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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FF94FF846663FED7646F723D" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Cnemodesmus riparius : Shelley &amp; Crawford 1996" docType="treatment" docVersion="2" lastPageNumber="259" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="258" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vohland, Katrin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamer, Michelle</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>251</mods:start>
<mods:end>251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039587C7FF94FF846663FED7646F723D" httpUri="http://treatment.plazi.org/id/039587C7FF94FF846663FED7646F723D" lastPageId="8" lastPageNumber="259" pageId="7" pageNumber="258">
<subSubSection box="[380,943,339,366]" pageId="7" pageNumber="258" type="nomenclature">
<paragraph blockId="7.[380,943,339,366]" box="[380,943,339,366]" pageId="7" pageNumber="258">
<heading box="[380,943,339,366]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="258" reason="3">
<taxonomicName ID-CoL="WH8F" authority="Shelley &amp; Crawford, 1996" authorityName=": Shelley &amp; Crawford" authorityYear="1996" box="[380,943,339,366]" class="Diplopoda" family="Paradoxosomatidae" genus="Cnemodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="riparius">
<emphasis box="[380,639,340,366]" italics="true" pageId="7" pageNumber="258">Cnemodesmus riparius</emphasis>
<bibRefCitation author="SHELLEY, R. M. &amp; CRAWFORD, C. S." box="[646,943,339,366]" pageId="7" pageNumber="258" pagination="1 - 8" refId="ref27034" refString="SHELLEY, R. M. &amp; CRAWFORD, C. S. 1996. Cnemodesmus riparius sp. nov., a riparian millipede from the Namib Desert, Africa (Polydesmida: Paradoxosomatidae). Myriapodologica 4 (1): 1 - 8." type="journal article" year="1996">Shelley &amp; Crawford, 1996</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[611,712,382,408]" pageId="7" pageNumber="258" type="description">
<paragraph blockId="7.[181,1142,382,884]" box="[611,712,382,408]" pageId="7" pageNumber="258">
<figureCitation box="[611,712,382,408]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="7" pageNumber="258">Figs 2–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[181,935,420,443]" pageId="7" pageNumber="258" type="reference_group">
<paragraph blockId="7.[181,1142,382,884]" box="[181,935,420,443]" pageId="7" pageNumber="258">
<treatmentCitationGroup box="[181,935,420,443]" pageId="7" pageNumber="258">
<treatmentCitation author="SHELLEY, R. M. &amp; CRAWFORD, C. S." box="[181,687,420,443]" page="4" pageId="7" pageNumber="258" year="1996">
<taxonomicName ID-CoL="WH8F" authority=": Shelley &amp; Crawford 1996: 2 - 4" authorityName=": Shelley &amp; Crawford" authorityPageNumber="2 - 4" authorityYear="1996" box="[181,687,420,443]" class="Diplopoda" family="Paradoxosomatidae" genus="Cnemodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="riparius">
<emphasis box="[181,391,421,442]" italics="true" pageId="7" pageNumber="258">Cnemodesmus riparius</emphasis>
:
<bibRefCitation author="SHELLEY, R. M. &amp; CRAWFORD, C. S." box="[403,687,420,443]" pageId="7" pageNumber="258" pagination="1 - 8" refId="ref27034" refString="SHELLEY, R. M. &amp; CRAWFORD, C. S. 1996. Cnemodesmus riparius sp. nov., a riparian millipede from the Namib Desert, Africa (Polydesmida: Paradoxosomatidae). Myriapodologica 4 (1): 1 - 8." type="journal article" year="1996">Shelley &amp; Crawford 1996: 2–4</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1, 2; Hamer 1998: 25.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="259" pageId="7" pageNumber="258" type="diagnosis">
<paragraph blockId="7.[181,1142,382,884]" pageId="7" pageNumber="258">
Diagnosis: Gonopod solenomere slender, long, coiled with laminate, folded process along inner margin near midlength, femoral process club-like. Tibiotarsus with broad, short basal process and acute, more distal process (
<figureCitation box="[752,821,521,547]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="7" pageNumber="258">Fig. 2</figureCitation>
).
</paragraph>
<paragraph blockId="7.[181,1142,382,884]" pageId="7" pageNumber="258">
The closest relatives of this species are
<taxonomicName authority="(Cook &amp; Collins, 1893)" baseAuthorityName="Cook &amp; Collins" baseAuthorityYear="1893" box="[657,1093,553,579]" class="Diplopoda" family="Paradoxosomatidae" genus="Paradesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="thysanopus">
<emphasis box="[657,816,553,579]" italics="true" pageId="7" pageNumber="258">C. thysanopus</emphasis>
(
<bibRefCitation author="COOK, O. F. &amp; COLLINS, G. N." box="[832,1085,553,579]" pageId="7" pageNumber="258" pagination="24 - 40" refId="ref24346" refString="COOK, O. F. &amp; COLLINS, G. N. 1893. The Myriapods collected by the Eclipse Expedition to West Africa 1889 - 1890. Annals of the New York Academy of Sciences 8: 24 - 40." type="journal article" year="1893">Cook &amp; Collins, 1893</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[181,499,585,612]" class="Diplopoda" family="Paradoxosomatidae" genus="Cnemodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="calundensis">
<emphasis box="[181,348,585,611]" italics="true" pageId="7" pageNumber="258">C. calundensis</emphasis>
Kraus, 1958
</taxonomicName>
, from
<collectingCountry box="[575,661,585,611]" name="Angola" pageId="7" pageNumber="258">Angola</collectingCountry>
. In
<taxonomicName baseAuthorityName="Cook &amp; Collins" baseAuthorityYear="1893" box="[706,866,585,611]" class="Diplopoda" family="Paradoxosomatidae" genus="Paradesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="thysanopus">
<emphasis box="[706,866,585,611]" italics="true" pageId="7" pageNumber="258">C. thysanopus</emphasis>
</taxonomicName>
, the tibiotarsus is much longer and bifurcate, and the basal process more acute, while
<taxonomicName authorityName="Kraus" authorityYear="1958" box="[891,1058,617,643]" class="Diplopoda" family="Paradoxosomatidae" genus="Cnemodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="calundensis">
<emphasis box="[891,1058,617,643]" italics="true" pageId="7" pageNumber="258">C. calundensis</emphasis>
</taxonomicName>
can be distinguished mainly by the less curved solenomere.
</paragraph>
<paragraph blockId="7.[181,1142,382,884]" box="[181,321,689,715]" pageId="7" pageNumber="258">Description:</paragraph>
<paragraph blockId="7.[181,1142,382,884]" box="[181,459,729,756]" pageId="7" pageNumber="258">
<emphasis box="[181,226,729,755]" italics="true" pageId="7" pageNumber="258">Size</emphasis>
: Width 1.1–1.4 mm.
</paragraph>
<paragraph blockId="7.[181,1142,382,884]" pageId="7" pageNumber="258">
<emphasis box="[181,260,761,787]" italics="true" pageId="7" pageNumber="258">Colour</emphasis>
: Dark brown, suture dirty white posterior to metatergites, dark spot medially giving animal appearance of having dorsal longitudinal band.
</paragraph>
<paragraph blockId="7.[181,1142,382,884]" pageId="7" pageNumber="258">
<emphasis box="[181,242,825,851]" italics="true" pageId="7" pageNumber="258">Head</emphasis>
: Frons setose. Antenna becoming thicker distally, microsensilla on distal part of antennomeres 5+6 (
<figureCitation box="[406,475,857,884]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="7" pageNumber="258">Fig. 7</figureCitation>
,
<emphasis box="[487,518,858,883]" italics="true" pageId="7" pageNumber="258">ms</emphasis>
).
</paragraph>
<caption pageId="7" pageNumber="258" startId="7.[181,220,1558,1580]" targetBox="[192,1129,934,1534]" targetPageId="7" targetType="figure">
<paragraph blockId="7.[181,1142,1558,1676]" pageId="7" pageNumber="258">
Figs 2–7.
<taxonomicName authority="Shelley &amp; Crawford" authorityName=": Shelley &amp; Crawford" authorityYear="1996" box="[268,661,1558,1580]" class="Diplopoda" family="Paradoxosomatidae" genus="Cnemodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="riparius">
<emphasis box="[268,474,1558,1580]" italics="true" pageId="7" pageNumber="258">Cnemodesmus riparius</emphasis>
Shelley &amp; Crawford
</taxonomicName>
, ♂: (2) left gonopod (mesal aspect); (3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5
<superScript attach="left" box="[862,872,1582,1594]" fontSize="5" pageId="7" pageNumber="258">th</superScript>
leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph blockId="8.[144,1105,204,1110]" box="[144,527,204,230]" pageId="8" pageNumber="259">
<emphasis box="[144,226,204,230]" italics="true" pageId="8" pageNumber="259">Collum</emphasis>
: Broadly rounded (
<figureCitation box="[445,513,204,230]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="8" pageNumber="259">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
<emphasis box="[144,385,236,262]" italics="true" pageId="8" pageNumber="259">Tergites and sternites</emphasis>
: Metatergal sulcus smooth and long (
<figureCitation box="[814,884,236,262]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="8" pageNumber="259">Fig. 4</figureCitation>
). Paraterga narrow, projecting slightly backwards up to segment 19, distally rounded. Pleurotergal ridges thin, reaching to the mid-body. Surface smooth. Progonopodial sternite setose, postgonopodial sternite smooth. Sternite 5 with cones fused together, forming a subrectangular process (
<figureCitation box="[152,220,364,390]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="8" pageNumber="259">Fig. 6</figureCitation>
).
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
<emphasis box="[144,197,396,421]" italics="true" pageId="8" pageNumber="259">Legs</emphasis>
: Femur of legs on segments 4–6 with digitiform process (
<figureCitation box="[842,910,395,422]" captionStart="Figs 2–7" captionStartId="7.[181,220,1558,1580]" captionTargetBox="[192,1129,934,1534]" captionTargetId="figure-269@7.[181,1141,930,1540]" captionTargetPageId="7" captionText="Figs 2–7.Cnemodesmus riparius Shelley &amp; Crawford,♂:(2) left gonopod(mesal aspect);(3) collum (lateral aspect); (4) segment 16 (dorsal aspect); (5) adenostyle on femur of 5th leg pair; (6) conus on sternite 5; (7) antenna with microsensilla on antennomeres 6 and 7. Abbreviations: (col) collum, (cx) coxa, (fp) femoral process, (ms) microsensilla on antennae, (msu) metatergal sulcus, (mz) metazonite, (o) ozopore, (pn) paratergite, (pz) prozonite, (s) solenomere. Scale bars = 1 mm." pageId="8" pageNumber="259">Fig. 5</figureCitation>
,
<emphasis box="[921,943,396,422]" italics="true" pageId="8" pageNumber="259">fp</emphasis>
). Tibia of legs without brushes.
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" box="[144,452,460,486]" pageId="8" pageNumber="259">
<emphasis box="[144,259,460,486]" italics="true" pageId="8" pageNumber="259">Gonopods</emphasis>
: As in diagnosis.
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
<materialsCitation collectingDate="1972-04-19" collectionCode="NMNW" collectorName="P. G. Olivier" country="Namibia" latitude="-22.570002" location="Windhoek" longLatPrecision="21" longitude="17.083612" municipality="Material" pageId="8" pageNumber="259" specimenCode="SMN 21695" specimenCount="1" specimenCount-male="1" stateProvince="Khomas">
<collectingMunicipality box="[144,221,495,517]" pageId="8" pageNumber="259">Material</collectingMunicipality>
examined:
<collectingCountry box="[325,428,495,516]" name="Namibia" pageId="8" pageNumber="259">NAMIBIA</collectingCountry>
:
<specimenCount box="[436,464,495,516]" pageId="8" pageNumber="259" type="male">1♂</specimenCount>
<collectingRegion box="[469,615,495,517]" country="Namibia" name="Khomas" pageId="8" pageNumber="259">Khomas district</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:039587C7FF94FF846663FED7646F723D:8EE3600AFF9BFF846570FE6C629E746E" box="[623,717,495,517]" country="Namibia" latitude="-22.570002" longLatPrecision="21" longitude="17.083612" municipality="Material" name="Windhoek" pageId="8" pageNumber="259" stateProvince="Khomas">Windhoek</location>
[
<geoCoordinate box="[727,832,495,516]" degrees="22" direction="south" minutes="34" orientation="latitude" pageId="8" pageNumber="259" precision="15" seconds="12" value="-22.570002">22°34'12&quot;S</geoCoordinate>
<geoCoordinate box="[837,944,495,517]" degrees="17" direction="east" minutes="05" orientation="longitude" pageId="8" pageNumber="259" precision="15" seconds="01" value="17.083612">17°05'01&quot;E</geoCoordinate>
],
<date box="[960,1057,495,517]" pageId="8" pageNumber="259" value="1972-04-19">
<collectingDate box="[960,1057,495,517]" pageId="8" pageNumber="259" value="1972-04-19">19.iv.1972</collectingDate>
</date>
,
<collectorName pageId="8" pageNumber="259">P.G. Olivier</collectorName>
, under stone [anterior part crushed] (
<specimenCode box="[544,658,519,541]" collectionCode="SMN" country="China" lsid="urn:lsid:biocol.org:col:14058" name="Simao District National Medical and Pharmaceutical Institute" pageId="8" pageNumber="259" type="Herbarium">SMN 21695</specimenCode>
) (
<collectionCode box="[676,756,519,540]" country="Namibia" httpUri="http://grbio.org/cool/y6s1-e7mm" name="National Museum of Namibia" pageId="8" pageNumber="259">NMNW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
Distribution:
<collectingCountry box="[295,394,556,582]" name="Namibia" pageId="8" pageNumber="259">Namibia</collectingCountry>
, Kuiseb [
<geoCoordinate box="[500,591,556,582]" degrees="23" direction="south" minutes="12" orientation="latitude" pageId="8" pageNumber="259" precision="925" value="-23.2">23°12'S</geoCoordinate>
<geoCoordinate box="[598,690,555,582]" degrees="15" direction="east" minutes="37" orientation="longitude" pageId="8" pageNumber="259" precision="925" value="15.616667">15°37'E</geoCoordinate>
] and Gaub [
<geoCoordinate box="[829,918,555,582]" degrees="23" direction="south" minutes="25" orientation="latitude" pageId="8" pageNumber="259" precision="925" value="-23.416666">23°25'S</geoCoordinate>
<geoCoordinate box="[926,1018,555,581]" degrees="16" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="259" precision="925" value="16.016666">16°01'E</geoCoordinate>
] rivers, as well as Windhoek.
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
Habitat: In
<collectingCountry box="[271,370,628,654]" name="Namibia" pageId="8" pageNumber="259">Namibia</collectingCountry>
, the species is distributed along the ephemeral beds of the Kuiseb and Gaub Rivers. Specimens were located in wet silt along river banks and a distribution throughout the river system therefore seems probable. Windhoek, where the examined specimen was collected, is part of the Swakop River catchment area [
<geoCoordinate box="[903,992,724,750]" degrees="22" direction="south" minutes="16" orientation="latitude" pageId="8" pageNumber="259" precision="925" value="-22.266666">22°16'S</geoCoordinate>
<geoCoordinate box="[998,1091,723,750]" degrees="15" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="259" precision="925" value="15.75">15°45'E</geoCoordinate>
], but only a few kilometres southwest of Windhoek, the Kuiseb River catchment area begins (
<bibRefCitation author="JACOBSON, P. J. &amp; JACOBSON, K. M. &amp; SEELY, M." box="[230,461,787,814]" pageId="8" pageNumber="259" refId="ref25501" refString="JACOBSON, P. J., JACOBSON, K. M. &amp; SEELY, M. 1995. Ephemeral rivers and their catchments. Sustaining people and development in western Namibia. Windhoek: Desert Research Foundation of Namibia." type="book" year="1995">
Jacobson
<emphasis box="[341,389,788,814]" italics="true" pageId="8" pageNumber="259">et al</emphasis>
. 1995
</bibRefCitation>
). The animals can only survive in this dry environment by selecting more humid places like deep silt deposits close to the river banks, from where they are presumed to emerge and aggregate on patches of green cyanobacteria under favourable conditions (
<bibRefCitation author="SHELLEY, R. M. &amp; CRAWFORD, C. S." box="[397,686,884,910]" pageId="8" pageNumber="259" pagination="1 - 8" refId="ref27034" refString="SHELLEY, R. M. &amp; CRAWFORD, C. S. 1996. Cnemodesmus riparius sp. nov., a riparian millipede from the Namib Desert, Africa (Polydesmida: Paradoxosomatidae). Myriapodologica 4 (1): 1 - 8." type="journal article" year="1996">Shelley &amp; Crawford 1996</bibRefCitation>
). Experiments concerning desiccation resistance showed great variability between individuals.
</paragraph>
<paragraph blockId="8.[144,1105,204,1110]" pageId="8" pageNumber="259">
Remarks: The specimen examined is smaller than the measurements given for the
<typeStatus pageId="8" pageNumber="259">holotype</typeStatus>
, which is in the National Museum of Natural History, Smithsonian Institution,
<collectingRegion box="[144,278,1020,1046]" country="United States of America" name="Washington" pageId="8" pageNumber="259">Washington</collectingRegion>
, DC,
<collectingCountry box="[337,391,1020,1046]" name="United States of America" pageId="8" pageNumber="259">USA</collectingCountry>
(
<bibRefCitation author="SHELLEY, R. M. &amp; CRAWFORD, C. S." box="[404,691,1020,1046]" pageId="8" pageNumber="259" pagination="1 - 8" refId="ref27034" refString="SHELLEY, R. M. &amp; CRAWFORD, C. S. 1996. Cnemodesmus riparius sp. nov., a riparian millipede from the Namib Desert, Africa (Polydesmida: Paradoxosomatidae). Myriapodologica 4 (1): 1 - 8." type="journal article" year="1996">Shelley &amp; Crawford 1996</bibRefCitation>
). Moreover, the solenomere is shorter, with the tips of the tibiotarsus more acute in the specimen examined. These differences are small enough to be regarded as falling within the range of variation of a species.
</paragraph>
</subSubSection>
</treatment>
</document>
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