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<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583433851" checkinUser="felipe" docAuthor="Vohland, Katrin &amp; Hamer, Michelle" docDate="2013" docId="039587C7FF80FF9866FCFB7C61627348" docLanguage="en" docName="AfrInvertebr.54.1.251-304.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Brevitibius : Attems 1950" docType="treatment" docVersion="2" lastPageNumber="271" masterDocId="FFACFFBFFF93FF8C671FFF836053766B" masterDocTitle="A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species" masterLastPageNumber="251" masterPageNumber="251" pageNumber="270" updateTime="1683642714112" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vohland, Katrin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamer, Michelle</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>251</mods:start>
<mods:end>251</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:039587C7FF80FF9866FCFB7C61627348" httpUri="http://treatment.plazi.org/id/039587C7FF80FF9866FCFB7C61627348" lastPageId="20" lastPageNumber="271" pageId="19" pageNumber="270">
<subSubSection box="[483,840,1278,1305]" pageId="19" pageNumber="270" type="nomenclature">
<paragraph blockId="19.[483,840,1278,1305]" box="[483,840,1278,1305]" pageId="19" pageNumber="270">
<heading box="[483,840,1278,1305]" centered="true" fontSize="36" level="2" pageId="19" pageNumber="270" reason="6">
Genus
<taxonomicName ID-CoL="7NTYW" authority="Attems, 1950" authorityName=": Attems" authorityYear="1950" box="[562,840,1278,1305]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis box="[562,684,1278,1304]" italics="true" pageId="19" pageNumber="270">Brevitibius</emphasis>
Attems, 1950
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[181,1060,1316,1339]" pageId="19" pageNumber="270" type="reference_group">
<paragraph blockId="19.[181,1142,1316,1676]" box="[181,1060,1316,1339]" pageId="19" pageNumber="270">
<treatmentCitationGroup box="[181,1060,1316,1339]" pageId="19" pageNumber="270">
<taxonomicName ID-CoL="7NTYW" authority=": Attems 1950: 249" authorityName=": Attems" authorityPageNumber="249" authorityYear="1950" box="[181,455,1316,1338]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis box="[181,280,1317,1338]" italics="true" pageId="19" pageNumber="270">Brevitibius</emphasis>
: Attems 1950: 249
</taxonomicName>
, fig. 93;
<treatmentCitation author="DEMANGE, J. - M." box="[535,721,1317,1338]" page="394" pageId="19" pageNumber="270" year="1970">
<bibRefCitation author="DEMANGE, J. - M." box="[535,721,1317,1338]" pageId="19" pageNumber="270" pagination="366 - 411" refId="ref24551" refString="DEMANGE, J. - M. 1970. Elements d'une revivision des Spirostreptidae. I. Etude de quelques caracteres taxonomiques des Spirostreptinae. Bulletin de l'Institut Fondamental d'Afrique Noire A 32: 366 - 411." type="journal article" year="1970">Demange 1970: 394</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KRABBE, E." box="[730,896,1317,1339]" page="131" pageId="19" pageNumber="270" year="1982">
<bibRefCitation author="KRABBE, E." box="[730,896,1317,1339]" pageId="19" pageNumber="270" pagination="1 - 476" refId="ref25792" refString="KRABBE, E. 1982. Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg 24: 1 - 476." type="journal article" year="1982">Krabbe 1982: 131</bibRefCitation>
</treatmentCitation>
; Hamer 1999: 10.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="271" pageId="19" pageNumber="270" type="materials_examined">
<paragraph blockId="19.[181,1142,1316,1676]" box="[181,1047,1353,1379]" pageId="19" pageNumber="270">
<typeStatus box="[181,237,1353,1379]" pageId="19" pageNumber="270">Type</typeStatus>
species:
<taxonomicName authority="Attems, 1934" authorityName="Attems" authorityYear="1934" box="[340,770,1353,1379]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="angolanus">
<emphasis box="[340,611,1353,1379]" italics="true" pageId="19" pageNumber="270">Spirostreptus angolanus</emphasis>
Attems, 1934
</taxonomicName>
, by original designation.
</paragraph>
<paragraph blockId="19.[181,1142,1316,1676]" pageId="19" pageNumber="270">
Diagnosis: Ozopores from the 6
<superScript attach="left" box="[549,562,1393,1407]" fontSize="6" pageId="19" pageNumber="270">th</superScript>
segment onwards. Prozonites with punctures, metazonites with distinct striae below ozopores. Limbus smooth. Sternites striated. Legs with much reduced pads (
<figureCitation box="[469,552,1457,1483]" captionStart="Figs 3843" captionStartId="21.[181,220,1534,1556]" captionTargetBox="[216,1095,340,1509]" captionTargetId="figure-135@21.[200,1119,331,1518]" captionTargetPageId="21" captionText="Figs 3843. Brevitibius oongongololo sp.n., ♂: (38) aboral aspect of gonopod; (39) oral aspect of gonopod; (40) mid­body leg; (41) gonopod aperture; (42) gnathochilarium; (43) head and first segments (ventral aspect).Abbreviations: (c) cardo, (col) collum, (cx) coxa, (f) femur, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of telocoxite, (m) mentum, (ml) median lamella of telocoxite, (pb) prebasilare, (prf) prefemur, (ptf) postfemur, (st) stipes, (ta) tarsus, (ti) tibia, (tp) tibial process. Scale bars = 1 mm." pageId="19" pageNumber="270">Fig. 40</figureCitation>
), hypoproct surface porous; with fold(s) and rows of punctures on collum. Gonopod aperture open distally, segments not fused ventrally (
<figureCitation box="[190,273,1521,1547]" captionStart="Figs 3843" captionStartId="21.[181,220,1534,1556]" captionTargetBox="[216,1095,340,1509]" captionTargetId="figure-135@21.[200,1119,331,1518]" captionTargetPageId="21" captionText="Figs 3843. Brevitibius oongongololo sp.n., ♂: (38) aboral aspect of gonopod; (39) oral aspect of gonopod; (40) mid­body leg; (41) gonopod aperture; (42) gnathochilarium; (43) head and first segments (ventral aspect).Abbreviations: (c) cardo, (col) collum, (cx) coxa, (f) femur, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of telocoxite, (m) mentum, (ml) median lamella of telocoxite, (pb) prebasilare, (prf) prefemur, (ptf) postfemur, (st) stipes, (ta) tarsus, (ti) tibia, (tp) tibial process. Scale bars = 1 mm." pageId="19" pageNumber="270">Fig. 41</figureCitation>
).
</paragraph>
<paragraph blockId="19.[181,1142,1316,1676]" pageId="19" pageNumber="270">
Gonopod telopodite with tibial process originating distad of femoral knee (
<figureCitation captionStart="Figs 3843" captionStartId="21.[181,220,1534,1556]" captionTargetBox="[216,1095,340,1509]" captionTargetId="figure-135@21.[200,1119,331,1518]" captionTargetPageId="21" captionText="Figs 3843. Brevitibius oongongololo sp.n., ♂: (38) aboral aspect of gonopod; (39) oral aspect of gonopod; (40) mid­body leg; (41) gonopod aperture; (42) gnathochilarium; (43) head and first segments (ventral aspect).Abbreviations: (c) cardo, (col) collum, (cx) coxa, (f) femur, (h) hypostomum, (l) lamella lingualis, (ll) lateral lamella of telocoxite, (m) mentum, (ml) median lamella of telocoxite, (pb) prebasilare, (prf) prefemur, (ptf) postfemur, (st) stipes, (ta) tarsus, (ti) tibia, (tp) tibial process. Scale bars = 1 mm." pageId="19" pageNumber="270">Figs 38, 39</figureCitation>
,
<figureCitation box="[223,254,1585,1611]" captionStart="Figs 4446" captionStartId="22.[144,183,1582,1604]" captionTargetBox="[157,1089,810,1558]" captionTargetId="figure-190@22.[144,1104,796,1569]" captionTargetPageId="22" captionText="Figs 4446. Brevitibius ondundu sp. n., ♂: (44) aboral aspect of gonopod; (45) oral aspect of gonopod; (46) tip of solenomere in detail. Abbreviations: (lc) lateral conus on median lamella, (ll) lateral lamella of gonocoxite, (ml) median lamella of gonocoxite, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="19" pageNumber="270">46</figureCitation>
,
<emphasis box="[265,287,1586,1611]" italics="true" pageId="19" pageNumber="270">tp</emphasis>
), its tip covered in fold of tibiotarsus, which is wrinkled at the outer side; tip of solenomere acute, with two small folds before end of seminal duct; lateral lamella with lateral conus (
<figureCitation box="[395,524,1649,1676]" captionStart="Figs 4446" captionStartId="22.[144,183,1582,1604]" captionTargetBox="[157,1089,810,1558]" captionTargetId="figure-190@22.[144,1104,796,1569]" captionTargetPageId="22" captionText="Figs 4446. Brevitibius ondundu sp. n., ♂: (44) aboral aspect of gonopod; (45) oral aspect of gonopod; (46) tip of solenomere in detail. Abbreviations: (lc) lateral conus on median lamella, (ll) lateral lamella of gonocoxite, (ml) median lamella of gonocoxite, (s) solenomere, (tp) tibial process. Scale bar = 1 mm." pageId="19" pageNumber="270">Figs 44, 45</figureCitation>
,
<emphasis box="[536,556,1649,1675]" italics="true" pageId="19" pageNumber="270">lc</emphasis>
).
</paragraph>
<caption box="[284,965,1032,1054]" pageId="20" pageNumber="271" startId="20.[284,317,1032,1054]" targetBox="[173,1074,226,1012]" targetPageId="20" targetType="figure">
<paragraph blockId="20.[284,965,1032,1054]" box="[284,965,1032,1054]" pageId="20" pageNumber="271">
Fig. 37. Distribution of the family
<taxonomicName authorityName="Brandt" authorityYear="1833" box="[597,734,1032,1054]" class="Diplopoda" family="Spirostreptidae" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
(
<taxonomicName authorityName="de Blainville in Gervais" authorityYear="1844" box="[746,845,1032,1054]" class="Diplopoda" kingdom="Animalia" pageId="20" pageNumber="271" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
) in Namibia.
</paragraph>
</caption>
<paragraph blockId="20.[144,1104,1096,1315]" pageId="20" pageNumber="271">
Remarks:
<taxonomicName authorityName=": Attems" authorityYear="1950" box="[258,380,1096,1122]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis box="[258,380,1096,1122]" italics="true" pageId="20" pageNumber="271">Brevitibius</emphasis>
</taxonomicName>
is a small genus, containing only two species:
<taxonomicName authority="(Attems, 1934)" baseAuthorityName="Attems" baseAuthorityYear="1934" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="angolanus">
<emphasis box="[903,1051,1096,1122]" italics="true" pageId="20" pageNumber="271">B. angolanus</emphasis>
(Attems, 1934)
</taxonomicName>
and
<taxonomicName authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[329,592,1128,1155]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis box="[329,446,1128,1154]" italics="true" pageId="20" pageNumber="271">B. obtusus</emphasis>
Kraus, 1958
</taxonomicName>
, both known from
<collectingCountry box="[801,887,1128,1154]" name="Angola" pageId="20" pageNumber="271">Angola</collectingCountry>
.
<bibRefCitation author="KRABBE, E." box="[897,1061,1128,1155]" pageId="20" pageNumber="271" pagination="1 - 476" refId="ref25792" refString="KRABBE, E. 1982. Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg 24: 1 - 476." type="journal article" year="1982">Krabbe (1982)</bibRefCitation>
described a species from
<collectingCountry box="[400,504,1160,1186]" name="Tanzania" pageId="20" pageNumber="271">Tanzania</collectingCountry>
, which she tentatively placed in this genus as
<taxonomicName authorityName=": Attems" authorityYear="1950" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="271">Brevitibius</emphasis>
</taxonomicName>
(?)
<emphasis box="[235,363,1192,1218]" italics="true" pageId="20" pageNumber="271">bulbiferans</emphasis>
.
</paragraph>
<paragraph blockId="20.[144,1104,1096,1315]" pageId="20" pageNumber="271">
Attems (1950) erected the genus based on the abruptly shortened tibial part of the telopodite, the anteriorly directed gonocoel and the large lateral conus (which is missing in
<taxonomicName authorityName="Kraus" authorityYear="1958" box="[172,290,1288,1314]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis box="[172,290,1288,1314]" italics="true" pageId="20" pageNumber="271">B. obtusus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>