treatments-xml/data/03/95/87/039587DBFFB22802FEA16195436DFEDC.xml

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<mods:title id="ED7C30796B799200A7568EAA6BB2618D">A revision of the Neotropical genus Motonerus Hansen (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
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<mods:namePart id="BA61B700B66281B62318AFAC96FDDC51">Ek, Martin Fiká Č</mods:namePart>
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<mods:namePart id="CE9FEBC924786FA714C74B03E72FB55D">Short, Andrew Edward Z.</mods:namePart>
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<paragraph id="8B8336CDFFB22839FEA161954588F92C" blockId="25.[264,687,1684,1745]" box="[264,687,1684,1710]" pageId="25" pageNumber="26">
<heading id="D0CB81A1FFB22839FEA161954588F92C" bold="true" box="[264,687,1684,1710]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="4C3C4D4EFFB22839FEA161954588F92C" ID-CoL="6RR83" authority="Hansen, 1989" authorityName="Hansen" authorityYear="1989" box="[264,687,1684,1710]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB22839FEA161954588F92C" bold="true" box="[264,687,1684,1710]" pageId="25" pageNumber="26">
<emphasis id="B948EADFFFB22839FEA161954527F92C" bold="true" box="[264,512,1684,1710]" italics="true" pageId="25" pageNumber="26">Motonerus obscurus</emphasis>
<bibRefCitation id="EFAD4B3CFFB22839FDA161954588F92C" author="Hansen" box="[520,687,1684,1710]" pageId="25" pageNumber="38" refString="Hansen, M. (1989). New genera of Sphaeridiinae (Coleoptera: Hydrophilidae). Entomologica Scandinavica, 20, 251 - 262." type="journal article" year="1989">Hansen, 1989</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8336CDFFB22839FEA161B645B8F953" blockId="25.[264,687,1684,1745]" box="[264,671,1719,1745]" pageId="25" pageNumber="26">
(
<figureCitation id="13072A48FFB22839FEB861B64645F953" box="[273,354,1719,1745]" captionStart="FIGURES 1 – 4" captionStartId="5.[264,383,1567,1591]" captionTargetBox="[283,1297,323,1547]" captionTargetId="figure@5.[264,1323,309,1557]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Dorsal habitus of Motonerus. 1: M. obscurus Hansen; 2: M. apterus sp. n.; 3: M. andersoni sp. n.; 4: M. depressus sp. n. Scale bar = 1 mm." httpUri="https://zenodo.org/record/173210/files/figure.png" pageId="25" pageNumber="26">Figs. 1</figureCitation>
, 
<figureCitation id="13072A48FFB22839FED961B6469BF953" box="[368,444,1719,1745]" captionStart="FIGURES 13 – 20" captionStartId="19.[264,382,1761,1785]" captionTargetBox="[305,1289,326,1738]" captionTargetId="figure@19.[305,1289,326,1738]" captionTargetPageId="19" captionText="FIGURES 13 – 20. Male genitalia of Motonerus. 13 – 14: M. andersoni sp. n., paratype; 15 – 16: M. hanseni sp. n., holotype; 17 – 18: M. obscurus Hansen, paratype; 19 – 20: M. problematicus sp. n., holotype. 13, 15, 17, 19 — aedeagus (scale bar = 0.1 mm); 14, 16, 18, 20 — apical part of median lobe (scale bar = 0.05 mm)." httpUri="https://zenodo.org/record/173212/files/figure.png" pageId="25" pageNumber="26">17–18</figureCitation>
, 
<figureCitation id="13072A48FFB22839FE6161B64533F953" box="[456,532,1719,1745]" captionStart="FIGURES 21 – 31" captionStartId="20.[264,382,1715,1739]" captionTargetBox="[264,1323,326,1691]" captionTargetId="figure@20.[264,1323,326,1691]" captionTargetPageId="20" captionText="FIGURES 21 – 31. Lateral habitus of Motonerus. 21: M. apterus sp. n.; 22: M. oosternoides sp. n.; 23: M. depressus sp. n.; 24: M. nublado sp. n.; 25: M. andersoni sp. n.; 26: M. hanseni sp. n.; 27: M. problematicus sp. n.; 28: M. obscurus Hansen (El Salvador); 29: M. obscurus Hansen (Oaxaca prov., Mexico); 30: M. sp. B; 31: M. sp. A. Scale bar = 1 mm." httpUri="https://zenodo.org/record/173213/files/figure.png" pageId="25" pageNumber="26">28–29</figureCitation>
, 
<figureCitation id="13072A48FFB22839FDB661B64518F953" box="[543,575,1719,1745]" captionStart="FIGURES 32 – 40" captionStartId="21.[264,383,1661,1685]" captionTargetBox="[306,1288,326,1631]" captionTargetId="figure@21.[291,1296,326,1637]" captionTargetPageId="21" captionText="FIGURES 32 – 40. Preepisternal elevation of mesothorax and median part of metaventrite of Motonerus. 32: M. apterus sp. n.; 33: M. oosternoides sp. n.; 34: M. depressus sp. n.; 35: M. andersoni sp. n.; 36: M. hanseni sp. n.; 37: M. nublado sp. n.; 38: M. obscurus Hansen; 39: M. problematicus sp. n.; 40: M. sp. A. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173214/files/figure.png" pageId="25" pageNumber="26">38</figureCitation>
, 
<figureCitation id="13072A48FFB22839FDE561B6454BF953" box="[588,620,1719,1745]" captionStart="FIGURES 41 – 53" captionStartId="22.[264,382,1710,1734]" captionTargetBox="[264,1323,326,1686]" captionTargetId="figure@22.[264,1323,326,1686]" captionTargetPageId="22" captionText="FIGURES 41 – 53. Details of of Motonerus. 41, 49: M. sp. B; 42, 52: M. andersoni sp. n.; 43, 53: M. apterus sp. n.; 44: M. depressus sp. n.; 45: M. hanseni sp. n.; 46: M. oosternoides sp. n.; 47: M. nublado sp. n.; 48: M. obscurus Hansen; 50: M. problematicus sp. n.; 51: M. sp. A. 41 — preepisternal elevation of mesothorax and median part of metaventrite (scale bar = 0.2 mm); 42 – 51 — median part of prosternum; 52 – 53 — sternite 9 of male." httpUri="https://zenodo.org/record/173215/files/figure.png" pageId="25" pageNumber="26">48</figureCitation>
, 
<figureCitation id="13072A48FFB22839FDDE61B645B2F953" box="[631,661,1719,1745]" captionStart="FIGURES 54 – 64" captionStartId="23.[264,382,1763,1787]" captionTargetBox="[264,1323,326,1738]" captionTargetId="figure@23.[264,1323,326,1738]" captionTargetPageId="23" captionText="FIGURES 54 – 64. Details of Motonerus. 54, 60, 63: M. apterus sp. n.; 55, 64: M. oosternoides sp. n.; 56: M. depressus sp. n.; 57, 62: M. nublado sp. n.; 59: M. problematicus sp. n.; 61: M. obscurus Hansen. 54 – 59 — epipleuron and lateral part of elytron; 60 – 62 — abdominal ventrites; 63 – 64 — head, lateral view. Scale bar = 0.5 mm (54 – 59), 0.25 mm (60 – 62); 0.1 mm (63 – 64)." httpUri="https://zenodo.org/record/173216/files/figure.png" pageId="25" pageNumber="26">61</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8B8336CDFFB22839FEA1600645D9F89F" blockId="25.[264,766,1798,1822]" box="[264,766,1798,1822]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB2C11E3FFB22839FEA1600645D9F89F" box="[264,766,1798,1822]" pageId="25" pageNumber="26">
<taxonomicName id="4C3C4D4EFFB22839FEA160064595F89F" ID-CoL="6RR83" authority="Hansen, 1989: 257" authorityName="Hansen" authorityPageNumber="257" authorityYear="1989" box="[264,690,1798,1822]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB22839FEA1600646C5F89C" box="[264,482,1799,1822]" italics="true" pageId="25" pageNumber="26">Motonerus obscurus</emphasis>
<treatmentCitation id="0A9D10DCFFB22839FE4060074595F89F" author="Hansen" box="[489,690,1798,1821]" page="257" pageId="25" pageNumber="26" year="1989">
<bibRefCitation id="EFAD4B3CFFB22839FE40600745A7F89F" author="Hansen" box="[489,640,1798,1821]" pageId="25" pageNumber="38" refString="Hansen, M. (1989). New genera of Sphaeridiinae (Coleoptera: Hydrophilidae). Entomologica Scandinavica, 20, 251 - 262." type="journal article" year="1989">Hansen, 1989</bibRefCitation>
: 257
</treatmentCitation>
</taxonomicName>
(pars.)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB2283BFEA1604F4643FBAC" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8B8336CDFFB22839FEA1604F4317F8EB" blockId="25.[264,1072,1870,1897]" box="[264,1072,1870,1897]" pageId="25" pageNumber="26">
<emphasis id="B948EADFFFB22839FEA1604F4696F8EA" bold="true" box="[264,433,1870,1896]" pageId="25" pageNumber="26">
<typeStatus id="5487886FFFB22839FEA1604F4662F8EA" box="[264,325,1870,1896]" pageId="25" pageNumber="26">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F32B765DFFB22839FE10604E45A6F8EB" box="[441,641,1871,1897]" name="El Salvador" pageId="25" pageNumber="26">EL SALVADOR</collectingCountry>
, Montecristo near Metapán, 
<quantity id="4CC49B28FFB22839FC79604E430BF8EB" box="[976,1068,1871,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="25" pageNumber="26" unit="m" value="2300.0">2300 m</quantity>
.
</paragraph>
<paragraph id="8B8336CDFFB1283AFE91661A4580FE7C" blockId="26.[264,1326,283,1870]" pageId="26" pageNumber="27">
<emphasis id="B948EADFFFB1283AFE91661A454AFEB7" bold="true" box="[312,621,283,309]" pageId="26" pageNumber="27">
<typeStatus id="5487886FFFB1283AFE91661A4652FEB7" box="[312,373,283,309]" pageId="26" pageNumber="27">Type</typeStatus>
Material. 
<typeStatus id="5487886FFFB1283AFE52661A454AFEB7" box="[507,621,283,309]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
♂ (
<collectionCode id="ED2DAE08FFB1283AFD30661D45FBFEB4" box="[665,732,284,310]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="26" pageNumber="27">CNC</collectionCode>
): “EL SAL., nr. Metapan / Montecristo, 
<quantity id="4CC49B28FFB1283AFB69661D423DFEB4" box="[1216,1306,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="26" pageNumber="27" unit="m" value="2300.0">2300 m</quantity>
/ 
<date id="FF82100DFFB1283AFEA166454687FEDC" box="[264,416,324,350]" pageId="26" pageNumber="27" value="1971-05-08" valueMax="1971-05-10" valueMin="1971-05-08">8–10.V.1971</date>
, S. Peck [printed label] // Ber. 199 / cloud forest [printed label] // 
<typeStatus id="5487886FFFB1283AFEA1666D4685FE04" box="[264,418,364,390]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C3C4D4EFFB1283AFE6B666D4566FE04" box="[450,577,364,390]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Motonerus</taxonomicName>
/ 
<taxonomicName id="4C3C4D4EFFB1283AFDC9666D45EFFE04" box="[608,712,364,390]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obscurus">obscurus</taxonomicName>
/ M. Hansen [red printed label] // 
<typeStatus id="5487886FFFB1283AFBD5666D4231FE04" box="[1148,1302,364,390]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
/ 
<collectionCode id="ED2DAE08FFB1283AFEA166954663FE2C" box="[264,324,404,430]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="26" pageNumber="27">CNC</collectionCode>
No. 20749 [red printed label, number hand­written]”. 
<emphasis id="B948EADFFFB1283AFB9966924697FE57" bold="true" pageId="26" pageNumber="27">
<typeStatus id="5487886FFFB1283AFB996692438AFE2F" box="[1072,1197,403,429]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
(8): 
<collectingCountry id="F32B765DFFB1283AFAAA6692468CFE57" name="El Salvador" pageId="26" pageNumber="27">EL SALVADOR</collectingCountry>
:
</emphasis>
same data as 
<typeStatus id="5487886FFFB1283AFDF866BD459CFE54" box="[593,699,444,470]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
, only with “ 
<typeStatus id="5487886FFFB1283AFCEC66BD44F9FE54" box="[837,990,444,470]" pageId="26" pageNumber="27" type="paratype">PARATYPE</typeStatus>
” on the first 
<typeStatus id="5487886FFFB1283AFBDD66BD4381FE54" box="[1140,1190,444,470]" pageId="26" pageNumber="27">type</typeStatus>
label [both 
<typeStatus id="5487886FFFB1283AFEA166E5461DFE7C" box="[264,314,484,510]" pageId="26" pageNumber="27">type</typeStatus>
labels yellow], 8 spec. (
<collectionCode id="ED2DAE08FFB1283AFDFE66E545BDFE7C" box="[599,666,484,510]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="26" pageNumber="27">CNC</collectionCode>
).
</paragraph>
<paragraph id="8B8336CDFFB1283AFE91650D46E1FCBC" blockId="26.[264,1326,283,1870]" pageId="26" pageNumber="27">
The original 
<typeStatus id="5487886FFFB1283AFE74650D4528FDA4" box="[477,527,524,550]" pageId="26" pageNumber="27">type</typeStatus>
series housed in 
<collectionCode id="ED2DAE08FFB1283AFD5C650D4416FDA4" box="[757,817,524,550]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="26" pageNumber="27">CNC</collectionCode>
consists of two different species as the specimen from 
<collectingCountry id="F32B765DFFB1283AFE6D65354560FDCC" box="[452,583,564,590]" name="Costa Rica" pageId="26" pageNumber="27">Costa Rica</collectingCountry>
was found not to be conspecific with the 
<typeStatus id="5487886FFFB1283AFBE265354396FDCC" box="[1099,1201,564,590]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
(this new species is described as 
<taxonomicName id="4C3C4D4EFFB1283AFD82655D4594FDF7" box="[555,691,604,629]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hanseni" status="sp. nov.">
<emphasis id="B948EADFFFB1283AFD82655D4594FDF7" box="[555,691,604,629]" italics="true" pageId="26" pageNumber="27">M. hanseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57A4FFB1283AFD16655D4424FDF4" box="[703,771,604,630]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
above). We did not have the opportunity to examine the 
<typeStatus id="5487886FFFB1283AFE3665854523FD1C" box="[415,516,644,670]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
from 
<collectingCountry id="F32B765DFFB1283AFDE465854581FD1C" box="[589,678,644,670]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
(Hidalgo Prov.) mentioned in the original description. As both specimens from Oaxaca province in 
<collectingCountry id="F32B765DFFB1283AFC9565AD44B1FD44" box="[828,918,684,710]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
examined by us (see Additional material examined) are slightly aberrant from all other specimens of 
<taxonomicName id="4C3C4D4EFFB1283AFBF365D543D7FD6F" box="[1114,1264,724,749]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB1283AFBF365D543D7FD6F" box="[1114,1264,724,749]" italics="true" pageId="26" pageNumber="27">M. obscurus</emphasis>
</taxonomicName>
(see Discussion for details), we cannot conclude if the third Mexican 
<typeStatus id="5487886FFFB1283AFBBE65FD435BFC94" box="[1047,1148,764,790]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
belongs to 
<taxonomicName id="4C3C4D4EFFB1283AFAA265FD4655FCBF" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB1283AFAA265FD4655FCBF" italics="true" pageId="26" pageNumber="27">M. obscurus</emphasis>
</taxonomicName>
or not.
</paragraph>
<paragraph id="8B8336CDFFB1283BFE91644D4431FE2C" blockId="26.[264,1326,283,1870]" lastBlockId="27.[264,1325,283,1870]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B948EADFFFB1283AFE91644D448EFCE4" bold="true" box="[312,937,844,870]" pageId="26" pageNumber="27">
Additional material examined (46): 
<collectingCountry id="F32B765DFFB1283AFD51644D4483FCE4" box="[760,932,844,870]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
:
</emphasis>
“ 
<collectingCountry id="F32B765DFFB1283AFC12644D4342FCE4" box="[955,1125,844,870]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: 
<collectionCode id="ED2DAE08FFB1283AFBD9644D43F1FCE4" box="[1136,1238,844,870]" pageId="26" pageNumber="27">FRANC</collectionCode>
. 
<collectionCode id="ED2DAE08FFB1283AFB49644D4200FCE4" box="[1248,1319,844,870]" httpUri="http://grbio.org/cool/3ry3-9mkc" name="Museum of the Rockies" pageId="26" pageNumber="27">MOR</collectionCode>
. / P.N. La Tigra 23.2 lm N. / Tegucigalpa, 
<quantity id="4CC49B28FFB1283AFD546475447CFC0C" box="[765,859,884,910]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="26" pageNumber="27" unit="m" value="1950.0">1950 m</quantity>
, 
<date id="FF82100DFFB1283AFCC16475432EFC0C" box="[872,1033,884,910]" pageId="26" pageNumber="27" value="1994-06-29">29.VI. / 1994</date>
­139C / oak/cloud forest litt. berl.”, ♂ (
<collectionCode id="ED2DAE08FFB1283AFE1D649D4536FC34" LSID="urn:lsid:biocol.org:col:33144" box="[436,529,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFD9B649D45FBFC34" box="[562,732,924,950]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: 
<collectionCode id="ED2DAE08FFB1283AFD42649D4476FC34" box="[747,849,924,950]" pageId="26" pageNumber="27">FRANC</collectionCode>
. 
<collectionCode id="ED2DAE08FFB1283AFCF6649D4481FC34" box="[863,934,924,950]" httpUri="http://grbio.org/cool/3ry3-9mkc" name="Museum of the Rockies" pageId="26" pageNumber="27">MOR</collectionCode>
. / Cerro Uyuca, ca. 
<quantity id="4CC49B28FFB1283AFB36649D43CBFC34" box="[1183,1260,924,950]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="km" value="10.0">10 km</quantity>
W. / Zamorano, 
<quantity id="4CC49B28FFB1283AFE3964C546CDFC5C" box="[400,490,964,990]" metricMagnitude="3" metricUnit="m" metricValue="1.8399999999999999" pageId="26" pageNumber="27" unit="m" value="1840.0">1840 m</quantity>
/ 
<date id="FF82100DFFB1283AFDAA64C5455BFC5C" box="[515,636,964,990]" pageId="26" pageNumber="27" value="1994-06-09">9.VI.1994</date>
­105, R. Anderson / cloud forest litter berlese”, 1 unsexed specimen (
<collectionCode id="ED2DAE08FFB1283AFE2264ED46CFFB84" LSID="urn:lsid:biocol.org:col:33144" box="[395,488,1004,1030]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFDA264ED4591FB84" box="[523,694,1004,1030]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: LA PAZ / Titule, Res. Biol. Guajiquiro / N 
<geoCoordinate id="EE08500AFFB1283AFB4D64ED4600FBAC" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="-14.166667">14°10' W</geoCoordinate>
87°50', 
<quantity id="4CC49B28FFB1283AFE2D631546C4FBAC" box="[388,483,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="26" pageNumber="27" unit="m" value="2130.0">2130 m</quantity>
/ 
<date id="FF82100DFFB1283AFDAD63154549FBAC" box="[516,622,1044,1070]" pageId="26" pageNumber="27" value="2002-05-07">7.V.2002</date>
R. Anderson cloud forest litter, 2002­010H”, 1 unsexed specimen (
<collectionCode id="ED2DAE08FFB1283AFE24633D46CDFBD4" LSID="urn:lsid:biocol.org:col:33144" box="[397,490,1084,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); same data, only number of the sample “2002­010E”, 2 unsexed specimens (
<collectionCode id="ED2DAE08FFB1283AFE38636546F8FBFC" box="[401,479,1124,1150]" pageId="26" pageNumber="27">AEZS</collectionCode>
, 
<collectionCode id="ED2DAE08FFB1283AFE4263654560FBFC" LSID="urn:lsid:biocol.org:col:33144" box="[491,583,1124,1150]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFDCC63654437FBFC" box="[613,784,1124,1150]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: LA PAZ, Cord. / Montecillos, via 
<quantity id="4CC49B28FFB1283AFB02636543D2FBFC" box="[1195,1269,1124,1150]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="km" value="25.0">25 km</quantity>
S.E. &amp; / 
<quantity id="4CC49B28FFB1283AFEE8638D46B6FB24" box="[321,401,1164,1190]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="km" value="18.0">18 km</quantity>
N.W. La Paz, 
<date id="FF82100DFFB1283AFDFA638D4436FB24" box="[595,785,1164,1190]" pageId="26" pageNumber="27" value="1994-08-21">21.VIII. / 1994</date>
­217C, 
<quantity id="4CC49B28FFB1283AFCC5638D44E9FB24" box="[876,974,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="2000.0">2000 m</quantity>
, R. Anderson / mixed pine/ hardwood forest litt. berl.”, 3ΨΨ (
<collectionCode id="ED2DAE08FFB1283AFD3863B545C9FB4C" LSID="urn:lsid:biocol.org:col:33144" box="[657,750,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFCA563B54491FB4C" box="[780,950,1204,1230]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: STA BARBARA / Cerro Santa Barbara, 
<quantity id="4CC49B28FFB1283AFED063DD46EFFB74" box="[377,456,1244,1270]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="km" value="11.0">11 km</quantity>
S. / &amp; 
<quantity id="4CC49B28FFB1283AFD9C63DD45AAFB74" box="[565,653,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="26" pageNumber="27" unit="km" value="5.6">5.6 km</quantity>
W Penas Blancas / 
<quantity id="4CC49B28FFB1283AFC3C63DD44D0FB74" box="[917,1015,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="26" pageNumber="27" unit="m" value="1870.0">1870 m</quantity>
, 
<date id="FF82100DFFB1283AFBA063DD4387FB74" box="[1033,1184,1244,1270]" pageId="26" pageNumber="27" value="1994-08-24">24.VIII.1994</date>
­220D / R. Anderson cloud for. litt. berl.”, ♂, Ψ, 4 unsexed specimens (
<collectionCode id="ED2DAE08FFB1283AFBA962054369FA9C" box="[1024,1102,1284,1310]" pageId="26" pageNumber="27">AEZS</collectionCode>
, 
<collectionCode id="ED2DAE08FFB1283AFBF66205439FFA9C" LSID="urn:lsid:biocol.org:col:33144" box="[1119,1208,1284,1310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
, 
<collectionCode id="ED2DAE08FFB1283AFB6262054238FA9C" box="[1227,1311,1284,1310]" pageId="26" pageNumber="27">MFPC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFEBB622D469BFAC4" box="[274,444,1324,1350]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: 
<collectionCode id="ED2DAE08FFB1283AFE63622D450DFAC4" box="[458,554,1324,1350]" pageId="26" pageNumber="27">SANTA</collectionCode>
BARBARA / Cerro Santa Barbara, 
<quantity id="4CC49B28FFB1283AFC49622D430CFAC4" box="[992,1067,1324,1350]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="km" value="15.0">15 km</quantity>
S. E. Penas Blancas, 
<quantity id="4CC49B28FFB1283AFEA16255464DFAEC" box="[264,362,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="26" pageNumber="27" unit="m" value="1850.0">1850 m</quantity>
, 
<date id="FF82100DFFB1283AFED2625546D0FAEC" box="[379,503,1364,1390]" pageId="26" pageNumber="27" value="1994-08-24">24.VIII.94</date>
/ S. Peck, 94­58, wet mont.for.litt.”, Ψ (
<collectionCode id="ED2DAE08FFB1283AFC526255437FFAEC" LSID="urn:lsid:biocol.org:col:33144" box="[1019,1112,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFBD562554201FAEC" box="[1148,1318,1364,1390]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: LEMPIRA / P.N. Celaque, nr. Gracias /Above Camp. Don Tomas / 
<quantity id="4CC49B28FFB1283AFB88627D435AFA14" box="[1057,1149,1404,1430]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="26" pageNumber="27" unit="m" value="2250.0">2250 m</quantity>
, N 
<geoCoordinate id="EE08500AFFB1283AFB0F627D420CFA14" box="[1190,1323,1404,1430]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="92" value="-14.545">14°32.7' W</geoCoordinate>
88°39.7' / 
<date id="FF82100DFFB1283AFE2562A54515FA3C" box="[396,562,1444,1470]" pageId="26" pageNumber="27" value="2002-05-12" valueMax="2002-05-13" valueMin="2002-05-12">12–13.V.2002</date>
, R. Anderson / mixed oak for. litter 2002­021A”, 1 unsexed specimen; “ 
<collectingCountry id="F32B765DFFB1283AFE3962CD451CFA64" box="[400,571,1484,1510]" name="Honduras" pageId="26" pageNumber="27">HONDURAS</collectingCountry>
: STA. BARBARA / Cerro Santa Barbara, 
<quantity id="4CC49B28FFB1283AFB8462CD43AFFA64" box="[1069,1160,1484,1510]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="26" pageNumber="27" unit="km" value="11.5">11.5km</quantity>
, S. / &amp; 
<quantity id="4CC49B28FFB1283AFB4862CD420CFA64" box="[1249,1323,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="26" pageNumber="27" unit="km" value="5.6">5.6km</quantity>
W. Penas Blancas / 
<quantity id="4CC49B28FFB1283AFE5562F5457CF98C" box="[508,603,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="26" pageNumber="27" unit="m" value="1870.0">1870 m</quantity>
, 
<date id="FF82100DFFB1283AFDC262F54425F98C" box="[619,770,1524,1550]" pageId="26" pageNumber="27" value="1994-08-24">24.VIII.1994</date>
­220H / R. Anderson, cloud for. litt. berl.”, 1 unsexed specimen (
<collectionCode id="ED2DAE08FFB1283AFE46611D456DF9B4" box="[495,586,1564,1590]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
); same data, but sample no. “220I”, 2ΨΨ (
<collectionCode id="ED2DAE08FFB1283AFB97611D43BEF9B4" box="[1086,1177,1564,1590]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
); same data, but sample no. “220J”, ♂ (
<collectionCode id="ED2DAE08FFB1283AFDE8614545BAF9DC" box="[577,669,1604,1630]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
). 
<emphasis id="B948EADFFFB1283AFD1961454412F9DC" bold="true" box="[688,821,1604,1630]" pageId="26" pageNumber="27">
<collectingCountry id="F32B765DFFB1283AFD1961454417F9DC" box="[688,816,1604,1630]" name="Mexico" pageId="26" pageNumber="27">MEXICO</collectingCountry>
:
</emphasis>
“ 
<collectingCountry id="F32B765DFFB1283AFCE1614544E4F9DC" box="[840,963,1604,1630]" name="Mexico" pageId="26" pageNumber="27">MEXICO</collectingCountry>
: CHIAPAS: 8.9 / km E Rayon Elev. / 
<quantity id="4CC49B28FFB1283AFEC8616D46E7F904" box="[353,448,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">1500 m</quantity>
, cloud forest / litter R. Anderson / 91­109, 
<date id="FF82100DFFB1283AFC70616D434FF904" box="[985,1128,1644,1670]" pageId="26" pageNumber="27" value="1991-09-19">19­IX­1991</date>
”, Ψ (
<collectionCode id="ED2DAE08FFB1283AFB00616D4221F904" LSID="urn:lsid:biocol.org:col:33144" box="[1193,1286,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFEA16195466AF92C" box="[264,333,1684,1710]" name="Mexico" pageId="26" pageNumber="27">MEX</collectingCountry>
. Rte. 131 / 
<quantity id="4CC49B28FFB1283AFE4861954512F92C" box="[481,565,1684,1710]" metricMagnitude="5" metricUnit="m" metricValue="1.8699999999999999" pageId="26" pageNumber="27" unit="km" value="187.0">187km</quantity>
. S. Oaxaca / Oax. 
<date id="FF82100DFFB1283AFC89619544B9F92C" box="[800,926,1684,1710]" pageId="26" pageNumber="27" value="1971-05-12">V­12­1971</date>
/ J. M. Campbell, 5800’ “, 2♂♂ (
<collectionCode id="ED2DAE08FFB1283AFEB961BD4675F954" box="[272,338,1724,1750]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="26" pageNumber="27">CNC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB1283AFED861BD46CBF954" box="[369,492,1724,1750]" name="Mexico" pageId="26" pageNumber="27">MEXICO</collectingCountry>
: Hidalgo / 
<quantity id="4CC49B28FFB1283AFDC261BD459BF954" box="[619,700,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="26" pageNumber="27" unit="km" value="4.4">4.4 km</quantity>
N Tlanchinol / Hwy. 105, 
<date id="FF82100DFFB1283AFC5D61BD435AF954" box="[1012,1149,1724,1750]" pageId="26" pageNumber="27" value="1992-07-06">6 July 1992</date>
/ 
<quantity id="4CC49B28FFB1283AFB3B61BD43C9F954" box="[1170,1262,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="26" pageNumber="27" unit="m" value="1420.0">1420 m</quantity>
, J. S. Ashe / H. Frania #11 / ex. treefall litter”, ♂ (
<collectionCode id="ED2DAE08FFB1283AFC8361E544A2F97C" box="[810,901,1764,1790]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="27">ZMUC</collectionCode>
). 
<emphasis id="B948EADFFFB1283AFC3261E5437CF97C" bold="true" box="[923,1115,1764,1790]" pageId="26" pageNumber="27">
<collectingCountry id="F32B765DFFB1283AFC3261E54371F97C" box="[923,1110,1764,1790]" name="Nicaragua" pageId="26" pageNumber="27">NICARAGUA</collectingCountry>
:
</emphasis>
“ 
<collectingCountry id="F32B765DFFB1283AFBC661E54200F97C" box="[1135,1319,1764,1790]" name="Nicaragua" pageId="26" pageNumber="27">NICARAGUA</collectingCountry>
: MATAGALPA / 
<quantity id="4CC49B28FFB1283AFE75600D4538F8A4" box="[476,543,1804,1830]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="27" unit="km" value="6.0">6 km</quantity>
N Matagalpa, Selva Negra / 
<quantity id="4CC49B28FFB1283AFC34600D4327F8A4" box="[925,1024,1804,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="26" pageNumber="27" unit="m" value="1550.0">1550 m</quantity>
, N 
<geoCoordinate id="EE08500AFFB1283AFB80600D4392F8A4" box="[1065,1205,1804,1830]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="92" value="-12.998333">12°59.9' W</geoCoordinate>
85°54.6' / 
<date id="FF82100DFFB1283AFEA1603546A5F8CC" box="[264,386,1844,1870]" pageId="26" pageNumber="27" value="2002-05-19">19.V.2002</date>
, R. Anderson, ridge– / top cloud for., litter, 2002­024H”, ♂ (
<collectionCode id="ED2DAE08FFB1283AFB24603543CDF8CC" LSID="urn:lsid:biocol.org:col:33144" box="[1165,1258,1844,1870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="26" pageNumber="27">CNMC</collectionCode>
). 
<emphasis id="B948EADFFFB1283BFAAA60354697FEB7" bold="true" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<collectingCountry id="F32B765DFFB1283BFAAA6035468CFEB7" lastPageId="27" lastPageNumber="28" name="El Salvador" pageId="26" pageNumber="27">EL SALVADOR</collectingCountry>
:
</emphasis>
“EL SAL., nr. Metapan / Montecristo, 
<quantity id="4CC49B28FFB0283BFC2D661D44F9FEB4" box="[900,990,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="2300.0">2300 m</quantity>
/ 
<date id="FF82100DFFB0283BFC5E661D43B1FEB4" box="[1015,1174,284,310]" pageId="27" pageNumber="28" value="1971-05-08" valueMax="1971-05-10" valueMin="1971-05-08">8.–10.V.1971</date>
, S. Peck”, ♂ (
<collectionCode id="ED2DAE08FFB0283BFEB966454675FEDC" box="[272,338,324,350]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="27" pageNumber="28">CNC</collectionCode>
); “ 
<collectingCountry id="F32B765DFFB0283BFED86645451BFEDC" box="[369,572,324,350]" name="El Salvador" pageId="27" pageNumber="28">EL SALVADOR</collectingCountry>
: STA ANA / Cerro Montechristo, 
<quantity id="4CC49B28FFB0283BFC6366454304FEDC" box="[970,1059,324,350]" metricMagnitude="4" metricUnit="m" metricValue="2.17" pageId="27" pageNumber="28" unit="km" value="21.7">21.7km</quantity>
/ N. E. Metapan, 
<quantity id="4CC49B28FFB0283BFB5866454605FE04" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="28" unit="m" value="2100.0">2100 m</quantity>
, 
<date id="FF82100DFFB0283BFE9D666D46D4FE04" box="[308,499,364,390]" pageId="27" pageNumber="28" value="1994-08-29">29.VIII. / 1994</date>
­228A, R. Anderson / hardwood forest litter, berlese” 3♂♂, Ψ, 16 unsexed specimens (
<collectionCode id="ED2DAE08FFB0283BFE5066954560FE2C" box="[505,583,404,430]" pageId="27" pageNumber="28">AEZS</collectionCode>
, 
<collectionCode id="ED2DAE08FFB0283BFDFB66954584FE2C" box="[594,675,404,430]" pageId="27" pageNumber="28">MFPC</collectionCode>
, 
<collectionCode id="ED2DAE08FFB0283BFD066695442EFE2C" box="[687,777,404,430]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="27" pageNumber="28">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="8B8336CDFFB0283BFE9166BA4660FD6C" blockId="27.[264,1325,283,1870]" pageId="27" pageNumber="28">
<emphasis id="B948EADFFFB0283BFE9166BA4573FE57" bold="true" box="[312,596,443,469]" pageId="27" pageNumber="28">Differential Diagnosis.</emphasis>
Body larger than 
<quantity id="4CC49B28FFB0283BFC9766BD4487FE54" box="[830,928,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="mm" value="1.8">1.8 mm</quantity>
; elytra nearly evenly convex in lateral view; elytral intervals 2, 4 and 6 usually slightly higher than remaining ones basally; lateral margins of elytra with denticles usually not overlapping 0.66 of elytral length; anteromedian prosternal excision deep and narrow; preepisternal elevation 1.6–2.0× longer than wide, with deep concave median area; metaventrite with more or less developed depression; abdominal ventrite 1 with submedian longitudinal ridges; median lobe widened apicad, abruptly narrowing subapically, bearing small circular excision on apex.
</paragraph>
<paragraph id="8B8336CDFFB0283BFE9165FD4643FBAC" blockId="27.[264,1325,283,1870]" pageId="27" pageNumber="28">
This species, by the combination of a wider preepisternal elevation of mesothorax with median concave area, longer body size and abdominal ventrite 1 with submedian longitudinal ridges, somewhat resembles 
<taxonomicName id="4C3C4D4EFFB0283BFD47644D44ACFCE7" box="[750,907,844,869]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="B948EADFFFB0283BFD47644D44ACFCE7" box="[750,907,844,869]" italics="true" pageId="27" pageNumber="28">M. andersoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D4EFFB0283BFC33644D433BFCE7" box="[922,1052,844,869]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="hanseni">
<emphasis id="B948EADFFFB0283BFC33644D433BFCE7" box="[922,1052,844,869]" italics="true" pageId="27" pageNumber="28">M. hanseni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D4EFFB0283BFBFC644D4203FCE7" box="[1109,1316,844,869]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B948EADFFFB0283BFBFC644D4203FCE7" box="[1109,1316,844,869]" italics="true" pageId="27" pageNumber="28">M. problematicus</emphasis>
</taxonomicName>
. From the latter two mentioned species it differs by having nearly evenly convex elytra in lateral view (they have distinctly depressed in the elytral midlength), moreover, from 
<taxonomicName id="4C3C4D4EFFB0283BFAA2649D4689FC5F" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B948EADFFFB0283BFAA2649D4689FC5F" italics="true" pageId="27" pageNumber="28">M. problematicus</emphasis>
</taxonomicName>
it can be easily distinguished by much wider preepisternal elevation with very distinctly developed median concavity. For comparison with 
<taxonomicName id="4C3C4D4EFFB0283BFB8E64ED43E1FB87" box="[1063,1222,1004,1029]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="B948EADFFFB0283BFB8E64ED43E1FB87" box="[1063,1222,1004,1029]" italics="true" pageId="27" pageNumber="28">M. andersoni</emphasis>
</taxonomicName>
see this species.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB0283DFE91633D459DFEB4" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8B8336CDFFB0283BFE91633D465DFB4C" blockId="27.[264,1325,283,1870]" pageId="27" pageNumber="28">
<emphasis id="B948EADFFFB0283BFE91633D46F5FBD4" bold="true" box="[312,466,1084,1110]" pageId="27" pageNumber="28">Description:</emphasis>
Body elongate oval, maximum width on posterior margin of pronotum; elytra narrowed posteriad, evenly convex in lateral view, with only very indistinct depression in basal 0.2. Length: 
<quantity id="4CC49B28FFB0283BFD34638D4417FB24" box="[669,816,1164,1190]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="27" pageNumber="28" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 mm</quantity>
(HT: 
<quantity id="4CC49B28FFB0283BFC29638D44C4FB24" box="[896,995,1164,1190]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="28" unit="mm" value="2.1">2.1 mm</quantity>
); width: 
<quantity id="4CC49B28FFB0283BFBFD638D43C0FB24" box="[1108,1255,1164,1190]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="27" pageNumber="28" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
(HT: 
<quantity id="4CC49B28FFB0283BFEA163B54649FB4C" box="[264,366,1204,1230]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="27" pageNumber="28" unit="mm" value="1.35">1.35mm</quantity>
).
</paragraph>
<paragraph id="8B8336CDFFB0283BFE9163DD444CFAC4" blockId="27.[264,1325,283,1870]" pageId="27" pageNumber="28">Coloration: Dorsal side brown, with slightly paler anterior margin of clypeus, lateral margins of pronotum and lateral margins of elytra. Ventral side reddish brown; mouthparts and antenna except antennal club paler, ferruginous.</paragraph>
<paragraph id="8B8336CDFFB0283BFE91625543B3F97C" blockId="27.[264,1325,283,1870]" pageId="27" pageNumber="28">Head: Clypeus with moderately coarse and not very densely setiferous punctation, interstices without microsculpture; anterior margin straight, finely rimmed. Lines dividing clypeus from vertex distinct even paramesally. Vertex with coarse and dense setiferous punctation, interstices smooth. Eyes quite small, divided by 9× width of one eye. Mentum slightly wider than long, anterior margin slightly bisinuate; surface with fine and sparse setiferous punctation, interstices smooth. Antenna with 9 antennomeres; scapus ca. as long as antennal club; antennomeres 2–6 shorter than scapus when combined; antennal club elongate oval, antennomere 7 slightly longer than antennomere 8; antennomere 9 slightly longer than antennomere 7, with acutely pointed apex and distinct subapical strangulation. Maxillary palpomere 2 ca. as long as palpomere 4, slightly widened apically; palpomere 3 shorter than palpomere 2, slightly swollen apically; palpomere 4 spindle­like.</paragraph>
<paragraph id="8B8336CDFFB0283CFE91600D44B5FE54" blockId="27.[264,1325,283,1870]" lastBlockId="28.[264,1324,284,1870]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Prothorax: Pronotum strongly arcuately narrowed anteriad, maximum width in posterior 0.25, Lateral margins slightly explanate; posterior margin slightly bisinuate; posterolateral corners only indistinctly pronounced. Posterior margin with slight but distinct impression in lateral 0.25 at posterior margin. Punctation coarse and dense, bearing long pubescence, interstices without microsculpture. Prosternum distinctly carinate medially; anteromaedian excision deep and narrow; lateral parts indistinctly divided from median area, dividing line nearly straight.</paragraph>
<paragraph id="8B8336CDFFB7283CFE9166E5445FFC0C" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">Mesothorax: Scutellar shield in form of equilateral triangle, bearing a few coarse, setiferous punctures, interstices smooth. Elytron with 9 punctural series; series 1–5 and 8–9 reaching elytral base; series 7 and 8 arising subbasally. Intervals strongly elevate, costate, bearing very fine and sparse hair­bearing punctation. Intervals lowly costate, evenly elevate at elytral base; intervals 2, 4 and 6 becoming slightly higher than adjacent intervals posteriad. Lateral margins with moderately large obtuse denticles basally, becoming finer and more distant apicad, reaching 0.75 of elytral length. Epipleura wider than pseudepipleura basally, reaching posterior margin of metaventrite; pseudepipleura narrowing posteriad, reaching elytral apex. Preepisternal elevation with subrhomboid plate 2.0× longer than wide, with elevate narrow lateral edge and deeply concave median area. Grooves for reception of procoxae shallow and wide.</paragraph>
<paragraph id="8B8336CDFFB7283CFE91649D46D3FBFC" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Metathorax: Metaventrite with pentagonal elevate median area bearing very sparse and fine setiferous punctation, interstices without microsculpture; median part with very distinct circular depression. Lateral parts of metaventrite with fine microreticulation and sparse, very fine hair­bearing punctures. Postcoxal ridges bent posteriad medially, forming a longitudinal ridge reaching anterior 0.2 of metaventrite length. Femoral lines missing. 
<collectingCountry id="F32B765DFFB7283CFEA163654664FBFC" box="[264,323,1124,1150]" name="India" pageId="28" pageNumber="29">Hind</collectingCountry>
wings present.
</paragraph>
<paragraph id="8B8336CDFFB7283CFE91638D44A3FB74" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">Legs short; femora with moderately strong but very sparse setiferous punctation, interstices without microsculpture; tibiae bearing small and fine spines; tarsi distincly shorter than femora, bearing numerous hairs ventrally.</paragraph>
<paragraph id="8B8336CDFFB7283CFE9162054485FAEC" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">Abdominal ventrite 1 carinate medially, bearing additional longitudinal ridges paramesally. Ventrites 2–5 finely microsculptured in apical part, smooth and very indistinctly and sparsely punctured on posterior margins.</paragraph>
<paragraph id="8B8336CDFFB7283CFE91627D4655F98C" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">Male genitalia: Parameres ca. 1.5× longer than phallobase, continuously narrowing apicad, apices slightly bent outwards, bearing two small setae. Median lobe slightly widened apicad, with wide and obtusely angulate apical part; apex with small circular excision.</paragraph>
<paragraph id="8B8336CDFFB7283DFE91611D459DFEB4" blockId="28.[264,1324,284,1870]" lastBlockId="29.[264,1323,284,830]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B948EADFFFB7283CFE91611D46E4F9B4" bold="true" box="[312,451,1564,1590]" pageId="28" pageNumber="29">Variability.</emphasis>
Moderately variable species in some external characters. Pronotum bears very shallow median longitudinal furrow in some specimens. Elytra are quite constant in most characters, only with lateral denticulation reaching to the subapical area (then small and rather distant denticles can be observed near elytral apex). Preepisternal elevation 1.6–2.0× longer than wide. Median depression on the metaventrite can have two much smaller lateral impressions without any punctures on their bottom, or can be slightly elongate rather than circular in some specimens; it is very shallow in some specimens. The shape of elytra in lateral view is constant in all examined specimens except of two males from 
<collectingCountry id="F32B765DFFB6283DFEE1661D4685FEB4" box="[328,418,284,310]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
(see discussion below).
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB6283DFE916642423CFDF4" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8B8336CDFFB6283DFE916642423CFDF4" blockId="29.[264,1323,284,830]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFE91664246E4FEDF" bold="true" box="[312,451,323,349]" pageId="29" pageNumber="30">Discussion.</emphasis>
Two males from prov. Oaxaca (
<collectingCountry id="F32B765DFFB6283DFCCE664544E3FEDC" box="[871,964,324,350]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
) listed under the Additional material examined above differ from the other examined specimens of 
<taxonomicName id="4C3C4D4EFFB6283DFBC4666D4224FE07" box="[1133,1283,364,389]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB6283DFBC4666D4224FE07" box="[1133,1283,364,389]" italics="true" pageId="29" pageNumber="30">M. obscurus</emphasis>
</taxonomicName>
by slightly depressed elytra in lateral view and slightly more elongate apex of the median lobe. As both these differences from usual specimens of 
<taxonomicName id="4C3C4D4EFFB6283DFC1966BD4364FE57" box="[944,1091,444,469]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB6283DFC1966BD4364FE57" box="[944,1091,444,469]" italics="true" pageId="29" pageNumber="30">M. obscurus</emphasis>
</taxonomicName>
are very slight, we decided to treat these Mexican specimens as 
<taxonomicName id="4C3C4D4EFFB6283DFCB566E54489FE7F" box="[796,942,484,509]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB6283DFCB566E54489FE7F" box="[796,942,484,509]" italics="true" pageId="29" pageNumber="30">M. obscurus</emphasis>
</taxonomicName>
for the time being. However, as the lateral shape of elytra is very constant in all 
<taxonomicName id="4C3C4D4EFFB6283DFC28650D4327FDA7" box="[897,1024,524,549]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFFB6283DFC28650D4327FDA7" box="[897,1024,524,549]" italics="true" pageId="29" pageNumber="30">Motonerus</emphasis>
</taxonomicName>
species, these Mexican specimens may represent a separate species closely allied to 
<taxonomicName id="4C3C4D4EFFB6283DFC7665354356FDCF" box="[991,1137,564,589]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFFB6283DFC7665354356FDCF" box="[991,1137,564,589]" italics="true" pageId="29" pageNumber="30">M. obscurus</emphasis>
</taxonomicName>
. More material from 
<collectingCountry id="F32B765DFFB6283DFEE1655D4686FDF4" box="[328,417,604,630]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
needs to be examined before a determination can be made with confidence.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB6283DFE9165854302FD6C" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="8B8336CDFFB6283DFE9165854302FD6C" blockId="29.[264,1323,284,830]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFE9165854687FD1C" bold="true" box="[312,416,644,670]" pageId="29" pageNumber="30">Biology:</emphasis>
The 
<typeStatus id="5487886FFFB6283DFE4D65854531FD1C" box="[484,534,644,670]" pageId="29" pageNumber="30">type</typeStatus>
series including the 
<typeStatus id="5487886FFFB6283DFCBE65854459FD1C" box="[791,894,644,670]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
was collected in a broadleaf cloud forest with 
<taxonomicName id="4C3C4D4EFFB6283DFE2665AD46D6FD47" box="[399,497,684,709]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="B948EADFFFB6283DFE2665AD46D6FD47" box="[399,497,684,709]" italics="true" pageId="29" pageNumber="30">Cyathea</emphasis>
</taxonomicName>
tree ferns and 
<taxonomicName id="4C3C4D4EFFB6283DFD0A65AD4416FD47" box="[675,817,684,709]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="B948EADFFFB6283DFD0A65AD4416FD47" box="[675,817,684,709]" italics="true" pageId="29" pageNumber="30">Podocarpus</emphasis>
</taxonomicName>
conifers, in which the litter was sifted and carrion and dung baited pitfall traps were set (S. Peck, pers.comm.).
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB6283DFE9165FD4543FC5B" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8B8336CDFFB6283DFE9165FD45C5FCBC" blockId="29.[264,1323,284,830]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFE9165FD46FDFC94" bold="true" box="[312,474,764,790]" pageId="29" pageNumber="30">Distribution:</emphasis>
Confirmed from 
<collectingCountry id="F32B765DFFB6283DFD0765FD440DFC94" box="[686,810,764,790]" name="Nicaragua" pageId="29" pageNumber="30">Nicaragua</collectingCountry>
, 
<collectingCountry id="F32B765DFFB6283DFC9E65FD448EFC94" box="[823,937,764,790]" name="Honduras" pageId="29" pageNumber="30">Honduras</collectingCountry>
and 
<collectingCountry id="F32B765DFFB6283DFC4E65FD4351FC94" box="[999,1142,764,790]" name="El Salvador" pageId="29" pageNumber="30">El Salvador</collectingCountry>
; its occurrence in 
<collectingCountry id="F32B765DFFB6283DFE8F64254658FCBC" box="[294,383,804,830]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
requires further confirmation.
</paragraph>
<paragraph id="8B8336CDFFB6283DFEA1649D4543FC5B" blockId="29.[264,851,923,985]" pageId="29" pageNumber="30">
<taxonomicName id="4C3C4D4EFFB6283DFEA1649D454CFC34" authority="Fika" authorityName="Fika" box="[264,619,924,950]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oosternoides">
<emphasis id="B948EADFFFB6283DFEA1649D454CFC34" bold="true" box="[264,619,924,950]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFEA1649D450CFC34" bold="true" box="[264,555,924,950]" italics="true" pageId="29" pageNumber="30">Motonerus oosternoides</emphasis>
Fiká
</emphasis>
</taxonomicName>
č 
<emphasis id="B948EADFFFB6283DFDD1649D4474FC34" bold="true" box="[632,851,924,950]" pageId="29" pageNumber="30">ek &amp; Short, sp. n.</emphasis>
(
<figureCitation id="13072A48FFB6283DFEB864BE46A6FC5B" box="[273,385,959,985]" captionStart="FIGURES 5 – 12" captionStartId="18.[264,382,1767,1791]" captionTargetBox="[292,1300,326,1748]" captionTargetId="figure@18.[292,1300,326,1748]" captionTargetPageId="18" captionText="FIGURES 5 – 12. Male genitalia of Motonerus. 5 – 6: M. apterus sp. n., paratype; 7 – 8: M. oosternoides sp. n., paratype; 9 – 10: M. depressus sp. n., holotype; 11 – 12: M. nublado, holotype. 5, 7, 9, 11 — aedeagus (scale bar = 0.1 mm); 6, 8, 10, 12 — apical part of median lobe (scale bar = 0.05 mm)." httpUri="https://zenodo.org/record/173211/files/figure.png" pageId="29" pageNumber="30">Figs. 7–8</figureCitation>
, 
<figureCitation id="13072A48FFB6283DFE2464BE468AFC5B" box="[397,429,959,985]" captionStart="FIGURES 21 – 31" captionStartId="20.[264,382,1715,1739]" captionTargetBox="[264,1323,326,1691]" captionTargetId="figure@20.[264,1323,326,1691]" captionTargetPageId="20" captionText="FIGURES 21 – 31. Lateral habitus of Motonerus. 21: M. apterus sp. n.; 22: M. oosternoides sp. n.; 23: M. depressus sp. n.; 24: M. nublado sp. n.; 25: M. andersoni sp. n.; 26: M. hanseni sp. n.; 27: M. problematicus sp. n.; 28: M. obscurus Hansen (El Salvador); 29: M. obscurus Hansen (Oaxaca prov., Mexico); 30: M. sp. B; 31: M. sp. A. Scale bar = 1 mm." httpUri="https://zenodo.org/record/173213/files/figure.png" pageId="29" pageNumber="30">22</figureCitation>
, 
<figureCitation id="13072A48FFB6283DFE1064BE46FFFC5B" box="[441,472,959,985]" captionStart="FIGURES 32 – 40" captionStartId="21.[264,383,1661,1685]" captionTargetBox="[306,1288,326,1631]" captionTargetId="figure@21.[291,1296,326,1637]" captionTargetPageId="21" captionText="FIGURES 32 – 40. Preepisternal elevation of mesothorax and median part of metaventrite of Motonerus. 32: M. apterus sp. n.; 33: M. oosternoides sp. n.; 34: M. depressus sp. n.; 35: M. andersoni sp. n.; 36: M. hanseni sp. n.; 37: M. nublado sp. n.; 38: M. obscurus Hansen; 39: M. problematicus sp. n.; 40: M. sp. A. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173214/files/figure.png" pageId="29" pageNumber="30">33</figureCitation>
, 
<figureCitation id="13072A48FFB6283DFE4C64BE4522FC5B" box="[485,517,959,985]" captionStart="FIGURES 41 – 53" captionStartId="22.[264,382,1710,1734]" captionTargetBox="[264,1323,326,1686]" captionTargetId="figure@22.[264,1323,326,1686]" captionTargetPageId="22" captionText="FIGURES 41 – 53. Details of of Motonerus. 41, 49: M. sp. B; 42, 52: M. andersoni sp. n.; 43, 53: M. apterus sp. n.; 44: M. depressus sp. n.; 45: M. hanseni sp. n.; 46: M. oosternoides sp. n.; 47: M. nublado sp. n.; 48: M. obscurus Hansen; 50: M. problematicus sp. n.; 51: M. sp. A. 41 — preepisternal elevation of mesothorax and median part of metaventrite (scale bar = 0.2 mm); 42 – 51 — median part of prosternum; 52 – 53 — sternite 9 of male." httpUri="https://zenodo.org/record/173215/files/figure.png" pageId="29" pageNumber="30">46</figureCitation>
, 
<figureCitation id="13072A48FFB6283DFDB864BE457BFC5B" box="[529,604,959,985]" captionStart="FIGURES 54 – 64" captionStartId="23.[264,382,1763,1787]" captionTargetBox="[264,1323,326,1738]" captionTargetId="figure@23.[264,1323,326,1738]" captionTargetPageId="23" captionText="FIGURES 54 – 64. Details of Motonerus. 54, 60, 63: M. apterus sp. n.; 55, 64: M. oosternoides sp. n.; 56: M. depressus sp. n.; 57, 62: M. nublado sp. n.; 59: M. problematicus sp. n.; 61: M. obscurus Hansen. 54 – 59 — epipleuron and lateral part of elytron; 60 – 62 — abdominal ventrites; 63 – 64 — head, lateral view. Scale bar = 0.5 mm (54 – 59), 0.25 mm (60 – 62); 0.1 mm (63 – 64)." httpUri="https://zenodo.org/record/173216/files/figure.png" pageId="29" pageNumber="30">55, 64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB6283EFEA1630F43E4FEB4" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8B8336CDFFB6283DFEA1630F4361FBAB" blockId="29.[264,1323,1038,1905]" box="[264,1094,1038,1065]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFEA1630F4695FBAA" bold="true" box="[264,434,1038,1064]" pageId="29" pageNumber="30">
<typeStatus id="5487886FFFB6283DFEA1630F4662FBAA" box="[264,325,1038,1064]" pageId="29" pageNumber="30">Type</typeStatus>
Locality
</emphasis>
. 
<collectingCountry id="F32B765DFFB6283DFE69630E4567FBAB" box="[448,576,1039,1065]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
, Chiriqui Prov., 
<quantity id="4CC49B28FFB6283DFCA9630E4475FBAB" box="[768,850,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="29" pageNumber="30" unit="km" value="5.6">5.6 km</quantity>
N Boquete, 
<quantity id="4CC49B28FFB6283DFC4F630E4364FBAB" box="[998,1091,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800 m</quantity>
.
</paragraph>
<paragraph id="8B8336CDFFB6283DFE9163374599FAEB" blockId="29.[264,1323,1038,1905]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFE916337454BFBD2" bold="true" box="[312,620,1078,1104]" pageId="29" pageNumber="30">
<typeStatus id="5487886FFFB6283DFE9163374652FBD2" box="[312,373,1078,1104]" pageId="29" pageNumber="30">Type</typeStatus>
Material. 
<typeStatus id="5487886FFFB6283DFE536337454BFBD2" box="[506,620,1078,1104]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
♂ (
<collectionCode id="ED2DAE08FFB6283DFD3F633645D5FBD3" box="[662,754,1079,1105]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="29" pageNumber="30">ZMUC</collectionCode>
): “ 
<collectingCountry id="F32B765DFFB6283DFCB9633644B3FBD3" box="[784,916,1079,1105]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
: Chiriqui / 
<quantity id="4CC49B28FFB6283DFBB263364343FBD3" box="[1051,1124,1079,1105]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="29" pageNumber="30" unit="km" value="5.6">5.6km</quantity>
N Boquete, / La Culebra Trail, 
<quantity id="4CC49B28FFB6283DFE69635E4507FBFB" box="[448,544,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800 m</quantity>
/ 
<date id="FF82100DFFB6283DFDED635E45F3FBFB" box="[580,724,1119,1145]" pageId="29" pageNumber="30" value="1996-06-15">15.Jun.1996</date>
R. Anderson / wet cloud forest litt. 96­131B”. 
<emphasis id="B948EADFFFB6283DFEA1638746EDFB22" bold="true" box="[264,458,1158,1184]" pageId="29" pageNumber="30">
<typeStatus id="5487886FFFB6283DFEA1638746A2FB22" box="[264,389,1158,1184]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
(47):
</emphasis>
<collectingCountry id="F32B765DFFB6283DFE7D63864572FB23" box="[468,597,1159,1185]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
: same data as 
<typeStatus id="5487886FFFB6283DFCAC63864449FB23" box="[773,878,1159,1185]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, 2♂♂, Ψ, 41 unsexed specimens (
<collectionCode id="ED2DAE08FFB6283DFAAB6386460AFB4B" pageId="29" pageNumber="30">AE ZS</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFE9263AE46B3FB4B" box="[315,404,1199,1225]" pageId="29" pageNumber="30">CMNC</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFE0B63AE46CCFB4B" box="[418,491,1199,1225]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="29" pageNumber="30">CUIC</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFE5163AE456EFB4B" box="[504,585,1199,1225]" pageId="29" pageNumber="30">MFPC</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFDFE63AE4585FB4B" box="[599,674,1199,1225]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="29" pageNumber="30">NMW</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFD1963AE442FFB4B" LSID="urn:lsid:biocol.org:col:34871" box="[688,776,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30">USNM</collectionCode>
, 
<collectionCode id="ED2DAE08FFB6283DFCBC63AE4448FB4B" box="[789,879,1199,1225]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="29" pageNumber="30">ZMUC</collectionCode>
); 
<collectingCountry id="F32B765DFFB6283DFC2C63AE4321FB4B" box="[901,1030,1199,1225]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
: CHIRIQUI / 
<quantity id="4CC49B28FFB6283DFB1963AE4224FB4B" box="[1200,1283,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="29" pageNumber="30" unit="km" value="5.6">5.6 km</quantity>
N. Bouquete / La Culebra Trail / 
<quantity id="4CC49B28FFB6283DFDDC63D645F7FB73" box="[629,720,1239,1265]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800 m</quantity>
/ 
<date id="FF82100DFFB6283DFD4263D64457FB73" box="[747,880,1239,1265]" pageId="29" pageNumber="30" value="1996-06-15">15.VI.1996</date>
/ R. Anderson, 96­131A / wet cloud forest litter”, 1 unsexed specimen (
<collectionCode id="ED2DAE08FFB6283DFD0B63FE45D8FA9B" LSID="urn:lsid:biocol.org:col:33144" box="[674,767,1279,1305]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33144" name="Colorado National Monument" pageId="29" pageNumber="30">CNMC</collectionCode>
); 
<collectingCountry id="F32B765DFFB6283DFCBD63FE44B2FA9B" box="[788,917,1279,1305]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
: Chiriqui / P. Int. La Amistad, Las Nubes / 
<quantity id="4CC49B28FFB6283DFEC26226469BFAC3" box="[363,444,1319,1345]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="29" pageNumber="30" unit="km" value="5.3">5.3 km</quantity>
W Cerro Punta, 
<quantity id="4CC49B28FFB6283DFD2D622645F9FAC3" box="[644,734,1319,1345]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="29" pageNumber="30" unit="m" value="2150.0">2150 m</quantity>
/ 
<date id="FF82100DFFB6283DFD5C622644A3FAC3" box="[757,900,1319,1345]" pageId="29" pageNumber="30" value="1995-06-15">15.Jun.1995</date>
R.Anderson / cloud forest litter 95­ 026”, 2 unsexed specimens (
<collectionCode id="ED2DAE08FFB6283DFDFF624E4595FAEB" box="[598,690,1359,1385]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="29" pageNumber="30">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="8B8336CDFFB6283DFE91627744E8F903" blockId="29.[264,1323,1038,1905]" pageId="29" pageNumber="30">
<emphasis id="B948EADFFFB6283DFE9162774576FA12" bold="true" box="[312,593,1398,1424]" pageId="29" pageNumber="30">Differential Diagnosis.</emphasis>
Body smaller than 
<quantity id="4CC49B28FFB6283DFCEB62764485FA13" box="[834,930,1399,1425]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="mm" value="1.8">1.8 mm</quantity>
; elytra evenly rounded in lateral view; elytral intervals almost evenly elevate thoughout; lateral margins with fine denticulation reaching elytral apex; anteromedian prosternal excision with obtuse “tooth” medially; preepisternal elevation of mesothorax lamellar, longer than metaventrite, without median concave area; metaventrite without median depression; abdominal ventrites without additional submedian longitudinal ridges; hind wings absent; median lobe with slightly widened serrate apex with apical excision.
</paragraph>
<paragraph id="8B8336CDFFB6283EFE91618E43E4FEB4" blockId="29.[264,1323,1038,1905]" lastBlockId="30.[264,1324,284,1870]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
Very similar to 
<taxonomicName id="4C3C4D4EFFB6283DFE5C618E4550F92A" box="[501,631,1679,1704]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="B948EADFFFB6283DFE5C618E4550F92A" box="[501,631,1679,1704]" italics="true" pageId="29" pageNumber="30">M. apterus</emphasis>
</taxonomicName>
(see under this species for comparison with it as well as with other 
<taxonomicName id="4C3C4D4EFFB6283DFE3B61B64536F952" box="[402,529,1719,1744]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFFB6283DFE3B61B64536F952" box="[402,529,1719,1744]" italics="true" pageId="29" pageNumber="30">Motonerus</emphasis>
</taxonomicName>
species) The combination of lamellar preepisternal elevation of mesothorax and missing longitudinal submedian ridges on abdominal ventrites can resemble 
<taxonomicName id="4C3C4D4EFFB6283DFE2D6006450EF8A2" box="[388,553,1799,1824]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B948EADFFFB6283DFE2D6006450EF8A2" box="[388,553,1799,1824]" italics="true" pageId="29" pageNumber="30">M. depressus</emphasis>
</taxonomicName>
, however both 
<taxonomicName id="4C3C4D4EFFB6283DFD56600644E0F8A2" box="[767,967,1799,1824]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oosternoides">
<emphasis id="B948EADFFFB6283DFD56600644E0F8A2" box="[767,967,1799,1824]" italics="true" pageId="29" pageNumber="30">M. oosternoides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D4EFFB6283DFBBD600643B9F8A2" box="[1044,1182,1799,1824]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="B948EADFFFB6283DFBBD600643B9F8A2" box="[1044,1182,1799,1824]" italics="true" pageId="29" pageNumber="30">M. apterus</emphasis>
</taxonomicName>
are easily distinguishable from it by evenly and highly convex elytra, much smaller body size, paler general coloration, distinctly developed anteromedian prosternal elevation, missing coarse crenulation on anterior part of the abdominal ventrite 1 and by the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB5283FFE916642446DFBD4" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="8B8336CDFFB5283EFE9166424695FE54" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">
<emphasis id="B948EADFFFB5283EFE91664246F7FEDF" bold="true" box="[312,464,323,349]" pageId="30" pageNumber="31">Description.</emphasis>
Body elongate oval, maximum width closely before posterior margin of pronotum; elytra narrowing posteriad, evenly highly convex in lateral view; pronotum slightly convex in lateral view. Length: 
<quantity id="4CC49B28FFB5283EFD4766954459FE2C" box="[750,894,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="30" pageNumber="31" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
(HT: 
<quantity id="4CC49B28FFB5283EFC626695430AFE2C" box="[971,1069,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="30" pageNumber="31" unit="mm" value="1.8">1.8 mm</quantity>
); width: 1.0– 
<quantity id="4CC49B28FFB5283EFB796695420CFE2C" box="[1232,1323,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="30" pageNumber="31" unit="mm" value="1.1">1.1 mm</quantity>
(HT: 
<quantity id="4CC49B28FFB5283EFEEE66BD4682FE54" box="[327,421,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="30" pageNumber="31" unit="mm" value="1.1">1.1 mm</quantity>
).
</paragraph>
<paragraph id="8B8336CDFFB5283EFE9166E54641FDF4" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">Coloration: Dorsal side brown; clypeus and lateral margins of pronotum and elytra slightly paler. Ventral side reddish brown; mentum, gula and legs slightly paler. Mouthparts and antennae yellowish brown, terminal antennomeres of antennal club whitish.</paragraph>
<paragraph id="8B8336CDFFB5283EFE916585434BFBD4" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">Head: Clypeus with fine and sparse punctation, interstices without microsculpture; anterior margin straight, narrowly rimmed. Lines dividing clypeus from vertex distinct even paramesally. Vertex with setiferous punctation much coarser and denser than on clypeus, interstices without microsculpture. Eyes small, separated by 10× width of one eye. Mentum slightly wider than long, with deeply bisinuate anterior margin; punctation coarse and moderately dense, interstices without microsculpture. Maxillary palpomere 2 as long as palpomere 4, distinctly swollen apically; palpomere 3 shorter than palpomere 2, short, stout, slightly swollen apically; palpomere 4 rather stout, spindle­like. Antenna with 9 antennomeres; scapus slightly longer than antennal club; antennomeres 2–6 small, slightly shorter than antennal club when combined; antennal club compact, elongate oval; antennomere 8 slightly shorter than antennomere 7; antennomere 9 slightly longer than antennomere 7, obtusely pointed, without distinct subapical strangulation.</paragraph>
<paragraph id="8B8336CDFFB5283EFE9163654412FAEC" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">Prothorax: Pronotum strongly arcuately narrowing anteriad, maximum width closely before posterior margin. Lateral margins slightly explanate; posterior margin bisinuate, posterolateral corners slightly pronounced. Sublaterally on posteior margin with very weak impression. Punctation moderately strong, sparse, consisting of setiferous punctures; interstices without microsculpture. Prosternum weakly carinate medially; anteromedian excision shallow and wide, with obtuse “tooth” medially; lateral parts indistinctly divided from median area, dividing line convex laterad.</paragraph>
<paragraph id="8B8336CDFFB5283EFE91627D4593F97C" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">Mesothorax: Scutellar shield in form of equilateral triangle, bearing a few fine punctures. Elytron with 9 punctural series; series 1–5 and 8 reaching elytral base; series 6, 7 a 9 arising subbasally. Elytral intervals weakly costate, evenly elevate basally; intervals 2 and 4 slightly narrower than adjacent intervals apically. Intervals with fine punctation bearing protruding bent hairs. Lateral margin of elytron with fine, equidistant denticles reaching elytral apex. Epipleuron slightly narrower than pseudepipleuron basally, reaching level of metathorax; pseudepipleuron narrowing posteriad, reaching elytral apex. Preepisternal elevation as an elevate bulge with narrow longitudinal carina medially, distinctly developed in anterior 0.5, weakly developed posteriorly. Grooves for reception of procoxae deep and not very wide.</paragraph>
<paragraph id="8B8336CDFFB5283EFE91600D420CF8CC" blockId="30.[264,1324,284,1870]" pageId="30" pageNumber="31">Metathorax: Metaventrite shorter than preepisternal elevation, without median depression, bearing fine setiferous punctation, interstices without microsculpture.</paragraph>
<paragraph id="8B8336CDFFB4283FFEA1661D4565FEDC" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">
Postcoxal ridges bent posteriad medially, forming ridge reaching 0.25 of metaventrite length. 
<collectingCountry id="F32B765DFFB4283FFEC9664546BCFEDC" box="[352,411,324,350]" name="India" pageId="31" pageNumber="32">Hind</collectingCountry>
wings absent.
</paragraph>
<paragraph id="8B8336CDFFB4283FFE91666D45CAFE54" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">Legs short; femora bearing very sparse and fine setiferous punctures, interstices shiny, without microsculpture. Tibiae slightly longer than tarsi, bearing numerous very fine spines. Tarsi densely pubescent ventrally.</paragraph>
<paragraph id="8B8336CDFFB4283FFE9166E5464CFDCC" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">Abdominal ventrite 1 bearing median longitudinal carina, additional submedian longitudinal ridges missing on all ventrites; fine crenulation of anterior part of ventrite 1 missing.</paragraph>
<paragraph id="8B8336CDFFB4283FFE91655D43B5FD6C" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">Male genitalia: Parameres ca. 1.6 longer than phalobase, gradually narrowing apicad, slightly bent outwards subapically, bearing a few small setae apically. Median lobe slightly longer than parameres, considerably narrowing in apical 0.15, slightly widened and bearing serrate rim apically; the rim interrupted by a distinct incision at apex.</paragraph>
<paragraph id="8B8336CDFFB4283FFE9165FD446DFBD4" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE9165FD46E4FC94" bold="true" box="[312,451,764,790]" pageId="31" pageNumber="32">Variability.</emphasis>
Very uniform species in most external characters. The only observed variability concerns the morphology of the preepisternal elevation of the mesothorax and the metaventrite. In some specimens, the posterior part of the preepisternal elevation is vaguely defined and the median ridge (formed by postcoxal ridges bent posteriad mesally) reaches nearly the posterior margin of metaventrite. The aedeagus of these specimens is, however, completely identical to that of the 
<typeStatus id="5487886FFFB4283FFCB364C544A3FC5C" box="[794,900,964,990]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
. Moreover, intermediate specimens with the metasternal median ridge only slightly protracted posteriad and only indistinctly narrowed preepisternal elevation were found. Thus, we consider this variability as intraspecific, without any taxonomic importance.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB4283FFE916365455BFA9C" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8B8336CDFFB4283FFE91636545D7FB4C" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE91636546E3FBFC" bold="true" box="[312,452,1124,1150]" pageId="31" pageNumber="32">Etymology.</emphasis>
<taxonomicName id="4C3C4D4EFFB4283FFE7A63654549FBFC" box="[467,622,1124,1150]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="36" phylum="Arthropoda" rank="species" species="oosternoides">Oosternoides</taxonomicName>
= resembling 
<taxonomicName id="4C3C4D4EFFB4283FFC8563654489FBFF" box="[812,942,1124,1149]" class="Insecta" family="Hydrophilidae" genus="Oosternum" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFFB4283FFC8563654489FBFF" box="[812,942,1124,1149]" italics="true" pageId="31" pageNumber="32">Oosternum</emphasis>
</taxonomicName>
, reflecting the small size and 
<taxonomicName id="4C3C4D4EFFB4283FFEA1638D46ADFB27" box="[264,394,1164,1189]" class="Insecta" family="Hydrophilidae" genus="Oosternum" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFFB4283FFEA1638D46ADFB27" box="[264,394,1164,1189]" italics="true" pageId="31" pageNumber="32">Oosternum</emphasis>
</taxonomicName>
­like habitus of the species. This name was originally given to this species in notes written by the late Michael Hansen.
</paragraph>
<paragraph id="8B8336CDFFB4283FFE9163DD455BFA9C" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE9163DD46B8FB74" bold="true" box="[312,415,1244,1270]" pageId="31" pageNumber="32">Habitat.</emphasis>
All specimens examined were sifted from the cloud forest litter in elevations between 
<date id="FF82100DFFB4283FFED862054507FA9C" box="[369,544,1284,1310]" pageId="31" pageNumber="32" value="1800" valueMax="2150">
1800 and 
<quantity id="4CC49B28FFB4283FFE4C62054507FA9C" box="[485,544,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="31" pageNumber="32" unit="m" value="2150.0">2150</quantity>
</date>
m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB4283FFE91622D4572F98B" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8B8336CDFFB4283FFE91622D464BFAEC" blockId="31.[264,1323,284,1390]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE91622D46FFFAC4" bold="true" box="[312,472,1324,1350]" pageId="31" pageNumber="32">Distribution.</emphasis>
The species is known from two closely situated localities in western 
<collectingCountry id="F32B765DFFB4283FFEA16255464FFAEC" box="[264,360,1364,1390]" name="Panama" pageId="31" pageNumber="32">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B8336CDFFB4283FFEA162CD4572F98B" blockId="31.[264,871,1483,1545]" pageId="31" pageNumber="32">
<taxonomicName id="4C3C4D4EFFB4283FFEA162CD4559FA64" authority="Fika" authorityName="Fika" box="[264,638,1484,1510]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B948EADFFFB4283FFEA162CD4559FA64" bold="true" box="[264,638,1484,1510]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFEA162CD4519FA64" bold="true" box="[264,574,1484,1510]" italics="true" pageId="31" pageNumber="32">Motonerus problematicus</emphasis>
Fiká
</emphasis>
</taxonomicName>
č 
<emphasis id="B948EADFFFB4283FFD2562CD4440FA64" bold="true" box="[652,871,1484,1510]" pageId="31" pageNumber="32">ek &amp; Short, sp. n.</emphasis>
(
<figureCitation id="13072A48FFB4283FFEB862EE46B9F98B" box="[273,414,1519,1545]" captionStart="FIGURES 13 – 20" captionStartId="19.[264,382,1761,1785]" captionTargetBox="[305,1289,326,1738]" captionTargetId="figure@19.[305,1289,326,1738]" captionTargetPageId="19" captionText="FIGURES 13 – 20. Male genitalia of Motonerus. 13 – 14: M. andersoni sp. n., paratype; 15 – 16: M. hanseni sp. n., holotype; 17 – 18: M. obscurus Hansen, paratype; 19 – 20: M. problematicus sp. n., holotype. 13, 15, 17, 19 — aedeagus (scale bar = 0.1 mm); 14, 16, 18, 20 — apical part of median lobe (scale bar = 0.05 mm)." httpUri="https://zenodo.org/record/173212/files/figure.png" pageId="31" pageNumber="32">Figs. 19–20</figureCitation>
, 
<figureCitation id="13072A48FFB4283FFE0362EE46EDF98B" box="[426,458,1519,1545]" captionStart="FIGURES 21 – 31" captionStartId="20.[264,382,1715,1739]" captionTargetBox="[264,1323,326,1691]" captionTargetId="figure@20.[264,1323,326,1691]" captionTargetPageId="20" captionText="FIGURES 21 – 31. Lateral habitus of Motonerus. 21: M. apterus sp. n.; 22: M. oosternoides sp. n.; 23: M. depressus sp. n.; 24: M. nublado sp. n.; 25: M. andersoni sp. n.; 26: M. hanseni sp. n.; 27: M. problematicus sp. n.; 28: M. obscurus Hansen (El Salvador); 29: M. obscurus Hansen (Oaxaca prov., Mexico); 30: M. sp. B; 31: M. sp. A. Scale bar = 1 mm." httpUri="https://zenodo.org/record/173213/files/figure.png" pageId="31" pageNumber="32">27</figureCitation>
, 
<figureCitation id="13072A48FFB4283FFE7E62EE46D0F98B" box="[471,503,1519,1545]" captionStart="FIGURES 32 – 40" captionStartId="21.[264,383,1661,1685]" captionTargetBox="[306,1288,326,1631]" captionTargetId="figure@21.[291,1296,326,1637]" captionTargetPageId="21" captionText="FIGURES 32 – 40. Preepisternal elevation of mesothorax and median part of metaventrite of Motonerus. 32: M. apterus sp. n.; 33: M. oosternoides sp. n.; 34: M. depressus sp. n.; 35: M. andersoni sp. n.; 36: M. hanseni sp. n.; 37: M. nublado sp. n.; 38: M. obscurus Hansen; 39: M. problematicus sp. n.; 40: M. sp. A. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/173214/files/figure.png" pageId="31" pageNumber="32">39</figureCitation>
, 
<figureCitation id="13072A48FFB4283FFDAB62EE4505F98B" box="[514,546,1519,1545]" captionStart="FIGURES 41 – 53" captionStartId="22.[264,382,1710,1734]" captionTargetBox="[264,1323,326,1686]" captionTargetId="figure@22.[264,1323,326,1686]" captionTargetPageId="22" captionText="FIGURES 41 – 53. Details of of Motonerus. 41, 49: M. sp. B; 42, 52: M. andersoni sp. n.; 43, 53: M. apterus sp. n.; 44: M. depressus sp. n.; 45: M. hanseni sp. n.; 46: M. oosternoides sp. n.; 47: M. nublado sp. n.; 48: M. obscurus Hansen; 50: M. problematicus sp. n.; 51: M. sp. A. 41 — preepisternal elevation of mesothorax and median part of metaventrite (scale bar = 0.2 mm); 42 – 51 — median part of prosternum; 52 – 53 — sternite 9 of male." httpUri="https://zenodo.org/record/173215/files/figure.png" pageId="31" pageNumber="32">50</figureCitation>
, 
<figureCitation id="13072A48FFB4283FFD8762EE456AF98B" box="[558,589,1519,1545]" captionStart="FIGURES 54 – 64" captionStartId="23.[264,382,1763,1787]" captionTargetBox="[264,1323,326,1738]" captionTargetId="figure@23.[264,1323,326,1738]" captionTargetPageId="23" captionText="FIGURES 54 – 64. Details of Motonerus. 54, 60, 63: M. apterus sp. n.; 55, 64: M. oosternoides sp. n.; 56: M. depressus sp. n.; 57, 62: M. nublado sp. n.; 59: M. problematicus sp. n.; 61: M. obscurus Hansen. 54 – 59 — epipleuron and lateral part of elytron; 60 – 62 — abdominal ventrites; 63 – 64 — head, lateral view. Scale bar = 0.5 mm (54 – 59), 0.25 mm (60 – 62); 0.1 mm (63 – 64)." httpUri="https://zenodo.org/record/173216/files/figure.png" pageId="31" pageNumber="32">59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3266546FFB42800FEA1613F46F5FD6C" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8336CDFFB4283FFEA1613F43FBF9DB" blockId="31.[264,1323,1598,1905]" box="[264,1244,1598,1625]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFEA1613F468AF9DA" bold="true" box="[264,429,1598,1624]" pageId="31" pageNumber="32">
<typeStatus id="5487886FFFB4283FFEA1613F4662F9DA" box="[264,325,1598,1624]" pageId="31" pageNumber="32">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32B765DFFB4283FFE1C613E4544F9DB" box="[437,611,1599,1625]" name="Costa Rica" pageId="31" pageNumber="32">COSTA RICA</collectingCountry>
, Puntarenas Prov., Monteverde Cloud Forest Reserve.
</paragraph>
<paragraph id="8B8336CDFFB4283FFE9161674505F8A3" blockId="31.[264,1323,1598,1905]" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE9161674550F902" bold="true" box="[312,631,1638,1664]" pageId="31" pageNumber="32">
<typeStatus id="5487886FFFB4283FFE9161674654F902" box="[312,371,1638,1664]" pageId="31" pageNumber="32">Type</typeStatus>
Material. 
<typeStatus id="5487886FFFB4283FFDAC61674550F902" box="[517,631,1638,1664]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
♂ (
<collectionCode id="ED2DAE08FFB4283FFD046166442EF903" box="[685,777,1639,1665]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="31" pageNumber="32">ZMUC</collectionCode>
): “ 
<collectingCountry id="F32B765DFFB4283FFC84616644C0F903" box="[813,999,1639,1665]" name="Costa Rica" pageId="31" pageNumber="32">COSTA RICA</collectingCountry>
: Puntarenas / Monteverde Reserve / 
<date id="FF82100DFFB4283FFE2A618E450FF92B" box="[387,552,1679,1705]" pageId="31" pageNumber="32" value="1993-05-25">25 May 1993</date>
, C. Michalski / ex; flight intercept trap // 
<taxonomicName id="4C3C4D4EFFB4283FFB80618E438FF92B" box="[1065,1192,1679,1705]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus" status="sp. nov.">Motonerus</taxonomicName>
<taxonomicNameLabel id="A27B57A4FFB4283FFB1A618E43CDF92B" box="[1203,1258,1679,1705]" pageId="31" pageNumber="32" rank="species">n.sp.</taxonomicNameLabel>
/ M. Hansen det.”. 
<emphasis id="B948EADFFFB4283FFE1461B74429F952" bold="true" box="[445,782,1718,1744]" pageId="31" pageNumber="32">
<typeStatus id="5487886FFFB4283FFE1461B74514F952" box="[445,563,1718,1744]" pageId="31" pageNumber="32" type="paratype">Paratype</typeStatus>
: 
<collectingCountry id="F32B765DFFB4283FFDE161B7442EF952" box="[584,777,1718,1744]" name="Costa Rica" pageId="31" pageNumber="32">COSTA RICA</collectingCountry>
:
</emphasis>
“Quebrada Segunda, P.N. Tapanti A. C. / Amistad, Prov. Carta 
<collectingCountry id="F32B765DFFB4283FFDAA61DE4589F97B" box="[515,686,1759,1785]" name="Costa Rica" pageId="31" pageNumber="32">COSTA RICA</collectingCountry>
1150 / m, 
<date id="FF82100DFFB4283FFC8061DE44B0F97B" box="[809,919,1759,1785]" pageId="31" pageNumber="32" value="1994-07">Jul. 1994</date>
, G. Mora, L.N. / 194000­559800 # 3010”, female (
<collectionCode id="ED2DAE08FFB4283FFE1460064532F8A3" box="[445,533,1799,1825]" pageId="31" pageNumber="32">INBIO</collectionCode>
).
</paragraph>
<paragraph id="8B8336CDFFB42800FE91602F4457FE54" blockId="31.[264,1323,1598,1905]" lastBlockId="32.[264,1325,284,1870]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B948EADFFFB4283FFE91602F4572F8CA" bold="true" box="[312,597,1838,1864]" pageId="31" pageNumber="32">Differential Diagnosis.</emphasis>
Body larger than 2.0 mm; elytra depressed, not abruptly elevate behind pronotum in lateral view; elytral intervals evenly elevate throughout; elytral margin with fine and more or less equidistant denticles reaching elytral apex; preepisternal elevation of mesothorax 3.0–4.0× longer than wide; metaventrite with shallow median depression; abdominal ventrites 1–3 with longitudinal ridges (not as distinct on ventrites 2–3 as on ventrite 1); apex of median lobe gradually narrowing apicad from apical 0.25, with only minute incision on apex.
</paragraph>
<paragraph id="8B8336CDFF8B2800FE9166E546F5FD6C" blockId="32.[264,1325,284,1870]" pageId="32" pageNumber="33">
Most easily recognized by the combination of depressed elytra in lateral view (in contrast to 
<taxonomicName id="4C3C4D4EFF8B2800FE3F650D451EFDA7" box="[406,569,524,549]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="B948EADFFF8B2800FE3F650D451EFDA7" box="[406,569,524,549]" italics="true" pageId="32" pageNumber="33">M. andersoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D4EFF8B2800FDD4650D4433FDA7" box="[637,788,524,549]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFF8B2800FDD4650D4433FDA7" box="[637,788,524,549]" italics="true" pageId="32" pageNumber="33">M. obscurus</emphasis>
</taxonomicName>
) not abruptly elevate behind pronotum (in contrast to 
<taxonomicName id="4C3C4D4EFF8B2800FE3E65354503FDCF" box="[407,548,564,589]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nublado">
<emphasis id="B948EADFFF8B2800FE3E65354503FDCF" box="[407,548,564,589]" italics="true" pageId="32" pageNumber="33">M. nublado</emphasis>
</taxonomicName>
), elytral intervals evenly elevate thoughout (in contrast to 
<taxonomicName id="4C3C4D4EFF8B2800FAA265354655FDF7" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFF8B2800FAA265354655FDF7" italics="true" pageId="32" pageNumber="33">M. obscurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D4EFF8B2800FE05655D4508FDF7" box="[428,559,604,629]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hanseni">
<emphasis id="B948EADFFF8B2800FE05655D4508FDF7" box="[428,559,604,629]" italics="true" pageId="32" pageNumber="33">M. hanseni</emphasis>
</taxonomicName>
), preepisternal elevation 3.0–4.0× longer than wide and in some cases with weak, very shallow median concavity, and denticulation of elytral margins equidistant or nearly so and reaching elytral apex (in contrast to 
<taxonomicName id="4C3C4D4EFF8B2800FBAF65AD43AEFD47" box="[1030,1161,684,709]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hanseni">
<emphasis id="B948EADFFF8B2800FBAF65AD43AEFD47" box="[1030,1161,684,709]" italics="true" pageId="32" pageNumber="33">M. hanseni</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D4EFF8B2800FB3065AD420CFD47" box="[1177,1323,684,709]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B948EADFFF8B2800FB3065AD420CFD47" box="[1177,1323,684,709]" italics="true" pageId="32" pageNumber="33">M. obscurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D4EFF8B2800FE9365D546E6FD6F" box="[314,449,724,749]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nublado">
<emphasis id="B948EADFFF8B2800FE9365D546E6FD6F" box="[314,449,724,749]" italics="true" pageId="32" pageNumber="33">M. nublado</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3266546FF8B2801FE9165FD4448FA9C" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8B8336CDFF8B2800FE9165FD4561FC0C" blockId="32.[264,1325,284,1870]" pageId="32" pageNumber="33">
<emphasis id="B948EADFFF8B2800FE9165FD46F7FC94" bold="true" box="[312,464,764,790]" pageId="32" pageNumber="33">Description.</emphasis>
Body elongate oval, maximum width closely before posterior margin of pronotum; elytra slightly narrowed posteriad, depressed in lateral view, not abruptly elevate behind posterior margin of pronotum. Length: 
<quantity id="4CC49B28FF8B2800FC38644D4307FCE4" box="[913,1056,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="32" pageNumber="33" unit="mm" value="2.25" valueMax="2.4" valueMin="2.1">2.1–2.4 mm</quantity>
(HT: 
<quantity id="4CC49B28FF8B2800FBC0644D43EDFCE4" box="[1129,1226,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="32" pageNumber="33" unit="mm" value="2.1">2.1 mm</quantity>
); width: 
<quantity id="4CC49B28FF8B2800FEA1647546B2FC0C" box="[264,405,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="32" pageNumber="33" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
(HT: 
<quantity id="4CC49B28FF8B2800FE726475451EFC0C" box="[475,569,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="mm" value="1.5">1.5 mm</quantity>
).
</paragraph>
<paragraph id="8B8336CDFF8B2800FE91649D467DFBAC" blockId="32.[264,1325,284,1870]" pageId="32" pageNumber="33">Coloration: Dorsal side black, with anterior margin of clypeus, lateral margins of pronotum and lateral margins of elytra paler, reddish brown. Ventral side pitchy brown, lateral portions slightly paler; mouthparts, antennae, mentum, gula and legs reddish brown.</paragraph>
<paragraph id="8B8336CDFF8B2800FE91633D4319F98C" blockId="32.[264,1325,284,1870]" pageId="32" pageNumber="33">Head: Clypeus with fine but rather dense punctation, interstices without microsculpture; anterior margin slightly convex, narrowly rimmed. Lines dividing clypeus from vertex distinct also paramesally. Vertex with coarse and rather dense setiferous punctation, interstices narrow, without microsculpture. Eyes small, separated by 10× width of one eye. Mentum slightly wider than long, anterior margin shallowly bisinuate; surface with moderately coarse and not very dense punctation, intertices with fine mesh­like microsculpture. Maxillary palpomere 2 ca. as long as palpomere 4, strongly swollen apically; palpomere 3 shorter than palpomere 2, slightly swollen apically; palpomere 4 spindle­like. Antenna with 9 antennomeres; scapus ca. as long as antennal club; antennomeres 2–6 shorter than scapus when combined; antennal club elongate oval; antennomere 7 slightly longer than antennomere 8; antennomere 9 slightly longer than antennomere 7, with acute apex and distinct strangulation subapically.</paragraph>
<paragraph id="8B8336CDFF8B2800FE91611D45B2F97C" blockId="32.[264,1325,284,1870]" pageId="32" pageNumber="33">Prothorax: Pronotum strongly arcuately narrowed anteriad, maximum width in posterior 0.25. Lateral margins slightly explanate; posterior margin slightly bisinuate, with weakly pronounced posterolateral corners. Punctation moderately coarse but very dense, interstices very narrow, without microsculpture. Prosternum with fine median carina; anteromedian excision deep and narrow; lateral parts indistinctly divided from median area, dividing line nearly straight.</paragraph>
<paragraph id="8B8336CDFF8B2801FE91600D4597FDF4" blockId="32.[264,1325,284,1870]" lastBlockId="33.[264,1324,284,1870]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">Mesothorax: Scutellar shield in form of equilateral triangle, slightly depressed in the middle, with a few fine punctures. Elytron with 9 punctural series; series 1–7 reaching elytral base, series 8–9 arising slightly subbasally. Intervals lowly costate, evenly elevate basally; intervals 2, 4 and 6 slightly more elevate than adjacent intervals posteriorly; intervals with fine pubescent punctation, interstices without microsculpture. Elytral margin with moderately large blunt denticles basally, becoming finer and more distant apicad, nearly reaching elytral apex. Epipleuron wider than pseudepipleuron basally, reaching posterior margin of metaventrite; pseudepipleuron narrowing posteriad, reaching elytral apex. Preepisternal elevation with plate 4.0× longer than wide, bearing slightly elevate narrow lateral edges and not very distinct median concave area. Grooves for reception of procoxae shalow, wide.</paragraph>
<paragraph id="8B8336CDFF8A2801FE91658544C0FCBC" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">
Metathorax: Metaventrite with elevate median pentagonal area bearing moderately strong and not very dense punctation, interstices without microsculpture; medially with quite deep elongate depression. Lateral portions with pubescent microsculpture. Postcoxal ridges bent medially, forming median longitudinal ridge reaching apical 0.15 of metaventrite length. Femoral lines absent. 
<collectingCountry id="F32B765DFF8A2801FD5264254411FCBC" box="[763,822,804,830]" name="India" pageId="33" pageNumber="34">Hind</collectingCountry>
wings present.
</paragraph>
<paragraph id="8B8336CDFF8A2801FE91644D442AFC0C" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">Legs short. Femora with very fine and sparse setiferous punctation. Tibiae longer than tarsi. Tarsi with dense pubescence ventrally.</paragraph>
<paragraph id="8B8336CDFF8A2801FE91649D444FFB84" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">Abdomen: Abdominal ventrite 1 with strong median longitudinal carina and with additional submedian longitudinal ridges. Ventrites 2 and 3 with distinct but not so strong additonal longitudinal ridges. Ventrites 4–5 smooth.</paragraph>
<paragraph id="8B8336CDFF8A2801FE91631544C7FB24" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">Male genitalia: Parameres 2.0× longer than phallobase, gradually narrowing apicad, apices slightly bent outwards, bearing two long setae. Median lobe considerably longer than parameres, rather wide subapically, narrowing apicad from apical 0.25. Apical grooved rim present in apical 0.25, apex with minute incision.</paragraph>
<paragraph id="8B8336CDFF8A2801FE9163B54448FA9C" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">
<emphasis id="B948EADFFF8A2801FE9163B54696FB4C" bold="true" box="[312,433,1204,1230]" pageId="33" pageNumber="34">Variation.</emphasis>
Because only two specimens are known to us, we cannot discuss the variability of this species. We observed a slight variability in the preepisternal elevation of mesothorax—this can be 3.0–4.0× longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FF8A2801FE91622D4510F904" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8B8336CDFF8A2801FE91622D4510F904" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">
<emphasis id="B948EADFFF8A2801FE91622D46E4FAC4" bold="true" box="[312,451,1324,1350]" pageId="33" pageNumber="34">Discussion.</emphasis>
Distinguishing of this species from 
<taxonomicName id="4C3C4D4EFF8A2801FCD9622D44C8FAC7" box="[880,1007,1324,1349]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFF8A2801FCD9622D44C8FAC7" box="[880,1007,1324,1349]" italics="true" pageId="33" pageNumber="34">Motonerus</emphasis>
</taxonomicName>
sp. B (
<collectingCountry id="F32B765DFF8A2801FBEC622D43B9FAC4" box="[1093,1182,1324,1350]" name="Bolivia" pageId="33" pageNumber="34">Bolivia</collectingCountry>
) is possible only on the basis of a slight difference in the shape of the preepisternal elevation (3.0–4.0× longer than wide in 
<taxonomicName id="4C3C4D4EFF8A2801FDAB627D45F3FA17" box="[514,724,1404,1429]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B948EADFFF8A2801FDAB627D45F3FA17" box="[514,724,1404,1429]" italics="true" pageId="33" pageNumber="34">M. problematicus</emphasis>
</taxonomicName>
and 5.5× longer than wide in 
<taxonomicName id="4C3C4D4EFF8A2801FBE6627D43E9FA17" box="[1103,1230,1404,1429]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFF8A2801FBE6627D43E9FA17" box="[1103,1230,1404,1429]" italics="true" pageId="33" pageNumber="34">Motonerus</emphasis>
</taxonomicName>
sp. B). Because of the low number of examined specimens of both species, we cannot exclude that this difference refers the intraspecific variability of both species. In this case, 
<taxonomicName id="4C3C4D4EFF8A2801FAA262CD4689F98F" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B948EADFFF8A2801FAA262CD4689F98F" italics="true" pageId="33" pageNumber="34">M. problematicus</emphasis>
</taxonomicName>
would be externally indistinguishable from 
<taxonomicName id="4C3C4D4EFF8A2801FC6162F54360F98F" box="[968,1095,1524,1549]" class="Insecta" family="Hydrophilidae" genus="Motonerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B948EADFFF8A2801FC6162F54360F98F" box="[968,1095,1524,1549]" italics="true" pageId="33" pageNumber="34">Motonerus</emphasis>
</taxonomicName>
sp. B for the time being. However, because of the large distance of localities where both species were found, we decided to treat them as two different species until the male of the putative separate Bolivian species is found.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FF8A2801FE91619543DBF8CC" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8B8336CDFF8A2801FE9161954687F97C" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">
<emphasis id="B948EADFFF8A2801FE91619546E3F92C" bold="true" box="[312,452,1684,1710]" pageId="33" pageNumber="34">Etymology.</emphasis>
<taxonomicName id="4C3C4D4EFF8A2801FE6461954598F92C" authority="(Lat.)" baseAuthorityName="Lat." box="[461,703,1684,1710]" class="Insecta" family="Hydrophilidae" genus="Problematicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="37" phylum="Arthropoda" rank="genus">Problematicus (Lat.)</taxonomicName>
= problematic, referring the fact that the solution of the status of this species remained as the last and most difficult problem during preparation of this paper.
</paragraph>
<paragraph id="8B8336CDFF8A2801FE91600D43DBF8CC" blockId="33.[264,1324,284,1870]" pageId="33" pageNumber="34">
<emphasis id="B948EADFFF8A2801FE91600D46B8F8A4" bold="true" box="[312,415,1804,1830]" pageId="33" pageNumber="34">Habitat.</emphasis>
One specimen was collected at light, the collecting circumstances of the second are not known to us. The elevation 
<quantity id="4CC49B28FF8A2801FD5660354470F8CC" box="[767,855,1844,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="34" unit="m" value="1150.0">1150 m</quantity>
a.s.l. is mentioned for the 
<typeStatus id="5487886FFF8A2801FB26603543DFF8CC" box="[1167,1272,1844,1870]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3266546FF892802FE91661A436DFEDC" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8B8336CDFF892802FE91661A436DFEDC" blockId="34.[264,1323,283,350]" pageId="34" pageNumber="35">
<emphasis id="B948EADFFF892802FE91661A46FFFEB7" bold="true" box="[312,472,283,309]" pageId="34" pageNumber="35">Distribution.</emphasis>
The species is known from two ca. 
<quantity id="4CC49B28FF892802FCD2661D44F3FEB4" box="[891,980,284,310]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="35" unit="km" value="130.0">130 km</quantity>
distant cloud forest localities in 
<collectingCountry id="F32B765DFF892802FE8F66454681FEDC" box="[294,422,324,350]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
(Cordillera de Talamanca and Cordillera de Guanacaste).
</paragraph>
</subSubSection>
</treatment>
</document>