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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FF84FF9B66C9FC53619470E7" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Doratogonus : Attems 1914" docType="treatment" docVersion="2" lastPageNumber="274" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="274" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vohland, Katrin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamer, Michelle</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>251</mods:start>
<mods:end>251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039587C7FF84FF9B66C9FC53619470E7" httpUri="http://treatment.plazi.org/id/039587C7FF84FF9B66C9FC53619470E7" lastPageNumber="274" pageId="23" pageNumber="274">
<subSubSection box="[470,853,976,1002]" pageId="23" pageNumber="274" type="nomenclature">
<paragraph blockId="23.[470,853,976,1002]" box="[470,853,976,1002]" pageId="23" pageNumber="274">
<heading box="[470,853,976,1002]" centered="true" fontSize="36" level="2" pageId="23" pageNumber="274" reason="6">
Genus
<taxonomicName ID-CoL="7PC7H" ID-ENA="241661" authority="Attems, 1914" authorityName=": Attems" authorityYear="1914" box="[549,853,976,1002]" class="Diplopoda" family="Spirostreptidae" genus="Doratogonus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[549,697,976,1001]" italics="true" pageId="23" pageNumber="274">Doratogonus</emphasis>
Attems, 1914
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="274" type="reference_group">
<paragraph blockId="23.[181,1142,1014,1676]" pageId="23" pageNumber="274">
<treatmentCitationGroup pageId="23" pageNumber="274">
<treatmentCitation author="PORAT, C. O." box="[181,419,1014,1035]" page="43" pageId="23" pageNumber="274" year="1872">
<taxonomicName ID-CoL="SJ4" authority="Porat, 1872: 43" authorityName="Porat" authorityPageNumber="43" authorityYear="1872" box="[181,419,1014,1035]" class="Diplopoda" family="Spirostreptidae" genus="Alloporus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[181,270,1014,1035]" italics="true" pageId="23" pageNumber="274">Alloporus</emphasis>
<bibRefCitation author="PORAT, C. O." box="[277,419,1014,1035]" pageId="23" pageNumber="274" pagination="3 - 45" refId="ref26674" refString="PORAT, C. O. 1872. Myriapoda Africa Australis, in Museo Regio Holmiensi asservata, Pars II. Diplopoda. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar 29 (5): 3 - 45." type="journal article" year="1872">Porat, 1872: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; 1876: 45;
<treatmentCitation author="SAUSSURE, H. DE &amp; ZEHNTER, L." box="[521,801,1014,1036]" page="175" pageId="23" pageNumber="274" year="1902">
<bibRefCitation author="SAUSSURE, H. DE &amp; ZEHNTER, L." box="[521,801,1014,1036]" pageId="23" pageNumber="274" pagination="1 - 356" refId="ref26826" refString="SAUSSURE, H. DE &amp; ZEHNTER, L. 1902. Histoire naturelle des myriapodes. In: Grandidier, A., ed., Histoire physique, naturelle et politique de Madagascar. Paris: Imprimerie Nationale, pp. 1 - 356." type="book chapter" year="1902">Saussure &amp; Zehnter 1902: 175</bibRefCitation>
</treatmentCitation>
; Attems 1928: 346; 1950: 204; Schubart 1966: 90;
<treatmentCitation author="KRABBE, E." box="[410,576,1038,1060]" page="113" pageId="23" pageNumber="274" year="1982">
<bibRefCitation author="KRABBE, E." box="[410,576,1038,1060]" pageId="23" pageNumber="274" pagination="1 - 476" refId="ref25792" refString="KRABBE, E. 1982. Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg 24: 1 - 476." type="journal article" year="1982">Krabbe 1982: 113</bibRefCitation>
</treatmentCitation>
. Synonymised by Hoffman (1987).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="23.[181,1142,1014,1676]" pageId="23" pageNumber="274">
<taxonomicName authority=": Attems 1914 a: 109" authorityName=": Attems" authorityPageNumber="109" authorityYear="1914" box="[181,489,1062,1084]" class="Diplopoda" family="Spirostreptidae" genus="Doratogonus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[181,301,1063,1084]" italics="true" pageId="23" pageNumber="274">Doratogonus</emphasis>
: Attems 1914
<emphasis box="[429,440,1063,1084]" italics="true" pageId="23" pageNumber="274">a</emphasis>
: 109
</taxonomicName>
; 1928: 339; 1934
<emphasis box="[647,658,1062,1083]" italics="true" pageId="23" pageNumber="274">b</emphasis>
: 483; Schubart 1966: 90;
<bibRefCitation author="DEMANGE, J. - M." box="[895,1082,1062,1083]" pageId="23" pageNumber="274" pagination="366 - 411" refId="ref24551" refString="DEMANGE, J. - M. 1970. Elements d'une revivision des Spirostreptidae. I. Etude de quelques caracteres taxonomiques des Spirostreptinae. Bulletin de l'Institut Fondamental d'Afrique Noire A 32: 366 - 411." type="journal article" year="1970">Demange 1970: 401</bibRefCitation>
; Hoffman 1987: 13; Hamer 1998: 47; 1999: 10; 2000: 2;
<bibRefCitation author="HAMER, M. L. &amp; SLOTOW, R." box="[746,1001,1086,1108]" pageId="23" pageNumber="274" pagination="295 - 311" refId="ref24906" refString="HAMER, M. L. &amp; SLOTOW, R. 2000. Patterns of distribution and speciation in the genus Doratogonus (Diplopoda: Spirostreptidae). Fragmenta Faunistica, Supplement 43: 295 - 311." type="journal article" year="2000">Hamer &amp; Slotow 2000: 295</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="274" type="materials_examined">
<paragraph blockId="23.[181,1142,1014,1676]" box="[181,727,1122,1149]" pageId="23" pageNumber="274">
<typeStatus box="[181,237,1123,1149]" pageId="23" pageNumber="274">Type</typeStatus>
species:
<taxonomicName authority="Voges, 1878" authorityName="Voges" authorityYear="1878" box="[340,722,1122,1149]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="setosus">
<emphasis box="[340,575,1123,1149]" italics="true" pageId="23" pageNumber="274">Spirostreptus setosus</emphasis>
Voges, 1878
</taxonomicName>
.
</paragraph>
<paragraph blockId="23.[181,1142,1014,1676]" pageId="23" pageNumber="274">
Diagnosis: Mandible with 10–12 pectinate lamellae. Collum projects anteriorly (
<figureCitation captionStart="Figs47–49" captionStartId="24.[144,183,1606,1628]" captionTargetBox="[183,1074,692,1585]" captionTargetId="figure-189@24.[145,1105,686,1590]" captionTargetPageId="24" captionText="Figs47–49.Doratogonus rugifronsAttems, ♂: (47) oral viewof gonopod; (48)lateral view of gonopod; (49) head and first segments, lateral view.Abbreviations: (col) collum, (fp) femoral process, (ll) lateral lamella, (ml) median lamella, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="23" pageNumber="274">Fig. 49</figureCitation>
). Lateral lamella of telocoxite with a small spine-like projection on lateral edge, median lamella typically setose and extended distally as a slender, spinous projection (
<figureCitation captionStart="Figs47–49" captionStartId="24.[144,183,1606,1628]" captionTargetBox="[183,1074,692,1585]" captionTargetId="figure-189@24.[145,1105,686,1590]" captionTargetPageId="24" captionText="Figs47–49.Doratogonus rugifronsAttems, ♂: (47) oral viewof gonopod; (48)lateral view of gonopod; (49) head and first segments, lateral view.Abbreviations: (col) collum, (fp) femoral process, (ll) lateral lamella, (ml) median lamella, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="23" pageNumber="274">Fig. 47</figureCitation>
). Telopodite with long, slender shaft, curved proximally, with long, strongly curved femoral process, and with torsion of main branch of telopodite at position of origin of femoral process, distal to which main branch is looped (
<figureCitation box="[809,939,1322,1348]" captionStart="Figs47–49" captionStartId="24.[144,183,1606,1628]" captionTargetBox="[183,1074,692,1585]" captionTargetId="figure-189@24.[145,1105,686,1590]" captionTargetPageId="24" captionText="Figs47–49.Doratogonus rugifronsAttems, ♂: (47) oral viewof gonopod; (48)lateral view of gonopod; (49) head and first segments, lateral view.Abbreviations: (col) collum, (fp) femoral process, (ll) lateral lamella, (ml) median lamella, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="23" pageNumber="274">Figs 47, 48</figureCitation>
). Solenomere distally forked or apically bifid (
<figureCitation box="[511,595,1354,1380]" captionStart="Figs47–49" captionStartId="24.[144,183,1606,1628]" captionTargetBox="[183,1074,692,1585]" captionTargetId="figure-189@24.[145,1105,686,1590]" captionTargetPageId="24" captionText="Figs47–49.Doratogonus rugifronsAttems, ♂: (47) oral viewof gonopod; (48)lateral view of gonopod; (49) head and first segments, lateral view.Abbreviations: (col) collum, (fp) femoral process, (ll) lateral lamella, (ml) median lamella, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="23" pageNumber="274">Fig. 47</figureCitation>
).
</paragraph>
<paragraph blockId="23.[181,1142,1014,1676]" pageId="23" pageNumber="274">
Remarks: Attems (1914) erected
<taxonomicName authorityName=": Attems" authorityYear="1914" box="[558,706,1395,1420]" class="Diplopoda" family="Spirostreptidae" genus="Doratogonus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[558,706,1395,1420]" italics="true" pageId="23" pageNumber="274">Doratogonus</emphasis>
</taxonomicName>
as a new genus, but its differentiation from
<taxonomicName authority="Porat, 1876" authorityName="Porat" authorityYear="1876" box="[242,491,1426,1453]" class="Diplopoda" family="Spirostreptidae" genus="Alloporus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[242,352,1426,1452]" italics="true" pageId="23" pageNumber="274">Alloporus</emphasis>
Porat, 1876
</taxonomicName>
, remained unclear. The two genera were separated only by the position of the first ozopore.
<bibRefCitation author="KRABBE, E." box="[550,715,1458,1484]" pageId="23" pageNumber="274" pagination="1 - 476" refId="ref25792" refString="KRABBE, E. 1982. Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg 24: 1 - 476." type="journal article" year="1982">Krabbe (1982)</bibRefCitation>
combined the generic concepts under
<taxonomicName authorityName="Porat" authorityYear="1876" box="[181,291,1490,1516]" class="Diplopoda" family="Spirostreptidae" genus="Alloporus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[181,291,1490,1516]" italics="true" pageId="23" pageNumber="274">Alloporus</emphasis>
</taxonomicName>
, but Hoffman (1987) revived the name
<taxonomicName authorityName=": Attems" authorityYear="1914" box="[743,891,1491,1516]" class="Diplopoda" family="Spirostreptidae" genus="Doratogonus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[743,891,1491,1516]" italics="true" pageId="23" pageNumber="274">Doratogonus</emphasis>
</taxonomicName>
for medium- to largesized spirostreptids which differ from
<taxonomicName authorityName="Porat" authorityYear="1876" box="[610,720,1522,1548]" class="Diplopoda" family="Spirostreptidae" genus="Alloporus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[610,720,1522,1548]" italics="true" pageId="23" pageNumber="274">Alloporus</emphasis>
</taxonomicName>
in having more pectinate lamellae on their mandibles and more numerous small setae on the distal half of the stipes. The genus
<taxonomicName authorityName="Porat" authorityYear="1876" box="[181,291,1586,1612]" class="Diplopoda" family="Spirostreptidae" genus="Alloporus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[181,291,1586,1612]" italics="true" pageId="23" pageNumber="274">Alloporus</emphasis>
</taxonomicName>
was placed on the list of genera
<emphasis box="[661,816,1586,1612]" italics="true" pageId="23" pageNumber="274">incertae sedis</emphasis>
. Hoffman (1987) considered
<taxonomicName authorityName=": Attems" authorityYear="1914" box="[181,329,1619,1644]" class="Diplopoda" family="Spirostreptidae" genus="Doratogonus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis box="[181,329,1619,1644]" italics="true" pageId="23" pageNumber="274">Doratogonus</emphasis>
</taxonomicName>
to be heterogeneous and provided a diagnosis, which he believed was too inclusive (
<bibRefCitation author="HAMER, M. L. &amp; SLOTOW, R." box="[297,443,1650,1676]" pageId="23" pageNumber="274" pagination="295 - 311" refId="ref24906" refString="HAMER, M. L. &amp; SLOTOW, R. 2000. Patterns of distribution and speciation in the genus Doratogonus (Diplopoda: Spirostreptidae). Fragmenta Faunistica, Supplement 43: 295 - 311." type="journal article" year="2000">Hamer 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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