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<document id="4631C1D494FBE9BF1FCFE4484FD9D0CE" ID-CLB-Dataset="7307" ID-DOI="10.1051/acarologia/20162243" ID-GBIF-Dataset="eb34fd8d-6897-40b4-a149-73b6cb23af22" ID-ISSN="2107-7207" ID-Zenodo-Dep="4640224" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616776549132" checkinUser="carolina" docAuthor="Vázquez, M. M., de Araújo, M. S. &amp; Feres, R. J. F." docDate="2016" docId="039587F1FFFB805C9D5375085877FBD7" docLanguage="en" docName="Acarologia.56.3.269-278.pdf" docOrigin="Acarologia 56 (3)" docSource="http://dx.doi.org/10.1051/acarologia/20162243" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Diplothyrus lehtineni Vázquez &amp; de Araújo &amp; Feres 2016, n. sp." docType="treatment" docUuid="270B9290-A2EA-4581-AF09-1D2153C243E6" docUuidSource="ZooBank" docVersion="8" lastPageNumber="276" masterDocId="FFACFF89FFF880549C7B73045D4AFFBD" masterDocTitle="A new species of Diplothyrus (Parasitiformes: Neothyridae) from Brazil" masterLastPageNumber="278" masterPageNumber="269" pageNumber="271" updateTime="1698935180394" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="50EC431A6686A94B477D406F2EBF5B5B">A new species of Diplothyrus (Parasitiformes: Neothyridae) from Brazil</mods:title>
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<treatment id="039587F1FFFB805C9D5375085877FBD7" ID-DOI="http://doi.org/10.5281/zenodo.4697126" ID-GBIF-Taxon="181020864" ID-Zenodo-Dep="4697126" LSID="urn:lsid:zoobank.org:act:270B9290-A2EA-4581-AF09-1D2153C243E6" httpUri="http://treatment.plazi.org/id/039587F1FFFB805C9D5375085877FBD7" lastPageId="8" lastPageNumber="276" pageId="3" pageNumber="271">
<subSubSection id="C326656CFFFB80579D5375085F3BF998" box="[296,625,1548,1573]" pageId="3" pageNumber="271" type="nomenclature">
<paragraph id="8B8336E7FFFB80579D5375085F3BF998" blockId="3.[296,625,1548,1607]" box="[296,625,1548,1573]" pageId="3" pageNumber="271">
<heading id="D0CB818BFFFB80579D5375085F3BF998" bold="true" box="[296,625,1548,1573]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="271" reason="2">
<emphasis id="B948EAF5FFFB80579D5375085F3BF998" bold="true" box="[296,625,1548,1573]" pageId="3" pageNumber="271">
<taxonomicName id="4C3C4D64FFFB80579D5375085F62F999" ID-CoL="8N97H" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[296,552,1548,1572]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFFB80579D5375085F62F999" bold="true" box="[296,552,1548,1572]" italics="true" pageId="3" pageNumber="271">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B578EFFFB80579E5475095F3BF998" box="[559,625,1549,1573]" pageId="3" pageNumber="271" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B8336E7FFFB80579DFA752B5F5DF9FA" blockId="3.[296,625,1548,1607]" box="[385,535,1583,1607]" pageId="3" pageNumber="271">
<emphasis id="B948EAF5FFFB80579DFA752B5F5DF9FA" bold="true" box="[385,535,1583,1607]" pageId="3" pageNumber="271">
(
<figureCitation id="13072A62FFFB80579DF2752B5F45F9FA" box="[393,527,1583,1607]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStart-4="FIGURE" captionStartId-0="4.[159,171,1292,1311]" captionStartId-1="5.[159,171,1848,1867]" captionStartId-2="6.[159,171,1858,1877]" captionStartId-3="7.[159,171,1755,1774]" captionStartId-4="8.[338,350,759,778]" captionTargetBox-0="[227,1361,302,1260]" captionTargetBox-1="[315,1283,339,1811]" captionTargetBox-2="[269,1319,357,1807]" captionTargetBox-3="[277,1305,450,1713]" captionTargetBox-4="[234,1347,307,716]" captionTargetId-0="figure-4@4.[227,1361,302,1260]" captionTargetId-1="figure-6@5.[264,1323,319,1817]" captionTargetId-2="figure-4@6.[264,1323,337,1826]" captionTargetId-3="figure-6@7.[264,1323,440,1723]" captionTargetId-4="figure-4@8.[227,1361,302,726]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2: Diplothyrus lehtineni n. sp.: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male." captionText-1="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." captionText-2="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." captionText-3="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." captionText-4="FIGURE 6: Diplothyrus lehtineni n. sp.: A-B Phase contrast images of Leg I tip and Hallers organ" figureDoi-0="http://doi.org/10.5281/zenodo.4640228" figureDoi-1="http://doi.org/10.5281/zenodo.4640230" figureDoi-2="http://doi.org/10.5281/zenodo.4640234" figureDoi-3="http://doi.org/10.5281/zenodo.4640236" figureDoi-4="http://doi.org/10.5281/zenodo.4640238" httpUri-0="https://zenodo.org/record/4640228/files/figure.png" httpUri-1="https://zenodo.org/record/4640230/files/figure.png" httpUri-2="https://zenodo.org/record/4640234/files/figure.png" httpUri-3="https://zenodo.org/record/4640236/files/figure.png" httpUri-4="https://zenodo.org/record/4640238/files/figure.png" pageId="3" pageNumber="271">Figures 2-6</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B8336E7FFFB80579CE475685EEFFECF" blockId="3.[159,762,1644,1949]" lastBlockId="3.[826,1429,311,1949]" pageId="3" pageNumber="271">
Differential diagnosis — Thons organ with 2 orifices inserted parallel to dorsal shield border, similar as in
<taxonomicName id="4C3C4D64FFFB80579D7875B75CC0F977" authorityName="Lehtinen" authorityYear="1999" box="[259,394,1714,1738]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B948EAF5FFFB80579D7875B75CC0F977" box="[259,394,1714,1738]" italics="true" pageId="3" pageNumber="271">D. schubarti</emphasis>
</taxonomicName>
, but with two big and conspicuous membranous structures like funnels. Tibiotarsus with 6 modified serrated setae (5 inserted in line, plus 1 lateral). Two modified serrated setae on palp genu, but neither of them like the modified setae present on that of
<taxonomicName id="4C3C4D64FFFB80579DED74665F50F8C4" authorityName="Lehtinen" authorityYear="1999" box="[406,538,1889,1913]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B948EAF5FFFB80579DED74665F50F8C4" box="[406,538,1889,1913]" italics="true" pageId="3" pageNumber="271">D. schubarti</emphasis>
</taxonomicName>
. Chelicera without complex branched outgrowth. Leg I with Hallers organ composed of a group of 7 highly modified sensillae.
</paragraph>
<paragraph id="8B8336E7FFFB80579F1B72805804FE21" blockId="3.[826,1429,311,1949]" box="[864,1358,388,412]" pageId="3" pageNumber="271">
<emphasis id="B948EAF5FFFB80579F1B72805EBFFE21" bold="true" box="[864,1013,388,412]" pageId="3" pageNumber="271">Description.</emphasis>
Females (N=7), males (N=7).
</paragraph>
<paragraph id="8B8336E7FFFB80579F1B72AA580DFBA7" blockId="3.[826,1429,311,1949]" pageId="3" pageNumber="271">
Idiosoma — Dark brown, well-sclerotized, in shape of a large dome, with numerous long, serrated, terminally acute setae. Dorsal shield with light reticulation interrupted by numerous shallow, round indentations. Two pairs of dorso-lateral orifices (Thons organ), the posterior orifice larger, surrounded by an ornamented funnel, the anterior one with an inverted funnel under the cuticle; both orifices are connected by a cuticular strip, inserted parallel to the dorsal shield margin (
<figureCitation id="13072A62FFFB805798C371EE5870FCBF" box="[1208,1338,746,770]" captionStart="FIGURE" captionStartId="4.[159,171,1292,1311]" captionTargetBox="[227,1361,302,1260]" captionTargetId="figure-4@4.[227,1361,302,1260]" captionTargetPageId="4" captionText="FIGURE 2: Diplothyrus lehtineni n. sp.: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male." figureDoi="http://doi.org/10.5281/zenodo.4640228" httpUri="https://zenodo.org/record/4640228/files/figure.png" pageId="3" pageNumber="271">Figures 2A</figureCitation>
,
<figureCitation id="13072A62FFFB8057993F71EE5E15FC98" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="3" pageNumber="271">5B and 5D</figureCitation>
). Ventral shield lightly granulate; with reticulate patterning around genital plate in male. Peritremes well-developed, at the lateral border of the dorsal shield (
<figureCitation id="13072A62FFFB80579FF470725947FC33" box="[911,1037,886,910]" captionStart="FIGURE" captionStartId="4.[159,171,1292,1311]" captionTargetBox="[227,1361,302,1260]" captionTargetId="figure-4@4.[227,1361,302,1260]" captionTargetPageId="4" captionText="FIGURE 2: Diplothyrus lehtineni n. sp.: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male." figureDoi="http://doi.org/10.5281/zenodo.4640228" httpUri="https://zenodo.org/record/4640228/files/figure.png" pageId="3" pageNumber="271">Figures 2B</figureCitation>
,
<figureCitation id="13072A62FFFB8057986C707259DEFC33" box="[1047,1172,886,910]" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="3" pageNumber="271">5A and 5C</figureCitation>
), extending as a patch of papillate cuticle (
<figureCitation id="13072A62FFFB80579865709D59D6FC0C" box="[1054,1180,921,945]" captionStart="FIGURE" captionStartId="4.[159,171,1292,1311]" captionTargetBox="[227,1361,302,1260]" captionTargetId="figure-4@4.[227,1361,302,1260]" captionTargetPageId="4" captionText="FIGURE 2: Diplothyrus lehtineni n. sp.: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male." figureDoi="http://doi.org/10.5281/zenodo.4640228" httpUri="https://zenodo.org/record/4640228/files/figure.png" pageId="3" pageNumber="271">Figures 2B</figureCitation>
,
<figureCitation id="13072A62FFFB805798DC709D5869FC0C" box="[1191,1315,921,945]" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="3" pageNumber="271">5A and 5C</figureCitation>
) from anterior to coxa I to coxa IV. Stigma inserted beyond level of coxa III, in peritrematal area (
<figureCitation id="13072A62FFFB8057996870DB58C7FC4A" box="[1299,1421,991,1015]" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="3" pageNumber="271">Figure 5C</figureCitation>
) which is extended between coxa I to coxa III.
</paragraph>
<paragraph id="8B8336E7FFFB80579F1B77295966FB13" blockId="3.[826,1429,311,1949]" pageId="3" pageNumber="271">
Gnathosoma — Deutosternum well developed with numerous small teeth. Cuticular patterning limited to papillate zones on each side of deutosternum (
<figureCitation id="13072A62FFFB80579F0477925EA1FB13" box="[895,1003,1174,1198]" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="3" pageNumber="271">Figures 5</figureCitation>
C-D).
</paragraph>
<paragraph id="8B8336E7FFFB80579F1B77C45EF8F9C0" blockId="3.[826,1429,311,1949]" pageId="3" pageNumber="271">
Subcapitulum (
<figureCitation id="13072A62FFFB8057986077C459C5FB65" box="[1051,1167,1216,1240]" captionStart="FIGURE" captionStartId="4.[159,171,1292,1311]" captionTargetBox="[227,1361,302,1260]" captionTargetId="figure-4@4.[227,1361,302,1260]" captionTargetPageId="4" captionText="FIGURE 2: Diplothyrus lehtineni n. sp.: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male." figureDoi="http://doi.org/10.5281/zenodo.4640228" httpUri="https://zenodo.org/record/4640228/files/figure.png" pageId="3" pageNumber="271">Figures 2</figureCitation>
C-D) with 8-11 setae (males with 8, females with 11), hypostomal lobes with two median long setae. Corniculus horn-like inserted dorso-lateral. Lateral lips well developed with numerous small projections. Labrum well developed. Chelicera (
<figureCitation id="13072A62FFFB80579850766B59D2FA3A" box="[1067,1176,1391,1415]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="3" pageNumber="271">Figures 3</figureCitation>
A-B) well developed, 2 basal segments without setae; fixed digit with a small seta. Movable digit with one large tooth, a large terminal hook and numerous small, median teeth; fixed digit with two large and numerous small teeth. Membranous outgrowth not present, chelicerae covered by a membranous structure (
<figureCitation id="13072A62FFFB8057991E75465E39F9C0" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="3" pageNumber="271">Figure 3</figureCitation>
A-B).
</paragraph>
<paragraph id="8B8336E7FFFB80579F1B758B59F5F820" blockId="3.[826,1429,311,1949]" pageId="3" pageNumber="271">
Palp (
<figureCitation id="13072A62FFFB80579FDE758B5958F91A" box="[933,1042,1679,1703]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="3" pageNumber="271">Figures 3</figureCitation>
C-D) — Trochanter with 2 setae, one with a bifurcate tip; femur with 20-21 setae and papillate cuticle; genu with 8-10 setae, two of them wide and serrate, the others acuminate; tibia with 60 curved, simple setae, and two
<emphasis id="B948EAF5FFFB805798B5741F59ABF88E" box="[1230,1249,1819,1843]" italics="true" pageId="3" pageNumber="271">al</emphasis>
serrated setae, 6v strong and serrated setae and one sensilla. The modified setae on the genu are like a doubled setae, with serrated border (
<figureCitation id="13072A62FFFB80579841748059F8F820" box="[1082,1202,1924,1949]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="3" pageNumber="271">Figure 3D</figureCitation>
).
</paragraph>
<paragraph id="8B8336E7FFFC80509CE473FE5C2EFEB0" blockId="4.[159,356,250,269]" box="[159,356,250,269]" pageId="4" pageNumber="272">
VAEzquez Ma.M.
<emphasis id="B948EAF5FFFC80509D4273FF5C2EFEB0" box="[313,356,250,269]" italics="true" pageId="4" pageNumber="272">et al.</emphasis>
</paragraph>
<caption id="DF43666FFFFC80509CE47608597CFA87" ID-DOI="http://doi.org/10.5281/zenodo.4640228" ID-Zenodo-Dep="4640228" httpUri="https://zenodo.org/record/4640228/files/figure.png" pageId="4" pageNumber="272" startId="4.[159,171,1292,1311]" targetBox="[227,1361,302,1260]" targetPageId="4">
<paragraph id="8B8336E7FFFC80509CE47608597CFA87" blockId="4.[159,1429,1291,1339]" pageId="4" pageNumber="272">
FIGURE 2:
<taxonomicName id="4C3C4D64FFFC80509D78760F5CF7FAA3" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[259,445,1291,1310]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFFC80509D78760F5CF7FAA3" box="[259,445,1291,1310]" italics="true" pageId="4" pageNumber="272">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<emphasis id="B948EAF5FFFC80509DBA76085CBFFAA2" bold="true" box="[449,501,1292,1311]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="A27B578EFFFC80509DBA76085CBFFAA2" box="[449,501,1292,1311]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A — Detail of the Thons organ and the internal connection by the membranous funnel; B — Detail of the peritreme and stigma; C — Subcapitulum of the female; D — Subcapitulum of the male.
</paragraph>
</caption>
<paragraph id="8B8336E7FFFC80509CBE76785F90F8C2" blockId="4.[159,762,1404,1919]" pageId="4" pageNumber="272">
Genital region (
<figureCitation id="13072A62FFFC80509D0476785CA1FA29" box="[383,491,1404,1428]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="4" pageNumber="272">Figures 3</figureCitation>
E-G and 5E-F) — Sternal lyrifissures not observed. Female genital shields include four plates (
<figureCitation id="13072A62FFFC80509DF476C65CB1FA67" box="[399,507,1474,1498]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="4" pageNumber="272">Figures 3</figureCitation>
E-G and 5E): progenital valve (
<emphasis id="B948EAF5FFFC80509D5C76E25C05FA40" box="[295,335,1510,1533]" italics="true" pageId="4" pageNumber="272">pgv</emphasis>
) with 15-16 acuminate setae (
<figureCitation id="13072A62FFFC80509EB176E15DB8F99D" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="4" pageNumber="272">Figure 3E</figureCitation>
); two laterogenital valves (
<emphasis id="B948EAF5FFFC80509E3E750C5F2DF99D" box="[581,615,1544,1568]" italics="true" pageId="4" pageNumber="272">lgv</emphasis>
) (
<figureCitation id="13072A62FFFC80509EFF750C5FB3F99D" box="[644,761,1544,1568]" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="4" pageNumber="272">Figure 3F</figureCitation>
<emphasis id="B948EAF5FFFC80509CE4752F5D8BF9FE" box="[159,193,1579,1603]" italics="true" pageId="4" pageNumber="272">lgv</emphasis>
) with 4 setae each, large trapezoid mesogenital valve (
<emphasis id="B948EAF5FFFC80509D63754B5C0DF9DB" box="[280,327,1615,1638]" italics="true" pageId="4" pageNumber="272">mgv</emphasis>
) with numerous acute setae (
<figureCitation id="13072A62FFFC80509ED7754A5D89F934" captionStart="FIGURE" captionStartId="5.[159,171,1848,1867]" captionTargetBox="[315,1283,339,1811]" captionTargetId="figure-6@5.[264,1323,319,1817]" captionTargetPageId="5" captionText="FIGURE 3: Diplothyrus lehtineni n. sp.: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate." figureDoi="http://doi.org/10.5281/zenodo.4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="4" pageNumber="272">Figure 3G</figureCitation>
). Under mesogenital valve a globose hyaline sac with long slim channels is visible and interpreted as a part of the ovipositor. One female was observed with two eggs. Male genital area (
<figureCitation id="13072A62FFFC80509EDA75DE5D8AF8AB" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figures 4A</figureCitation>
and
<figureCitation id="13072A62FFFC80509C8475F95C51F8A8" box="[255,283,1789,1813]" captionStart="FIGURE" captionStartId="7.[159,171,1755,1774]" captionTargetBox="[277,1305,450,1713]" captionTargetId="figure-6@7.[264,1323,440,1723]" captionTargetPageId="7" captionText="FIGURE 5: Diplothyrus lehtineni n. sp. SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates." figureDoi="http://doi.org/10.5281/zenodo.4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="4" pageNumber="272">5F</figureCitation>
) consisting of two plates positioned between coxae IV, anterior one with 17-18 large, lanceolate setae and posterior one without setae but with papillate cuticle. Males with distinct epiandrum.
</paragraph>
<paragraph id="8B8336E7FFFC80509F1B7678592FFA67" blockId="4.[826,1429,1404,1498]" pageId="4" pageNumber="272">
Anal region (
<figureCitation id="13072A62FFFC8050987976785939FA29" box="[1026,1139,1404,1428]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figures 4</figureCitation>
B-C) — Each anal plate with large and acute setae,
<quantity id="4CC49B02FFFC805098F1769B59F3FA0A" box="[1162,1209,1439,1463]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="272" unit="in" value="9.0">9 in</quantity>
females (
<figureCitation id="13072A62FFFC80509954769B58C7FA0A" box="[1327,1421,1439,1463]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Fig. 4B</figureCitation>
) and
<quantity id="4CC49B02FFFC80509F1576C65EECFA67" box="[878,934,1474,1498]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="272" unit="in" value="10.0">10 in</quantity>
males (
<figureCitation id="13072A62FFFC8050987B76C65912FA67" box="[1024,1112,1474,1498]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8B8336E7FFFC80509F1B750759D1F821" blockId="4.[826,1429,1539,1948]" pageId="4" pageNumber="272">
Legs (
<figureCitation id="13072A62FFFC80509FDC7507595FF9A6" box="[935,1045,1539,1563]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figures 4</figureCitation>
D-F and 6A-B) — Leg I Hallers organ (
<figureCitation id="13072A62FFFC80509FEB7522594AF983" box="[912,1024,1574,1598]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figures 4</figureCitation>
D-E and 6A-B) in the distodorsal part, comprising seven sensillae (
<figureCitation id="13072A62FFFC805098B3754D5866F9DC" box="[1224,1324,1609,1633]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figure 4</figureCitation>
D-E). All legs with a distinct basifemur. Leg I without acro- or basitarsus. Leg setation on all segments based on whorls of 8 or 9 setae each. Tarsus I: The subterminal Hallers organ is in a depression with dense cluster of highly modified sensillae (
<figureCitation id="13072A62FFFC805098A775FC581EF8AD" box="[1244,1364,1784,1808]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figure 4D</figureCitation>
). Pretarsus I with almost sessile claws (
<figureCitation id="13072A62FFFC805098A2741F581AF88E" box="[1241,1360,1819,1843]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figure 4E</figureCitation>
); pretarsi II-IV (
<figureCitation id="13072A62FFFC80509FC0743A5967F8EB" box="[955,1069,1854,1878]" captionStart="FIGURE" captionStartId="6.[159,171,1858,1877]" captionTargetBox="[269,1319,357,1807]" captionTargetId="figure-4@6.[264,1323,337,1826]" captionTargetPageId="6" captionText="FIGURE 4: Diplothyrus lehtineni n. sp.: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male." figureDoi="http://doi.org/10.5281/zenodo.4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="4" pageNumber="272">Figure 4F</figureCitation>
) with an ambulacral stalk carrying 1 pair of small setae, with well developed claws, and a large empodium.
</paragraph>
</subSubSection>
<caption id="DF43666FFFFD80519CE4743C5FEAF83F" ID-DOI="http://doi.org/10.5281/zenodo.4640230" ID-Zenodo-Dep="4640230" httpUri="https://zenodo.org/record/4640230/files/figure.png" pageId="5" pageNumber="273" startId="5.[159,171,1848,1867]" targetBox="[315,1283,339,1811]" targetPageId="5">
<paragraph id="8B8336E7FFFD80519CE4743C5FEAF83F" blockId="5.[159,1429,1848,1923]" pageId="5" pageNumber="273">
FIGURE 3:
<taxonomicName id="4C3C4D64FFFD80519D71743C5C8CF8F6" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[266,454,1848,1867]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFFD80519D71743C5C8CF8F6" box="[266,454,1848,1867]" italics="true" pageId="5" pageNumber="273">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<emphasis id="B948EAF5FFFD80519DB6743C5F4CF8F6" bold="true" box="[461,518,1848,1867]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="A27B578EFFFD80519DB6743C5F4CF8F6" box="[461,518,1848,1867]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A — The chelicera, right side, lateral view, detail with visible hyaline membrane; B — whole chelicera; C — The palp, antero and posterodorsal views; D — Detail of the modified setae of the genu; E — Female anterogynal plate; F — laterogynal plate; G — mesogynal plate.
</paragraph>
</caption>
<caption id="DF43666FFFFE80529CE474465FEEF8CD" ID-DOI="http://doi.org/10.5281/zenodo.4640234" ID-Zenodo-Dep="4640234" httpUri="https://zenodo.org/record/4640234/files/figure.png" pageId="6" pageNumber="274" startId="6.[159,171,1858,1877]" targetBox="[269,1319,357,1807]" targetPageId="6">
<paragraph id="8B8336E7FFFE80529CE474465FEEF8CD" blockId="6.[159,1428,1857,1904]" pageId="6" pageNumber="274">
FIGURE 4:
<taxonomicName id="4C3C4D64FFFE80529D7D74455C8BF8E9" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[262,449,1857,1876]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="6" pageNumber="274" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFFE80529D7D74455C8BF8E9" box="[262,449,1857,1876]" italics="true" pageId="6" pageNumber="274">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<emphasis id="B948EAF5FFFE80529DBD74455CB6F8E9" bold="true" box="[454,508,1857,1876]" pageId="6" pageNumber="274">
<taxonomicNameLabel id="A27B578EFFFE80529DBD74455CB6F8E9" box="[454,508,1857,1876]" pageId="6" pageNumber="274" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A male genital plates; Anal plates: B female; C male; D Set of modified sensillae (Hallers organ); E Leg I tips of the female; F Lleg II male.
</paragraph>
</caption>
<caption id="DF43666FFFFF80539CE475DF59FCF8B4" ID-DOI="http://doi.org/10.5281/zenodo.4640236" ID-Zenodo-Dep="4640236" httpUri="https://zenodo.org/record/4640236/files/figure.png" pageId="7" pageNumber="275" startId="7.[159,171,1755,1774]" targetBox="[277,1305,450,1713]" targetPageId="7">
<paragraph id="8B8336E7FFFF80539CE475DF59FCF8B4" blockId="7.[159,1429,1754,1801]" pageId="7" pageNumber="275">
FIGURE 5:
<taxonomicName id="4C3C4D64FFFF80539D7C75DE5C88F950" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[263,450,1754,1773]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="7" pageNumber="275" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFFF80539D7C75DE5C88F950" box="[263,450,1754,1773]" italics="true" pageId="7" pageNumber="275">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<emphasis id="B948EAF5FFFF80539DB375DE5CB5F950" bold="true" box="[456,511,1754,1773]" pageId="7" pageNumber="275">
<taxonomicNameLabel id="A27B578EFFFF80539DB375DE5CB5F950" box="[456,511,1754,1773]" pageId="7" pageNumber="275" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
SEM images: A Thons organ, peritreme and stigma; B Thons organ; C detail of peritreme and stigma; D details of funnel structure of Thons organ; E female genital plates; F male genital plates.
</paragraph>
</caption>
<caption id="DF43666FFFF0805C9D2971F359A8FCB7" ID-DOI="http://doi.org/10.5281/zenodo.4640238" ID-Zenodo-Dep="4640238" box="[338,1250,758,778]" httpUri="https://zenodo.org/record/4640238/files/figure.png" pageId="8" pageNumber="276" startId="8.[338,350,759,778]" targetBox="[234,1347,307,716]" targetPageId="8">
<paragraph id="8B8336E7FFF0805C9D2971F359A8FCB7" blockId="8.[338,1250,758,778]" box="[338,1250,758,778]" pageId="8" pageNumber="276">
FIGURE 6:
<taxonomicName id="4C3C4D64FFF0805C9DCC71F25F3BFCB4" authority="Vázquez &amp; de Araújo &amp; Feres, 2016" authorityName="Vázquez &amp; de Araújo &amp; Feres" authorityYear="2016" box="[439,625,758,777]" class="Arachnida" family="Neothyridae" genus="Diplothyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="8" pageNumber="276" phylum="Arthropoda" rank="species" species="lehtineni" status="sp. nov.">
<emphasis id="B948EAF5FFF0805C9DCC71F25F3BFCB4" box="[439,625,758,777]" italics="true" pageId="8" pageNumber="276">Diplothyrus lehtineni</emphasis>
</taxonomicName>
<emphasis id="B948EAF5FFF0805C9E0D71F25FE0FCB4" bold="true" box="[630,682,758,777]" pageId="8" pageNumber="276">
<taxonomicNameLabel id="A27B578EFFF0805C9E0D71F25FE0FCB4" box="[630,682,758,777]" pageId="8" pageNumber="276" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A-B Phase contrast images of Leg I tip and Hallers organ
</paragraph>
</caption>
<subSubSection id="C326656CFFF0805C9CBE704F5877FBD7" pageId="8" pageNumber="276" type="materials_examined">
<paragraph id="8B8336E7FFF0805C9CBE704F5C49FC3B" blockId="8.[159,762,843,1949]" pageId="8" pageNumber="276">
<emphasis id="B948EAF5FFF0805C9CBE704F5FB0FCDE" bold="true" box="[197,762,843,867]" pageId="8" pageNumber="276">
<typeStatus id="54878845FFF0805C9CBE704F5C7DFCDE" box="[197,311,843,867]" pageId="8" pageNumber="276" type="holotype">Holotype</typeStatus>
and male
<typeStatus id="54878845FFF0805C9DB4704F5F73FCDE" box="[463,569,843,867]" pageId="8" pageNumber="276" type="paratype">paratype</typeStatus>
measurements
</emphasis>
(Table 1)
</paragraph>
<paragraph id="8B8336E7FFF0805C9CBE709F5F95FAAC" blockId="8.[159,762,843,1949]" pageId="8" pageNumber="276">Female: Idiosoma. Length 1851, width 1257. Palps (total length 655), trochanter 77, femur 213, genu 145, tibiotarsus 218. Leg I (total length 2009), coxa 167, trochanter 200, femur 495, genu 286, tibia 362, tarsus 498. Leg II (total length 1624), coxa 138, trochanter 212, femur 357, genu 235, tíbia 265, basitarsus 157, telotarsus 257. Leg III (total length 1710), coxa 137, trochanter 214, femur 384, genu 253, tibia 276, basitarsus 164, telotarsus 279. Leg IV (total length 2192), coxa 127, trochanter 288, femur 510, genu 323, tibia 367, basitarsus 207, telotarsus 367.</paragraph>
<paragraph id="8B8336E7FFF0805C9CBE76225F95F920" blockId="8.[159,762,843,1949]" pageId="8" pageNumber="276">Male: Idiosoma — Length 1852, width 1230. Palps (total length 663), trochanter 80, femur 222, genu 137, tibiotarsus 222. Leg I (total length 2068), coxa 158, trochanter 197, femur 501, genu 324, tibia 392, tarsus 494. Leg II (total length 1699), coxa 145, trochanter 178, femur 397, genu 253, tibia 302, basitarsus 172, telotarsus 250. Leg III (total length 1738), coxa 134, trochanter 225, femur 390, genu 244, tibia 296, basitarsus 180, telotarsus 267. Leg IV (total length 2228), coxa 133, trochanter 271, femur 486, genu 335, tibia 380, basitarsus 245, telotarsus 375.</paragraph>
<paragraph id="8B8336E7FFF0805C9CBE75B65873FC3B" blockId="8.[159,762,843,1949]" lastBlockId="8.[826,1429,843,1130]" pageId="8" pageNumber="276">
Material examined —
<typeStatus id="54878845FFF0805C9D9B75B65F05F977" box="[480,591,1714,1738]" pageId="8" pageNumber="276" type="holotype">Holotype</typeStatus>
<materialsCitation id="3B543CBAFFF0805C9E2675B75CB5F88E" ID-GBIF-Occurrence="3077738303" collectingDate="2011-11-03" collectorName="Serra das Andorinhas &amp; A. B. Bonaldo" country="Brazil" latitude="-6.2249722" location="Geraldo do Araguaia" longLatPrecision="1" longitude="-48.4345" pageId="8" pageNumber="276" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D3AFD6EFFF0805C9E2675B75F21F977" box="[605,619,1715,1738]" pageId="8" pageNumber="276" type="female"></specimenCount>
.
<collectingCountry id="F32B7677FFF0805C9EEA75B65FBFF977" box="[657,757,1714,1738]" name="Brazil" pageId="8" pageNumber="276">BRAZIL</collectingCountry>
, ParAE, Sªo
<location id="8EE3603CFFF0805C9D5E75D15F78F950" LSID="urn:lsid:plazi:treatment:039587F1FFFB805C9D5375085877FBD7:8EE3603CFFF0805C9D5E75D15F78F950" box="[293,562,1749,1773]" country="Brazil" latitude="-6.2249722" longLatPrecision="1" longitude="-48.4345" name="Geraldo do Araguaia" pageId="8" pageNumber="276">Geraldo do Araguaia</location>
,
<collectorName id="26C95331FFF0805C9E3375D15C40F8AD" pageId="8" pageNumber="276">Serra das Andorinhas</collectorName>
, coll.
<collectorName id="26C95331FFF0805C9D1075FC5F47F8AD" box="[363,525,1784,1808]" pageId="8" pageNumber="276">A.B. Bonaldo</collectorName>
<emphasis id="B948EAF5FFF0805C9E6075FD5F1DF8AD" box="[539,599,1784,1808]" italics="true" pageId="8" pageNumber="276">et al.</emphasis>
,
<geoCoordinate id="EE085020FFF0805C9E1775FC5FB0F8AD" box="[620,762,1784,1808]" degrees="06" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="276" precision="1" seconds="29.9" value="-6.2249722">06°1329.9&quot;S</geoCoordinate>
<geoCoordinate id="EE085020FFF0805C9CE4741F5C72F88E" box="[159,312,1819,1843]" degrees="48" direction="west" minutes="26" orientation="longitude" pageId="8" pageNumber="276" precision="1" seconds="04.2" value="-48.4345">48°2604.2&quot;W</geoCoordinate>
,
<date id="FF821027FFF0805C9D30741F5CB5F88E" box="[331,511,1819,1843]" pageId="8" pageNumber="276" value="2011-11-03">
<collectingDate id="EFC6E9CFFFF0805C9D30741F5CB5F88E" box="[331,511,1819,1843]" pageId="8" pageNumber="276" value="2011-11-03">03 Nov. 2011</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B543CBAFFF0805C9E69741F5D9FF8C7" ID-GBIF-Occurrence="3077738304" collectingDate="2011-11-03" collectorName="Serra das Andorinhas &amp; A. B. Bonaldo" country="Brazil" latitude="-6.2249722" location="Winkler" longLatPrecision="1" longitude="-48.4345" pageId="8" pageNumber="276" specimenCount="2" typeStatus="paratype">
<location id="8EE3603CFFF0805C9E69741F5F3AF88E" LSID="urn:lsid:plazi:treatment:039587F1FFFB805C9D5375085877FBD7:8EE3603CFFF0805C9E69741F5F3AF88E" box="[530,624,1819,1843]" name="Winkler" pageId="8" pageNumber="276">Winkler</location>
extraction.
<typeStatus id="54878845FFF0805C9CE4743B5C5CF8EA" box="[159,278,1855,1879]" pageId="8" pageNumber="276" type="paratype">Paratypes</typeStatus>
, same location and collectors as for
<typeStatus id="54878845FFF0805C9EC7743A5D9FF8C7" pageId="8" pageNumber="276" type="holotype">holotype</typeStatus>
</materialsCitation>
,
<specimenCount id="9D3AFD6EFFF0805C9C9E74665C4BF8C4" box="[229,257,1890,1914]" pageId="8" pageNumber="276" type="female">4♀</specimenCount>
,
<specimenCount id="9D3AFD6EFFF0805C9D6874655C7CF8C4" box="[275,310,1889,1913]" pageId="8" pageNumber="276" type="male">5♂</specimenCount>
,
<geoCoordinate id="EE085020FFF0805C9D3374655CA9F8C4" box="[328,483,1889,1913]" degrees="06" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="276" precision="1" seconds="10.47" value="-6.219575">06°1310.47&quot;S</geoCoordinate>
<geoCoordinate id="EE085020FFF0805C9D9574655FDCF8C4" box="[494,662,1889,1913]" degrees="48" direction="west" minutes="26" orientation="longitude" pageId="8" pageNumber="276" precision="1" seconds="17.97" value="-48.438328">48°2617.97&quot;W</geoCoordinate>
,
<date id="FF821027FFF0805C9EDF74665D9DF820" pageId="8" pageNumber="276" value="2011-10-30">30 Oct. 2011</date>
;
<specimenCount id="9D3AFD6EFFF0805C9C9674815C42F821" box="[237,264,1925,1949]" pageId="8" pageNumber="276" type="female">1♀</specimenCount>
,
<specimenCount id="9D3AFD6EFFF0805C9D6474815C0BF821" box="[287,321,1925,1949]" pageId="8" pageNumber="276" type="male">1♂</specimenCount>
,
<geoCoordinate id="EE085020FFF0805C9D2374805CACF821" box="[344,486,1924,1948]" degrees="06" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="276" precision="1" seconds="08.1" value="-6.218917">06°1308.1&quot;S</geoCoordinate>
<geoCoordinate id="EE085020FFF0805C9D8F74805FC4F821" box="[500,654,1924,1948]" degrees="48" direction="west" minutes="26" orientation="longitude" pageId="8" pageNumber="276" precision="1" seconds="17.4" value="-48.438168">48°2617.4&quot;W</geoCoordinate>
,
<date id="FF821027FFF0805C9EDA74815E38FCDE" box="[673,882,843,1949]" pageId="8" pageNumber="276" value="2011-10-31">31 Oct. 2011</date>
;
<specimenCount id="9D3AFD6EFFF0805C9FF9704F5EEEFCDF" box="[898,932,843,867]" pageId="8" pageNumber="276" type="male">1♂</specimenCount>
,
<geoCoordinate id="EE085020FFF0805C9FCD704F590EFCDE" box="[950,1092,843,867]" degrees="06" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="276" precision="1" seconds="29.9" value="-6.2249722">06°1329.9&quot;S</geoCoordinate>
<geoCoordinate id="EE085020FFF0805C9835704F59ADFCDE" box="[1102,1255,843,867]" degrees="48" direction="west" minutes="26" orientation="longitude" pageId="8" pageNumber="276" precision="1" seconds="04.2" value="-48.4345">48°2604.2&quot;W</geoCoordinate>
,
<date id="FF821027FFF0805C988D704F58DBFCDE" box="[1270,1425,843,867]" pageId="8" pageNumber="276" value="2011-11-03">03 Nov 2011</date>
;
<specimenCount id="9D3AFD6EFFF0805C9F41706A5E1FFC38" box="[826,853,878,902]" pageId="8" pageNumber="276" type="female">1♀</specimenCount>
,
<geoCoordinate id="EE085020FFF0805C9F18706A5EBAFC3B" box="[867,1008,878,902]" degrees="06" direction="south" minutes="13" orientation="latitude" pageId="8" pageNumber="276" precision="1" seconds="36.9" value="-6.226917">06°1336.9&quot;S</geoCoordinate>
<geoCoordinate id="EE085020FFF0805C9F8C706A59DBFC3B" box="[1015,1169,878,902]" degrees="48" direction="west" minutes="30" orientation="longitude" pageId="8" pageNumber="276" precision="1" seconds="02.1" value="-48.500584">48°3002.1&quot;W</geoCoordinate>
,
<date id="FF821027FFF0805C98E7706A587FFC3B" box="[1180,1333,878,902]" pageId="8" pageNumber="276" value="2011-11-05">05 Nov. 2011</date>
.
</paragraph>
<paragraph id="8B8336E7FFF0805C9F1B709E5ED7FC45" blockId="8.[826,1429,843,1130]" pageId="8" pageNumber="276">
<materialsCitation id="3B543CBAFFF0805C9F1B709E5EC1FC68" ID-GBIF-Occurrence="3077738302" collectionCode="MPEG" pageId="8" pageNumber="276" specimenCount="1" typeStatus="holotype">
Deposition of types —
<typeStatus id="54878845FFF0805C98E0709E5840FC0F" box="[1179,1290,922,946]" pageId="8" pageNumber="276" type="holotype">Holotype</typeStatus>
female in
<collectionCode id="ED2DAE22FFF0805C9F4170B95EC1FC68" box="[826,907,957,981]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="8" pageNumber="276">MPEG</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B543CBAFFF0805C9FED70B95ED3FC45" ID-GBIF-Occurrence="3077738301" collectionCode="MPEG, DZSJRP" pageId="8" pageNumber="276" specimenCount="1" typeStatus="paratype">
<typeStatus id="54878845FFF0805C9FED70B95940FC68" box="[918,1034,957,981]" pageId="8" pageNumber="276" type="paratype">Paratypes</typeStatus>
females and males in
<collectionCode id="ED2DAE22FFF0805C996970B9582AFC68" box="[1298,1376,957,981]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="8" pageNumber="276">MPEG</collectionCode>
and
<collectionCode id="ED2DAE22FFF0805C9F4170E45ED3FC45" box="[826,921,992,1016]" pageId="8" pageNumber="276">DZSJRP</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8336E7FFF0805C9F1B77085877FBD7" blockId="8.[826,1429,843,1130]" pageId="8" pageNumber="276">
Etymology — The species is dedicated to Dr Pekka
<collectionCode id="ED2DAE22FFF0805C9FF077345ED7FBFA" box="[907,925,1072,1095]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="276">T</collectionCode>
. Lehtinen, who has made significant contributions to the knowledge of
<taxonomicName id="4C3C4D64FFF0805C98DE77565870FBD7" authorityName="Thon" authorityYear="1905" box="[1189,1338,1106,1130]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Holothyrida" pageId="8" pageNumber="276" phylum="Arthropoda" rank="order">Holothyrida</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>