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<document id="5BE53BBBE906B2728AF1E6362A67F114" ID-DOI="10.5281/zenodo.169954" ID-GBIF-Dataset="8f46c5d2-179a-4553-a1ee-3a190b807c5d" ID-ISSN="1175­5326" ID-Zenodo-Dep="169954" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459748806134" checkinUser="plazi" docAuthor="Willassen, Endre" docDate="2005" docId="039587E1FFFB576D887C6AC4FC91F8E8" docLanguage="en" docName="zt01049p032.pdf" docOrigin="Zootaxa 1049" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Diamesa aculeata Willassen, 2005, new species" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FFACFF99FFFD576A89746B62FFC9FFBC" masterDocTitle="New species of Diamesa (Diptera: Chironomidae) from Tibet: conspecific males and females associated with mitochondrial DNA" masterLastPageNumber="32" masterPageNumber="19" pageNumber="25" updateTime="1698211182002" updateUser="plazi">
<mods:mods id="871DA1F8BAB1DA18552CB8D901F25EF2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CF96CE61A3A44957BA7BEAA824DB8D2C">
<mods:title id="3126E7F1E56DB810344446B0F16906DC">New species of Diamesa (Diptera: Chironomidae) from Tibet: conspecific males and females associated with mitochondrial DNA</mods:title>
</mods:titleInfo>
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<mods:namePart id="7E6F790193C97ABDF63FBC5DC6793C9A">Willassen, Endre</mods:namePart>
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<mods:title id="D9E7B4FECAC73B04AFBEE6C7ACA5D0C9">Zootaxa</mods:title>
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<mods:part id="E0E125EE245F4E62F13FFF4CEE832CC8">
<mods:date id="F02701BF6BC17FDD2CDE37C4B0C2005C">2005</mods:date>
<mods:detail id="C0D94F1DCF0A4DFC72B0A509ECEBECAC" type="volume">
<mods:number id="7534291F69A422DCC120482C5B3969EF">1049</mods:number>
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<mods:classification id="B5A4ACA36BE9A8E97BA213FF552206E6">journal article</mods:classification>
<mods:identifier id="D40734CF7017EAFC0E3E9F0E7DD6397F" type="DOI">10.5281/zenodo.169954</mods:identifier>
<mods:identifier id="F45DED33354E7F548F1A220ACC385024" type="GBIF-Dataset">8f46c5d2-179a-4553-a1ee-3a190b807c5d</mods:identifier>
<mods:identifier id="82CD44B4108BA4908C0D21DC9B03FC54" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="E600F6A85FD3379ED6D00FDE1BF5B61B" type="Zenodo-Dep">169954</mods:identifier>
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<treatment id="039587E1FFFB576D887C6AC4FC91F8E8" ID-DOI="http://doi.org/10.5281/zenodo.6266254" ID-GBIF-Taxon="119337182" ID-Zenodo-Dep="6266254" LSID="urn:lsid:plazi:treatment:039587E1FFFB576D887C6AC4FC91F8E8" httpUri="http://treatment.plazi.org/id/039587E1FFFB576D887C6AC4FC91F8E8" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
<subSubSection id="C326657CFFFB576C887C6AC4FE57FE5F" pageId="6" pageNumber="25" type="nomenclature">
<paragraph id="8B8336F7FFFB576C887C6AC4FDBEFE7C" blockId="6.[264,631,422,483]" box="[264,631,422,448]" pageId="6" pageNumber="25">
<heading id="D0CB819BFFFB576C887C6AC4FDBEFE7C" bold="true" box="[264,631,422,448]" fontSize="11" level="1" pageId="6" pageNumber="25" reason="1">
<emphasis id="B948EAE5FFFB576C887C6AC4FDBEFE7C" bold="true" box="[264,631,422,448]" pageId="6" pageNumber="25">
<taxonomicName id="4C3C4D74FFFB576C887C6AC4FE16FE7C" ID-CoL="359CN" box="[264,479,422,448]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="aculeata" status="sp. nov.">
<emphasis id="B948EAE5FFFB576C887C6AC4FE16FE7C" bold="true" box="[264,479,422,448]" italics="true" pageId="6" pageNumber="25">Diamesa aculeata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B579EFFFB576C88936AC4FDBEFE7C" box="[487,631,422,448]" pageId="6" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8336F7FFFB576C887C6AABFE57FE5F" blockId="6.[264,631,422,483]" box="[264,414,457,483]" pageId="6" pageNumber="25">
(
<figureCitation id="13072A72FFFB576C88656AABFE5FFE5F" box="[273,406,457,483]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="6" pageNumber="25">Figs. 2A–E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326657CFFFB576C887C696EFB9BFDF2" pageId="6" pageNumber="25" type="materials_examined">
<paragraph id="8B8336F7FFFB576C887C696EFB9BFDF2" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
<typeStatus id="54878855FFFB576C887C696EFEBCFD9A" box="[264,373,524,550]" pageId="6" pageNumber="25" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F32B7667FFFB576C88BB696EFDD3FD9A" box="[463,538,524,550]" name="China" pageId="6" pageNumber="25">China</collectingCountry>
: Tibet, Rongbuk,
<quantity id="4CC49B12FFFB576C8B8D696EFC9EFD9A" box="[761,855,524,550]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="25" unit="m" value="5000.0">5000 m</quantity>
a.sl.,
<date id="FF821037FFFB576C8AD2696EFBE6FD9A" box="[934,1071,524,550]" pageId="6" pageNumber="25" value="1993-07-07">7.Jul. 1993</date>
, leg. T. Solhøy, det. E.Willassen, EW36. In Academica Sinica, Zoological Institute, Beijing.
</paragraph>
</subSubSection>
<subSubSection id="C326657CFFFB576C884C693EFDBAFD52" pageId="6" pageNumber="25" type="etymology">
<paragraph id="8B8336F7FFFB576C884C693EFBCDFD22" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
<emphasis id="B948EAE5FFFB576C884C693EFE7CFDC9" box="[312,437,604,629]" italics="true" pageId="6" pageNumber="25">Etymology</emphasis>
: The species name is from Latin
<emphasis id="B948EAE5FFFB576C8A32693EFC7FFDC9" box="[838,950,604,629]" italics="true" pageId="6" pageNumber="25">aculeatus</emphasis>
, sharp­pointed, referring to the cuspidate microtrichia on the mesal edge of the male gonocoxite.
</paragraph>
<paragraph id="8B8336F7FFFB576C884C69CEFDBAFD52" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
<emphasis id="B948EAE5FFFB576C884C69CEFE61FD79" box="[312,424,684,709]" italics="true" pageId="6" pageNumber="25">Genetics:</emphasis>
Genbank accession number
<accessionNumber id="946FAB14FFFB576C8A6B69CEFC60FD7A" box="[799,937,684,710]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM051233" pageId="6" pageNumber="25">
AM
<date id="FF821037FFFB576C8A3B69CEFC60FD7A" box="[847,937,684,710]" pageId="6" pageNumber="25" value="33-05-12">051233</date>
</accessionNumber>
, mitochondrial COII gene for cytochrome oxidase subunit II.
</paragraph>
</subSubSection>
<subSubSection id="C326657CFFFB576D884C699EFD0BFA20" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25" type="description">
<paragraph id="8B8336F7FFFB576C884C699EFDD5FCAA" blockId="6.[264,1324,524,1406]" box="[312,540,764,790]" pageId="6" pageNumber="25">
<emphasis id="B948EAE5FFFB576C884C699EFE0DFCA9" box="[312,452,764,789]" italics="true" pageId="6" pageNumber="25">Male imago</emphasis>
(
<emphasis id="B948EAE5FFFB576C88A1699EFE2DFCA9" box="[469,484,764,789]" italics="true" pageId="6" pageNumber="25">n</emphasis>
= 1)
</paragraph>
<paragraph id="8B8336F7FFFB576C884C6846FD71FBBA" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
Head: Antenna plumose with 13 flagellomeres; longest antennal seta longer than ultimate flagellomere; AR 1.2; pedicel large, with 3 setae. Eyes moderately projecting dorsomesally; with pubescence between ommatidia. Coronal suture complete. Temporal setae including 3 orbitals, 17 verticals, and 9? postorbitals. Clypeus
<quantity id="4CC49B12FFFB576C8D5D68FEFB4CFC0A" box="[1065,1157,924,950]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="25" unit="m" value="130.0">130 m</quantity>
long,
<quantity id="4CC49B12FFFB576C8DBB68FEFAE2FC0A" box="[1231,1323,924,950]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="25" unit="m" value="107.0">107 m</quantity>
wide; with 12 setae. Palpomere lengths (m): 37, 85, 133, 126, 211. Tentorium with conspicuous anterolateral projection.
</paragraph>
<paragraph id="8B8336F7FFFB576C884C6F76FD3EFB92" blockId="6.[264,1324,524,1406]" box="[312,759,1044,1070]" pageId="6" pageNumber="25">Thorax damaged in DNA preparation.</paragraph>
<paragraph id="8B8336F7FFFB576C884C6F5EFDEDFB40" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
Wing: Length
<quantity id="4CC49B12FFFB576C88886F5EFDBCFBEA" box="[508,629,1084,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.17" pageId="6" pageNumber="25" unit="mm" value="3.17">3.17 mm</quantity>
. VR 0.88. Membrane without setae, punctuation of microtrichia distinct at 100x magnification. Costa produced beyond R4+5. R with 15 setae, R1 with 12 setae and 2 sensillae campaniformia, R2+3 with 2 sensillae campaniformia, R4+5 with 3 setae and 1 sensilla campaniformia. Alula without setae. Squama with marginal fringe of about 35 setae.
</paragraph>
<paragraph id="8B8336F7FFFB576C884C6E68FB44FAC6" blockId="6.[264,1324,524,1406]" pageId="6" pageNumber="25">
Legs: Comb on ti3 with 15 setae. Tarsi 1–3 respectively with following numbers of apical/preapical pseudospurs: 2/0, 2/0, ­/­ on p1; 2/7, 2/5, 2/0 on p2; 2/10, 2/5, 2/0 on p3.
<collectingCountry id="F32B7667FFFB576C887C6E02FE8AFAC6" box="[264,323,1376,1402]" name="India" pageId="6" pageNumber="25">Hind</collectingCountry>
leg with about 25 sensillae chaetica distributed from 0.45 to 0.52 of ta1.
</paragraph>
<caption id="DF43667FFFFB576C887C6ED0FB10FA76" ID-Table-UUID="DF43667FFFFB576C887C6ED0FB10FA76" box="[264,1241,1458,1482]" httpUri="http://table.plazi.org/id/DF43667FFFFB576C887C6ED0FB10FA76" pageId="6" pageNumber="25" targetBox="[272,1290,1515,1660]" targetIsTable="true" targetPageId="6">
<paragraph id="8B8336F7FFFB576C887C6ED0FB10FA76" blockId="6.[264,1241,1458,1482]" box="[264,1241,1458,1482]" pageId="6" pageNumber="25">
<emphasis id="B948EAE5FFFB576C887C6ED0FEB2FA76" bold="true" box="[264,379,1458,1482]" pageId="6" pageNumber="25">TABLE 4.</emphasis>
Lengths (in µm) and proportions of legs in male
<taxonomicName id="4C3C4D74FFFB576C8AFD6ED1FB82FA76" box="[905,1099,1459,1482]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="aculeata" status="sp. nov.">
<emphasis id="B948EAE5FFFB576C8AFD6ED1FB82FA76" box="[905,1099,1459,1482]" italics="true" pageId="6" pageNumber="25">Diamesa aculeata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B579EFFFB576C8D256ED1FB1DFA76" box="[1105,1236,1459,1482]" pageId="6" pageNumber="25" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8B8336F7FFFB576C88026E89FAC3F9CE" pageId="6" pageNumber="25">
<table id="F93CC457FFFBA89588646E89FAC3F9C0" box="[272,1290,1515,1660]" gridcols="11" gridrows="4" pageId="6" pageNumber="25">
<tr id="350C34B5FFFBA89588646E89FAC3F9BB" box="[272,1290,1515,1543]" gridrow="0" pageId="6" pageNumber="25" rowspan-0="1">
<th id="76DD5DC9FFFBA895881F6E89FE5FF9BB" box="[363,406,1515,1543]" gridcol="1" gridrow="0" pageId="6" pageNumber="25">fe</th>
<th id="76DD5DC9FFFBA89588AD6E89FDCDF9BB" box="[473,516,1515,1543]" gridcol="2" gridrow="0" pageId="6" pageNumber="25">ti</th>
<th id="76DD5DC9FFFBA8958B336E89FDBBF9BB" box="[583,626,1515,1543]" gridcol="3" gridrow="0" pageId="6" pageNumber="25">ta1</th>
<th id="76DD5DC9FFFBA8958BC56E89FD18F9BB" box="[689,721,1515,1543]" gridcol="4" gridrow="0" pageId="6" pageNumber="25">ta2</th>
<th id="76DD5DC9FFFBA8958A7F6E89FCE2F9BB" box="[779,811,1515,1543]" gridcol="5" gridrow="0" pageId="6" pageNumber="25">ta3</th>
<th id="76DD5DC9FFFBA8958A116E89FC4DF9BB" box="[869,900,1515,1543]" gridcol="6" gridrow="0" pageId="6" pageNumber="25">ta4</th>
<th id="76DD5DC9FFFBA8958ACA6E89FC17F9BB" box="[958,990,1515,1543]" gridcol="7" gridrow="0" pageId="6" pageNumber="25">ta5</th>
<th id="76DD5DC9FFFBA8958D6F6E89FB89F9BB" box="[1051,1088,1515,1543]" gridcol="8" gridrow="0" pageId="6" pageNumber="25">LR</th>
<th id="76DD5DC9FFFBA8958DF46E89FB6CF9BB" box="[1152,1189,1515,1543]" gridcol="9" gridrow="0" pageId="6" pageNumber="25">BV</th>
<th id="76DD5DC9FFFBA8958D916E89FAC3F9BB" box="[1253,1290,1515,1543]" gridcol="10" gridrow="0" pageId="6" pageNumber="25">SV</th>
</tr>
<tr id="350C34B5FFFBA89588646D73FAC3F992" box="[272,1290,1553,1582]" gridrow="1" pageId="6" pageNumber="25">
<th id="76DD5DC9FFFBA89588646D73FEEDF992" box="[272,292,1553,1582]" gridcol="0" gridrow="1" pageId="6" pageNumber="25">p1</th>
<td id="76DD5DC9FFFBA895881F6D73FE5FF992" box="[363,406,1553,1582]" gridcol="1" gridrow="1" pageId="6" pageNumber="25">1229</td>
<td id="76DD5DC9FFFBA89588AD6D73FDCDF992" box="[473,516,1553,1582]" gridcol="2" gridrow="1" pageId="6" pageNumber="25">1536</td>
<td id="76DD5DC9FFFBA8958B336D73FDBBF992" box="[583,626,1553,1582]" gridcol="3" gridrow="1" pageId="6" pageNumber="25">1087</td>
<td id="76DD5DC9FFFBA8958BC56D73FD18F992" box="[689,721,1553,1582]" gridcol="4" gridrow="1" pageId="6" pageNumber="25">521</td>
<td id="76DD5DC9FFFBA8958A7F6D73FCE2F992" box="[779,811,1553,1582]" gridcol="5" gridrow="1" pageId="6" pageNumber="25">­</td>
<td id="76DD5DC9FFFBA8958A116D73FC4DF992" box="[869,900,1553,1582]" gridcol="6" gridrow="1" pageId="6" pageNumber="25">­</td>
<td id="76DD5DC9FFFBA8958ACA6D73FC17F992" box="[958,990,1553,1582]" gridcol="7" gridrow="1" pageId="6" pageNumber="25">­</td>
<td id="76DD5DC9FFFBA8958D6F6D73FB89F992" box="[1051,1088,1553,1582]" gridcol="8" gridrow="1" pageId="6" pageNumber="25">0.71</td>
<td id="76DD5DC9FFFBA8958DF46D73FB6CF992" box="[1152,1189,1553,1582]" gridcol="9" gridrow="1" pageId="6" pageNumber="25">­</td>
<td id="76DD5DC9FFFBA8958D916D73FAC3F992" box="[1253,1290,1553,1582]" gridcol="10" gridrow="1" pageId="6" pageNumber="25">2.54</td>
</tr>
<tr id="350C34B5FFFBA89588646D5AFAC3F9E9" box="[272,1290,1592,1621]" gridrow="2" pageId="6" pageNumber="25">
<th id="76DD5DC9FFFBA89588646D5AFEEDF9E9" box="[272,292,1592,1621]" gridcol="0" gridrow="2" pageId="6" pageNumber="25">p2</th>
<td id="76DD5DC9FFFBA895881F6D5AFE5FF9E9" box="[363,406,1592,1621]" gridcol="1" gridrow="2" pageId="6" pageNumber="25">1252</td>
<td id="76DD5DC9FFFBA89588AD6D5AFDCDF9E9" box="[473,516,1592,1621]" gridcol="2" gridrow="2" pageId="6" pageNumber="25">1370</td>
<td id="76DD5DC9FFFBA8958B336D5AFDBBF9E9" box="[583,626,1592,1621]" gridcol="3" gridrow="2" pageId="6" pageNumber="25">685</td>
<td id="76DD5DC9FFFBA8958BC56D5AFD18F9E9" box="[689,721,1592,1621]" gridcol="4" gridrow="2" pageId="6" pageNumber="25">369</td>
<td id="76DD5DC9FFFBA8958A7F6D5AFCE2F9E9" box="[779,811,1592,1621]" gridcol="5" gridrow="2" pageId="6" pageNumber="25">208</td>
<td id="76DD5DC9FFFBA8958A116D5AFC4DF9E9" box="[869,900,1592,1621]" gridcol="6" gridrow="2" pageId="6" pageNumber="25">90</td>
<td id="76DD5DC9FFFBA8958ACA6D5AFC17F9E9" box="[958,990,1592,1621]" gridcol="7" gridrow="2" pageId="6" pageNumber="25">95</td>
<td id="76DD5DC9FFFBA8958D6F6D5AFB89F9E9" box="[1051,1088,1592,1621]" gridcol="8" gridrow="2" pageId="6" pageNumber="25">0.50</td>
<td id="76DD5DC9FFFBA8958DF46D5AFB6CF9E9" box="[1152,1189,1592,1621]" gridcol="9" gridrow="2" pageId="6" pageNumber="25">4.34</td>
<td id="76DD5DC9FFFBA8958D916D5AFAC3F9E9" box="[1253,1290,1592,1621]" gridcol="10" gridrow="2" pageId="6" pageNumber="25">3.83</td>
</tr>
<tr id="350C34B5FFFBA89588646D3DFAC3F9C0" box="[272,1290,1631,1660]" gridrow="3" pageId="6" pageNumber="25">
<th id="76DD5DC9FFFBA89588646D3DFEEDF9C0" box="[272,292,1631,1660]" gridcol="0" gridrow="3" pageId="6" pageNumber="25">p3</th>
<td id="76DD5DC9FFFBA895881F6D3DFE5FF9C0" box="[363,406,1631,1660]" gridcol="1" gridrow="3" pageId="6" pageNumber="25">1465</td>
<td id="76DD5DC9FFFBA89588AD6D3DFDCDF9C0" box="[473,516,1631,1660]" gridcol="2" gridrow="3" pageId="6" pageNumber="25">1583</td>
<td id="76DD5DC9FFFBA8958B336D3DFDBBF9C0" box="[583,626,1631,1660]" gridcol="3" gridrow="3" pageId="6" pageNumber="25">1087</td>
<td id="76DD5DC9FFFBA8958BC56D3DFD18F9C0" box="[689,721,1631,1660]" gridcol="4" gridrow="3" pageId="6" pageNumber="25">568</td>
<td id="76DD5DC9FFFBA8958A7F6D3DFCE2F9C0" box="[779,811,1631,1660]" gridcol="5" gridrow="3" pageId="6" pageNumber="25">284</td>
<td id="76DD5DC9FFFBA8958A116D3DFC4DF9C0" box="[869,900,1631,1660]" gridcol="6" gridrow="3" pageId="6" pageNumber="25">114</td>
<td id="76DD5DC9FFFBA8958ACA6D3DFC17F9C0" box="[958,990,1631,1660]" gridcol="7" gridrow="3" pageId="6" pageNumber="25">123</td>
<td id="76DD5DC9FFFBA8958D6F6D3DFB89F9C0" box="[1051,1088,1631,1660]" gridcol="8" gridrow="3" pageId="6" pageNumber="25">0.69</td>
<td id="76DD5DC9FFFBA8958DF46D3DFB6CF9C0" box="[1152,1189,1631,1660]" gridcol="9" gridrow="3" pageId="6" pageNumber="25">3.79</td>
<td id="76DD5DC9FFFBA8958D916D3DFAC3F9C0" box="[1253,1290,1631,1660]" gridcol="10" gridrow="3" pageId="6" pageNumber="25">2.80</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8336F7FFFB576D884C6DCCFD9EFD9A" blockId="6.[264,1321,1710,1896]" lastBlockId="7.[264,1324,284,1876]" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
Hypopygium: Sternapodeme arced. Pars ventralis moderately large (
<figureCitation id="13072A72FFFB576C8D0D6DCCFB13F974" box="[1145,1242,1710,1736]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="6" pageNumber="25">Fig. 2B</figureCitation>
); with longitudinal striae, but without microtrichia. Aedeagal lobes sickle­shaped, mesal margin well sclerotised. Basal plate of gonocoxite weakly developed. Basimedial setal cluster absent. Medial field almost completely devoid of microtrichia dorsally; dorsal edge of mesal margin (
<figureCitation id="13072A72FFFB576C88CE6C2CFDC6F8D4" box="[442,527,1870,1896]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="6" pageNumber="25">Fig.2A</figureCitation>
) with ornamentation of thornlike microtrichia that are particularly dense on the ventral side (
<figureCitation id="13072A72FFFA576D8B4E6A7EFD46FE8A" box="[570,655,284,310]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="7" pageNumber="26">Fig.2C</figureCitation>
); ventromesal margin of gonocoxite with setae and weak microtrichia (
<figureCitation id="13072A72FFFA576D88DF6A26FDC0FEE2" box="[427,521,324,350]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="7" pageNumber="26">Fig. 2B</figureCitation>
) Gonostyli somewhat flattened and twisted, apex with sclerotised point and 1 megaseta. Tergite IX divided into 2 oval protrusions, each with about 15 moderately long setae. Tergal bands relatively distinct, running from anterior margin of tergite IX to the free base of anal point. Anal point inserted between tergal protrusions; base broad with microtrichia; distal part slender and nude, apex slightly expanded with 1 distal peg­sensilla (
<figureCitation id="13072A72FFFA576D889F696EFD80FD9A" box="[491,585,524,550]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="7" pageNumber="26">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8B8336F7FFFA576D884C6956FCE7FDF2" blockId="7.[264,1324,284,1876]" box="[312,814,564,590]" pageId="7" pageNumber="26">
<emphasis id="B948EAE5FFFA576D884C6956FE29FDF1" box="[312,480,564,589]" italics="true" pageId="7" pageNumber="26">Female imago</emphasis>
(
<emphasis id="B948EAE5FFFA576D88856956FDC9FDF1" box="[497,512,564,589]" italics="true" pageId="7" pageNumber="26">n</emphasis>
= 1, putative association)
</paragraph>
<paragraph id="8B8336F7FFFA576D884C693EFB57FD52" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
Head: Antennae partly lost. Pedicel with 5 or 6 setae. Coronal suture complete. Temporal setae more or less contiguous, about 40 on each side, including 9 postorbitals. Eyes pubescent, moderately projecting dorsomesally. Clypeus
<quantity id="4CC49B12FFFA576D8A9069CEFBF6FD7A" box="[996,1087,684,710]" metricMagnitude="2" metricUnit="m" metricValue="1.67" pageId="7" pageNumber="26" unit="m" value="167.0">167 m</quantity>
long,
<quantity id="4CC49B12FFFA576D8DFC69CEFB2AFD7A" box="[1160,1251,684,710]" metricMagnitude="2" metricUnit="m" metricValue="1.52" pageId="7" pageNumber="26" unit="m" value="152.0">152 m</quantity>
wide, with 24 setae. Palps lost. Tentorium with conspicuous anterolateral projection.
</paragraph>
<paragraph id="8B8336F7FFFA576D884C699EFD27FCDA" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
Thorax: Antepronotum with 9 lateral setae. Acrostichals absent. Dorsocentrals
<quantity id="4CC49B12FFFA576D8DAD699EFADCFCAA" box="[1241,1301,764,790]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="7" pageNumber="26" unit="in" value="16.0">16 in</quantity>
1 row, except anteriorly; 9 prealars behind callus, and about setae on 40 scutellum. Protuberance of epimeron II with 8 setae.
</paragraph>
<paragraph id="8B8336F7FFFA576D884C6816FE83FB88" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
Wing: Length
<quantity id="4CC49B12FFFA576D888D6816FDA6FC32" box="[505,623,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="4.11" pageId="7" pageNumber="26" unit="mm" value="4.11">4.11 mm</quantity>
. VR 0.91. Punctuation of membrane visible at 125x magnification. Costa well produced beyond R4+5. R with 18 setae. R1 with 19 setae and 2 sensillae campaniformia. R2+3 with 2 sensillae campaniformia. R4+5 with 18 setae and 3 sensillae campaniformia. Alula with 5 setae. Squama with marginal fringe of about 60 setae.
</paragraph>
<paragraph id="8B8336F7FFFA576D884C6F20FB1AFBE0" blockId="7.[264,1324,284,1876]" box="[312,1235,1090,1116]" pageId="7" pageNumber="26">Legs: Lost except right hind femur and tibia. Comb of hind tibia with 15 setae.</paragraph>
<paragraph id="8B8336F7FFFA576D884C6F08FD0BFA20" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
Genitalia (
<figureCitation id="13072A72FFFA576D88B56F08FDE3FB38" box="[449,554,1130,1156]" captionStart="FIGURES 2 A – E" captionStartId="8.[264,370,1746,1767]" captionTargetBox="[274,1314,330,1702]" captionTargetId="figure@8.[269,1317,325,1708]" captionTargetPageId="8" captionText="FIGURES 2 A – E. Diamesa aculeata new species. A, male genitalia in dorsal view. B, ventral view of pars ventralis, aedeagal lobe, basal plate and ventromesal margin of gonocoxite IX. C, ventral view of cuspidate microtrichia along dorsomesal edge of medial field. D, apex of anal point. E, female genitalia in lateral view." httpUri="https://zenodo.org/record/169956/files/figure.png" pageId="7" pageNumber="26">Fig. 2E</figureCitation>
): Sternite VIII with about 20 short setae on each side. Gonocoxapodeme conspicuous with distinct anteromesal curve. Gonapophysis VIII with flap covering mesal part of ventolateral lobe. Seminal capsules elongate with long neck, capsule surface with weak granulation. Seminal ducts with few bends. Gonocoxite IX with laterally directed knoblike projection; about 15 weak setae confined to knob. Tergite IX completely divided into 2 plates, of which broader lateral parts are slightly more sclerotised; each plate with about 20 relatively short setae. Segment X slightly protruding laterad. Cerci rounded in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C326657CFFFA576D884C6EC8FC41F8B8" pageId="7" pageNumber="26" type="discussion">
<paragraph id="8B8336F7FFFA576D884C6EC8FC41F8B8" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
<emphasis id="B948EAE5FFFA576D884C6EC8FE57FA7F" box="[312,414,1450,1475]" italics="true" pageId="7" pageNumber="26">Remarks</emphasis>
: The male of
<taxonomicName id="4C3C4D74FFFA576D8B166EC8FD3AFA7F" box="[610,755,1450,1475]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B948EAE5FFFA576D8B166EC8FD3AFA7F" box="[610,755,1450,1475]" italics="true" pageId="7" pageNumber="26">D. aculeata</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="4C3C4D74FFFA576D8D386EC8FEA6FA50" authority="Edwards" authorityName="Edwards" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="bertrami">
<emphasis id="B948EAE5FFFA576D8D386EC8FAE2FA7F" box="[1100,1323,1450,1475]" italics="true" pageId="7" pageNumber="26">Diamesa bertrami</emphasis>
Edwards
</taxonomicName>
and also to the Himalayan
<taxonomicName id="4C3C4D74FFFA576D8BCA6EB0FC31FA50" authority="Pagast" authorityName="Pagast" box="[702,1016,1490,1516]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="kasaulica">
<emphasis id="B948EAE5FFFA576D8BCA6EB0FC68FA57" box="[702,929,1490,1515]" italics="true" pageId="7" pageNumber="26">Diamesa kasaulica</emphasis>
Pagast
</taxonomicName>
described by
<bibRefCitation id="EFAD4B06FFFA576D8DD76EB0FE93F9A8" author="Serra-Tosio" pageId="7" pageNumber="32" refString="Serra-Tosio, B. (1983) Nouveaux Diamesinae de la Palearctide meridionale et orientale (Diptera, Chironomidae). Spixiana, 6, 1 - 26." type="journal article" year="1983">Serra­Tosio (1983)</bibRefCitation>
. The ornamentation of the mesal edge and otherwise smooth medial field may serve as diagnostic features of male
<taxonomicName id="4C3C4D74FFFA576D8B056D40FD32F987" box="[625,763,1570,1595]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B948EAE5FFFA576D8B056D40FD32F987" box="[625,763,1570,1595]" italics="true" pageId="7" pageNumber="26">D. aculeata</emphasis>
</taxonomicName>
. Both
<taxonomicName id="4C3C4D74FFFA576D8A3F6D40FC1EF987" box="[843,983,1570,1595]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="bertrami">
<emphasis id="B948EAE5FFFA576D8A3F6D40FC1EF987" box="[843,983,1570,1595]" italics="true" pageId="7" pageNumber="26">D. bertrami</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4D74FFFA576D8D656D40FB6FF987" box="[1041,1190,1570,1595]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="kasaulica">
<emphasis id="B948EAE5FFFA576D8D656D40FB6FF987" box="[1041,1190,1570,1595]" italics="true" pageId="7" pageNumber="26">D. kasaulica</emphasis>
</taxonomicName>
have hairy eyes as opposed to the weakly pubescent eyes in
<taxonomicName id="4C3C4D74FFFA576D8A216D28FC29F9DF" box="[853,992,1610,1635]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B948EAE5FFFA576D8A216D28FC29F9DF" box="[853,992,1610,1635]" italics="true" pageId="7" pageNumber="26">D. aculeata</emphasis>
</taxonomicName>
. The only female specimen with pubescent eyes in the Rongbuk material presumably also belongs to
<taxonomicName id="4C3C4D74FFFA576D8DE26D10FAEAF937" box="[1174,1315,1650,1675]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B948EAE5FFFA576D8DE26D10FAEAF937" box="[1174,1315,1650,1675]" italics="true" pageId="7" pageNumber="26">D. aculeata</emphasis>
</taxonomicName>
. Antennae, palps, and most of the legs are lacking and the wings were preserved in a folded state. A short, partial Cox II sequence was recovered from the male of
<taxonomicName id="4C3C4D74FFFA576D8D1C6DA0FB3DF967" box="[1128,1268,1730,1755]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B948EAE5FFFA576D8D1C6DA0FB3DF967" box="[1128,1268,1730,1755]" italics="true" pageId="7" pageNumber="26">D. aculeata</emphasis>
</taxonomicName>
, but sequences were not obtained from the putative female.
</paragraph>
</subSubSection>
<subSubSection id="C326657CFFFA576D884C6C70FC91F8E8" pageId="7" pageNumber="26" type="materials_examined">
<paragraph id="8B8336F7FFFA576D884C6C70FC91F8E8" blockId="7.[264,1324,284,1876]" pageId="7" pageNumber="26">
<emphasis id="B948EAE5FFFA576D884C6C70FDE3F897" box="[312,554,1810,1835]" italics="true" pageId="7" pageNumber="26">Additional material:</emphasis>
<collectingCountry id="F32B7667FFFA576D8B466C70FDB4F890" box="[562,637,1810,1836]" name="China" pageId="7" pageNumber="26">China</collectingCountry>
: Tibet, Rongbuk,
<quantity id="4CC49B12FFFA576D8A266C70FC64F890" box="[850,941,1810,1836]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="26" unit="m" value="5000.0">5000 m</quantity>
a.sl.,
<date id="FF821037FFFA576D8A816C70FBB2F890" box="[1013,1147,1810,1836]" pageId="7" pageNumber="26" value="1993-07-09">9.Jul. 1993</date>
, Malaise Trap, leg. T. Solhøy, det. E. Willassen;
<specimenCount id="9D3AFD7EFFFA576D8BFA6C58FD3DF8E8" box="[654,756,1850,1876]" pageId="7" pageNumber="26" type="female">1 female</specimenCount>
(
<collectionCode id="ED2DAE32FFFA576D8A706C58FC82F8E8" box="[772,843,1850,1876]" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="7" pageNumber="26">IZAS</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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