480 lines
40 KiB
XML
480 lines
40 KiB
XML
<document ID-DOI="10.11646/zootaxa.5239.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7624101" ID-ZooBank="4AFDDAD0-55C4-4921-8A82-455DFB01E437" checkinTime="1675933316800" checkinUser="plazi" docAuthor="Spratt, David M." docDate="2023" docId="039587DFFF95FFA0FF68F8C5FDE2B7EC" docLanguage="en" docName="zootaxa.5239.2.2.pdf" docOrigin="Zootaxa 5239 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gongylonema bettongiae Spratt 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="216" masterDocId="FFACFFA7FF9CFFACFFFFFF94FFC5B521" masterDocTitle="Redescription of species of Gongylonema Molin, 1857 (Nematoda: Spiruroidea Gongylonematidae) parasitic in some Australian vertebrate hosts and description of three new species" masterLastPageNumber="220" masterPageNumber="204" pageNumber="213" updateTime="1676305748254" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Redescription of species of Gongylonema Molin, 1857 (Nematoda: Spiruroidea Gongylonematidae) parasitic in some Australian vertebrate hosts and description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Spratt, David M.</mods:namePart>
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<mods:affiliation>Australian National Wildlife Collection, CSIRO National Research Collections, Canberra, ACT, Australia.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5239</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>204</mods:start>
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<mods:end>220</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5239.2.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5239.2.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7624101</mods:identifier>
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<mods:identifier type="ZooBank">4AFDDAD0-55C4-4921-8A82-455DFB01E437</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7624115" ID-Zenodo-Dep="7624115" LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC" httpUri="http://treatment.plazi.org/id/039587DFFF95FFA0FF68F8C5FDE2B7EC" lastPageId="12" lastPageNumber="216" pageId="9" pageNumber="213">
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<subSubSection box="[151,554,1873,1900]" pageId="9" pageNumber="213" type="nomenclature">
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<paragraph blockId="9.[151,554,1873,1900]" box="[151,554,1873,1900]" pageId="9" pageNumber="213">
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||
<heading bold="true" box="[151,554,1873,1900]" fontSize="11" level="1" pageId="9" pageNumber="213" reason="1">
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<emphasis bold="true" box="[151,554,1873,1900]" pageId="9" pageNumber="213">
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<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[151,453,1873,1900]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="213" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov.">
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<emphasis bold="true" box="[151,453,1873,1900]" italics="true" pageId="9" pageNumber="213">Gongylonema bettongiae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[461,554,1873,1899]" pageId="9" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="214" pageId="9" pageNumber="213" type="materials_examined">
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<paragraph blockId="9.[151,1436,1940,2039]" pageId="9" pageNumber="213">
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<emphasis bold="true" box="[151,344,1940,1966]" pageId="9" pageNumber="213">Type specimens.</emphasis>
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<materialsCitation box="[349,622,1940,1968]" collectionCode="AHC" pageId="9" pageNumber="213" specimenCode="AHC 49241" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[349,453,1940,1966]" pageId="9" pageNumber="213">Holotype</typeStatus>
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<specimenCount box="[459,480,1941,1968]" pageId="9" pageNumber="213">♁</specimenCount>
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<specimenCode box="[484,622,1940,1966]" pageId="9" pageNumber="213">AHC 49241</specimenCode>
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</materialsCitation>
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,
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<typeStatus box="[632,722,1940,1966]" pageId="9" pageNumber="213">allotype</typeStatus>
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<specimenCount box="[728,749,1941,1967]" pageId="9" pageNumber="213" type="female">♀</specimenCount>
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AHC 49242, 10 ♁♁,
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<typeStatus box="[989,1096,1941,1967]" pageId="9" pageNumber="213">paratypes</typeStatus>
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AHC 49243,
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<specimenCount box="[1248,1323,1940,1967]" pageId="9" pageNumber="213" type="female">15 ♀♀</specimenCount>
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<typeStatus box="[1329,1436,1941,1967]" pageId="9" pageNumber="213">paratypes</typeStatus>
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AHC 49244, 10 ♁♁
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<typeStatus box="[385,492,1977,2003]" pageId="9" pageNumber="213">paratypes</typeStatus>
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QM G240161,
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<specimenCount box="[670,747,1976,2003]" pageId="9" pageNumber="213" type="female">10 ♀♀</specimenCount>
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<typeStatus box="[754,861,1977,2003]" pageId="9" pageNumber="213">paratypes</typeStatus>
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QM G240162, 9 ♁♁,
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<specimenCount box="[1116,1193,1976,2003]" pageId="9" pageNumber="213" type="female">22 ♀♀</specimenCount>
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<typeStatus box="[1200,1307,1977,2003]" pageId="9" pageNumber="213">paratypes</typeStatus>
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(N1337).
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</paragraph>
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<paragraph blockId="9.[151,1436,1940,2039]" box="[199,528,2012,2039]" pageId="9" pageNumber="213">
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<emphasis bold="true" box="[199,358,2012,2038]" pageId="9" pageNumber="213">
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<typeStatus box="[199,258,2012,2038]" pageId="9" pageNumber="213">Type</typeStatus>
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locality.
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</emphasis>
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<materialsCitation box="[364,524,2013,2039]" pageId="9" pageNumber="213">Kempton, Tas</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="10.[151,1437,150,2013]" box="[199,1181,150,176]" pageId="10" pageNumber="214">
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<emphasis bold="true" box="[199,322,150,176]" pageId="10" pageNumber="214">
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<typeStatus box="[199,258,150,176]" pageId="10" pageNumber="214">Type</typeStatus>
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host.
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</emphasis>
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<taxonomicName authority="(Desmarest)" baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[329,689,150,176]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="gaimardi">
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<emphasis box="[329,546,150,176]" italics="true" pageId="10" pageNumber="214">Bettongia gaimardi</emphasis>
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(Desmarest)
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</taxonomicName>
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(Marsupialia:
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<taxonomicName authorityName="Gray" authorityYear="1821" box="[855,978,150,176]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
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) (eastern bettong)
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</paragraph>
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<paragraph blockId="10.[151,1437,150,2013]" box="[199,645,186,213]" pageId="10" pageNumber="214">
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<emphasis bold="true" box="[199,393,186,212]" pageId="10" pageNumber="214">Site of infection.</emphasis>
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Oesophageal mucosa.
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</paragraph>
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<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
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<emphasis bold="true" box="[199,403,222,248]" pageId="10" pageNumber="214">Additional hosts.</emphasis>
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<taxonomicName authority="(Kerr)" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[412,721,222,248]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="tridactylus">
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<emphasis box="[412,642,222,248]" italics="true" pageId="10" pageNumber="214">Potorous tridactylus</emphasis>
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(Kerr)
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</taxonomicName>
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(Marsupialia:
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<taxonomicName authorityName="Gray" authorityYear="1821" box="[889,1012,222,248]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
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) (long-nosed potoroo);
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<taxonomicName authority="(J. Gray)" baseAuthorityName="J. Gray" class="Mammalia" family="Potoroidae" genus="Aepyprymnus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="rufescens">
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<emphasis italics="true" pageId="10" pageNumber="214">Aepyprymnus rufescens</emphasis>
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(J. Gray)
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</taxonomicName>
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(Marsupialia:
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<taxonomicName authorityName="Gray" authorityYear="1821" box="[540,663,258,284]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
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) (rufous rat kangaroo);
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<taxonomicName authority="(Kerr)" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[943,1276,258,284]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="vulpecula">
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<emphasis box="[943,1196,258,284]" italics="true" pageId="10" pageNumber="214">Trichosurus vulpecula</emphasis>
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(Kerr)
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</taxonomicName>
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(Marsupialia:
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<taxonomicName authorityName="Thomas" authorityYear="1888" box="[151,312,294,320]" class="Mammalia" family="Phalangeridae" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="family">Phalangeridae</taxonomicName>
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) (brushtail possum).
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</paragraph>
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||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
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<emphasis bold="true" box="[199,437,330,356]" pageId="10" pageNumber="214">Material examined.</emphasis>
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<materialsCitation box="[447,1431,330,359]" location="Tas" pageId="10" pageNumber="214" specimenCount="32" specimenCount-female="32">
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||
<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FE40FEDFFE2FB444" box="[447,490,331,357]" name="Tas" pageId="10" pageNumber="214">Tas</location>
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: from
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<taxonomicName baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[570,793,330,356]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="gaimardi">
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<emphasis box="[570,793,330,356]" italics="true" pageId="10" pageNumber="214">Bettongia gaimardi</emphasis>
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</taxonomicName>
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,
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<typeStatus box="[810,873,331,357]" pageId="10" pageNumber="214">types</typeStatus>
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; oesophageal mucosa, 29 ♁♁,
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<specimenCount box="[1241,1322,330,357]" pageId="10" pageNumber="214" type="female">32 ♀♀</specimenCount>
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(N1363)
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</materialsCitation>
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;
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<materialsCitation box="[151,623,366,395]" location="Bridgenorth" pageId="10" pageNumber="214" specimenCount="2" specimenCount-female="2">
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<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FF68FEFAFEE4B4A9" box="[151,289,366,392]" name="Bridgenorth" pageId="10" pageNumber="214">Bridgenorth</location>
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, 3 ♁♁,
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<specimenCount box="[387,453,366,393]" pageId="10" pageNumber="214" type="female">2 ♀♀</specimenCount>
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(AHC 16543)
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</materialsCitation>
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;
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<materialsCitation box="[638,1432,366,395]" location="Tamar" pageId="10" pageNumber="214" specimenCount="1" specimenCount-female="1">
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adjacent to
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<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FCFAFEFBFC89B4A8" box="[773,844,367,393]" name="Tamar" pageId="10" pageNumber="214">Tamar</location>
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estuary, 13 ♁♁ (AHC 46903);
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<specimenCount box="[1216,1261,366,393]" pageId="10" pageNumber="214" type="female">1 ♀</specimenCount>
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(AHC 46904)
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</materialsCitation>
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;
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<materialsCitation location="Margate" pageId="10" pageNumber="214" specimenCount="2" specimenCount-female="2">
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<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FF68FE07FF32B48C" box="[151,247,403,429]" name="Margate" pageId="10" pageNumber="214">Margate</location>
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, 2 ♁♁ (N4273), 3 ♁♁ (N4293). From
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<taxonomicName authority=", Margate" authorityName="Margate" box="[700,1040,402,429]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="tridactylus">
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<emphasis box="[700,938,402,428]" italics="true" pageId="10" pageNumber="214">Potorous tridactylus,</emphasis>
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Margate
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</taxonomicName>
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2 ♁♁, 7 contracted & coiled
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<specimenCount box="[1387,1430,403,429]" pageId="10" pageNumber="214" type="female">♀♀</specimenCount>
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,
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<specimenCount box="[151,186,438,464]" pageId="10" pageNumber="214" type="female">1♀</specimenCount>
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</materialsCitation>
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anterior and posterior ends (AHC 49245), 3 ♁♁ (AHC 00000); locality unknown, 1 ♁ (AHC 11916). From
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<taxonomicName authority=", Icena" authorityName="Icena" box="[151,482,474,501]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="vulpecula">
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<emphasis box="[151,403,474,500]" italics="true" pageId="10" pageNumber="214">Trichosurus vulpecula</emphasis>
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, Icena
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</taxonomicName>
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, 2 ♁♁,
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<specimenCount box="[577,642,475,501]" pageId="10" pageNumber="214" type="female">7 ♀♀</specimenCount>
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(N966),
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<materialsCitation box="[749,1432,474,503]" location="Mt. Wellington" pageId="10" pageNumber="214" specimenCount="5" specimenCount-female="5">
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||
1 ♁ (N1055);
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||
<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FC6DFE4FFB84B4D5" box="[914,1089,474,501]" name="Mt. Wellington" pageId="10" pageNumber="214">Mt. Wellington</location>
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, 16 ♁♁,
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<specimenCount box="[1197,1262,474,501]" pageId="10" pageNumber="214" type="female">5 ♀♀</specimenCount>
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||
(AHC 49247)
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||
</materialsCitation>
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||
;
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||
<materialsCitation box="[151,360,510,539]" location="Kempton" pageId="10" pageNumber="214" specimenCount="1" specimenCount-female="1">
|
||
<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FF68FE6BFEC7B738" box="[151,258,511,537]" name="Kempton" pageId="10" pageNumber="214">Kempton</location>
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||
, ♁♁,
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||
<specimenCount box="[325,360,510,536]" pageId="10" pageNumber="214" type="female">1♀</specimenCount>
|
||
</materialsCitation>
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||
(N1327).
|
||
</paragraph>
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||
<paragraph blockId="10.[151,1437,150,2013]" box="[199,1348,546,575]" pageId="10" pageNumber="214">
|
||
<collectingRegion box="[199,293,546,572]" country="Australia" name="Victoria" pageId="10" pageNumber="214">Victoria</collectingRegion>
|
||
; from
|
||
<taxonomicName box="[365,594,546,572]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="tridactylus">
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||
<emphasis box="[365,594,546,572]" italics="true" pageId="10" pageNumber="214">Potorous tridactylus</emphasis>
|
||
</taxonomicName>
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||
oesophageal mucosa, Serendip Sanctuary Lara, 1 ♁ (AHC 10981).
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<materialsCitation box="[199,1171,582,611]" location="New South Wales" pageId="10" pageNumber="214" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales">
|
||
<collectingRegion box="[199,403,582,609]" country="Australia" name="New South Wales" pageId="10" pageNumber="214">New South Wales</collectingRegion>
|
||
; from
|
||
<taxonomicName authority=", Taronga Zoo" authorityName="Taronga Zoo" box="[477,904,582,609]" class="Mammalia" family="Potoroidae" genus="Aepyprymnus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis box="[477,743,582,609]" italics="true" pageId="10" pageNumber="214">Aepyprymnus rufescens</emphasis>
|
||
, Taronga Zoo
|
||
</taxonomicName>
|
||
, 1 ♁ posterior end;
|
||
<specimenCount box="[1128,1171,582,609]" pageId="10" pageNumber="214" type="female">1 ♀</specimenCount>
|
||
</materialsCitation>
|
||
<materialsCitation location="Grafton" pageId="10" pageNumber="214" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales">
|
||
anterior (AHC 49246);
|
||
<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FF68FDFEFF34B7A5" box="[151,241,618,644]" name="Grafton" pageId="10" pageNumber="214" stateProvince="New South Wales">Grafton</location>
|
||
,
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||
<specimenCount box="[252,294,618,645]" pageId="10" pageNumber="214" type="female">1 ♀</specimenCount>
|
||
</materialsCitation>
|
||
<materialsCitation county="From" location="Quart Pot Rd." municipality="State Forest" pageId="10" pageNumber="214" specimenCount="3" specimenCount-female="3" stateProvince="New South Wales">
|
||
unfertilised (N3387).
|
||
<collectingCounty box="[544,605,619,645]" pageId="10" pageNumber="214">From</collectingCounty>
|
||
<taxonomicName authority=", Buckenboura" authorityName="Buckenboura" box="[611,1024,618,645]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="vulpecula">
|
||
<emphasis box="[611,861,618,644]" italics="true" pageId="10" pageNumber="214">Trichosurus vulpecula</emphasis>
|
||
, Buckenboura
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||
</taxonomicName>
|
||
<collectingMunicipality box="[1031,1166,618,645]" pageId="10" pageNumber="214">State Forest</collectingMunicipality>
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,
|
||
<location LSID="urn:lsid:plazi:treatment:039587DFFF95FFA0FF68F8C5FDE2B7EC:8EE36012FF96FFA6FB66FDFEFAF3B7A5" box="[1177,1334,618,645]" county="From" municipality="State Forest" name="Quart Pot Rd." pageId="10" pageNumber="214" stateProvince="New South Wales">Quart Pot Rd.</location>
|
||
, 5 ♁♁,
|
||
<specimenCount pageId="10" pageNumber="214" type="female">3 ♀♀</specimenCount>
|
||
(N1241)
|
||
</materialsCitation>
|
||
, 8 ♁♁,
|
||
<specimenCount box="[386,449,654,681]" pageId="10" pageNumber="214" type="female">8 ♀♀</specimenCount>
|
||
(N1352).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="214" type="etymology">
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<emphasis bold="true" box="[199,334,690,716]" pageId="10" pageNumber="214">Etymology.</emphasis>
|
||
The specific name is derived from the generic and common name, eastern bettong, the most commonly infected of the hosts encountered in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="214" type="description">
|
||
<paragraph blockId="10.[151,1437,150,2013]" box="[199,1029,762,789]" pageId="10" pageNumber="214">
|
||
<emphasis bold="true" box="[199,337,762,788]" pageId="10" pageNumber="214">Description</emphasis>
|
||
(based on specimens from
|
||
<taxonomicName baseAuthorityName="Desmarest" baseAuthorityYear="1822" box="[645,863,762,788]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="gaimardi">
|
||
<emphasis box="[645,863,762,788]" italics="true" pageId="10" pageNumber="214">Bettongia gaimardi</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation box="[887,1018,762,788]" captionStart="FIGURE 4" captionStartId="11.[151,249,1885,1909]" captionTargetBox="[158,1430,418,1862]" captionTargetId="figure-154@11.[151,1436,417,1862]" captionTargetPageId="11" captionText="FIGURE 4A–H. Gongylonema bettongiae sp. nov. A, Cephalic end female, en face view. B, Anterior end holotype male, ventrolateral view. C, Excretory pore, lateral view. D, Posterior end holotype male, lateral view. E, Left spicule tip with membranous terminal ending. F, Right spicule tip with membranous terminal ending. G. Gubernaculum accommodating both right and left spicules. H, Posterior end female, ventral view. Scale bars: Figs. 4A, 4E, 4F = 20 µm. Figs. 4B–4D, 4G, 4H = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.7624113" httpUri="https://zenodo.org/record/7624113/files/figure.png" pageId="10" pageNumber="214">Fig. 4 A–H</figureCitation>
|
||
.)
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<emphasis box="[199,291,798,824]" italics="true" pageId="10" pageNumber="214">General</emphasis>
|
||
: Nematodes with marked sexual dimorphism, cuticular bosses present on anterior end, prominent transverse cuticular striae present. Cephalic end with cuticular peri-buccal ring, buccal capsule present. Oral opening not elongated dorso-ventrally, 2 cuticular formations at outer margins of dorso-ventral axis and extending over opening, 6 small lateral lips, 3 on each side, on inside rim of buccal capsule, 2 pairs of internal papillae, 2 pairs of cephalic papillae and 2 large, lateral amphids. Deirids situated anteriorly near origin of lateral alae. Oesophagus long, divided into anterior muscular and posterior glandular regions.
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<emphasis box="[199,257,1014,1040]" italics="true" pageId="10" pageNumber="214">Male</emphasis>
|
||
(
|
||
<typeStatus box="[273,373,1014,1040]" pageId="10" pageNumber="214">holotype</typeStatus>
|
||
in italics, mean
|
||
<specimenCount box="[563,706,1014,1041]" pageId="10" pageNumber="214" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
|
||
+ range in parentheses): Total length
|
||
<emphasis box="[1140,1189,1014,1040]" italics="true" pageId="10" pageNumber="214">20.7</emphasis>
|
||
21.1(19.5–24.2) mm. Maximum width
|
||
<emphasis box="[351,393,1050,1076]" italics="true" pageId="10" pageNumber="214">156</emphasis>
|
||
177 (154–208). Cuticular bosses on all surfaces anteriorly, restricted to dorsal and ventral surfaces posteriorly, extending
|
||
<emphasis box="[512,554,1086,1112]" italics="true" pageId="10" pageNumber="214">848</emphasis>
|
||
964 (655–1192) from anterior end. Buccal capsule
|
||
<emphasis box="[1163,1191,1086,1112]" italics="true" pageId="10" pageNumber="214">52</emphasis>
|
||
51 (42–54) long,
|
||
<emphasis box="[1408,1436,1086,1112]" italics="true" pageId="10" pageNumber="214">18</emphasis>
|
||
16(13–20) wide. Deirids
|
||
<emphasis box="[437,479,1122,1148]" italics="true" pageId="10" pageNumber="214">125</emphasis>
|
||
136 (125–146) from anterior end. Lateral alae present entire body length, widening posteriorly to form asymmetric caudal alae, right wider than left, extending to tail tip. Nerve ring
|
||
<emphasis box="[1224,1266,1158,1184]" italics="true" pageId="10" pageNumber="214">208</emphasis>
|
||
227 (208–260) and excretory pore
|
||
<emphasis box="[367,409,1194,1220]" italics="true" pageId="10" pageNumber="214">353</emphasis>
|
||
374 (353–395) from anterior end. Muscular oesophagus
|
||
<emphasis box="[1046,1088,1194,1220]" italics="true" pageId="10" pageNumber="214">416</emphasis>
|
||
522 (416–613) long, glandular oesophagus
|
||
<emphasis box="[293,349,1230,1256]" italics="true" pageId="10" pageNumber="214">5565</emphasis>
|
||
5749 (5380–6431) long. Spicules dissimilar, unequal, right spicule
|
||
<emphasis box="[1145,1187,1230,1256]" italics="true" pageId="10" pageNumber="214">139</emphasis>
|
||
140 (135–145) long, left spicule finely striated
|
||
<emphasis box="[449,505,1266,1292]" italics="true" pageId="10" pageNumber="214">4749</emphasis>
|
||
5054 (4500–7179) long, proximal end highly ornate; gubernaculum bipartite, left side accommodating left spicule, boat-shaped
|
||
<emphasis box="[671,699,1302,1328]" italics="true" pageId="10" pageNumber="214">52</emphasis>
|
||
60 (52–62) long, right side accommodating right spicule striated,
|
||
<emphasis box="[151,179,1338,1364]" italics="true" pageId="10" pageNumber="214">21</emphasis>
|
||
32 (21–42) long. Caudal end generally with 5, rarely 4 pairs of pedunculated pre-cloacal papillae, 3 pairs of pedunculated post-cloacal papillae and 3 pairs of sessile papillae near tail tip, anterior pair larger than posterior pairs. Pairs of pre-cloacal papillae variable distances anterior to cloacal aperture. Most anterior one or two pairs often difficult to observe because tail twisted or cork-screwed, 4
|
||
<superScript attach="left" box="[896,909,1446,1460]" fontSize="6" pageId="10" pageNumber="214">th</superScript>
|
||
and 5
|
||
<superScript attach="left" box="[983,996,1446,1460]" fontSize="6" pageId="10" pageNumber="214">th</superScript>
|
||
pairs ranging 191 (166–270) and 250 (188–312) anterior to cloacal aperture. In
|
||
<specimenCount box="[627,781,1482,1509]" pageId="10" pageNumber="214" type="generic">one specimen</specimenCount>
|
||
manipulated to a near ventral view, 5 pairs of pre-cloacal papillae 25, 56, 106, 156 and 291 from anterior lip of cloaca, 3 pairs of post-cloacal papillae 31, 75 and 135 from the posterior lip. Distal end of tail curved ventrally,
|
||
<emphasis box="[732,774,1554,1580]" italics="true" pageId="10" pageNumber="214">156</emphasis>
|
||
166 (154–187) long. Phasmids not observed.
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<emphasis box="[199,313,1590,1617]" italics="true" pageId="10" pageNumber="214">Female: (</emphasis>
|
||
<typeStatus box="[313,403,1590,1616]" pageId="10" pageNumber="214">allotype</typeStatus>
|
||
in italics, mean
|
||
<specimenCount box="[606,752,1590,1617]" pageId="10" pageNumber="214" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
|
||
+ range in parentheses): Total length
|
||
<emphasis box="[1207,1235,1590,1616]" italics="true" pageId="10" pageNumber="214">49</emphasis>
|
||
41 (32–49) mm. Maximum width
|
||
<emphasis box="[345,387,1626,1652]" italics="true" pageId="10" pageNumber="214">237</emphasis>
|
||
257 (186–318) at tail end. Cuticular bosses dense, occurring on all surfaces initially, confined to dorsal and ventral surfaces over most of length, extending
|
||
<emphasis box="[834,890,1662,1688]" italics="true" pageId="10" pageNumber="214">1367</emphasis>
|
||
1858 (1367–2491) from anterior end. Prominent transverse cuticular striae present. Buccal capsule
|
||
<emphasis box="[721,749,1698,1724]" italics="true" pageId="10" pageNumber="214">52</emphasis>
|
||
54 (52–62) long,
|
||
<emphasis box="[954,982,1698,1724]" italics="true" pageId="10" pageNumber="214">16</emphasis>
|
||
15 (11–17) wide. Deirid
|
||
<emphasis box="[1271,1313,1698,1724]" italics="true" pageId="10" pageNumber="214">144</emphasis>
|
||
144 (129– 156) from anterior end. Nerve ring
|
||
<emphasis box="[556,598,1734,1760]" italics="true" pageId="10" pageNumber="214">262</emphasis>
|
||
271 (258–302) and excretory pore
|
||
<emphasis box="[1004,1046,1734,1760]" italics="true" pageId="10" pageNumber="214">478</emphasis>
|
||
452 (426–478) from anterior end. Muscular oesophagus
|
||
<emphasis box="[398,440,1770,1796]" italics="true" pageId="10" pageNumber="214">593</emphasis>
|
||
639 (562–770) long, glandular oesophagus
|
||
<emphasis box="[923,989,1770,1796]" italics="true" pageId="10" pageNumber="214">11182</emphasis>
|
||
6926 (4112–12140) long. Exceptionally long (11182, 12140) glandular oesophagus in
|
||
<specimenCount box="[682,820,1806,1833]" pageId="10" pageNumber="214" type="female">two females</specimenCount>
|
||
Position of vulva highly variable,
|
||
<emphasis box="[1225,1281,1806,1832]" italics="true" pageId="10" pageNumber="214">6890</emphasis>
|
||
4801 (2703– 6890) from posterior extremity. Tail conical,
|
||
<emphasis box="[656,698,1842,1868]" italics="true" pageId="10" pageNumber="214">177</emphasis>
|
||
211 (177–234) long. Phasmids
|
||
<emphasis box="[1055,1083,1842,1868]" italics="true" pageId="10" pageNumber="214">17</emphasis>
|
||
20 (17–21) from posterior end. Larvated eggs
|
||
<emphasis box="[316,344,1878,1904]" italics="true" pageId="10" pageNumber="214">54</emphasis>
|
||
58 (52–63) long,
|
||
<emphasis box="[545,573,1878,1904]" italics="true" pageId="10" pageNumber="214">32</emphasis>
|
||
32 (31–33) wide, with thick smooth shells.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="216" pageId="10" pageNumber="214" type="discussion">
|
||
<paragraph blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="214">
|
||
<emphasis bold="true" box="[199,315,1914,1940]" pageId="10" pageNumber="214">Remarks.</emphasis>
|
||
The glandular oesophagus in females of
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[782,935,1914,1941]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="10" pageNumber="214" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov.">
|
||
<emphasis box="[782,935,1914,1941]" italics="true" pageId="10" pageNumber="214">G. bettongiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[943,1035,1914,1941]" pageId="10" pageNumber="214" rank="species">
|
||
<emphasis bold="true" box="[943,1029,1914,1940]" pageId="10" pageNumber="214">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Johnston & Mawson" authorityYear="1942" box="[1091,1232,1914,1941]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="10" pageNumber="214" phylum="Nematoda" rank="species" species="alecturae">
|
||
<emphasis box="[1091,1232,1914,1941]" italics="true" pageId="10" pageNumber="214">G. alecturae</emphasis>
|
||
</taxonomicName>
|
||
are the longest of the species described herein and those in
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[614,766,1950,1977]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="10" pageNumber="214" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov">
|
||
<emphasis box="[614,766,1950,1977]" italics="true" pageId="10" pageNumber="214">G. bettongiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[773,858,1950,1976]" pageId="10" pageNumber="214">
|
||
<taxonomicNameLabel box="[773,858,1950,1976]" pageId="10" pageNumber="214" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
the most variable by as much as a factor of three.
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,1437,150,2013]" lastBlockId="11.[151,1437,151,393]" lastPageId="11" lastPageNumber="215" pageId="10" pageNumber="214">
|
||
Specimens of
|
||
<taxonomicName authorityName="Spratt" authorityYear="2023" box="[354,504,1986,2013]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="10" pageNumber="214" phylum="Nematoda" rank="species" species="bettongiae">
|
||
<emphasis box="[354,504,1986,2013]" italics="true" pageId="10" pageNumber="214">G. bettongiae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName box="[569,703,1986,2012]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="214" phylum="Chordata" rank="species" species="vulpecula">
|
||
<emphasis box="[569,703,1986,2012]" italics="true" pageId="10" pageNumber="214">T. vulpecula</emphasis>
|
||
</taxonomicName>
|
||
differ from those from the three species of potoroids in the slightly longer right and particularly left spicules and tail in males, slightly greater distance of the vulva from the tail tip and the shorter tail in females. Nevertheless, the basic morphological features are the same especially in
|
||
<emphasis box="[1252,1331,187,213]" italics="true" pageId="11" pageNumber="215">en face</emphasis>
|
||
view, the ornamentation on the proximal end of the left spicule and the morphology of the bipartite gubernaculum in males.
|
||
</paragraph>
|
||
<paragraph blockId="11.[151,1437,151,393]" pageId="11" pageNumber="215">
|
||
The left spicule of this species is more markedly flexed, bent, or curved throughout its length than in other species in this study and has the greatest variability in length ranging from almost 4.5 to
|
||
<quantity box="[1128,1192,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="215" unit="mm" value="7.0">7 mm</quantity>
|
||
in potoroids and even more in specimens from
|
||
<taxonomicName box="[431,568,331,357]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="11" pageNumber="215" phylum="Chordata" rank="species" species="vulpecula">
|
||
<emphasis box="[431,568,331,357]" italics="true" pageId="11" pageNumber="215">T. vulpecula</emphasis>
|
||
</taxonomicName>
|
||
. Only the left spicule of
|
||
<taxonomicName authorityName="Johnston & Mawson" authorityYear="1942" box="[849,990,331,357]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="11" pageNumber="215" phylum="Nematoda" rank="species" species="alecturae">
|
||
<emphasis box="[849,990,331,357]" italics="true" pageId="11" pageNumber="215">G. alecturae</emphasis>
|
||
</taxonomicName>
|
||
with its highly characteristic barbed tip is greater in length,>
|
||
<quantity box="[391,472,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="215" unit="mm" value="18.0">18 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7624113" ID-Zenodo-Dep="7624113" httpUri="https://zenodo.org/record/7624113/files/figure.png" pageId="11" pageNumber="215" startId="11.[151,249,1885,1909]" targetBox="[158,1430,418,1862]" targetPageId="11">
|
||
<paragraph blockId="11.[151,1437,1885,2017]" pageId="11" pageNumber="215">
|
||
<emphasis bold="true" box="[151,327,1885,1909]" pageId="11" pageNumber="215">FIGURE 4 A–H.</emphasis>
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[332,584,1885,1909]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="11" pageNumber="215" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov.">
|
||
<emphasis box="[332,584,1885,1909]" italics="true" pageId="11" pageNumber="215">Gongylonema bettongiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[589,691,1885,1909]" pageId="11" pageNumber="215">
|
||
<taxonomicNameLabel box="[589,667,1885,1909]" pageId="11" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
, Cephalic end female,
|
||
<emphasis box="[918,989,1885,1909]" italics="true" pageId="11" pageNumber="215">en face</emphasis>
|
||
view.
|
||
<emphasis bold="true" box="[1052,1069,1885,1909]" pageId="11" pageNumber="215">B</emphasis>
|
||
, Anterior end holotype male, ventrolateral view.
|
||
<emphasis bold="true" box="[281,299,1921,1945]" pageId="11" pageNumber="215">C</emphasis>
|
||
, Excretory pore, lateral view.
|
||
<emphasis bold="true" box="[606,630,1921,1945]" pageId="11" pageNumber="215">D,</emphasis>
|
||
Posterior end holotype male, lateral view.
|
||
<emphasis bold="true" box="[1065,1082,1921,1945]" pageId="11" pageNumber="215">E</emphasis>
|
||
, Left spicule tip with membranous terminal ending.
|
||
<emphasis bold="true" box="[325,340,1957,1981]" pageId="11" pageNumber="215">F</emphasis>
|
||
, Right spicule tip with membranous terminal ending.
|
||
<emphasis bold="true" box="[893,913,1957,1981]" pageId="11" pageNumber="215">G</emphasis>
|
||
. Gubernaculum accommodating both right and left spicules.
|
||
<emphasis bold="true" box="[245,265,1993,2017]" pageId="11" pageNumber="215">H</emphasis>
|
||
, Posterior end female, ventral view. Scale bars: Figs. 4A, 4E, 4F = 20 µm. Figs. 4B–4D, 4G, 4H = 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="12.[151,1436,151,717]" pageId="12" pageNumber="216">
|
||
The number of pedunculated pre-cloacal papillae was more variable (4–6) in specimens from
|
||
<taxonomicName box="[1307,1436,151,177]" class="Mammalia" family="Phalangeridae" genus="Trichosurus" kingdom="Animalia" order="Diprotodontia" pageId="12" pageNumber="216" phylum="Chordata" rank="species" species="vulpecula">
|
||
<emphasis box="[1307,1436,151,177]" italics="true" pageId="12" pageNumber="216">T.vulpecula</emphasis>
|
||
</taxonomicName>
|
||
although the lower number is most likely due to the twisted or cork-screwed form of the tail masking the most anterior one or two pairs.
|
||
</paragraph>
|
||
<paragraph blockId="12.[151,1436,151,717]" pageId="12" pageNumber="216">
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[199,482,259,285]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov.">
|
||
<emphasis box="[199,482,259,285]" italics="true" pageId="12" pageNumber="216">Gongylonema bettongiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[491,583,259,285]" pageId="12" pageNumber="216" rank="species">
|
||
<emphasis bold="true" box="[491,576,259,285]" pageId="12" pageNumber="216">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName authorityName="Johnston & Mawson" authorityYear="1942" box="[838,980,259,285]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="species" species="alecturae">
|
||
<emphasis box="[838,980,259,285]" italics="true" pageId="12" pageNumber="216">G. alecturae</emphasis>
|
||
</taxonomicName>
|
||
by the generally shorter right and much shorter left spicule, the smaller number of pre-and post-cloacal pedunculated papillae, the much shorter tail in the males, and the generally shorter muscular and longer glandular oesophagus, the shorter distance of the vulva from the posterior extremity and the shorter tail in females.
|
||
</paragraph>
|
||
<paragraph blockId="12.[151,1436,151,717]" pageId="12" pageNumber="216">
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[199,481,403,429]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="species" species="bettongiae" status="sp. nov.">
|
||
<emphasis box="[199,481,403,429]" italics="true" pageId="12" pageNumber="216">Gongylonema bettongiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[487,576,403,429]" pageId="12" pageNumber="216">
|
||
<taxonomicNameLabel box="[487,576,403,429]" pageId="12" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName authority="Spratt, 2023" authorityName="Spratt" authorityYear="2023" box="[825,989,403,429]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="species" species="aguilarense" status="sp. nov.">
|
||
<emphasis box="[825,989,403,429]" italics="true" pageId="12" pageNumber="216">G. aguilarense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[996,1087,403,429]" pageId="12" pageNumber="216" rank="species">
|
||
<emphasis bold="true" box="[996,1080,403,429]" pageId="12" pageNumber="216">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
described herein by the shorter body lengths of males and generally longer lengths of females, the much longer extent of bosses and the lengths of the muscular and glandular oesophagus in both sexes, the longer right and much longer left spicule, the fewer number of post-cloacal papillae, arranged symmetrically, the longer tails in the males and the generally shorter tails in the females.
|
||
</paragraph>
|
||
<paragraph blockId="12.[151,1436,151,717]" pageId="12" pageNumber="216">
|
||
The intermediate hosts of species of
|
||
<taxonomicName authorityName="Molin" authorityYear="1857" box="[592,749,583,609]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="genus">
|
||
<emphasis box="[592,749,583,609]" italics="true" pageId="12" pageNumber="216">Gongylonema</emphasis>
|
||
</taxonomicName>
|
||
are beetles and cockroaches (
|
||
<bibRefCitation author="Anderson, R. C." box="[1068,1250,583,609]" pageId="12" pageNumber="216" refId="ref10757" refString="Anderson, R. C. (1992) Nematode parasites of vertebrates, their development and transmission. Commonwealth Agricultural Bureaux International. Wallingford, Oxon, 578 pp." type="book" year="1992">Anderson, 1992</bibRefCitation>
|
||
).The occurrence of
|
||
<taxonomicName authorityName="Spratt" authorityYear="2023" box="[180,332,619,645]" class="Secernentea" family="Gongylonematidae" genus="Gongylonema" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="216" phylum="Nematoda" rank="species" species="bettongiae">
|
||
<emphasis box="[180,332,619,645]" italics="true" pageId="12" pageNumber="216">G. bettongiae</emphasis>
|
||
</taxonomicName>
|
||
in brushtail possums is not surprising as they may be lured to the ground by the smell of the fruiting bodies of underground fungal hyphae, a major component of the diet of bettongs and potoroos (Eldridge, 2006, in van Dyck & Strahan, 2008, p. 284).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |