38" phylum="Chordata" rank="species" species="orcus">Pristimantis orcus</taxonomicName>
;
</emphasis>
ranges followed by means and one standard deviation in parentheses.
</paragraph>
</caption>
<paragraph id="8BB636B1FFBBEC4F75BFFC03FDBDFC34" blockId="8.[151,1436,850,963]" box="[411,606,940,963]" pageId="8" pageNumber="38">
<taxonomicName id="4C094D32FFBBEC4F75BFFC03FDBDFC34" box="[411,606,940,963]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="38" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFBBEC4F75BFFC03FDBDFC34" box="[411,606,940,963]" italics="true" pageId="8" pageNumber="38">Pristimantis orcus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<paragraph id="8BB636B1FFBBEC4F748CFC79FBE2F8EF" pageId="8" pageNumber="38">
<table id="F909C411FFBB13B874B3FC79FA7FF8EF" box="[151,1436,982,1816]" gridcols="3" gridrows="20" pageId="8" pageNumber="38">
<tr id="353934F3FFBB13B874B3FC79FA7FFC1A" box="[151,1436,982,1005]" gridrow="0" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FC79FEF9FC1A" box="[151,282,982,1005]" gridcol="0" gridrow="0" pageId="8" pageNumber="38">Characters</th>
<th id="76E85D8FFFBB13B875BFFC79FD68FC1A" box="[411,651,982,1005]" gridcol="1" gridrow="0" pageId="8" pageNumber="38">
Females (
<emphasis id="B97DEAA3FFBBEC4F7620FC79FDD4FC1A" box="[516,567,982,1005]" italics="true" pageId="8" pageNumber="38">n =3</emphasis>
)
</th>
<th id="76E85D8FFFBB13B877B2FC79FA7FFC1A" box="[918,1436,982,1005]" gridcol="2" gridrow="0" pageId="8" pageNumber="38">
Males (
<emphasis id="B97DEAA3FFBBEC4F77C3FC79FBC2FC1A" box="[999,1057,982,1005]" italics="true" pageId="8" pageNumber="38">n = 4</emphasis>
)
</th>
</tr>
<tr id="353934F3FFBB13B874B3FBAEFA7FFBEF" box="[151,1436,1025,1048]" gridrow="1" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FBAEFEF9FBEF" box="[151,282,1025,1048]" gridcol="0" gridrow="1" pageId="8" pageNumber="38">SVL</th>
<td id="76E85D8FFFBB13B875BFFBAEFD68FBEF" box="[411,651,1025,1048]" gridcol="1" gridrow="1" pageId="8" pageNumber="38">32.636.5 (34.0 ± 2.1)</td>
<td id="76E85D8FFFBB13B877B2FBAEFA7FFBEF" box="[918,1436,1025,1048]" gridcol="2" gridrow="1" pageId="8" pageNumber="38">20.025.1 (22.9 ± 2.2)</td>
</tr>
<tr id="353934F3FFBB13B874B3FB83FA7FFBB4" box="[151,1436,1068,1091]" gridrow="2" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FB83FEF9FBB4" box="[151,282,1068,1091]" gridcol="0" gridrow="2" pageId="8" pageNumber="38">TL</th>
<td id="76E85D8FFFBB13B875BFFB83FD68FBB4" box="[411,651,1068,1091]" gridcol="1" gridrow="2" pageId="8" pageNumber="38">15.016.1 (15.7 ± 0.6)</td>
<td id="76E85D8FFFBB13B877B2FB83FA7FFBB4" box="[918,1436,1068,1091]" gridcol="2" gridrow="2" pageId="8" pageNumber="38">9.912.6 (11.3 ± 1.1)</td>
</tr>
<tr id="353934F3FFBB13B874B3FBF9FA7FFB9A" box="[151,1436,1110,1133]" gridrow="3" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FBF9FEF9FB9A" box="[151,282,1110,1133]" gridcol="0" gridrow="3" pageId="8" pageNumber="38">FL</th>
<td id="76E85D8FFFBB13B875BFFBF9FD68FB9A" box="[411,651,1110,1133]" gridcol="1" gridrow="3" pageId="8" pageNumber="38">13.315.1 (14.3 ± 0.9)</td>
<td id="76E85D8FFFBB13B877B2FBF9FA7FFB9A" box="[918,1436,1110,1133]" gridcol="2" gridrow="3" pageId="8" pageNumber="38">8.110.6 (9.6 ± 1.1)</td>
</tr>
<tr id="353934F3FFBB13B874B3FB2EFA7FFB6F" box="[151,1436,1153,1176]" gridrow="4" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FB2EFEF9FB6F" box="[151,282,1153,1176]" gridcol="0" gridrow="4" pageId="8" pageNumber="38">HL</th>
<td id="76E85D8FFFBB13B875BFFB2EFD68FB6F" box="[411,651,1153,1176]" gridcol="1" gridrow="4" pageId="8" pageNumber="38">12.714.1 (13.3 ± 0.7)</td>
<td id="76E85D8FFFBB13B877B2FB2EFA7FFB6F" box="[918,1436,1153,1176]" gridcol="2" gridrow="4" pageId="8" pageNumber="38">8.810.1 (9.1 ± 0.8)</td>
</tr>
<tr id="353934F3FFBB13B874B3FB03FA7FFB34" box="[151,1436,1196,1219]" gridrow="5" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FB03FEF9FB34" box="[151,282,1196,1219]" gridcol="0" gridrow="5" pageId="8" pageNumber="38">HW</th>
<td id="76E85D8FFFBB13B875BFFB03FD68FB34" box="[411,651,1196,1219]" gridcol="1" gridrow="5" pageId="8" pageNumber="38">12.513.5 (13.0 ± 0.5)</td>
<td id="76E85D8FFFBB13B877B2FB03FA7FFB34" box="[918,1436,1196,1219]" gridcol="2" gridrow="5" pageId="8" pageNumber="38">7.89.2 (8.5 ± 0.6)</td>
</tr>
<tr id="353934F3FFBB13B874B3FB79FA7FFB1A" box="[151,1436,1238,1261]" gridrow="6" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FB79FEF9FB1A" box="[151,282,1238,1261]" gridcol="0" gridrow="6" pageId="8" pageNumber="38">ED</th>
<td id="76E85D8FFFBB13B875BFFB79FD68FB1A" box="[411,651,1238,1261]" gridcol="1" gridrow="6" pageId="8" pageNumber="38">3.43.6 (3.5 ± 0.1)</td>
<td id="76E85D8FFFBB13B877B2FB79FA7FFB1A" box="[918,1436,1238,1261]" gridcol="2" gridrow="6" pageId="8" pageNumber="38">2.43.7 (3.1 ± 0.5)</td>
</tr>
<tr id="353934F3FFBB13B874B3FAAEFA7FFAEF" box="[151,1436,1281,1304]" gridrow="7" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FAAEFEF9FAEF" box="[151,282,1281,1304]" gridcol="0" gridrow="7" pageId="8" pageNumber="38">TY</th>
<td id="76E85D8FFFBB13B875BFFAAEFD68FAEF" box="[411,651,1281,1304]" gridcol="1" gridrow="7" pageId="8" pageNumber="38">1.51.9 (1.8 ± 0.2)</td>
<td id="76E85D8FFFBB13B877B2FAAEFA7FFAEF" box="[918,1436,1281,1304]" gridcol="2" gridrow="7" pageId="8" pageNumber="38">1.11.5 (1.3 ± 0.2)</td>
</tr>
<tr id="353934F3FFBB13B874B3FA83FA7FFAB4" box="[151,1436,1324,1347]" gridrow="8" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FA83FEF9FAB4" box="[151,282,1324,1347]" gridcol="0" gridrow="8" pageId="8" pageNumber="38">IOD</th>
<td id="76E85D8FFFBB13B875BFFA83FD68FAB4" box="[411,651,1324,1347]" gridcol="1" gridrow="8" pageId="8" pageNumber="38">3.43.8 (2.7 ± 0.2)</td>
<td id="76E85D8FFFBB13B877B2FA83FA7FFAB4" box="[918,1436,1324,1347]" gridcol="2" gridrow="8" pageId="8" pageNumber="38">2.33.1 (2.6 ± 0.4)</td>
</tr>
<tr id="353934F3FFBB13B874B3FAF9FA7FFA9A" box="[151,1436,1366,1389]" gridrow="9" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FAF9FEF9FA9A" box="[151,282,1366,1389]" gridcol="0" gridrow="9" pageId="8" pageNumber="38">EW</th>
<td id="76E85D8FFFBB13B875BFFAF9FD68FA9A" box="[411,651,1366,1389]" gridcol="1" gridrow="9" pageId="8" pageNumber="38">3.13.3 (3.2 ± 0.1)</td>
<td id="76E85D8FFFBB13B877B2FAF9FA7FFA9A" box="[918,1436,1366,1389]" gridcol="2" gridrow="9" pageId="8" pageNumber="38">1.82.8 (2.2 ± 0.5)</td>
</tr>
<tr id="353934F3FFBB13B874B3FA2EFA7FFA6F" box="[151,1436,1409,1432]" gridrow="10" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FA2EFEF9FA6F" box="[151,282,1409,1432]" gridcol="0" gridrow="10" pageId="8" pageNumber="38">IND</th>
<td id="76E85D8FFFBB13B875BFFA2EFD68FA6F" box="[411,651,1409,1432]" gridcol="1" gridrow="10" pageId="8" pageNumber="38">2.63.0 (2.8 ± 0.2)</td>
<td id="76E85D8FFFBB13B877B2FA2EFA7FFA6F" box="[918,1436,1409,1432]" gridcol="2" gridrow="10" pageId="8" pageNumber="38">1.82.2 (2.1 ± 0.2)</td>
</tr>
<tr id="353934F3FFBB13B874B3FA03FA7FFA34" box="[151,1436,1452,1475]" gridrow="11" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FA03FEF9FA34" box="[151,282,1452,1475]" gridcol="0" gridrow="11" pageId="8" pageNumber="38">EN</th>
<td id="76E85D8FFFBB13B875BFFA03FD68FA34" box="[411,651,1452,1475]" gridcol="1" gridrow="11" pageId="8" pageNumber="38">3.84.7 (4.2 ± 0.5)</td>
<td id="76E85D8FFFBB13B877B2FA03FA7FFA34" box="[918,1436,1452,1475]" gridcol="2" gridrow="11" pageId="8" pageNumber="38">2.73.0 (2.9 ± 0.1)</td>
</tr>
<tr id="353934F3FFBB13B874B3FA79FA7FFA1A" box="[151,1436,1494,1517]" gridrow="12" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3FA79FEF9FA1A" box="[151,282,1494,1517]" gridcol="0" gridrow="12" pageId="8" pageNumber="38">TL/SVL</th>
<td id="76E85D8FFFBB13B875BFFA79FD68FA1A" box="[411,651,1494,1517]" gridcol="1" gridrow="12" pageId="8" pageNumber="38">0.440.48</td>
<td id="76E85D8FFFBB13B877B2FA79FA7FFA1A" box="[918,1436,1494,1517]" gridcol="2" gridrow="12" pageId="8" pageNumber="38">0.470.50</td>
</tr>
<tr id="353934F3FFBB13B874B3F9AEFA7FF9EF" box="[151,1436,1537,1560]" gridrow="13" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F9AEFEF9F9EF" box="[151,282,1537,1560]" gridcol="0" gridrow="13" pageId="8" pageNumber="38">FL/SVL</th>
<td id="76E85D8FFFBB13B875BFF9AEFD68F9EF" box="[411,651,1537,1560]" gridcol="1" gridrow="13" pageId="8" pageNumber="38">0.410.44</td>
<td id="76E85D8FFFBB13B877B2F9AEFA7FF9EF" box="[918,1436,1537,1560]" gridcol="2" gridrow="13" pageId="8" pageNumber="38">0.410.42</td>
</tr>
<tr id="353934F3FFBB13B874B3F983FA7FF9B4" box="[151,1436,1580,1603]" gridrow="14" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F983FEF9F9B4" box="[151,282,1580,1603]" gridcol="0" gridrow="14" pageId="8" pageNumber="38">HL/SVL</th>
<td id="76E85D8FFFBB13B875BFF983FD68F9B4" box="[411,651,1580,1603]" gridcol="1" gridrow="14" pageId="8" pageNumber="38">0.390.40</td>
<td id="76E85D8FFFBB13B877B2F983FA7FF9B4" box="[918,1436,1580,1603]" gridcol="2" gridrow="14" pageId="8" pageNumber="38">0.370.42</td>
</tr>
<tr id="353934F3FFBB13B874B3F9F9FA7FF99A" box="[151,1436,1622,1645]" gridrow="15" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F9F9FEF9F99A" box="[151,282,1622,1645]" gridcol="0" gridrow="15" pageId="8" pageNumber="38">HW/SVL</th>
<td id="76E85D8FFFBB13B875BFF9F9FD68F99A" box="[411,651,1622,1645]" gridcol="1" gridrow="15" pageId="8" pageNumber="38">0.370.40</td>
<td id="76E85D8FFFBB13B877B2F9F9FA7FF99A" box="[918,1436,1622,1645]" gridcol="2" gridrow="15" pageId="8" pageNumber="38">0.360.39</td>
</tr>
<tr id="353934F3FFBB13B874B3F92EFA7FF96F" box="[151,1436,1665,1688]" gridrow="16" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F92EFEF9F96F" box="[151,282,1665,1688]" gridcol="0" gridrow="16" pageId="8" pageNumber="38">HW/HL</th>
<td id="76E85D8FFFBB13B875BFF92EFD68F96F" box="[411,651,1665,1688]" gridcol="1" gridrow="16" pageId="8" pageNumber="38">0.951.02</td>
<td id="76E85D8FFFBB13B877B2F92EFA7FF96F" box="[918,1436,1665,1688]" gridcol="2" gridrow="16" pageId="8" pageNumber="38">0.850.99</td>
</tr>
<tr id="353934F3FFBB13B874B3F903FA7FF934" box="[151,1436,1708,1731]" gridrow="17" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F903FEF9F934" box="[151,282,1708,1731]" gridcol="0" gridrow="17" pageId="8" pageNumber="38">EN/ED</th>
<td id="76E85D8FFFBB13B875BFF903FD68F934" box="[411,651,1708,1731]" gridcol="1" gridrow="17" pageId="8" pageNumber="38">1.121.34</td>
<td id="76E85D8FFFBB13B877B2F903FA7FF934" box="[918,1436,1708,1731]" gridcol="2" gridrow="17" pageId="8" pageNumber="38">0.811.17</td>
</tr>
<tr id="353934F3FFBB13B874B3F979FA7FF91A" box="[151,1436,1750,1773]" gridrow="18" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F979FEF9F91A" box="[151,282,1750,1773]" gridcol="0" gridrow="18" pageId="8" pageNumber="38">EW/IOD</th>
<td id="76E85D8FFFBB13B875BFF979FD68F91A" box="[411,651,1750,1773]" gridcol="1" gridrow="18" pageId="8" pageNumber="38">0.870.91</td>
<td id="76E85D8FFFBB13B877B2F979FA7FF91A" box="[918,1436,1750,1773]" gridcol="2" gridrow="18" pageId="8" pageNumber="38">0.751.22</td>
</tr>
<tr id="353934F3FFBB13B874B3F8AEFA7FF8EF" box="[151,1436,1793,1816]" gridrow="19" pageId="8" pageNumber="38">
<th id="76E85D8FFFBB13B874B3F8AEFEF9F8EF" box="[151,282,1793,1816]" gridcol="0" gridrow="19" pageId="8" pageNumber="38">TY/ED</th>
<td id="76E85D8FFFBB13B875BFF8AEFD68F8EF" box="[411,651,1793,1816]" gridcol="1" gridrow="19" pageId="8" pageNumber="38">0.420.56</td>
<td id="76E85D8FFFBB13B877B2F8AEFA7FF8EF" box="[918,1436,1793,1816]" gridcol="2" gridrow="19" pageId="8" pageNumber="38">0.390.46</td>
</tr>
</table>
</paragraph>
<subSubSection id="C313653AFFBBEC4E74E1F8CCFF19FE37" lastPageId="9" lastPageNumber="39" pageId="8" pageNumber="38" type="discussion">
<paragraph id="8BB636B1FFBBEC4F74E1F8CCFBA9F853" blockId="8.[151,1436,1891,2034]" pageId="8" pageNumber="38">
<emphasis id="B97DEAA3FFBBEC4F74E1F8CCFEA1F88A" bold="true" box="[197,322,1891,1917]" pageId="8" pageNumber="38">Remarks.</emphasis>
Because of its distribution in northern
<collectingCountry id="F31E7621FFBBEC4F770CF8CBFC83F889" box="[808,864,1892,1918]" name="Peru" pageId="8" pageNumber="38">Peru</collectingCountry>
relatively close to the borders of
<collectingCountry id="F31E7621FFBBEC4F7126F8CBFA8BF889" box="[1282,1384,1892,1918]" name="Ecuador" pageId="8" pageNumber="38">Ecuador</collectingCountry>
and
<collectingCountry id="F31E7621FFBBEC4F74B3F825FEEDF853" box="[151,270,1930,1956]" name="Colombia" pageId="8" pageNumber="38">Colombia</collectingCountry>
,
<taxonomicName id="4C094D32FFBBEC4F753DF824FE13F853" box="[281,496,1931,1956]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="38" phylum="Chordata" rank="species" species="orcus">
<emphasis id="B97DEAA3FFBBEC4F753DF824FE13F853" box="[281,496,1931,1956]" italics="true" pageId="8" pageNumber="38">Pristimantis orcus</emphasis>
</taxonomicName>
is very likely distributed in these countries as well.
</paragraph>
<paragraph id="8BB636B1FFBBEC4F74E1F81EFA7FF805" blockId="8.[151,1436,1891,2034]" pageId="8" pageNumber="38">
Several species of
<taxonomicName id="4C094D32FFBBEC4F7583F81EFDD9F83D" box="[423,570,1969,1994]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFBBEC4F7583F81EFDD9F83D" box="[423,570,1969,1994]" italics="true" pageId="8" pageNumber="38">Pristimantis</emphasis>
</taxonomicName>
from the Amazonian lowlands have the belly, groin, anterior surface of thighs, and concealed surfaces of tibia with a contrasting coloration pattern of black and white (
<tableCitation id="C68B030AFFBBEC4F7109F877FA6AF805" box="[1325,1417,2008,2034]" captionStart="TABLE 1" captionStartId="5.[151,240,151,175]" captionTargetBox="[151,1436,241,1557]" captionTargetPageId="5" captionText="TABLE 1. Selected characters (+ = character present; — = character absent) and character conditions among selected" httpUri="http://table.plazi.org/id/DF766639FFB6EC4274B3FF38FA7FFF58" pageId="8" pageNumber="38" tableUuid="DF766639FFB6EC4274B3FF38FA7FFF58">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BB636B1FFBAEC4E74B3FF37FF19FE37" blockId="9.[151,1437,152,448]" pageId="9" pageNumber="39">
Currently all these species are assigned to the
<taxonomicName id="4C094D32FFBAEC4E7697FF37FCDCFF46" box="[691,831,152,177]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B97DEAA3FFBAEC4E7697FF37FCDCFF46" box="[691,831,152,177]" italics="true" pageId="9" pageNumber="39">unistrigatus</emphasis>
</taxonomicName>
Group which is the largest species group (
<bibRefCitation id="EF984B40FFBAEC4E7116FF37FEADFF2F" author="Lynch" pageId="9" pageNumber="39" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus in western Ecuador. Systematics, ecology, and biogeography. Natural History Museum Special Publication, The University of Kansas, 23, 1 - 236." type="journal article" year="1997">Lynch &amp; Duellman 1997</bibRefCitation>
) and known to be paraphyletic (
<bibRefCitation id="EF984B40FFBAEC4E76E3FF11FC4AFF2F" author="Hedges" box="[711,937,190,216]" pageId="9" pageNumber="39" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">
Hedges
<emphasis id="B97DEAA3FFBAEC4E770CFF10FC81FF2F" box="[808,866,191,216]" italics="true" pageId="9" pageNumber="39">et al.</emphasis>
2008
</bibRefCitation>
). Most members of the
<taxonomicName id="4C094D32FFBAEC4E709AFF10FAA9FF2F" box="[1214,1354,191,216]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B97DEAA3FFBAEC4E709AFF10FAA9FF2F" box="[1214,1354,191,216]" italics="true" pageId="9" pageNumber="39">unistrigatus</emphasis>
</taxonomicName>
Group were assigned to it because their morphological character set did not seem to fit into one of the other species groups of
<taxonomicName id="4C094D32FFBAEC4E7531FEA3FE48FED2" box="[277,427,268,293]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFBAEC4E7531FEA3FE48FED2" box="[277,427,268,293]" italics="true" pageId="9" pageNumber="39">Pristimantis</emphasis>
</taxonomicName>
. Because of the morphological similarities among
<emphasis id="B97DEAA3FFBAEC4E7001FEA3FCB0FEBC" italics="true" pageId="9" pageNumber="39">
<taxonomicName id="4C094D32FFBAEC4E7001FEA3FA9AFED2" box="[1061,1401,268,293]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="altamazonicus">Pristimantis altamazonicus</taxonomicName>
,
<taxonomicName id="4C094D32FFBAEC4E71A2FEA3FEC2FEBC" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="diadematus">P. diadematus</taxonomicName>
,
<taxonomicName id="4C094D32FFBAEC4E7509FE9DFE00FEBC" box="[301,483,306,331]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="eurydactylus">P. eurydactylus</taxonomicName>
,
<taxonomicName id="4C094D32FFBAEC4E75CBFE9DFDACFEBC" box="[495,591,306,331]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="orcus">P. orcus</taxonomicName>
,
<taxonomicName id="4C094D32FFBAEC4E767FFE9DFCACFEBC" box="[603,847,306,331]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="species" species="ventrimarmoratus">P. ventrimarmoratus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C094D32FFBAEC4E77A8FE9DFBF8FEBC" box="[908,1051,306,331]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="B97DEAA3FFBAEC4E77A8FE9DFBF8FEBC" box="[908,1051,306,331]" italics="true" pageId="9" pageNumber="39">Pristimantis</emphasis>
</taxonomicName>
sp., we assume that they share a close phylogenetic history and might be part of a distinct clade. Before assigning them to a new species group, the results of phylogenetic analyses currently being undertaken by different research groups should be awaited.
</paragraph>
</subSubSection>
</treatment>
</document>