treatments-xml/data/03/95/87/039587CB35556933FF25F9BBFE69F826.xml

<document id="261FEC3BE0409E0FB39287AE892E9C8B" ID-CLB-Dataset="6426" ID-DOI="10.11646/zootaxa.4970.1.4" ID-GBIF-Dataset="325d1b05-f970-44b3-b10f-b2db9c5e9da7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4756031" ID-ZooBank="3F7A60FB-7206-441E-94C6-AF718B9A5E56" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620889976977" checkinUser="plazi" docAuthor="Yoshino, Hiroki, Kambara, Yuriko, Kajiwara, Shota &amp; Ohsawa, Takeshi A." docDate="2021" docId="039587CB35556933FF25F9BBFE69F826" docLanguage="en" docName="zootaxa.4970.1.4.pdf" docOrigin="Zootaxa 4970 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnoria aspera Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="124" masterDocId="FFACFFB335546936FFB2FFA8FFA5FFB9" masterDocTitle="New species of Sargassum-boring Limnoria Leach, 1814 (Crustacea, Isopoda Limnoriidae) from Japan" masterLastPageNumber="130" masterPageNumber="119" pageNumber="120" updateTime="1698939191155" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4176CBF12C32BBE1D63E07E9C5C5B1D7">New species of Sargassum-boring Limnoria Leach, 1814 (Crustacea, Isopoda Limnoriidae) from Japan</mods:title>
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<mods:namePart id="8E1F819535DB8433FE56EC52FA41A0DE">Yoshino, Hiroki</mods:namePart>
<mods:affiliation id="E7781DBCB90E7F5FF325BDAE8A0CAB80">Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan.</mods:affiliation>
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<mods:namePart id="CDA7AA1BCFF12C73452921658FAAD3C0">Kambara, Yuriko</mods:namePart>
<mods:affiliation id="A40A63BEC32504328250B3AB71DA0892">Department of Biology, Graduate School of Science, Chiba University, Chiba, Japan</mods:affiliation>
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<mods:namePart id="7087904435895ABD99E7E8FE2F98D867">Kajiwara, Shota</mods:namePart>
<mods:affiliation id="FE3721EF38080A78A104054C713E1434">Department of Biology, Graduate School of Science, Chiba University, Chiba, Japan</mods:affiliation>
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<mods:namePart id="805DE926ECA2CA713D53F692E3143015">Ohsawa, Takeshi A.</mods:namePart>
<mods:affiliation id="75E84B2330B3D0F55340AB545235E8D6">Department of Biology, Graduate School of Science, Chiba University, Chiba, Japan</mods:affiliation>
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<subSubSection id="C326655635556937FF25F9BBFD66F994" box="[151,707,1554,1581]" pageId="1" pageNumber="120" type="nomenclature">
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<heading id="D0CB81B135556937FF25F9BBFD66F994" bold="true" box="[151,707,1554,1581]" fontSize="11" level="1" pageId="1" pageNumber="120" reason="1">
<emphasis id="B948EACF35556937FF25F9BBFD66F994" bold="true" box="[151,707,1554,1581]" pageId="1" pageNumber="120">
<taxonomicName id="4C3C4D5E35556937FF25F9BBFDFDF995" authority="Yoshino &amp; Ohsawa" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[151,600,1554,1581]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35556937FF25F9BBFEC4F994" bold="true" box="[151,353,1555,1581]" italics="true" pageId="1" pageNumber="120">Limnoria aspera</emphasis>
Yoshino &amp; Ohsawa
</taxonomicName>
, 
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<paragraph id="8B8336DD35556937FF25F99FFF5DF9E8" blockId="1.[151,707,1554,1617]" box="[151,248,1591,1617]" pageId="1" pageNumber="120">
<figureCitation id="13072A5835556937FF25F99FFF5DF9E8" box="[151,248,1591,1617]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="2.[151,250,1079,1103]" captionStartId-1="2.[151,250,1780,1804]" captionStartId-2="3.[151,250,1942,1966]" captionStartId-3="4.[151,250,1049,1073]" captionStartId-4="5.[151,250,1385,1409]" captionStartId-5="6.[151,250,1026,1050]" captionStartId-6="6.[151,250,1861,1885]" captionStartId-7="9.[151,250,977,1001]" captionTargetBox-0="[296,1368,450,1053]" captionTargetBox-1="[151,1436,1343,1756]" captionTargetBox-2="[163,1422,193,1909]" captionTargetBox-3="[155,1430,386,1022]" captionTargetBox-4="[157,1426,387,1352]" captionTargetBox-5="[169,1430,186,1002]" captionTargetBox-6="[155,1431,1116,1830]" captionTargetBox-7="[151,1436,181,954]" captionTargetId-0="figure-150@2.[151,1436,450,1053]" captionTargetId-1="figure-246@2.[151,1436,1343,1756]" captionTargetId-2="figure-17@3.[151,1436,181,1918]" captionTargetId-3="figure-108@4.[151,1436,381,1026]" captionTargetId-4="figure-109@5.[151,1436,381,1362]" captionTargetId-5="figure-28@6.[151,1436,181,1002]" captionTargetId-6="figure-53@6.[151,1436,1111,1837]" captionTargetId-7="figure-17@9.[151,1436,181,954]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="6" captionTargetPageId-7="9" captionText-0="FIGURE 1. Limnoria aspera sp. nov., paratype (male, 3.2 mm, CBM-ZC 16567). Body; dorsal view. Stereo microscope image (SZX10; Olympus). Scale bar: 1 mm." captionText-1="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." captionText-2="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." captionText-3="FIGURE 4. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm." captionText-4="FIGURE 5. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm." captionText-5="FIGURE 6. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A–C, pereopods 5–7. Scale bars: A–C = 0.1 mm." captionText-6="FIGURE 7. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleopod 1; B, pleopod 2; C, pleopod 5. Scale bars: A–C = 0.1 mm." captionText-7="FIGURE 8. Limnoria aspera sp. nov., paratype (male 2.8 mm, CBM-ZC 16565). A, left mandible, apical view, SEM image (JSM-6010LA; JEOL); B, pereopod 2, lateral view, SEM image (JSM-6010LA; JEOL); uropod, ventral view, SEM image (JSM-6010LA; JEOL). Scale bar: A = 0.02 mm, B, C = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4756033" figureDoi-1="http://doi.org/10.5281/zenodo.4756035" figureDoi-2="http://doi.org/10.5281/zenodo.4756037" figureDoi-3="http://doi.org/10.5281/zenodo.4756039" figureDoi-4="http://doi.org/10.5281/zenodo.4756043" figureDoi-5="http://doi.org/10.5281/zenodo.4756045" figureDoi-6="http://doi.org/10.5281/zenodo.4756053" figureDoi-7="http://doi.org/10.5281/zenodo.4756055" httpUri-0="https://zenodo.org/record/4756033/files/figure.png" httpUri-1="https://zenodo.org/record/4756035/files/figure.png" httpUri-2="https://zenodo.org/record/4756037/files/figure.png" httpUri-3="https://zenodo.org/record/4756039/files/figure.png" httpUri-4="https://zenodo.org/record/4756043/files/figure.png" httpUri-5="https://zenodo.org/record/4756045/files/figure.png" httpUri-6="https://zenodo.org/record/4756053/files/figure.png" httpUri-7="https://zenodo.org/record/4756055/files/figure.png" pageId="1" pageNumber="120">Figs 1–8</figureCitation>
</paragraph>
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<emphasis id="B948EACF35556937FF25F9D0FEDAF92B" bold="true" box="[151,383,1656,1682]" pageId="1" pageNumber="120">Material examined.</emphasis>
<materialsCitation id="3B543C8035556937FE36F9DFFE0CF962" ID-GBIF-Occurrence="3118985301" collectingDate="2020-09-09" collectorName="Takeshi A. Ohsawa &amp; Hiroki Yoshino" country="Japan" elevation="5" latitude="35.116665" location="Kamogawa-shi" longLatPrecision="1195" longitude="140.16667" pageId="1" pageNumber="120" specimenCount="1" stateProvince="Chiba" typeStatus="holotype">
<emphasis id="B948EACF35556937FE36F9DFFE50F92B" bold="true" box="[388,501,1655,1682]" pageId="1" pageNumber="120">
<emphasis id="B948EACF35556937FE36F9DFFE49F92B" bold="true" box="[388,492,1655,1682]" italics="true" pageId="1" pageNumber="120">
<typeStatus id="5487887F35556937FE36F9DFFE49F92B" box="[388,492,1655,1682]" pageId="1" pageNumber="120" type="holotype">Holotype</typeStatus>
</emphasis>
:
</emphasis>
male 
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, 
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, 
<collectingRegion id="49F8F83F35556937FCFFF9D0FBAFF92B" box="[845,1034,1656,1682]" country="Japan" name="Chiba" pageId="1" pageNumber="120">Chiba prefecture</collectingRegion>
, 
<collectingCountry id="F32B764D35556937FBA1F9D1FBF3F92A" box="[1043,1110,1657,1683]" name="Japan" pageId="1" pageNumber="120">Japan</collectingCountry>
, 
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, 
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, subtidal zone (
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in depth), 
<taxonomicName id="4C3C4D5E35556937FE12F934FD62F90F" authorityName="Harvey" authorityYear="1860" box="[416,711,1692,1718]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="120" phylum="Phaeophyta" rank="species" species="ringgoldianum">
<emphasis id="B948EACF35556937FE12F934FD62F90F" box="[416,711,1692,1718]" italics="true" pageId="1" pageNumber="120">Sargassum ringgoldianum</emphasis>
</taxonomicName>
holdfasts, 
<collectorName id="26C9530B35556937FCF1F934FBB9F90F" box="[835,1052,1692,1718]" pageId="1" pageNumber="120">Takeshi A. Ohsawa</collectorName>
and 
<collectorName id="26C9530B35556937FBE6F934FAA6F90F" box="[1108,1283,1692,1718]" pageId="1" pageNumber="120">Hiroki Yoshino</collectorName>
, 
<date id="FF82101D35556937FABDF934FF6AF963" pageId="1" pageNumber="120" value="2020-09-09">
<collectingDate id="EFC6E9F535556937FABDF934FF6AF963" pageId="1" pageNumber="120" value="2020-09-09">9 September 2020</collectingDate>
</date>
(CBM-ZC 16564).
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</paragraph>
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<emphasis id="B948EACF35556937FF75F94CFEE7F947" bold="true" box="[199,322,1764,1790]" pageId="1" pageNumber="120">
<emphasis id="B948EACF35556937FF75F94CFE9CF947" bold="true" box="[199,313,1764,1790]" italics="true" pageId="1" pageNumber="120">
<typeStatus id="5487887F35556937FF75F94CFE9CF947" box="[199,313,1764,1790]" pageId="1" pageNumber="120" type="paratype">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
male 
<quantity id="4CC49B3835556937FE3CF94CFE4FF946" box="[398,490,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="120" unit="mm" value="2.8">2.8 mm</quantity>
, Kamogawa-shi, 
<collectingRegion id="49F8F83F35556937FD06F94CFCD2F947" box="[692,887,1764,1790]" country="Japan" name="Chiba" pageId="1" pageNumber="120">Chiba prefecture</collectingRegion>
, 
<collectingCountry id="F32B764D35556937FC37F94DFC6DF946" box="[901,968,1765,1791]" name="Japan" pageId="1" pageNumber="120">Japan</collectingCountry>
, 
<geoCoordinate id="EE08501A35556937FC64F94CFB9CF946" box="[982,1081,1764,1791]" degrees="35" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="120" precision="925" value="35.116665">35°07’N</geoCoordinate>
, 
<geoCoordinate id="EE08501A35556937FBFBF94CFB12F947" box="[1097,1207,1764,1790]" degrees="140" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="120" precision="925" value="140.16667">140°10’E</geoCoordinate>
, subtidal zone (
<quantity id="4CC49B3835556937FAC0F94CFF08F89A" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="1" pageNumber="120" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
in depth), 
<taxonomicName id="4C3C4D5E35556937FE81F8A0FDFBF89B" authorityName="Harvey" authorityYear="1860" box="[307,606,1800,1826]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="120" phylum="Phaeophyta" rank="species" species="ringgoldianum">
<emphasis id="B948EACF35556937FE81F8A0FDFBF89B" box="[307,606,1800,1826]" italics="true" pageId="1" pageNumber="120">Sargassum ringgoldianum</emphasis>
</taxonomicName>
holdfasts, Yuriko Kambara, 
<date id="FF82101D35556937FC03F8A0FBEFF89B" box="[945,1098,1800,1826]" pageId="1" pageNumber="120" value="2014-07-14">14 July 2014</date>
(CBM-ZC 16565). male 
<quantity id="4CC49B3835556937FACBF8A0FF60F8FE" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="120" unit="mm" value="2.7">2.7 mm</quantity>
, Kamogawa-shi, 
<collectingRegion id="49F8F83F35556937FE22F884FDF1F8FF" box="[400,596,1836,1862]" country="Japan" name="Chiba" pageId="1" pageNumber="120">Chiba prefecture</collectingRegion>
, 
<collectingCountry id="F32B764D35556937FDD0F885FD00F8FE" box="[610,677,1837,1863]" name="Japan" pageId="1" pageNumber="120">Japan</collectingCountry>
, 
<geoCoordinate id="EE08501A35556937FD06F884FCB2F8FE" box="[692,791,1836,1863]" degrees="35" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="120" precision="925" value="35.116665">35°07’N</geoCoordinate>
, 
<geoCoordinate id="EE08501A35556937FC95F884FC30F8FF" box="[807,917,1836,1862]" degrees="140" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="120" precision="925" value="140.16667">140°10’E</geoCoordinate>
, subtidal zone (
<quantity id="4CC49B3835556937FBE3F884FB38F8FE" box="[1105,1181,1836,1863]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="1" pageNumber="120" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
in depth), 
<taxonomicName id="4C3C4D5E35556937FA90F884FE86F8D3" authorityName="C.Agardh" authorityYear="1820" baseAuthorityName="Mert. ex Turner" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="120" phylum="Phaeophyta" rank="species" species="siliquastrum">
<emphasis id="B948EACF35556937FA90F884FE86F8D3" italics="true" pageId="1" pageNumber="120">Sargassum siliquastrum</emphasis>
</taxonomicName>
holdfasts, Takeshi A. Ohsawa and Hiroki Yoshino, 
<date id="FF82101D35556937FCCCF8F8FBF5F8D3" box="[894,1104,1872,1898]" pageId="1" pageNumber="120" value="2020-09-09">9 September 2020</date>
(CBM-ZC 16566). male 
<quantity id="4CC49B3835556937FACBF8F8FF60F836" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="120" unit="mm" value="3.2">3.2 mm</quantity>
, Kamogawa-shi, 
<collectingRegion id="49F8F83F35556937FE22F8DCFDF1F837" box="[400,596,1908,1934]" country="Japan" name="Chiba" pageId="1" pageNumber="120">Chiba prefecture</collectingRegion>
, 
<collectingCountry id="F32B764D35556937FDD0F8DDFD00F836" box="[610,677,1909,1935]" name="Japan" pageId="1" pageNumber="120">Japan</collectingCountry>
, 
<geoCoordinate id="EE08501A35556937FD06F8DCFCB2F836" box="[692,791,1908,1935]" degrees="35" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="120" precision="925" value="35.116665">35°07’N</geoCoordinate>
, 
<geoCoordinate id="EE08501A35556937FC95F8DCFC30F837" box="[807,917,1908,1934]" degrees="140" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="120" precision="925" value="140.16667">140°10’E</geoCoordinate>
, subtidal zone (
<quantity id="4CC49B3835556937FBE3F8DCFB38F836" box="[1105,1181,1908,1935]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="1" pageNumber="120" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
in depth), 
<taxonomicName id="4C3C4D5E35556937FA90F8DCFE86F80B" authorityName="C.Agardh" authorityYear="1820" baseAuthorityName="Mert. ex Turner" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="1" pageNumber="120" phylum="Phaeophyta" rank="species" species="siliquastrum">
<emphasis id="B948EACF35556937FA90F8DCFE86F80B" italics="true" pageId="1" pageNumber="120">Sargassum siliquastrum</emphasis>
</taxonomicName>
holdfasts, Takeshi A. Ohsawa and Hiroki Yoshino, 
<date id="FF82101D35556937FCD7F830FB94F80B" box="[869,1073,1944,1970]" pageId="1" pageNumber="120" value="2020-09-09">9 September 2020</date>
(CBM-ZC 16567).
</paragraph>
<paragraph id="8B8336DD35556937FF75F814FC93F86E" blockId="1.[151,1437,1655,2043]" box="[199,822,1980,2007]" pageId="1" pageNumber="120">
<emphasis id="B948EACF35556937FF75F814FEC3F86F" bold="true" box="[199,358,1980,2006]" pageId="1" pageNumber="120">Type locality.</emphasis>
Kamogawa-shi, 
<collectingRegion id="49F8F83F35556937FD96F814FD41F86F" box="[548,740,1980,2006]" country="Japan" name="Chiba" pageId="1" pageNumber="120">Chiba Prefecture</collectingRegion>
, 
<collectingCountry id="F32B764D35556937FD5DF815FC97F86E" box="[751,818,1981,2007]" name="Japan" pageId="1" pageNumber="120">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C326655635556932FF75F848FC3CF92C" lastPageId="4" lastPageNumber="123" pageId="1" pageNumber="120" type="description">
<paragraph id="8B8336DD35556934FF75F848FD39FEF8" blockId="1.[151,1437,1655,2043]" lastBlockId="2.[151,1437,150,429]" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120">
<emphasis id="B948EACF35556937FF75F848FEFCF843" bold="true" box="[199,345,2016,2042]" pageId="1" pageNumber="120">Description.</emphasis>
<emphasis id="B948EACF35556937FED0F848FE39F843" box="[354,412,2016,2042]" italics="true" pageId="1" pageNumber="120">Body</emphasis>
oblong, dark yellow to light brown in 70% ethanol (
<figureCitation id="13072A5835556937FBB0F848FBF7F843" box="[1026,1106,2016,2042]" captionStart="FIGURE 1" captionStartId="2.[151,250,1079,1103]" captionTargetBox="[296,1368,450,1053]" captionTargetId="figure-150@2.[151,1436,450,1053]" captionTargetPageId="2" captionText="FIGURE 1. Limnoria aspera sp. nov., paratype (male, 3.2 mm, CBM-ZC 16567). Body; dorsal view. Stereo microscope image (SZX10; Olympus). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756033" httpUri="https://zenodo.org/record/4756033/files/figure.png" pageId="1" pageNumber="120">Figs. 1</figureCitation>
, 
<figureCitation id="13072A5835556937FBD1F848FB2CF843" box="[1123,1161,2016,2042]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="1" pageNumber="120">2A</figureCitation>
). 
<emphasis id="B948EACF35556937FB2DF848FB79F843" box="[1183,1244,2016,2042]" italics="true" pageId="1" pageNumber="120">Head</emphasis>
almost globular. 
<emphasis id="B948EACF35566934FF25FF3FFF69FF09" box="[151,204,151,176]" italics="true" pageId="2" pageNumber="121">Eyes</emphasis>
black in color, each with 8 ommatidia. 
<emphasis id="B948EACF35566934FD37FF3FFD47FF08" box="[645,738,151,177]" italics="true" pageId="2" pageNumber="121">Pleonite</emphasis>
composed of 5 distinct segments. Pereonal segment 1 longest, approximately 1.5–1.7 times longer than segment 2 (
<figureCitation id="13072A5835566934FD52FF13FC84FF6C" box="[736,801,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1079,1103]" captionTargetBox="[296,1368,450,1053]" captionTargetId="figure-150@2.[151,1436,450,1053]" captionTargetPageId="2" captionText="FIGURE 1. Limnoria aspera sp. nov., paratype (male, 3.2 mm, CBM-ZC 16567). Body; dorsal view. Stereo microscope image (SZX10; Olympus). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756033" httpUri="https://zenodo.org/record/4756033/files/figure.png" pageId="2" pageNumber="121">Fig. 1</figureCitation>
), with 1 thinly V-shaped groove. Segments 2–4 subequal length. Posterior pereonal segments 4–7 progressively shorten. Coxal plates of pereonal segments 2–4 rectangular in shape and those of posterior segments prolonged acutely at posterior angle. 
<emphasis id="B948EACF35566934FBBFFEABFB13FEA4" box="[1037,1206,258,285]" italics="true" pageId="2" pageNumber="121">Pereonites 6, 7</emphasis>
and pleonites 1 to 4 each with transverse row of many small setae.
</paragraph>
<paragraph id="8B8336DD35566934FF75FEE3FD10FE14" blockId="2.[151,1437,150,429]" pageId="2" pageNumber="121">
<emphasis id="B948EACF35566934FF75FEE3FE81FEDC" box="[199,292,331,357]" italics="true" pageId="2" pageNumber="121">Pleonite</emphasis>
5 approximately 0.6 times as long as pleotelson (
<figureCitation id="13072A5835566934FCE8FEE3FC01FEDC" box="[858,932,331,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121">Figs 2</figureCitation>
, 
<figureCitation id="13072A5835566934FC03FEE3FC73FEDC" box="[945,982,331,357]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">3A</figureCitation>
). Pleonite 5 dorsomedially with Y- or Ishaped indistinct longitudinal carina on which few scale spikes form line (
<figureCitation id="13072A5835566934FC5AFEC7FBE3FE30" box="[1000,1094,367,393]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121">Figs 2B</figureCitation>
, 
<figureCitation id="13072A5835566934FBE1FEC7FBDCFE30" box="[1107,1145,367,393]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">3A</figureCitation>
). Pleonite 5 covered with fused scales, which make tile mosaics (
<figureCitation id="13072A5835566934FDFDFE3BFD0DFE14" box="[591,680,403,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121">Fig. 2C</figureCitation>
).
</paragraph>
<caption id="DF43665535566934FF25FB9FFDB0FBCA" ID-DOI="http://doi.org/10.5281/zenodo.4756033" ID-Zenodo-Dep="4756033" httpUri="https://zenodo.org/record/4756033/files/figure.png" pageId="2" pageNumber="121" startId="2.[151,250,1079,1103]" targetBox="[296,1368,450,1053]" targetPageId="2">
<paragraph id="8B8336DD35566934FF25FB9FFDB0FBCA" blockId="2.[151,1436,1079,1139]" pageId="2" pageNumber="121">
<emphasis id="B948EACF35566934FF25FB9FFEB6FBF6" bold="true" box="[151,275,1079,1103]" pageId="2" pageNumber="121">FIGURE 1.</emphasis>
<taxonomicName id="4C3C4D5E35566934FEAAFB90FE64FBF6" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[280,449,1080,1103]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35566934FEAAFB90FE64FBF6" box="[280,449,1080,1103]" italics="true" pageId="2" pageNumber="121">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35566934FE75FB9FFDB2FBF6" bold="true" box="[455,535,1079,1103]" pageId="2" pageNumber="121">
<taxonomicNameLabel id="A27B57B435566934FE75FB9FFDB2FBF6" box="[455,535,1079,1103]" pageId="2" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype (male, 3.2 mm, CBM-ZC 16567). Body; dorsal view. Stereo microscope image (SZX10; Olympus). Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8B8336DD35566934FF75FB09FC56FA91" blockId="2.[151,1436,1185,1320]" pageId="2" pageNumber="121">
<emphasis id="B948EACF35566934FF75FB09FE98FB02" box="[199,317,1185,1211]" italics="true" pageId="2" pageNumber="121">Pleotelson</emphasis>
0.6–0.8 times as long as wide, medially with 1 tubercle on which few scale spikes follow its line (
<figureCitation id="13072A5835566934FF2DFB6DFF4CFB59" box="[159,233,1221,1248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121">Figs 2</figureCitation>
, 
<figureCitation id="13072A5835566934FF44FB6EFEBEFB59" box="[246,283,1222,1248]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">3A</figureCitation>
). Pleotelson covered with fused scales, which present tiled pattern on surface (
<figureCitation id="13072A5835566934FB25FB6DFB54FB59" box="[1175,1265,1221,1248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1780,1804]" captionTargetBox="[151,1436,1343,1756]" captionTargetId="figure-246@2.[151,1436,1343,1756]" captionTargetPageId="2" captionText="FIGURE 2. Limnoria aspera sp. nov., A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121">Fig. 2C</figureCitation>
). Lateral crests and posterior margin of pleotelson margined with sets of about 2–5 directed upward scale spikes. Posterior edge of pleotelson with fringe of long sheathed setae and many short setae (
<figureCitation id="13072A5835566934FC3CFAA5FC43FA91" box="[910,998,1293,1320]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">Fig. 3B</figureCitation>
).
</paragraph>
<caption id="DF43665535566934FF25F95CFF49F8C1" ID-DOI="http://doi.org/10.5281/zenodo.4756035" ID-Zenodo-Dep="4756035" httpUri="https://zenodo.org/record/4756035/files/figure.png" pageId="2" pageNumber="121" startId="2.[151,250,1780,1804]" targetBox="[151,1436,1343,1756]" targetPageId="2">
<paragraph id="8B8336DD35566934FF25F95CFF49F8C1" blockId="2.[151,1436,1780,1912]" pageId="2" pageNumber="121">
<emphasis id="B948EACF35566934FF25F95CFEB6F8B5" bold="true" box="[151,275,1780,1804]" pageId="2" pageNumber="121">FIGURE 2.</emphasis>
<taxonomicName id="4C3C4D5E35566934FEA8F95DFE61F8B5" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[282,452,1781,1804]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35566934FEA8F95DFE61F8B5" box="[282,452,1781,1804]" italics="true" pageId="2" pageNumber="121">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35566934FE79F95CFDB9F8B5" bold="true" box="[459,540,1780,1804]" pageId="2" pageNumber="121">
<taxonomicNameLabel id="A27B57B435566934FE79F95CFDB9F8B5" box="[459,540,1780,1804]" pageId="2" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A–B, holotype (male, 2.6 mm, CBM-ZC 16564); C, paratype (male 2.8 mm, CBM-ZC 16565). A, pleotelson, dorsal view, Optical microscope image (VHX-7000; Keyence); B, pleotelson, dorsal view, 3D digital microscope image (VHX-7000; Keyence); C, pleotelson, dorsal view, SEM image (JSM-6010LA; JEOL). Scale bars: A–C = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8336DD35566934FF75F80FFEFEF85D" blockId="2.[151,1436,1958,2020]" pageId="2" pageNumber="121">
<emphasis id="B948EACF35566934FF75F80FFE81F878" box="[199,292,1959,1985]" italics="true" pageId="2" pageNumber="121">Antenna</emphasis>
1 with 4 flagellar articles; second article with 7 aesthetascs (
<figureCitation id="13072A5835566934FC65F80EFB97F879" box="[983,1074,1958,1984]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">Fig. 3D</figureCitation>
). Flagellum of antenna 2 with 4 articles (
<figureCitation id="13072A5835566934FF45F862FEEAF85D" box="[247,335,1994,2020]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="2" pageNumber="121">Fig. 3E</figureCitation>
).
</paragraph>
<caption id="DF43665535576935FF25F83EFB66F86B" ID-DOI="http://doi.org/10.5281/zenodo.4756037" ID-Zenodo-Dep="4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="3" pageNumber="122" startId="3.[151,250,1942,1966]" targetBox="[163,1422,193,1909]" targetPageId="3">
<paragraph id="8B8336DD35576935FF25F83EFB66F86B" blockId="3.[151,1436,1942,2002]" pageId="3" pageNumber="122">
<emphasis id="B948EACF35576935FF25F83EFEB1F817" bold="true" box="[151,276,1942,1966]" pageId="3" pageNumber="122">FIGURE 3.</emphasis>
<taxonomicName id="4C3C4D5E35576935FEAEF83FFE62F817" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[284,455,1943,1966]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35576935FEAEF83FFE62F817" box="[284,455,1943,1966]" italics="true" pageId="3" pageNumber="122">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35576935FE7DF83EFD84F817" bold="true" box="[463,545,1942,1966]" pageId="3" pageNumber="122">
<taxonomicNameLabel id="A27B57B435576935FE7DF83EFD84F817" box="[463,545,1942,1966]" pageId="3" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm.
</paragraph>
</caption>
<paragraph id="8B8336DD35506932FF75FF3FFD28FEA4" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75FF3FFEE8FF08" box="[199,333,151,177]" italics="true" pageId="4" pageNumber="123">Mandibular</emphasis>
palp lacking, replaced by single long stout seta (
<figureCitation id="13072A5835506932FC32FF3FFBA7FF08" box="[896,1026,151,177]" captionStart="FIGURE 4" captionStartId="4.[151,250,1049,1073]" captionTargetBox="[155,1430,386,1022]" captionTargetId="figure-108@4.[151,1436,381,1026]" captionTargetPageId="4" captionText="FIGURE 4. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756039" httpUri="https://zenodo.org/record/4756039/files/figure.png" pageId="4" pageNumber="123">Figs 4A, B</figureCitation>
). Mandibular incisors lack rasp and file (
<figureCitation id="13072A5835506932FF79FF13FEECFF6C" box="[203,329,187,213]" captionStart="FIGURE 4" captionStartId="4.[151,250,1049,1073]" captionTargetBox="[155,1430,386,1022]" captionTargetId="figure-108@4.[151,1436,381,1026]" captionTargetPageId="4" captionText="FIGURE 4. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756039" httpUri="https://zenodo.org/record/4756039/files/figure.png" pageId="4" pageNumber="123">Figs 4A–B</figureCitation>
, 
<figureCitation id="13072A5835506932FEE6FF13FEDCFF6C" box="[340,377,187,213]" captionStart="FIGURE 8" captionStartId="9.[151,250,977,1001]" captionTargetBox="[151,1436,181,954]" captionTargetId="figure-17@9.[151,1436,181,954]" captionTargetPageId="9" captionText="FIGURE 8. Limnoria aspera sp. nov., paratype (male 2.8 mm, CBM-ZC 16565). A, left mandible, apical view, SEM image (JSM-6010LA; JEOL); B, pereopod 2, lateral view, SEM image (JSM-6010LA; JEOL); uropod, ventral view, SEM image (JSM-6010LA; JEOL). Scale bar: A = 0.02 mm, B, C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756055" httpUri="https://zenodo.org/record/4756055/files/figure.png" pageId="4" pageNumber="123">8A</figureCitation>
). Lacinia mobilis of right mandible branched at intermediate point, branches gradually curving 90-degrees and serrated on anterior side (
<figureCitation id="13072A5835506932FDD4FF77FD1AFF40" box="[614,703,223,249]" captionStart="FIGURE 4" captionStartId="4.[151,250,1049,1073]" captionTargetBox="[155,1430,386,1022]" captionTargetId="figure-108@4.[151,1436,381,1026]" captionTargetPageId="4" captionText="FIGURE 4. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756039" httpUri="https://zenodo.org/record/4756039/files/figure.png" pageId="4" pageNumber="123">Fig. 4B</figureCitation>
). Posterior branch almost same length as anterior branch. Lacinia mobilis of left mandible with 1 serrated seta.
</paragraph>
<paragraph id="8B8336DD35506932FF75FE8FFED0FEDC" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="123">
Epipod of 
<emphasis id="B948EACF35506932FE8EFE8FFE17FEF8" box="[316,434,295,321]" italics="true" pageId="4" pageNumber="123">maxilliped</emphasis>
, clavate, approximately 3 times as long as wide, reaching articulation of palp, with simple true setae (
<figureCitation id="13072A5835506932FEA3FEE3FECCFEDC" box="[273,361,331,357]" captionStart="FIGURE 4" captionStartId="4.[151,250,1049,1073]" captionTargetBox="[155,1430,386,1022]" captionTargetId="figure-108@4.[151,1436,381,1026]" captionTargetPageId="4" captionText="FIGURE 4. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756039" httpUri="https://zenodo.org/record/4756039/files/figure.png" pageId="4" pageNumber="123">Fig. 4E</figureCitation>
).
</paragraph>
<caption id="DF43665535506932FF25FBB1FC90FBEC" ID-DOI="http://doi.org/10.5281/zenodo.4756039" ID-Zenodo-Dep="4756039" httpUri="https://zenodo.org/record/4756039/files/figure.png" pageId="4" pageNumber="123" startId="4.[151,250,1049,1073]" targetBox="[155,1430,386,1022]" targetPageId="4">
<paragraph id="8B8336DD35506932FF25FBB1FC90FBEC" blockId="4.[151,1436,1049,1109]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF25FBB1FEB3FB88" bold="true" box="[151,278,1049,1073]" pageId="4" pageNumber="123">FIGURE 4.</emphasis>
<taxonomicName id="4C3C4D5E35506932FEADFBB2FE6EFB88" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[287,459,1050,1073]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35506932FEADFBB2FE6EFB88" box="[287,459,1050,1073]" italics="true" pageId="4" pageNumber="123">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35506932FE66FBB1FD8DFB88" bold="true" box="[468,552,1049,1073]" pageId="4" pageNumber="123">
<taxonomicNameLabel id="A27B57B435506932FE66FBB1FD8DFB88" box="[468,552,1049,1073]" pageId="4" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (male, 2.6 mm, CBM-ZC 16564). A, left mandible; B, right mandible; C, maxilla 1; D, maxilla 2; E, maxilliped. Scale bars: A–E = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8336DD35506932FF75FB2BFE60FA97" blockId="4.[151,1437,1155,2010]" pageId="4" pageNumber="123">
Secondary unguis on 
<emphasis id="B948EACF35506932FE08FB2BFD86FB24" box="[442,547,1155,1181]" italics="true" pageId="4" pageNumber="123">pereopod</emphasis>
1 undivided, without spinule or with 1–7 spinules (
<figureCitation id="13072A5835506932FBD4FB2BFB65FB27" box="[1126,1216,1155,1182]" captionStart="FIGURE 5" captionStartId="5.[151,250,1385,1409]" captionTargetBox="[157,1426,387,1352]" captionTargetId="figure-109@5.[151,1436,381,1362]" captionTargetPageId="5" captionText="FIGURE 5. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756043" httpUri="https://zenodo.org/record/4756043/files/figure.png" pageId="4" pageNumber="123">Fig. 5A</figureCitation>
). Secondary unguis on pereopods 2–6 undivided (
<figureCitation id="13072A5835506932FE46FB0FFDDFFB7B" box="[500,634,1191,1218]" captionStart="FIGURE 5" captionStartId="5.[151,250,1385,1409]" captionTargetBox="[157,1426,387,1352]" captionTargetId="figure-109@5.[151,1436,381,1362]" captionTargetPageId="5" captionText="FIGURE 5. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756043" httpUri="https://zenodo.org/record/4756043/files/figure.png" pageId="4" pageNumber="123">Figs 5B, D</figureCitation>
, 
<figureCitation id="13072A5835506932FD3BFB0FFD68FB7B" box="[649,717,1191,1218]" captionStart="FIGURE 6" captionStartId="6.[151,250,1026,1050]" captionTargetBox="[169,1430,186,1002]" captionTargetId="figure-28@6.[151,1436,181,1002]" captionTargetPageId="6" captionText="FIGURE 6. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A–C, pereopods 5–7. Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756045" httpUri="https://zenodo.org/record/4756045/files/figure.png" pageId="4" pageNumber="123">6A–B</figureCitation>
) or slightly bifid (
<figureCitation id="13072A5835506932FC1BFB0FFBA3FB7B" box="[937,1030,1191,1218]" captionStart="FIGURE 5" captionStartId="5.[151,250,1385,1409]" captionTargetBox="[157,1426,387,1352]" captionTargetId="figure-109@5.[151,1436,381,1362]" captionTargetPageId="5" captionText="FIGURE 5. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756043" httpUri="https://zenodo.org/record/4756043/files/figure.png" pageId="4" pageNumber="123">Fig. 5C</figureCitation>
). Secondary unguis on pereopod 7 undivided with 4–7 spinules (
<figureCitation id="13072A5835506932FE52FB63FD92FB5F" box="[480,567,1227,1254]" captionStart="FIGURE 6" captionStartId="6.[151,250,1026,1050]" captionTargetBox="[169,1430,186,1002]" captionTargetId="figure-28@6.[151,1436,181,1002]" captionTargetPageId="6" captionText="FIGURE 6. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A–C, pereopods 5–7. Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756045" httpUri="https://zenodo.org/record/4756045/files/figure.png" pageId="4" pageNumber="123">Fig. 6C</figureCitation>
). Ventral comb seta absent on merus and present on carpus of pereopods 6 and 7. Propodus of pereopods 3–5 with prominent barbed projection opposing secondary unguis, projection reduced on pereopod 2 (
<figureCitation id="13072A5835506932FE91FABBFEC9FA97" box="[291,364,1299,1326]" captionStart="FIGURE 5" captionStartId="5.[151,250,1385,1409]" captionTargetBox="[157,1426,387,1352]" captionTargetId="figure-109@5.[151,1436,381,1362]" captionTargetPageId="5" captionText="FIGURE 5. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756043" httpUri="https://zenodo.org/record/4756043/files/figure.png" pageId="4" pageNumber="123">Figs 5</figureCitation>
, 
<figureCitation id="13072A5835506932FECAFABBFE2DFA94" box="[376,392,1299,1325]" captionStart="FIGURE 6" captionStartId="6.[151,250,1026,1050]" captionTargetBox="[169,1430,186,1002]" captionTargetId="figure-28@6.[151,1436,181,1002]" captionTargetPageId="6" captionText="FIGURE 6. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A–C, pereopods 5–7. Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756045" httpUri="https://zenodo.org/record/4756045/files/figure.png" pageId="4" pageNumber="123">6</figureCitation>
, 
<figureCitation id="13072A5835506932FE26FABCFE12FA97" box="[404,439,1300,1326]" captionStart="FIGURE 8" captionStartId="9.[151,250,977,1001]" captionTargetBox="[151,1436,181,954]" captionTargetId="figure-17@9.[151,1436,181,954]" captionTargetPageId="9" captionText="FIGURE 8. Limnoria aspera sp. nov., paratype (male 2.8 mm, CBM-ZC 16565). A, left mandible, apical view, SEM image (JSM-6010LA; JEOL); B, pereopod 2, lateral view, SEM image (JSM-6010LA; JEOL); uropod, ventral view, SEM image (JSM-6010LA; JEOL). Scale bar: A = 0.02 mm, B, C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756055" httpUri="https://zenodo.org/record/4756055/files/figure.png" pageId="4" pageNumber="123">8B</figureCitation>
).
</paragraph>
<paragraph id="8B8336DD35506932FF75FA9FFBE2FA23" blockId="4.[151,1437,1155,2010]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75FA9FFE81FAE8" box="[199,292,1335,1361]" italics="true" pageId="4" pageNumber="123">Pleopod</emphasis>
2 with plumose setae up to 0.9 times length of exopod (
<figureCitation id="13072A5835506932FC12FA9FFC5CFAEB" box="[928,1017,1335,1362]" captionStart="FIGURE 7" captionStartId="6.[151,250,1861,1885]" captionTargetBox="[155,1431,1116,1830]" captionTargetId="figure-53@6.[151,1436,1111,1837]" captionTargetPageId="6" captionText="FIGURE 7. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleopod 1; B, pleopod 2; C, pleopod 5. Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756053" httpUri="https://zenodo.org/record/4756053/files/figure.png" pageId="4" pageNumber="123">Fig. 7B</figureCitation>
). Appendix masculina long, reaching beyond endopod tip, articulating near midlength of endopod. Endopod of pleopod 5, oval (
<figureCitation id="13072A5835506932FB17FAF3FAA5FACF" box="[1189,1280,1371,1398]" captionStart="FIGURE 7" captionStartId="6.[151,250,1861,1885]" captionTargetBox="[155,1431,1116,1830]" captionTargetId="figure-53@6.[151,1436,1111,1837]" captionTargetPageId="6" captionText="FIGURE 7. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleopod 1; B, pleopod 2; C, pleopod 5. Scale bars: A–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4756053" httpUri="https://zenodo.org/record/4756053/files/figure.png" pageId="4" pageNumber="123">Fig. 7C</figureCitation>
). Peduncle of pleopod 5 with simple seta laterally. Peduncles with coupling hook sequence 32220.
</paragraph>
<paragraph id="8B8336DD35506932FF75FA0BFCA4F9BC" blockId="4.[151,1437,1155,2010]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75FA0BFEB8FA04" box="[199,285,1443,1469]" italics="true" pageId="4" pageNumber="123">Uropod</emphasis>
exopod with laterally recurved apical claw (
<figureCitation id="13072A5835506932FCBEFA0BFCC3FA07" box="[780,870,1443,1470]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[163,1422,193,1909]" captionTargetId="figure-17@3.[151,1436,181,1918]" captionTargetPageId="3" captionText="FIGURE 3. Limnoria aspera sp. nov., holotype (male, 2.6 mm, CBM-ZC 16564). A, pleonite 5 and pleotelson; B, posterior margin of pleotelson; C, uropod; D, antenna 1; E, antenna 2. Scale bars: A, C–E = 0.1 mm, B = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756037" httpUri="https://zenodo.org/record/4756037/files/figure.png" pageId="4" pageNumber="123">Figs 3C</figureCitation>
, 
<figureCitation id="13072A5835506932FCC0FA0CFC33FA07" box="[882,918,1444,1470]" captionStart="FIGURE 8" captionStartId="9.[151,250,977,1001]" captionTargetBox="[151,1436,181,954]" captionTargetId="figure-17@9.[151,1436,181,954]" captionTargetPageId="9" captionText="FIGURE 8. Limnoria aspera sp. nov., paratype (male 2.8 mm, CBM-ZC 16565). A, left mandible, apical view, SEM image (JSM-6010LA; JEOL); B, pereopod 2, lateral view, SEM image (JSM-6010LA; JEOL); uropod, ventral view, SEM image (JSM-6010LA; JEOL). Scale bar: A = 0.02 mm, B, C = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4756055" httpUri="https://zenodo.org/record/4756055/files/figure.png" pageId="4" pageNumber="123">8C</figureCitation>
). Row of simple long setae on endopod placed apically. Uropod peduncle about 1.1 times as long as endopod, with many short simple setae and row of plumose setae. Exopod about 0.4–0.5 times as long as peduncle.
</paragraph>
<paragraph id="8B8336DD35506932FF75F9B8FD80F9C8" blockId="4.[151,1437,1155,2010]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75F9B8FE2CF993" bold="true" box="[199,393,1552,1578]" pageId="4" pageNumber="123">Molecular data.</emphasis>
<materialsCitation id="3B543C8035506932FE26F9B8FD84F9C8" ID-GBIF-Occurrence="3118985302" accessionNumber="LC610781, LC610782, LC612562, LC612563" pageId="4" pageNumber="123" specimenCount="2" typeStatus="holotype">
We deposited the nucleotide sequences of COI region of mitochondrial genes (Acc. no. 
<accessionNumber id="946FAB3E35506932FF25F99BFEB7F9F4" box="[151,274,1587,1613]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC610781" pageId="4" pageNumber="123">LC610781</accessionNumber>
, 
<accessionNumber id="946FAB3E35506932FE90F99BFE39F9F7" box="[290,412,1587,1614]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC610782" pageId="4" pageNumber="123">LC610782</accessionNumber>
) and 28S rRNA of nuclear genes (Acc. no. 
<accessionNumber id="946FAB3E35506932FC00F99BFB88F9F7" box="[946,1069,1587,1614]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC612562" pageId="4" pageNumber="123">LC612562</accessionNumber>
, 
<accessionNumber id="946FAB3E35506932FB8FF99BFB12F9F7" box="[1085,1207,1587,1614]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC612563" pageId="4" pageNumber="123">LC612563</accessionNumber>
) obtained from the 
<typeStatus id="5487887F35506932FF25F9FFFF5CF9C8" box="[151,249,1623,1649]" pageId="4" pageNumber="123" type="holotype">holotype</typeStatus>
and 
<typeStatus id="5487887F35506932FE82F9F0FE35F9CB" box="[304,400,1624,1650]" pageId="4" pageNumber="123" type="paratype">paratype</typeStatus>
in GenBank
</materialsCitation>
.
</paragraph>
<paragraph id="8B8336DD35506932FF75F9D3FC3CF92C" blockId="4.[151,1437,1155,2010]" box="[199,921,1659,1686]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75F9D3FEE4F92C" bold="true" box="[199,321,1659,1685]" pageId="4" pageNumber="123">Substrate.</emphasis>
<taxonomicName id="4C3C4D5E35506932FEFAF9D4FDCAF92C" authorityName="Harvey" authorityYear="1860" box="[328,623,1659,1686]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="4" pageNumber="123" phylum="Phaeophyta" rank="species" species="ringgoldianum">
<emphasis id="B948EACF35506932FEFAF9D4FDCAF92C" box="[328,623,1659,1686]" italics="true" pageId="4" pageNumber="123">Sargassum ringgoldianum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FDCCF9D4FC83F92C" authorityName="C.Agardh" authorityYear="1820" baseAuthorityName="Mert. ex Turner" box="[638,806,1659,1686]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fucales" pageId="4" pageNumber="123" phylum="Phaeophyta" rank="species" species="siliquastrum">
<emphasis id="B948EACF35506932FDCCF9D4FC83F92C" box="[638,806,1659,1686]" italics="true" pageId="4" pageNumber="123">S. siliquastrum</emphasis>
</taxonomicName>
holdfasts.
</paragraph>
</subSubSection>
<subSubSection id="C326655635506932FF75F937FD54F900" box="[199,753,1695,1722]" pageId="4" pageNumber="123" type="distribution">
<paragraph id="8B8336DD35506932FF75F937FD54F900" blockId="4.[151,1437,1155,2010]" box="[199,753,1695,1722]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75F937FEC4F900" bold="true" box="[199,353,1695,1721]" pageId="4" pageNumber="123">Distribution.</emphasis>
Known only from the 
<typeStatus id="5487887F35506932FDD2F908FD35F903" box="[608,656,1696,1722]" pageId="4" pageNumber="123">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C326655635506932FF75F96CFEF3F8B8" pageId="4" pageNumber="123" type="etymology">
<paragraph id="8B8336DD35506932FF75F96CFEF3F8B8" blockId="4.[151,1437,1155,2010]" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75F96CFEEBF967" bold="true" box="[199,334,1732,1758]" pageId="4" pageNumber="123">Etymology.</emphasis>
From the Latin 
<taxonomicName id="4C3C4D5E35506932FDB6F96CFDF5F964" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[516,592,1732,1757]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="aspera">
<emphasis id="B948EACF35506932FDB6F96CFDF5F964" box="[516,592,1732,1757]" italics="true" pageId="4" pageNumber="123">aspera</emphasis>
</taxonomicName>
(= rough; Gender feminine), referring to the rough exoskeleton on pleonite 5 and pleotelson.
</paragraph>
</subSubSection>
<subSubSection id="C326655635506933FF75F8A4FE69F826" lastPageId="5" lastPageNumber="124" pageId="4" pageNumber="123" type="discussion">
<paragraph id="8B8336DD35506933FF75F8A4FC0DFF40" blockId="4.[151,1437,1155,2010]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="124" pageId="4" pageNumber="123">
<emphasis id="B948EACF35506932FF75F8A4FE9EF89F" bold="true" box="[199,315,1804,1830]" pageId="4" pageNumber="123">Remarks.</emphasis>
Mandibular palp of 
<taxonomicName id="4C3C4D5E35506932FD9BF8A4FD42F89C" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[553,743,1804,1829]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35506932FD9BF8A4FD42F89C" box="[553,743,1804,1829]" italics="true" pageId="4" pageNumber="123">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35506932FD42F8A4FCE9F89F" bold="true" box="[752,844,1804,1830]" pageId="4" pageNumber="123">
<taxonomicNameLabel id="A27B57B435506932FD42F8A4FCE9F89F" box="[752,844,1804,1830]" pageId="4" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is reduced to a seta. This feature is also found in 7 species: 
<taxonomicName id="4C3C4D5E35506932FF4BF898FED9F8F0" baseAuthorityName="Ortiz &amp; Lalana" baseAuthorityYear="1988" box="[249,380,1839,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="bacescui">
<emphasis id="B948EACF35506932FF4BF898FED9F8F0" box="[249,380,1839,1865]" italics="true" pageId="4" pageNumber="123">L. bacescui</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FE3EF898FDE7F8F0" authorityName="Pillai" authorityYear="1957" box="[396,578,1839,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B948EACF35506932FE3EF898FDE7F8F0" box="[396,578,1839,1865]" italics="true" pageId="4" pageNumber="123">L. bituberculata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FDE1F898FD08F8F3" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[595,685,1840,1866]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35506932FDE1F898FD08F8F3" box="[595,685,1840,1866]" italics="true" pageId="4" pageNumber="123">L. furca</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FD0FF898FC91F8F0" authorityName="Nunomura" authorityYear="2012" box="[701,820,1840,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35506932FD0FF898FC91F8F0" box="[701,820,1840,1865]" italics="true" pageId="4" pageNumber="123">L. nagatai</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FCF6F898FC77F8F0" authorityName="Menzies" authorityYear="1957" box="[836,978,1839,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="segnoides">
<emphasis id="B948EACF35506932FCF6F898FC77F8F0" box="[836,978,1839,1865]" italics="true" pageId="4" pageNumber="123">L. segnoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FC51F898FBD0F8F0" authorityName="Cookson" authorityYear="1991" box="[995,1141,1839,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="uncapedis">
<emphasis id="B948EACF35506932FC51F898FBD0F8F0" box="[995,1141,1839,1865]" italics="true" pageId="4" pageNumber="123">L. uncapedis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D5E35506932FB02F898FA83F8F0" baseAuthorityName="Kussakin" baseAuthorityYear="1963" box="[1200,1318,1840,1865]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="zinovae">
<emphasis id="B948EACF35506932FB02F898FA83F8F0" box="[1200,1318,1840,1865]" italics="true" pageId="4" pageNumber="123">L. zinovae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAD4B2C35506932FA85F887FF77F8D7" author="Cookson, L. J." pageId="4" pageNumber="123" pagination="137 - 262" refId="ref4601" refString="Cookson, L. J. (1991) Australasian species of Limnoriidae (Crustacea: Isopoda.). Memoirs of Museum Victoria, 52 (2), 137 - 262. https: // doi. org / 10.24199 / j. mmv. 1991.52.02" type="journal article" year="1991">Cookson 1991</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FF50F8FBFEC6F8D7" author="Pillai, N. K." box="[226,355,1875,1902]" pageId="4" pageNumber="123" pagination="149 - 157" refId="ref5232" refString="Pillai, N. K. (1957) A new species of Limnoria from Kerala. Bulletin of the Central Research Institute, University of Kerala, 5 (2), 149 - 157." type="journal article" year="1957">Pillai 1957</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FEC1F8FBFD84F8D4" author="Kussakin, O. G." box="[371,545,1875,1901]" pageId="4" pageNumber="123" pagination="281 - 292" refId="ref4923" refString="Kussakin, O. G. (1963) Some data on the systematics of the family Limnoriidae (Isopoda) from northern and far-eastern seas of the USSR. Crustaceana, 5 (4), 281 - 292. https: // doi. org / 10.1163 / 156854063 x 00237" type="journal article" year="1963">Kussakin 1963</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FD83F8FBFD73F8D7" author="Menzies, R. J." box="[561,726,1875,1902]" pageId="4" pageNumber="123" pagination="101 - 200" refId="ref5096" refString="Menzies, R. J. (1957) The marine borer family Limnoriidae (Crustacea, Isopoda). Part I: Northern and Central America: Systematics, distribution, and ecology. Bulletin of Marine Science, 7 (2), 101 - 200." type="journal article" year="1957">Menzies 1957</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FD54F8FCFC00F8D7" author="Nunomura, N." box="[742,933,1876,1902]" pageId="4" pageNumber="123" pagination="77 - 85" refId="ref5145" refString="Nunomura, N. (2012) Marine isopod crustaceans of Seto Inland Sea deposited at Toyama Science Museum, 3. Suborder Cymothoida, Limnoriidea and Sphaeromatidea. Contributions from the Toyama Science Museum, 35, 77 - 85." type="journal article" year="2012">Nunomura 2012</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FC06F8FBFB0EF8D7" author="Ortiz, M. &amp; Lalana, R." box="[948,1195,1875,1902]" pageId="4" pageNumber="123" pagination="37 - 42" refId="ref5187" refString="Ortiz, M. &amp; Lalana, R. (1988) Una nueva especie del genero Phycolimnoria (Isopoda, Limnoriidae) de aguas cubanas. Revista de Investigaciones Marinas, 9 (2), 37 - 42. [in Spanish]" type="journal article" year="1988">Ortiz &amp; Lalana 1988</bibRefCitation>
; 
<bibRefCitation id="EFAD4B2C35506932FB08F8FBFF71F82B" author="Yoshino, H. &amp; Ohsawa, T. A." pageId="4" pageNumber="123" pagination="236 - 250" refId="ref5781" refString="Yoshino, H. &amp; Ohsawa, T. A. (2019) Description of new species of algal-boring Limnoria (Crustacea, Isopoda, Limnoriidae) from Japan and redescription of Limnoria segnoides Menzies, 1957 and L. nagatai Nunomura, 2012. Zootaxa, 4550 (2), 236 - 250. https: // doi. org / 10.11646 / zootaxa. 4550.2.5" type="journal article" year="2019">Yoshino &amp; Ohsawa 2019</bibRefCitation>
). 
<bibRefCitation id="EFAD4B2C35506932FF55F8DFFE2FF82B" author="Cookson, L. J." box="[231,394,1911,1938]" pageId="4" pageNumber="123" pagination="137 - 262" refId="ref4601" refString="Cookson, L. J. (1991) Australasian species of Limnoriidae (Crustacea: Isopoda.). Memoirs of Museum Victoria, 52 (2), 137 - 262. https: // doi. org / 10.24199 / j. mmv. 1991.52.02" type="journal article" year="1991">Cookson 1991</bibRefCitation>
stated that “ 
<taxonomicName id="4C3C4D5E35506932FDABF8D0FD0CF828" authorityName="Cookson" authorityYear="1991" box="[537,681,1911,1937]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="uncapedis">
<emphasis id="B948EACF35506932FDABF8D0FD0CF828" box="[537,681,1911,1937]" italics="true" pageId="4" pageNumber="123">L. uncapedis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35506932FD05F8D0FCE6F828" authorityName="Menzies" authorityYear="1957" box="[695,835,1911,1937]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="segnoides">
<emphasis id="B948EACF35506932FD05F8D0FCE6F828" box="[695,835,1911,1937]" italics="true" pageId="4" pageNumber="123">L. segnoides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D5E35506932FCCEF8D0FB95F828" authorityName="Pillai" authorityYear="1957" box="[892,1072,1911,1937]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B948EACF35506932FCCEF8D0FB95F828" box="[892,1072,1911,1937]" italics="true" pageId="4" pageNumber="123">L. bituberculata</emphasis>
</taxonomicName>
all share the following features: broad maxillipedal epipod, loss of mandibular palp, similar shape of the lacinia mobilis of the right mandible, and modification of the secondary unguis on pereopod 1”, and 
<bibRefCitation id="EFAD4B2C35506932FC9FF817FBFFF863" author="Yoshino, H. &amp; Ohsawa, T. A." box="[813,1114,1983,2010]" pageId="4" pageNumber="123" pagination="236 - 250" refId="ref5781" refString="Yoshino, H. &amp; Ohsawa, T. A. (2019) Description of new species of algal-boring Limnoria (Crustacea, Isopoda, Limnoriidae) from Japan and redescription of Limnoria segnoides Menzies, 1957 and L. nagatai Nunomura, 2012. Zootaxa, 4550 (2), 236 - 250. https: // doi. org / 10.11646 / zootaxa. 4550.2.5" type="journal article" year="2019">Yoshino &amp; Ohsawa (2019)</bibRefCitation>
said that the species of non- mandibular-palp group are algal-feeding and shared the broad epipod of maxilliped, except seagrass-feeding 
<taxonomicName id="4C3C4D5E35516933FA34FF3FFF48FF6D" baseAuthorityName="Kussakin" baseAuthorityYear="1963" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="zinovae">
<emphasis id="B948EACF35516933FA34FF3FFF48FF6D" italics="true" pageId="5" pageNumber="124">L. zinovae</emphasis>
</taxonomicName>
, the shape of epipod of maxilliped of which is unknown. 
<taxonomicName id="4C3C4D5E35516933FCC3FF13FC7FFF6D" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[881,986,187,212]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FCC3FF13FC7FFF6D" box="[881,986,187,212]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FC53FF13FB9EFF6C" bold="true" box="[993,1083,187,213]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FC53FF13FB9EFF6C" box="[993,1083,187,213]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
fits into this group, as an algalfeeding species that also has the broad epipod of maxilliped (
<tableCitation id="C6BE036635516933FCF1FF77FC32FF40" box="[835,919,223,249]" captionStart="TABLE 1" captionStartId="7.[151,239,640,666]" captionTargetPageId="7" captionText="TABLE 1. Comparative table of morphological characters of non-mandibular-palp group of Limnoria (Isopoda, Limnoriidae)." httpUri="http://table.plazi.org/id/DF43665535536931FF25FD28FA79FD23" pageId="5" pageNumber="124" tableUuid="DF43665535536931FF25FD28FA79FD23">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B8336DD35516933FF75FEABFEF3FEDC" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="124">
<taxonomicName id="4C3C4D5E35516933FF75FEABFEEDFEA4" baseAuthorityName="Ortiz &amp; Lalana" baseAuthorityYear="1988" box="[199,328,259,285]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="bacescui">
<emphasis id="B948EACF35516933FF75FEABFEEDFEA4" box="[199,328,259,285]" italics="true" pageId="5" pageNumber="124">L. bacescui</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D5E35516933FEE5FEABFE41FEA4" authorityName="Menzies" authorityYear="1957" box="[343,484,259,285]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="segnoides">
<emphasis id="B948EACF35516933FEE5FEABFE41FEA4" box="[343,484,259,285]" italics="true" pageId="5" pageNumber="124">L. segnoides</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3C4D5E35516933FD96FEABFD3CFEA5" baseAuthorityName="Kussakin" baseAuthorityYear="1963" box="[548,665,259,284]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="zinovae">
<emphasis id="B948EACF35516933FD96FEABFD3CFEA5" box="[548,665,259,284]" italics="true" pageId="5" pageNumber="124">L. zinovae</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="4C3C4D5E35516933FC97FEABFC35FEA5" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[805,912,259,284]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FC97FEABFC35FEA5" box="[805,912,259,284]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FC2AFEABFC56FEA4" bold="true" box="[920,1011,259,285]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FC2AFEABFC56FEA4" box="[920,1011,259,285]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the number of flagellar articles of antenna 2, the secondary unguis on the pereopods, lacinia mobilis of right mandible, and the sculpturing of pleonite 5 and pleotelson.
</paragraph>
<caption id="DF43665535516933FF25FAC1FCADFA1C" ID-DOI="http://doi.org/10.5281/zenodo.4756043" ID-Zenodo-Dep="4756043" httpUri="https://zenodo.org/record/4756043/files/figure.png" pageId="5" pageNumber="124" startId="5.[151,250,1385,1409]" targetBox="[157,1426,387,1352]" targetPageId="5">
<paragraph id="8B8336DD35516933FF25FAC1FCADFA1C" blockId="5.[151,1436,1385,1446]" pageId="5" pageNumber="124">
<emphasis id="B948EACF35516933FF25FAC1FEB0FA3B" bold="true" box="[151,277,1385,1410]" pageId="5" pageNumber="124">FIGURE 5.</emphasis>
<taxonomicName id="4C3C4D5E35516933FEACFAC2FE6EFA38" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[286,459,1386,1409]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FEACFAC2FE6EFA38" box="[286,459,1386,1409]" italics="true" pageId="5" pageNumber="124">Limnoria aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FE66FAC2FD82FA3B" bold="true" box="[468,551,1386,1410]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FE66FAC2FD82FA3B" box="[468,551,1386,1410]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (male, 2.6 mm, CBM-ZC 16564). A, pereopod 1 and secondary unguis of pereopod 1; B–D, pereopods 2–4. Scale bars: A–D = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8336DD35516933FF75FA7DFB62F9E3" blockId="5.[151,1437,1492,1951]" pageId="5" pageNumber="124">
<taxonomicName id="4C3C4D5E35516933FF75FA7DFE94FA57" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[199,305,1493,1518]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FF75FA7DFE94FA57" box="[199,305,1493,1518]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FE8BFA7CFE36FA57" bold="true" box="[313,403,1492,1518]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FE8BFA7CFE36FA57" box="[313,403,1492,1518]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="4C3C4D5E35516933FD95FA7DFD1DFA57" authorityName="Cookson" authorityYear="1991" box="[551,696,1492,1518]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="uncapedis">
<emphasis id="B948EACF35516933FD95FA7DFD1DFA57" box="[551,696,1492,1518]" italics="true" pageId="5" pageNumber="124">L. uncapedis</emphasis>
</taxonomicName>
in the number of flagellar articles of antennae 1 and 2, secondary unguis of pereopod 1, lacinia mobilis of right and left mandible, and pereopods 2–5 with barbed projection on the propodus. However, 
<taxonomicName id="4C3C4D5E35516933FE31F9B5FDB1F98F" authorityName="Cookson" authorityYear="1991" box="[387,532,1564,1590]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="uncapedis">
<emphasis id="B948EACF35516933FE31F9B5FDB1F98F" box="[387,532,1564,1590]" italics="true" pageId="5" pageNumber="124">L. uncapedis</emphasis>
</taxonomicName>
is pale yellow and has no carinae on pleonite 5 and pleotelson, while 
<taxonomicName id="4C3C4D5E35516933FA83F9B5FA39F98F" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[1329,1436,1565,1590]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FA83F9B5FA39F98F" box="[1329,1436,1565,1590]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FF25F9E8FF54F9E3" bold="true" box="[151,241,1600,1626]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FF25F9E8FF54F9E3" box="[151,241,1600,1626]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is dark yellow and has Y- or I-shaped carina on pleonite 5 and 1 tubercle on pleotelson.
</paragraph>
<paragraph id="8B8336DD35516933FF75F9CCFD72F953" blockId="5.[151,1437,1492,1951]" pageId="5" pageNumber="124">
Similarly to 
<taxonomicName id="4C3C4D5E35516933FEE1F9CDFE18F9C7" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[339,445,1637,1662]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FEE1F9CDFE18F9C7" box="[339,445,1637,1662]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FE76F9CCFDBBF9C7" bold="true" box="[452,542,1636,1662]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FE76F9CCFDBBF9C7" box="[452,542,1636,1662]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C3C4D5E35516933FD99F9CDFD7AF9C7" authorityName="Pillai" authorityYear="1957" box="[555,735,1636,1662]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B948EACF35516933FD99F9CDFD7AF9C7" box="[555,735,1636,1662]" italics="true" pageId="5" pageNumber="124">L. bituberculata</emphasis>
</taxonomicName>
has 4 flagellar articles of antenna 1, spinules on the secondary unguis of pereopod 1, and a dorsomedial longitudinal carina between 2 longitudinal grooves on pleonite 5. However, 
<taxonomicName id="4C3C4D5E35516933FF25F905FEE8F97F" authorityName="Pillai" authorityYear="1957" box="[151,333,1708,1734]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B948EACF35516933FF25F905FEE8F97F" box="[151,333,1708,1734]" italics="true" pageId="5" pageNumber="124">L. bituberculata</emphasis>
</taxonomicName>
is pale yellow and has 2 prominent tubercles on the pleotelson, while 
<taxonomicName id="4C3C4D5E35516933FBC5F905FB47F97F" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[1143,1250,1709,1734]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FBC5F905FB47F97F" box="[1143,1250,1709,1734]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FB59F904FAE3F97F" bold="true" box="[1259,1350,1708,1734]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FB59F904FAE3F97F" box="[1259,1350,1708,1734]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is dark yellow and has 1 medial tubercle on the pleotelson.
</paragraph>
<paragraph id="8B8336DD35516933FF75F95DFE69F826" blockId="5.[151,1437,1492,1951]" pageId="5" pageNumber="124">
<taxonomicName id="4C3C4D5E35516933FF75F95DFE95F8B7" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[199,304,1781,1806]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FF75F95DFE95F8B7" box="[199,304,1781,1806]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FE85F95CFE35F8B7" bold="true" box="[311,400,1780,1806]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FE85F95CFE35F8B7" box="[311,400,1780,1806]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to be most similar to 
<taxonomicName id="4C3C4D5E35516933FD7FF95DFC81F8B7" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[717,804,1780,1806]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35516933FD7FF95DFC81F8B7" box="[717,804,1780,1806]" italics="true" pageId="5" pageNumber="124">L. furca</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3C4D5E35516933FCD3F95DFC71F8B7" authorityName="Nunomura" authorityYear="2012" box="[865,980,1781,1806]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35516933FCD3F95DFC71F8B7" box="[865,980,1781,1806]" italics="true" pageId="5" pageNumber="124">L. nagatai</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C3C4D5E35516933FC53F95DFBEFF8B7" authority="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa, 2021" authorityName="Yoshino &amp; Kambara &amp; Kajiwara &amp; Ohsawa" authorityYear="2021" box="[993,1098,1781,1806]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B948EACF35516933FC53F95DFBEFF8B7" box="[993,1098,1781,1806]" italics="true" pageId="5" pageNumber="124">L. aspera</emphasis>
</taxonomicName>
<emphasis id="B948EACF35516933FBE3F95CFB0FF8B7" bold="true" box="[1105,1194,1780,1806]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A27B57B435516933FBE3F95CFB0FF8B7" box="[1105,1194,1780,1806]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from 
<taxonomicName id="4C3C4D5E35516933FF25F8B1FF4BF88B" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[151,238,1816,1842]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35516933FF25F8B1FF4BF88B" box="[151,238,1816,1842]" italics="true" pageId="5" pageNumber="124">L. furca</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3C4D5E35516933FE98F8B1FE38F88B" authorityName="Nunomura" authorityYear="2012" box="[298,413,1817,1842]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35516933FE98F8B1FE38F88B" box="[298,413,1817,1842]" italics="true" pageId="5" pageNumber="124">L. nagatai</emphasis>
</taxonomicName>
by dark yellow body color (
<taxonomicName id="4C3C4D5E35516933FD64F8B1FC88F88B" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[726,813,1816,1842]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35516933FD64F8B1FC88F88B" box="[726,813,1816,1842]" italics="true" pageId="5" pageNumber="124">L. furca</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3C4D5E35516933FCDBF8B1FC79F88B" authorityName="Nunomura" authorityYear="2012" box="[873,988,1817,1842]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35516933FCDBF8B1FC79F88B" box="[873,988,1817,1842]" italics="true" pageId="5" pageNumber="124">L. nagatai</emphasis>
</taxonomicName>
pale yellow), undivided or slightly bifid secondary unguis of pereopods (
<taxonomicName id="4C3C4D5E35516933FDB7F895FDF8F8EF" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[517,605,1852,1878]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35516933FDB7F895FDF8F8EF" box="[517,605,1852,1878]" italics="true" pageId="5" pageNumber="124">L. furca</emphasis>
</taxonomicName>
bifid or trifid, 
<taxonomicName id="4C3C4D5E35516933FCB4F895FCDFF8EF" authorityName="Nunomura" authorityYear="2012" box="[774,890,1853,1878]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35516933FCB4F895FCDFF8EF" box="[774,890,1853,1878]" italics="true" pageId="5" pageNumber="124">L. nagatai</emphasis>
</taxonomicName>
bifid), and the medial tubercle on the pleotelson lacks the attachment of two inverted V-shaped carinae (
<taxonomicName id="4C3C4D5E35516933FCB4F8C9FCF8F8C3" authorityName="Yoshino &amp; Ohsawa" authorityYear="2019" box="[774,861,1888,1914]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="furca">
<emphasis id="B948EACF35516933FCB4F8C9FCF8F8C3" box="[774,861,1888,1914]" italics="true" pageId="5" pageNumber="124">L. furca</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3C4D5E35516933FC29F8C9FBABF8C3" authorityName="Nunomura" authorityYear="2012" box="[923,1038,1889,1914]" class="Malacostraca" family="Limnoriidae" genus="Limnoria" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="nagatai">
<emphasis id="B948EACF35516933FC29F8C9FBABF8C3" box="[923,1038,1889,1914]" italics="true" pageId="5" pageNumber="124">L. nagatai</emphasis>
</taxonomicName>
with medial tubercle followed by a pair of subparallel carinae).
</paragraph>
</subSubSection>
</treatment>
</document>