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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FFBAFFA6667FFEEE640377BD" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Odontopygidae Attems 1909" docType="key" docVersion="2" lastPageNumber="293" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="292" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vohland, Katrin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamer, Michelle</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>251</mods:start>
<mods:end>251</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039587C7FFBAFFA6667FFEEE640377BD" httpUri="http://treatment.plazi.org/id/039587C7FFBAFFA6667FFEEE640377BD" lastPageId="42" lastPageNumber="293" pageId="41" pageNumber="292">
<subSubSection box="[352,971,365,391]" pageId="41" pageNumber="292" type="nomenclature">
<paragraph blockId="41.[352,971,365,391]" box="[352,971,365,391]" pageId="41" pageNumber="292">
<heading box="[352,971,365,391]" centered="true" fontSize="36" level="2" pageId="41" pageNumber="292" reason="5">
Key to species of
<taxonomicName ID-CoL="7NK6C" ID-ENA="244995" authorityName="Attems" authorityYear="1909" box="[552,725,365,391]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="41" pageNumber="292" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
occurring in
<collectingCountry box="[875,971,365,391]" name="Namibia" pageId="41" pageNumber="292">Namibia</collectingCountry>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="293" pageId="41" pageNumber="292" type="key">
<keyStep pageId="41" pageNumber="292">
<paragraph blockId="41.[181,1142,407,530]" pageId="41" pageNumber="292">
<keyLead pageId="41" pageNumber="292">
1 Minute species, width &lt;
<quantity box="[500,566,407,434]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="292" unit="mm" value="1.0">1 mm</quantity>
, paraprocts without spines (
<figureCitation box="[887,975,407,433]" captionStart="Figs 76–78" captionStartId="43.[181,220,790,812]" captionTargetBox="[221,1094,219,769]" captionTargetId="figure-334@43.[181,1141,204,773]" captionTargetPageId="43" captionText="Figs 76–78.Bandeirenica andara sp. n., ♂: (76) head and first segments, lateral view; (77) gonopod, aboral view; (78) last segments, lateral view.Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (lp) lateral process, (pa) paraproct, (pofp) postfemoral process, (prfp) prefemoral process, (tp1) tarsal process 1, (tp2) tarsal process 2. Scale bar = 1 mm." pageId="41" pageNumber="292">Figs 78</figureCitation>
,
<figureCitation box="[989,1020,407,433]" captionStart="Figs 88–90" captionStartId="49.[181,220,680,702]" captionTargetBox="[220,1117,222,654]" captionTargetId="figure-325@49.[181,1141,204,664]" captionTargetPageId="49" captionText="Figs 88–90. Syndesmogenus fragilis sp. n., ♂: (88) gonopod, aboral view; (89) last segments, lateral view; (90) head and first segments,lateral view.Abbreviations:(ap)apical process,(col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (prfp) prefemoral process, (s) solenomere, (sp2) small process of solenomere, (tc) telocoxite process. Scale bar = 1 mm." pageId="41" pageNumber="292">89</figureCitation>
); gonopod solenomere with distal spinous processes (
<figureCitation box="[697,783,439,465]" captionStart="Figs 76–78" captionStartId="43.[181,220,790,812]" captionTargetBox="[221,1094,219,769]" captionTargetId="figure-334@43.[181,1141,204,773]" captionTargetPageId="43" captionText="Figs 76–78.Bandeirenica andara sp. n., ♂: (76) head and first segments, lateral view; (77) gonopod, aboral view; (78) last segments, lateral view.Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (lp) lateral process, (pa) paraproct, (pofp) postfemoral process, (prfp) prefemoral process, (tp1) tarsal process 1, (tp2) tarsal process 2. Scale bar = 1 mm." pageId="41" pageNumber="292">Figs 78</figureCitation>
,
<figureCitation box="[795,824,439,465]" captionStart="Figs 88–90" captionStartId="49.[181,220,680,702]" captionTargetBox="[220,1117,222,654]" captionTargetId="figure-325@49.[181,1141,204,664]" captionTargetPageId="49" captionText="Figs 88–90. Syndesmogenus fragilis sp. n., ♂: (88) gonopod, aboral view; (89) last segments, lateral view; (90) head and first segments,lateral view.Abbreviations:(ap)apical process,(col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (prfp) prefemoral process, (s) solenomere, (sp2) small process of solenomere, (tc) telocoxite process. Scale bar = 1 mm." pageId="41" pageNumber="292">88</figureCitation>
) .........................................2
</keyLead>
</paragraph>
<paragraph blockId="41.[181,1142,407,530]" pageId="41" pageNumber="292">
<keyLead pageId="41" pageNumber="292">
– Medium-sized species, width&gt;
<quantity box="[589,661,471,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="41" pageNumber="292" unit="mm" value="2.0">2 mm</quantity>
, paraprocts with dorsal spines (
<figureCitation box="[1038,1127,471,497]" captionStart="Figs 79–82" captionStartId="45.[181,220,1464,1486]" captionTargetBox="[249,1092,214,1446]" captionTargetId="figure-42@45.[181,1141,204,1466]" captionTargetPageId="45" captionText="Figs 79–82. Chaleponcus limbatus Attems, ♂: (79) gonopod, aboral view; (80) epiproct, lateral view; (81) gonopodial aperture; (82) head and segments 1–8, lateral view. Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (s) solenomere, (ste) sternite, (tl1, tl2, tl3) tarsal lobes 1–3. Scale bar = 1 mm." pageId="41" pageNumber="292">Fig. 80</figureCitation>
); gonopod solenomere without distal processes (
<figureCitation box="[745,832,503,529]" captionStart="Figs 79–82" captionStartId="45.[181,220,1464,1486]" captionTargetBox="[249,1092,214,1446]" captionTargetId="figure-42@45.[181,1141,204,1466]" captionTargetPageId="45" captionText="Figs 79–82. Chaleponcus limbatus Attems, ♂: (79) gonopod, aboral view; (80) epiproct, lateral view; (81) gonopodial aperture; (82) head and segments 1–8, lateral view. Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (s) solenomere, (ste) sternite, (tl1, tl2, tl3) tarsal lobes 1–3. Scale bar = 1 mm." pageId="41" pageNumber="292">Figs 79</figureCitation>
,
<figureCitation box="[844,874,503,529]" captionStart="Figs 83, 84" captionStartId="46.[144,183,1606,1628]" captionTargetBox="[172,1081,1056,1581]" captionTargetId="figure-327@46.[144,1104,1050,1591]" captionTargetPageId="46" captionText="Figs 83, 84. Chaleponcus niger Attems, ♂: (83) gonopod, aboral view; (84) mid­body leg. Abbreviations: (cx) coxa, (f) femur,(fp) femoral process, (prf) prefemur, (ptf) postfemur,(s) solenomere,(ta) tarsus, (ti) tibia. Scale bar = 1 mm." pageId="41" pageNumber="292">83</figureCitation>
,
<figureCitation box="[886,915,503,530]" captionStart="Figs 85–87" captionStartId="47.[181,220,1606,1628]" captionTargetBox="[197,1122,1071,1591]" captionTargetId="figure-315@47.[181,1141,1067,1596]" captionTargetPageId="47" captionText="Figs 85–87. Spinotarsus xanthonotus Attems, ♂: (85) gonopod, aboral view; (86) telopodite, oral view; (87) telocoxite, oral view.Abbreviations: (s) solenomere, (prfp) prefemoral process, (ta) tarsal lobe, (tip) tibial process, (tl1) basal lobe of tarsus. Scale bar = 1 mm." pageId="41" pageNumber="292">85</figureCitation>
).............................3
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="41" pageNumber="292">
<paragraph blockId="41.[181,1142,546,669]" pageId="41" pageNumber="292">
<keyLead pageId="41" pageNumber="292">
2 Less than 60 segments; telocoxite without basally directed acute process (
<figureCitation box="[1048,1129,546,572]" captionStart="Figs 76–78" captionStartId="43.[181,220,790,812]" captionTargetBox="[221,1094,219,769]" captionTargetId="figure-334@43.[181,1141,204,773]" captionTargetPageId="43" captionText="Figs 76–78.Bandeirenica andara sp. n., ♂: (76) head and first segments, lateral view; (77) gonopod, aboral view; (78) last segments, lateral view.Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (lp) lateral process, (pa) paraproct, (pofp) postfemoral process, (prfp) prefemoral process, (tp1) tarsal process 1, (tp2) tarsal process 2. Scale bar = 1 mm." pageId="41" pageNumber="292">Fig. 77</figureCitation>
) .....................................................................................
<taxonomicName authority="Vohland &amp; Hamer, 2013" authorityName="Vohland &amp; Hamer" authorityYear="2013" box="[824,1075,578,604]" class="Diplopoda" family="Odontopygidae" genus="Bandeirenica" kingdom="Animalia" order="Spirostreptida" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="andara" status="sp. n.">
<emphasis bold="true" box="[824,1075,578,604]" pageId="41" pageNumber="292">Bandeirenica andara</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1141,578,604]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph blockId="41.[181,1142,546,669]" pageId="41" pageNumber="292">
<keyLead pageId="41" pageNumber="292">
– More than 70 segments; telocoxite distal-orally with basally directed acute process (
<figureCitation box="[230,312,642,668]" captionStart="Figs 88–90" captionStartId="49.[181,220,680,702]" captionTargetBox="[220,1117,222,654]" captionTargetId="figure-325@49.[181,1141,204,664]" captionTargetPageId="49" captionText="Figs 88–90. Syndesmogenus fragilis sp. n., ♂: (88) gonopod, aboral view; (89) last segments, lateral view; (90) head and first segments,lateral view.Abbreviations:(ap)apical process,(col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (prfp) prefemoral process, (s) solenomere, (sp2) small process of solenomere, (tc) telocoxite process. Scale bar = 1 mm." pageId="41" pageNumber="292">Fig. 88</figureCitation>
,
<emphasis box="[324,344,643,668]" italics="true" pageId="41" pageNumber="292">tc</emphasis>
)..............................................................
<taxonomicName authority="Vohland &amp; Hamer, 2013" authorityName="Vohland &amp; Hamer" authorityYear="2013" box="[794,1075,642,668]" class="Diplopoda" family="Odontopygidae" genus="Syndesmogenus" kingdom="Animalia" order="Spirostreptida" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="fragilis" status="sp. n.">
<emphasis bold="true" box="[794,1075,642,668]" pageId="41" pageNumber="292">Syndesmogenus fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1141,643,669]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="42" lastPageNumber="293" pageId="41" pageNumber="292">
<paragraph blockId="41.[181,1142,685,775]" pageId="41" pageNumber="292">
<keyLead pageId="41" pageNumber="292">
3 With red longitudinal line dorsally; solenomere spiralled distally, main part of tarsus comprising a single leaf and much enlarged (
<figureCitation box="[725,806,717,743]" captionStart="Figs 85–87" captionStartId="47.[181,220,1606,1628]" captionTargetBox="[197,1122,1071,1591]" captionTargetId="figure-315@47.[181,1141,1067,1596]" captionTargetPageId="47" captionText="Figs 85–87. Spinotarsus xanthonotus Attems, ♂: (85) gonopod, aboral view; (86) telopodite, oral view; (87) telocoxite, oral view.Abbreviations: (s) solenomere, (prfp) prefemoral process, (ta) tarsal lobe, (tip) tibial process, (tl1) basal lobe of tarsus. Scale bar = 1 mm." pageId="41" pageNumber="292">Fig. 86</figureCitation>
) .............................................. ..................................................................
<taxonomicName authority="Attems, 1909" authorityName=": Attems" authorityYear="1909" box="[689,1142,748,775]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="xanthonotus">
<emphasis bold="true" box="[689,985,748,775]" pageId="41" pageNumber="292">Spinotarsus xanthonotus</emphasis>
<bibRefCitation author="ATTEMS, C. G." box="[991,1142,749,775]" pageId="41" pageNumber="292" pagination="1 - 52" refId="ref23416" refString="ATTEMS, C. G. 1909 a. Myriapoda. In: Schultze, L., ed., Zoologische und Anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika, ausgefuhrt in den Jahren 1901 - 1903 von Dr. Leonhard Schultze. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 14: 1 - 52." type="journal article" year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<caption box="[320,1004,1654,1676]" pageId="41" pageNumber="292" startId="41.[320,353,1654,1676]" targetBox="[210,1111,843,1629]" targetPageId="41" targetType="figure">
<paragraph blockId="41.[320,1004,1654,1676]" box="[320,1004,1654,1676]" pageId="41" pageNumber="292">
Fig. 75. Distribution of the family
<taxonomicName authorityName="Attems" authorityYear="1909" box="[634,774,1654,1676]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="41" pageNumber="292" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
(
<taxonomicName authorityName="de Blainville in Gervais" authorityYear="1844" box="[785,884,1654,1676]" class="Diplopoda" kingdom="Animalia" pageId="41" pageNumber="292" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
) in Namibia.
</paragraph>
</caption>
<paragraph blockId="42.[144,1104,204,470]" pageId="42" pageNumber="293">
<keyLead pageId="42" pageNumber="293">
– Animal with transversely striped appearance; solenomere not distally spiralled, main part of tarsus divided into three leaves, the medium one beset with hairs (
<figureCitation box="[1011,1099,236,262]" captionStart="Figs 79–82" captionStartId="45.[181,220,1464,1486]" captionTargetBox="[249,1092,214,1446]" captionTargetId="figure-42@45.[181,1141,204,1466]" captionTargetPageId="45" captionText="Figs 79–82. Chaleponcus limbatus Attems, ♂: (79) gonopod, aboral view; (80) epiproct, lateral view; (81) gonopodial aperture; (82) head and segments 1–8, lateral view. Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (s) solenomere, (ste) sternite, (tl1, tl2, tl3) tarsal lobes 1–3. Scale bar = 1 mm." pageId="42" pageNumber="293">Figs 79</figureCitation>
,
<figureCitation box="[184,213,268,294]" captionStart="Figs 83, 84" captionStartId="46.[144,183,1606,1628]" captionTargetBox="[172,1081,1056,1581]" captionTargetId="figure-327@46.[144,1104,1050,1591]" captionTargetPageId="46" captionText="Figs 83, 84. Chaleponcus niger Attems, ♂: (83) gonopod, aboral view; (84) mid­body leg. Abbreviations: (cx) coxa, (f) femur,(fp) femoral process, (prf) prefemur, (ptf) postfemur,(s) solenomere,(ta) tarsus, (ti) tibia. Scale bar = 1 mm." pageId="42" pageNumber="293">83</figureCitation>
) (
<taxonomicName authorityName=": Attems" authorityYear="1914" box="[237,383,268,294]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="42" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis box="[237,383,268,294]" italics="true" pageId="42" pageNumber="293">Chaleponcus</emphasis>
</taxonomicName>
) ...................................................................................................4
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="293">
<paragraph blockId="42.[144,1104,204,470]" pageId="42" pageNumber="293">
<keyLead pageId="42" pageNumber="293">
4 Gonopod telocoxite apically sclerotised, with lateral spines (
<figureCitation box="[861,942,308,334]" captionStart="Figs 83, 84" captionStartId="46.[144,183,1606,1628]" captionTargetBox="[172,1081,1056,1581]" captionTargetId="figure-327@46.[144,1104,1050,1591]" captionTargetPageId="46" captionText="Figs 83, 84. Chaleponcus niger Attems, ♂: (83) gonopod, aboral view; (84) mid­body leg. Abbreviations: (cx) coxa, (f) femur,(fp) femoral process, (prf) prefemur, (ptf) postfemur,(s) solenomere,(ta) tarsus, (ti) tibia. Scale bar = 1 mm." pageId="42" pageNumber="293">Fig. 83</figureCitation>
) ..................... ...............................................................................................
<taxonomicName authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[851,1104,339,366]" class="Diplopoda" family="Paradoxosomatidae" genus="Habrodesmus" kingdom="Animalia" order="Polydesmida" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="niger">
<emphasis bold="true" box="[851,947,339,365]" pageId="42" pageNumber="293">C. niger</emphasis>
Attems, 1914
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="42.[144,1104,204,470]" box="[144,1104,371,398]" pageId="42" pageNumber="293">
<keyLead box="[144,1104,371,398]" pageId="42" pageNumber="293">
– Gonopod telocoxite not sclerotised apically, but broadly widened (
<figureCitation box="[928,1009,372,398]" captionStart="Figs 79–82" captionStartId="45.[181,220,1464,1486]" captionTargetBox="[249,1092,214,1446]" captionTargetId="figure-42@45.[181,1141,204,1466]" captionTargetPageId="45" captionText="Figs 79–82. Chaleponcus limbatus Attems, ♂: (79) gonopod, aboral view; (80) epiproct, lateral view; (81) gonopodial aperture; (82) head and segments 1–8, lateral view. Abbreviations: (col) collum, (e) epiproct, (hy) hypoproct, (pa) paraproct, (s) solenomere, (ste) sternite, (tl1, tl2, tl3) tarsal lobes 1–3. Scale bar = 1 mm." pageId="42" pageNumber="293">Fig. 79</figureCitation>
)..........5
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="42" pageNumber="293">
<paragraph blockId="42.[144,1104,204,470]" box="[144,1104,411,438]" pageId="42" pageNumber="293">
<keyLead box="[144,1104,411,438]" pageId="42" pageNumber="293">
5 Median lamella of telopodite tarsus prolonged..............
<taxonomicName authority="Attems, 1914" authorityName=": Attems" authorityYear="1914" box="[807,1104,411,438]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis bold="true" box="[807,946,411,437]" pageId="42" pageNumber="293">C. limbatus</emphasis>
Attems, 1914
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="42.[144,1104,204,470]" box="[144,1104,443,470]" pageId="42" pageNumber="293">
<keyLead box="[144,1104,443,470]" pageId="42" pageNumber="293">
– Median lamella of telopodite tarsus not prolonged.....
<taxonomicName authority="Attems, 1922" authorityName="Attems" authorityYear="1922" box="[785,1104,443,470]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="hereronius">
<emphasis bold="true" box="[785,947,443,469]" pageId="42" pageNumber="293">C. hereronius</emphasis>
Attems, 1922
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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