treatments-xml/data/03/95/87/039587C09E28F745FF6D6560FC381AEE.xml

<document id="C3B45A0160ACF71EE0A78F4337635FC2" ID-CLB-Dataset="46332" ID-DOI="10.11646/zootaxa.3768.5.4" ID-GBIF-Dataset="10b84292-5b52-413c-95b1-386da3a568ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="229083" ID-ZooBank="38F94125-5B01-46B4-BFAA-71A0D9BCFE73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460390182532" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2014" docId="039587C09E28F745FF6D6560FC381AEE" docLanguage="en" docName="zt03768p582.pdf" docOrigin="Zootaxa 3768 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malayopotamon weh Ng, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="582" masterDocId="FFACFFB89E28F743FFFA633FF9691901" masterDocTitle="A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae), a freshwater crab from northern Sumatra, Indonesia" masterLastPageNumber="582" masterPageNumber="576" pageNumber="576" updateTime="1698298122240" updateUser="ExternalLinkService">
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<mods:title id="4D38EA1202E73F225626289049E938EC">A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae), a freshwater crab from northern Sumatra, Indonesia</mods:title>
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<mods:namePart id="828ECE9559328207A96855D230CC9EC8">Ng, Peter K. L.</mods:namePart>
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<mods:date id="F8D1A62B657239B379B2D231445D6770">2014</mods:date>
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<treatment id="039587C09E28F745FF6D6560FC381AEE" ID-DOI="http://doi.org/10.5281/zenodo.6144872" ID-GBIF-Taxon="119416552" ID-Zenodo-Dep="6144872" LSID="urn:lsid:plazi:treatment:039587C09E28F745FF6D6560FC381AEE" httpUri="http://treatment.plazi.org/id/039587C09E28F745FF6D6560FC381AEE" lastPageId="6" lastPageNumber="582" pageId="0" pageNumber="576">
<subSubSection id="C326655D9E28F743FF6D6560F87B1F9D" pageId="0" pageNumber="576" type="nomenclature">
<paragraph id="8B8336D69E28F743FF6D6560F8801F79" blockId="0.[151,489,1631,1693]" box="[151,489,1631,1656]" pageId="0" pageNumber="576">
<heading id="D0CB81BA9E28F743FF6D6560F8801F79" bold="true" box="[151,489,1631,1656]" fontSize="10" level="2" pageId="0" pageNumber="576" reason="2">
<emphasis id="B948EAC49E28F743FF6D6560F8801F79" bold="true" box="[151,489,1631,1656]" pageId="0" pageNumber="576">
<taxonomicName id="4C3C4D559E28F743FF6D6560F8E11F79" ID-CoL="734SJ" box="[151,392,1631,1656]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="576" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E28F743FF6D6560F8E11F79" bold="true" box="[151,392,1631,1656]" italics="true" pageId="0" pageNumber="576">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E28F743FE756560F8801F79" box="[399,489,1631,1656]" pageId="0" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8336D69E28F743FF6D65BBF87B1F9D" blockId="0.[151,489,1631,1693]" box="[151,274,1668,1693]" pageId="0" pageNumber="576">
(
<figureCitation id="13072A539E28F743FF6565BBF8601F9D" box="[159,265,1668,1693]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1845,1867]" captionStartId-1="3.[151,250,1945,1967]" captionStartId-2="4.[151,250,1901,1923]" captionStartId-3="5.[151,250,1227,1249]" captionTargetBox-0="[151,1436,517,1823]" captionTargetBox-1="[151,1436,193,1923]" captionTargetBox-2="[151,1436,193,1878]" captionTargetBox-3="[174,1423,203,1214]" captionTargetId-0="figure@2.[151,1436,517,1824]" captionTargetId-1="figure@3.[151,1436,193,1924]" captionTargetId-2="figure@4.[151,1436,193,1879]" captionTargetId-3="figure@5.[151,1436,193,1224]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. Colours in life (photographs: Daniel Ng)." captionText-1="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." captionText-2="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." captionText-3="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri-0="https://zenodo.org/record/229084/files/figure.png" httpUri-1="https://zenodo.org/record/229085/files/figure.png" httpUri-2="https://zenodo.org/record/229086/files/figure.png" httpUri-3="https://zenodo.org/record/229087/files/figure.png" pageId="0" pageNumber="576">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326655D9E28F743FF6D65F3FA021E09" pageId="0" pageNumber="576" type="materials_examined">
<paragraph id="8B8336D69E28F743FF6D65F3FA021E09" blockId="0.[151,1436,1740,2017]" pageId="0" pageNumber="576">
<emphasis id="B948EAC49E28F743FF6D65F3F8171FE4" bold="true" box="[151,382,1740,1765]" pageId="0" pageNumber="576">Material examined</emphasis>
. 
<typeStatus id="548788749E28F743FE6B65F3F8971FE4" box="[401,510,1740,1765]" pageId="0" pageNumber="576" type="holotype">Holotype</typeStatus>
: male (15.5 × 
<quantity id="4CC49B339E28F743FD4A65F3FA731FE5" box="[688,794,1740,1764]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="0" pageNumber="576" unit="mm" value="12.1">12.1 mm</quantity>
) (
<collectionCode id="ED2DAE139E28F743FCCC65F3FA111FE5" box="[822,888,1740,1764]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="576">MZB</collectionCode>
), waterfall, 
<collectingCountry id="F32B76469E28F743FBFC65F3FD2F1FE4" box="[1030,1094,1740,1765]" name="Iceland" pageId="0" pageNumber="576">Pulau</collectingCountry>
<taxonomicName id="4C3C4D559E28F743FBAB65F3FDEE1FE4" box="[1105,1159,1740,1765]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
, 
<geoCoordinate id="EE0850119E28F743FB6D65F3FD8E1FE4" box="[1175,1255,1740,1765]" direction="north" orientation="latitude" pageId="0" pageNumber="576" precision="555" value="5.82">5.82°N</geoCoordinate>
<geoCoordinate id="EE0850119E28F743FB0965F3FC381FE4" box="[1267,1361,1740,1765]" direction="east" orientation="longitude" pageId="0" pageNumber="576" precision="555" value="95.28">95.28°E</geoCoordinate>
, Aceh Province, northern Sumatra, 
<collectingCountry id="F32B76469E28F743FE2065D0FB231E09" box="[474,586,1775,1800]" name="Indonesia" pageId="0" pageNumber="576">Indonesia</collectingCountry>
, coll. locals, 
<date id="FF8210169E28F743FD2165D0FA0E1E09" box="[731,871,1775,1800]" pageId="0" pageNumber="576" value="2013-03">March 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C326655D9E28F742FF3D642BFA791865" lastPageId="1" lastPageNumber="577" pageId="0" pageNumber="576" type="diagnosis">
<paragraph id="8B8336D69E28F742FF3D642BFA791865" blockId="0.[151,1436,1740,2017]" lastBlockId="1.[151,1436,151,2013]" lastPageId="1" lastPageNumber="577" pageId="0" pageNumber="576">
<emphasis id="B948EAC49E28F743FF3D642BF8531E2C" bold="true" box="[199,314,1812,1837]" pageId="0" pageNumber="576">Diagnosis</emphasis>
. Carapace with branchial, gastric regions gently convex, not distinctly inflated or swollen (
<figureCitation id="13072A539E28F743FAC4642BFCE81E2D" box="[1342,1409,1812,1837]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="0" pageNumber="576">Fig. 2</figureCitation>
A, B); frontal region with low granules; anterolateral regions with distinct low striae, most of branchial regions almost smooth; epibranchial regions with very low striae; mesogastric, urogastric, cardiac, intestinal regions almost smooth, orbital regions almost smooth (
<figureCitation id="13072A539E28F743FDAE6440FBFE1E99" box="[596,663,1919,1944]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="0" pageNumber="576">Fig. 2</figureCitation>
A); epigastric cristae not clearly demarcated, area marked by rows of transverse striae; postorbital cristae distinct but not sharp crest, demarcated by series of strong, adjacently arranged striae (
<figureCitation id="13072A539E28F743FF1E64F7F84E1EE1" box="[228,295,1992,2017]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="0" pageNumber="576">Fig. 2</figureCitation>
A, B); external orbital tooth broadly triangular, outer margin gently granulated, twice length of inner margin; separated from epibranchial tooth by small notch; confluent with supraorbital margin (
<figureCitation id="13072A539E29F742FAFD63A8FC3919B1" box="[1287,1360,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B); epibranchial tooth small, triangular, separated from external orbital tooth by small cleft, not contiguous with postorbital cristae (
<figureCitation id="13072A539E29F742FE9563E0F8DB19F9" box="[367,434,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B); male thoracic sternum relatively narrow transversely, sternites 1, 2 separated from sternite 3 by deep, sinuous, almost complete suture (
<figureCitation id="13072A539E29F742FD0A623BFA5D181D" box="[752,820,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
D); G1 subterminal part gently sinuous with proximal outer margin almost straight; terminal segment about half length of subterminal segment, cylindrical, lateral margins gently diverging to subtruncate tip (
<figureCitation id="13072A539E29F742FD726273FBA51865" box="[648,716,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="1" pageNumber="577">Fig. 4</figureCitation>
A–C).
</paragraph>
</subSubSection>
<subSubSection id="C326655D9E29F741FF3D6250F8F61841" lastPageId="2" lastPageNumber="578" pageId="1" pageNumber="577" type="description">
<paragraph id="8B8336D69E29F742FF3D6250FDE71D35" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
<emphasis id="B948EAC49E29F742FF3D6250FB471889" bold="true" box="[199,558,367,392]" pageId="1" pageNumber="577">
Description of male 
<typeStatus id="548788749E29F742FE3D6250FB471889" box="[455,558,367,392]" pageId="1" pageNumber="577" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace quadrate, smooth, branchial, gastric regions gently convex, not distinctly inflated or swollen (
<figureCitation id="13072A539E29F742FE1D62ABFB4318AD" box="[487,554,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Frontal region with low granules; anterolateral regions with distinct low striae, posterolateral regions with low striae, barely visible; rest of branchial regions almost smooth; epibranchial regions with low striae; mesogastric, urogastric, cardiac, intestinal regions almost smooth, epigastric regions with low transverse striae anteriorly, almost smooth posteriorly; orbital regions almost smooth; suborbital, subhepatic, pterygostomial regions with low granules, striae (
<figureCitation id="13072A539E29F742FD3D611BFA651B3D" box="[711,780,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A). Epigastric cristae not clearly demarcated, area marked by rows of transverse striae, medially separated by distinct Y-shaped furrow, not confluent with postorbital cristae; postorbital cristae distinct but not as distinct sharp crest, discernible by series of strong, adjacently arranged striae (
<figureCitation id="13072A539E29F742FF6561B0F98A1BA9" box="[159,227,655,680]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Cervical grooves distinct, relatively broad, joining prominent H-shaped median gastric groove (
<figureCitation id="13072A539E29F742FA9461B0F9CC1BCD" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Frontal margin divided into 2 broad, truncate lobes, separated by shallow concavity; margin of each lobe gently sinuous, separated from supraorbital margin by distinct rounded angle, no lobe discernible (
<figureCitation id="13072A539E29F742FAF461E8FC3D1BF1" box="[1294,1364,727,752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). External orbital tooth broadly triangular, outer margin gently granulated, twice length of inner margin; separated from epibranchial tooth by small notch; confluent with supraorbital margin (
<figureCitation id="13072A539E29F742FC0B6020FD5D1A39" box="[1009,1076,799,824]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Epibranchial tooth small, triangular, distinct, separated from external orbital tooth by small cleft, not contiguous with postorbital cristae, transverse striae (
<figureCitation id="13072A539E29F742FEA66058F8F61A81" box="[348,415,871,896]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Anterolateral margins convex, crested, lined with small sharp granules (
<figureCitation id="13072A539E29F742FAE96058FC3F1A81" box="[1299,1366,871,896]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Posterolateral margin gently concave, converging towards gently concave posterior carapace margin (
<figureCitation id="13072A539E29F742FAE860B3FC3F1AA5" box="[1298,1366,908,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A, B). Orbits subovate; eye filling up most of orbital space; eye short, peduncle stout; cornea large, pigmented (
<figureCitation id="13072A539E29F742FAB26090FCE71AC9" box="[1352,1422,943,968]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
). Supraorbital margin gently concave, entire, lined with round granules (
<figureCitation id="13072A539E29F742FC0B60EBFD571AED" box="[1009,1086,980,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
). Suborbital margin concave, complete, lined with roundgranules (
<figureCitation id="13072A539E29F742FDC160C8FBE91D11" box="[571,640,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
C). Posterior margin of epistome gently sinuous; with distinct acutely triangular, narrow median triangle, lateral margins with small submedian fissure (
<figureCitation id="13072A539E29F742FBD26723FD021D35" box="[1064,1131,1052,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="8B8336D69E29F742FF3D6700FD951DA1" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
Third maxilliped covering most of buccal cavity when closed; ischium subrectangular, with distinct submedian groove which is subparallel to inner margin; merus quadrate, wider than long, surface slightly rugose, anteroexternal angle rounded; exopod relatively slender, reaching to middle of merus, with long flagellum (
<figureCitation id="13072A539E29F742FB6C67B8FDBE1DA1" box="[1174,1239,1159,1184]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8B8336D69E29F742FF3D6793FB221CC1" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
Chelipeds slightly asymmetrical (
<figureCitation id="13072A539E29F742FDBF6793FBE31DC5" box="[581,650,1196,1221]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A). Anterior margin of basis-ischium with prominent sharp granules; outer surface of merus covered with distinct transverse striae, inner margins of merus lined sharp granules, upper margin with progressively larger granules with subdistal margin subspiniform; outer surface of carpus distinctly rugose, granular, inner margins distinctly granular, distal angle with short, sharp spine, small basal spiniform tubercle (
<figureCitation id="13072A539E29F742FEFE6603F8231C55" box="[260,330,1340,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A). Chelae relatively stout; outer surfaces distinctly rugose, granular, especially on upper parts; fingers gently curved, as long as palm; dorsal margin of dactylus gently serrated, outer surface lined with rows of pits; propodal finger with longitudinal groove lined with pits; cutting edges of both fingers with variously sized blade-like teeth, denticles (
<figureCitation id="13072A539E29F742FE3F6698FB6E1CC1" box="[453,519,1447,1472]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
B, C).
</paragraph>
<paragraph id="8B8336D69E29F742FF3D66F3F86F1F99" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
Ambulatory legs relatively short, second pair longest, last pair shortest (
<figureCitation id="13072A539E29F742FBED66F3FD361CE5" box="[1047,1119,1484,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A). Outer surface of merus rugose, dorsal margin uneven, appears gently serrated, subdistal angle distinct but not spiniform; carpus rugose, with dorsal margin cristae, outer surface with distinct submedian crista; outer surface of propodus rugose with low submedian longitudinal ridge, dorsal margin uneven, ventral margin distinctly serrated; dactylus gently curved (almost straight on last ambulatory leg), quadrate in cross section, margins with short, sharp spines, tip corneous (
<figureCitation id="13072A539E29F742FF656540F98A1F99" box="[159,227,1663,1688]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="1" pageNumber="577">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8B8336D69E29F742FF3D659BF8C61E71" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
Thoracic sternum relatively narrow transversely, surface pitted with lateral parts partially setose, but otherwise smooth, glabrous (
<figureCitation id="13072A539E29F742FE9265F8F8C21FE1" box="[360,427,1735,1760]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
D). Sternites 1, 2 completely fused to form distinct triangular plate with distinctly convex lateral margins; separated from sternite 3 by deep, sinuous, almost complete suture but lateralmost part distinctly shallow, appears interrupted; sternites 3, 4 completely fused; sternoabdominal cavity reaching to imaginary line connecting anterior edge of coxae of chelipeds (
<figureCitation id="13072A539E29F742FD28640BFA701E4D" box="[722,793,1844,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
D). Male abdominal locking tubercle low, rounded, on median part of sternite 5.
</paragraph>
<paragraph id="8B8336D69E29F742FF3D6443FD8D1EDD" blockId="1.[151,1436,151,2013]" pageId="1" pageNumber="577">
Abdomen triangular, all somites, telson free; somites 3–6 trapezoidal, gradually decreasing in width; somite 6 with length greater than half width, lateral margins gently convex; somites 1, 2 subrectangular, wide, reaching to bases of coxae of fourth ambulatory legs, sternite 8 not visible when abdomen closed (
<figureCitation id="13072A539E29F742FBA564FBFDC81EDD" box="[1119,1185,1988,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="1" pageNumber="577">Fig. 3</figureCitation>
D–F).
</paragraph>
<paragraph id="8B8336D69E2AF741FF3D63A7FDB0181D" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="578">
G1 relatively stout; subterminal part gently sinuous with proximal outer margin almost straight, lined with long setae; terminal segment about half length of subterminal segment, cylindrical, lateral margins gently diverging to subtruncate tip, margin of fold on ventral margin, lateral margins lined with short, simple setae (
<figureCitation id="13072A539E2AF741FB1D63E0FC4419F9" box="[1255,1325,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="2" pageNumber="578">Fig. 4</figureCitation>
A–C). G2 long, subequal to length of G1; distal segment, about one third length of basal segment (
<figureCitation id="13072A539E2AF741FB88623BFDDE181D" box="[1138,1207,260,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="2" pageNumber="578">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="8B8336D69E2AF741FF3D6218F8F61841" blockId="2.[151,1436,151,464]" box="[199,415,295,320]" pageId="2" pageNumber="578">Female not known.</paragraph>
</subSubSection>
<subSubSection id="C326655D9E2AF741FF3D6273FA0B18D1" pageId="2" pageNumber="578" type="etymology">
<paragraph id="8B8336D69E2AF741FF3D6273F87A1889" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="578">
<emphasis id="B948EAC49E2AF741FF3D6273F82F1864" bold="true" box="[199,326,332,357]" pageId="2" pageNumber="578">Etymology</emphasis>
. The species is named after the island where it was collected. The name is used as a noun in apposition.
</paragraph>
<paragraph id="8B8336D69E2AF741FF3D62ABFA0B18D1" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="578">
<emphasis id="B948EAC49E2AF741FF3D62ABF87318AC" bold="true" box="[199,282,404,429]" pageId="2" pageNumber="578">Colour</emphasis>
. The colour in life was orange-brown overall on dorsal surfaces (
<figureCitation id="13072A539E2AF741FC0462ABFD5418AD" box="[1022,1085,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1845,1867]" captionTargetBox="[151,1436,517,1823]" captionTargetId="figure@2.[151,1436,517,1824]" captionTargetPageId="2" captionText="FIGURE 1. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. Colours in life (photographs: Daniel Ng)." httpUri="https://zenodo.org/record/229084/files/figure.png" pageId="2" pageNumber="578">Fig. 1</figureCitation>
A), with the chelae orange and the third maxillipeds and ventral surfaces pale yellow (
<figureCitation id="13072A539E2AF741FD076288FA5418D1" box="[765,829,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1845,1867]" captionTargetBox="[151,1436,517,1823]" captionTargetId="figure@2.[151,1436,517,1824]" captionTargetPageId="2" captionText="FIGURE 1. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. Colours in life (photographs: Daniel Ng)." httpUri="https://zenodo.org/record/229084/files/figure.png" pageId="2" pageNumber="578">Fig. 1</figureCitation>
B).
</paragraph>
</subSubSection>
<caption id="DF43665E9E2AF741FF6D640AF8CB1E6B" httpUri="https://zenodo.org/record/229084/files/figure.png" pageId="2" pageNumber="578" targetBox="[151,1436,517,1823]" targetPageId="2">
<paragraph id="8B8336D69E2AF741FF6D640AF8CB1E6B" blockId="2.[151,1435,1845,1898]" pageId="2" pageNumber="578">
<emphasis id="B948EAC49E2AF741FF6D640AF8661E4B" bold="true" box="[151,271,1845,1867]" pageId="2" pageNumber="578">FIGURE 1</emphasis>
. 
<taxonomicName id="4C3C4D559E2AF741FEE5640AF8911E4A" box="[287,504,1845,1867]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2AF741FEE5640AF8911E4A" box="[287,504,1845,1867]" italics="true" pageId="2" pageNumber="578">Malayopotamon weh</emphasis>
</taxonomicName>
<emphasis id="B948EAC49E2AF741FDFB6409FB3D1E4A" bold="true" box="[513,596,1846,1867]" pageId="2" pageNumber="578">
<taxonomicNameLabel id="A27B57BF9E2AF741FDFB6409FB3D1E4A" box="[513,596,1846,1867]" pageId="2" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (15.5 × 12.1 mm) (MZB), Pulau 
<taxonomicName id="4C3C4D559E2AF741FB9B640AFDF81E4A" box="[1121,1169,1845,1867]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
, Sumatra. Colours in life (photographs: Daniel Ng).
</paragraph>
</caption>
<caption id="DF43665E9E2BF740FF6D64A6FA631ECF" httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="3" pageNumber="579" targetBox="[151,1436,193,1923]" targetPageId="3">
<paragraph id="8B8336D69E2BF740FF6D64A6FA631ECF" blockId="3.[151,1435,1945,1998]" pageId="3" pageNumber="579">
<emphasis id="B948EAC49E2BF740FF6D64A6F8671EAF" bold="true" box="[151,270,1945,1967]" pageId="3" pageNumber="579">FIGURE 2</emphasis>
. 
<taxonomicName id="4C3C4D559E2BF740FEE664A6F89D1EAE" box="[284,500,1945,1967]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="579" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2BF740FEE664A6F89D1EAE" box="[284,500,1945,1967]" italics="true" pageId="3" pageNumber="579">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2BF740FE0664A6FB271EAF" box="[508,590,1945,1966]" pageId="3" pageNumber="579" rank="species">
<emphasis id="B948EAC49E2BF740FE0664A6FB271EAF" bold="true" box="[508,590,1945,1966]" pageId="3" pageNumber="579">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male (15.5 × 12.1 mm) (MZB), Pulau 
<taxonomicName id="4C3C4D559E2BF740FBB564A6FDE91EAE" box="[1103,1152,1945,1967]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace.
</paragraph>
</caption>
<caption id="DF43665E9E2CF747FF6D6452F9961EDE" httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="4" pageNumber="580" targetBox="[151,1436,193,1878]" targetPageId="4">
<paragraph id="8B8336D69E2CF747FF6D6452F9961EDE" blockId="4.[151,1436,1901,2015]" pageId="4" pageNumber="580">
<emphasis id="B948EAC49E2CF747FF6D6452F8781E83" bold="true" box="[151,273,1901,1923]" pageId="4" pageNumber="580">FIGURE 3</emphasis>
. 
<taxonomicName id="4C3C4D559E2CF747FED86452F8951E82" box="[290,508,1901,1923]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="580" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2CF747FED86452F8951E82" box="[290,508,1901,1923]" italics="true" pageId="4" pageNumber="580">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2CF747FDFD6452FB351E83" box="[519,604,1901,1922]" pageId="4" pageNumber="580" rank="species">
<emphasis id="B948EAC49E2CF747FDFD6452FB351E83" bold="true" box="[519,604,1901,1922]" pageId="4" pageNumber="580">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male (15.5 × 12.1 mm) (MZB), Pulau 
<taxonomicName id="4C3C4D559E2CF747FB8C6452FDCE1E82" box="[1142,1191,1901,1923]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace).
</paragraph>
</caption>
<caption id="DF43665E9E2DF746FF6D67F4F8001C1E" httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="5" pageNumber="581" targetBox="[174,1423,203,1214]" targetPageId="5">
<paragraph id="8B8336D69E2DF746FF6D67F4F8001C1E" blockId="5.[151,1436,1227,1311]" pageId="5" pageNumber="581">
<emphasis id="B948EAC49E2DF746FF6D67F4F8641DE1" bold="true" box="[151,269,1227,1249]" pageId="5" pageNumber="581">FIGURE 4</emphasis>
. 
<taxonomicName id="4C3C4D559E2DF746FEE167F4F8981DE0" box="[283,497,1227,1249]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2DF746FEE167F4F8981DE0" box="[283,497,1227,1249]" italics="true" pageId="5" pageNumber="581">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2DF746FE0267F3FB201DE0" box="[504,585,1228,1249]" pageId="5" pageNumber="581" rank="species">
<emphasis id="B948EAC49E2DF746FE0267F3FB201DE0" bold="true" box="[504,585,1228,1249]" pageId="5" pageNumber="581">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male (15.5 × 12.1 mm) (MZB), Pulau 
<taxonomicName id="4C3C4D559E2DF746FBB867F4FD1A1DE0" box="[1090,1139,1227,1249]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
, Sumatra. A, left G1, ventral view; B, terminal segment of left G1, ventral view; C, terminal segment of left G1, dorsal view; D, left G2. Scales: A, D = 1.0 mm; B, C = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C326655D9E2DF745FF3D6674FC381AEE" lastPageId="6" lastPageNumber="582" pageId="5" pageNumber="581" type="discussion">
<paragraph id="8B8336D69E2DF746FF3D6674FB821FA9" blockId="5.[151,1437,1355,2028]" pageId="5" pageNumber="581">
<emphasis id="B948EAC49E2DF746FF3D6674F85D1C65" bold="true" box="[199,308,1355,1380]" pageId="5" pageNumber="581">Remarks</emphasis>
. As reviewed by Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FD866673FA6A1C65" box="[636,771,1356,1380]" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article">Tan (1999)</bibRefCitation>
, species of 
<taxonomicName id="4C3C4D559E2DF746FC5E6674FD351C65" box="[932,1116,1355,1380]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAC49E2DF746FC5E6674FD351C65" box="[932,1116,1355,1380]" italics="true" pageId="5" pageNumber="581">Malayopotamon</emphasis>
</taxonomicName>
can be divided into two morphological groups on the basis of their carapace form (see also 
<bibRefCitation id="EFAD4B279E2DF746FC60664FFDEA1C89" author="Wowor" box="[922,1155,1392,1416]" pageId="5" pageNumber="581" refString="Wowor, D. &amp; Tan, S. H. (2010) Description of a new species of Malayopotamon Bott, 1968 (Decapoda, Brachyura, Potamidae) from Gunung Slamet, Central Java, Indonesia. Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Crustaceana Monographs, 14, 727 - 734. http: // dx. doi. org / 10.1163 / 9789047427759 _ 054" type="journal article" year="2010">Wowor &amp; Tan 2010</bibRefCitation>
). Members of one group (
<taxonomicName id="4C3C4D559E2DF746FF5A66AAF8831CAD" authority="Ng &amp; Wowor, 1991" authorityName="Ng &amp; Wowor" authorityYear="1991" box="[160,490,1427,1452]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2DF746FF5A66AAF86A1CAD" box="[160,259,1427,1452]" italics="true" pageId="5" pageNumber="581">M. batak</emphasis>
Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FEAB66ABF8831CAD" author="Wowor" box="[337,490,1428,1452]" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Wowor, D. (1991) A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from northern Sumatra. Treubia, 30 (2), 165 - 170." type="journal article" year="1991">Wowor, 1991</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FE0C66AAFA321CAD" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[502,859,1427,1452]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B948EAC49E2DF746FE0C66AAFBF71CAD" box="[502,670,1427,1452]" italics="true" pageId="5" pageNumber="581">M. granulatum</emphasis>
(De Man, 1892)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FC9D66AAFDD01CAD" authority="Wowor &amp; Tan, 2010" authorityName="Wowor &amp; Tan" authorityYear="2010" box="[871,1209,1427,1452]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="lipkei">
<emphasis id="B948EAC49E2DF746FC9D66AAFAA31CAD" box="[871,970,1427,1452]" italics="true" pageId="5" pageNumber="581">M. lipkei</emphasis>
<bibRefCitation id="EFAD4B279E2DF746FC2866ABFDD01CAD" author="Wowor" box="[978,1209,1428,1452]" pageId="5" pageNumber="581" refString="Wowor, D. &amp; Tan, S. H. (2010) Description of a new species of Malayopotamon Bott, 1968 (Decapoda, Brachyura, Potamidae) from Gunung Slamet, Central Java, Indonesia. Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Crustaceana Monographs, 14, 727 - 734. http: // dx. doi. org / 10.1163 / 9789047427759 _ 054" type="journal article" year="2010">Wowor &amp; Tan, 2010</bibRefCitation>
</taxonomicName>
) have the branchial, gastric, orbital and frontal regions strongly rugose and granular, whereas members of the second group (
<taxonomicName id="4C3C4D559E2DF746FA876686FB721CF5" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="brevimarginatum">
<emphasis id="B948EAC49E2DF746FA876686F83E1CF5" italics="true" pageId="5" pageNumber="581">M. brevimarginatum</emphasis>
(De Man, 1892)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FDD066E2FA241CF5" authority="Nobili, 1900" authorityName="Nobili" authorityYear="1900" box="[554,845,1499,1524]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B948EAC49E2DF746FDD066E2FBF71CF5" box="[554,670,1501,1524]" italics="true" pageId="5" pageNumber="581">M. gestroi</emphasis>
(Nobili, 1900)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FCA166E2FDC51CF5" authority="Balss, 1937" authorityName="Balss" authorityYear="1937" box="[859,1196,1499,1524]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="granulosum">
<emphasis id="B948EAC49E2DF746FCA166E2FD601CF5" box="[859,1033,1499,1524]" italics="true" pageId="5" pageNumber="581">M. granulosum</emphasis>
(Balss, 1937)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FB4066E2F9B31F19" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="javanense">
<emphasis id="B948EAC49E2DF746FB4066E2FC3B1CF5" box="[1210,1362,1501,1524]" italics="true" pageId="5" pageNumber="581">M. javanense</emphasis>
(Bott, 1968)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FF1D653EFB721F19" authority="Ng &amp; Tan, 1999" authorityName="Ng &amp; Tan" authorityYear="1999" box="[231,539,1535,1560]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B948EAC49E2DF746FF1D653EF83E1F19" box="[231,343,1535,1560]" italics="true" pageId="5" pageNumber="581">M. similis</emphasis>
Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FE52653FFB721F19" author="Tan" box="[424,539,1536,1560]" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan, 1999</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FDD2653EFA1A1F19" authority="Miers, 1880" authorityName="Miers" authorityYear="1880" box="[552,883,1535,1560]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="sumatrense">
<emphasis id="B948EAC49E2DF746FDD2653EFBA41F19" box="[552,717,1537,1560]" italics="true" pageId="5" pageNumber="581">M. sumatrense</emphasis>
(Miers, 1880)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FC7A653EFDF71F19" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[896,1182,1535,1560]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="tobaense">
<emphasis id="B948EAC49E2DF746FC7A653EFD631F19" box="[896,1034,1535,1560]" italics="true" pageId="5" pageNumber="581">M. tobaense</emphasis>
(Bott, 1968)
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FB56653EF9B81F3D" authority="Ng &amp; Tan, 1999" authorityName="Ng &amp; Tan" authorityYear="1999" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="turgeo">
<emphasis id="B948EAC49E2DF746FB56653EFC721F19" box="[1196,1307,1537,1560]" italics="true" pageId="5" pageNumber="581">M. turgeo</emphasis>
Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FA91653FF9B81F3D" author="Tan" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan, 1999</bibRefCitation>
</taxonomicName>
) have these carapace regions lightly rugose to almost smooth. The new species belongs to the second group. The G1 structure of 
<taxonomicName id="4C3C4D559E2DF746FE856576F8BA1F61" box="[383,467,1607,1632]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2DF746FE856576F8BA1F61" box="[383,467,1607,1632]" italics="true" pageId="5" pageNumber="581">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2DF746FE266578FB5E1F61" box="[476,567,1607,1632]" pageId="5" pageNumber="581" rank="species">
<emphasis id="B948EAC49E2DF746FE266578FB5E1F61" bold="true" box="[476,567,1607,1632]" pageId="5" pageNumber="581">sp. nov.</emphasis>
</taxonomicNameLabel>
is very distinctive and does not resemble that of any congener, being gently curved outwards, the terminal segment long and cylindrical with the tip subtruncate, and the subterminal segment relatively slender (see discussion in Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FD8865AFFBB61FA9" author="Tan" box="[626,735,1680,1704]" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B8336D69E2DF746FF3D658BF8791E5D" blockId="5.[151,1437,1355,2028]" pageId="5" pageNumber="581">
Ng &amp; 
<bibRefCitation id="EFAD4B279E2DF746FEEA658BF8E51FCD" box="[272,396,1716,1740]" pageId="5" pageNumber="581" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article">Tan (1999)</bibRefCitation>
discussed the variation in the G1 structure with growth and noted that while it is relatively more slender in smaller males, the general shape and form remains about the same. The degree of bending and/or curvature, and the general structure of the terminal segment has not been observed to vary significantly in species for which there is a good sample size (e.g., 
<taxonomicName id="4C3C4D559E2DF746FD80641EFA661E39" box="[634,783,1825,1848]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="javanense">
<emphasis id="B948EAC49E2DF746FD80641EFA661E39" box="[634,783,1825,1848]" italics="true" pageId="5" pageNumber="581">M. javanense</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FCE6641EFD681E39" box="[796,1025,1823,1848]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="brevimarginatum">
<emphasis id="B948EAC49E2DF746FCE6641EFD681E39" box="[796,1025,1823,1848]" italics="true" pageId="5" pageNumber="581">M. brevimarginatum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2DF746FBF5641EFDDE1E39" box="[1039,1207,1823,1848]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B948EAC49E2DF746FBF5641EFDDE1E39" box="[1039,1207,1823,1848]" italics="true" pageId="5" pageNumber="581">M. granulatum</emphasis>
</taxonomicName>
) (see also 
<bibRefCitation id="EFAD4B279E2DF746FAD4641FF86D1E5D" author="Wowor" pageId="5" pageNumber="581" refString="Wowor, D. &amp; Tan, S. H. (2010) Description of a new species of Malayopotamon Bott, 1968 (Decapoda, Brachyura, Potamidae) from Gunung Slamet, Central Java, Indonesia. Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Crustaceana Monographs, 14, 727 - 734. http: // dx. doi. org / 10.1163 / 9789047427759 _ 054" type="journal article" year="2010">Wowor &amp; Tan 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8B8336D69E2DF746FF3D6458F8801EEA" blockId="5.[151,1437,1355,2028]" pageId="5" pageNumber="581">
<taxonomicName id="4C3C4D559E2DF746FF3D6458F8D11E81" box="[199,440,1895,1920]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2DF746FF3D6458F8D11E81" box="[199,440,1895,1920]" italics="true" pageId="5" pageNumber="581">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2DF746FE3E6458FB4B1E81" box="[452,546,1895,1920]" pageId="5" pageNumber="581" rank="species">
<emphasis id="B948EAC49E2DF746FE3E6458FB4B1E81" bold="true" box="[452,546,1895,1920]" pageId="5" pageNumber="581">sp. nov.</emphasis>
</taxonomicNameLabel>
seems to be a relatively small-size species. Although it is only 
<quantity id="4CC49B339E2DF746FADC6458FC131E7E" box="[1318,1402,1895,1920]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="581" unit="mm" value="15.0">15 mm</quantity>
in carapace width, a size in which larger species like 
<taxonomicName id="4C3C4D559E2DF746FD2B64B2FADE1EA5" box="[721,951,1931,1956]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="brevimarginatum">
<emphasis id="B948EAC49E2DF746FD2B64B2FADE1EA5" box="[721,951,1931,1956]" italics="true" pageId="5" pageNumber="581">M. brevimarginatum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D559E2DF746FC1564B2FDFE1EA5" box="[1007,1175,1931,1956]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B948EAC49E2DF746FC1564B2FDFE1EA5" box="[1007,1175,1931,1956]" italics="true" pageId="5" pageNumber="581">M. granulatum</emphasis>
</taxonomicName>
are juvenile, the G1 of the 
<typeStatus id="548788749E2DF746FF3E6490F84F1EC9" box="[196,294,1967,1992]" pageId="5" pageNumber="581" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C3C4D559E2DF746FEA9648EF8C31EC9" box="[339,426,1967,1992]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2DF746FEA9648EF8C31EC9" box="[339,426,1967,1992]" italics="true" pageId="5" pageNumber="581">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2DF746FE4E6490FB781EC9" box="[436,529,1967,1992]" pageId="5" pageNumber="581" rank="species">
<emphasis id="B948EAC49E2DF746FE4E6490FB781EC9" bold="true" box="[436,529,1967,1992]" pageId="5" pageNumber="581">sp. nov.</emphasis>
</taxonomicNameLabel>
is relatively well developed, although the reduced setation along the margins suggests it is not fully mature.
</paragraph>
<paragraph id="8B8336D69E2EF745FF3D63A8FB5C1A5C" blockId="6.[151,1436,151,1007]" pageId="6" pageNumber="582">
Three other species are known from Aceh Province in northern Sumatra, 
<taxonomicName id="4C3C4D559E2EF745FC0263A6FD3219B1" box="[1016,1115,151,176]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2EF745FC0263A6FD3219B1" box="[1016,1115,151,176]" italics="true" pageId="6" pageNumber="582">M. batak</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2EF745FB9363A6FD9B19B1" box="[1129,1266,151,176]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="tobaense">
<emphasis id="B948EAC49E2EF745FB9363A6FD9B19B1" box="[1129,1266,151,176]" italics="true" pageId="6" pageNumber="582">M. tobaense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D559E2EF745FAD263A6FCFC19B1" box="[1320,1429,153,176]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="turgeo">
<emphasis id="B948EAC49E2EF745FAD263A6FCFC19B1" box="[1320,1429,153,176]" italics="true" pageId="6" pageNumber="582">M. turgeo</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C3C4D559E2EF745FF6D6383F8EA19D4" box="[151,387,188,213]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FF6D6383F8EA19D4" box="[151,387,188,213]" italics="true" pageId="6" pageNumber="582">Malayopotamon weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FE706383F88D19D4" box="[394,484,188,213]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FE706383F88D19D4" bold="true" box="[394,484,188,213]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
can immediately be separated from 
<taxonomicName id="4C3C4D559E2EF745FC876382FA8919D4" box="[893,992,188,213]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2EF745FC876382FA8919D4" box="[893,992,188,213]" italics="true" pageId="6" pageNumber="582">M. batak</emphasis>
</taxonomicName>
by its slightly rugose to smooth gastric and cardiac and frontal regions, which in 
<taxonomicName id="4C3C4D559E2EF745FD9063DEFBA419F9" box="[618,717,223,248]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2EF745FD9063DEFBA419F9" box="[618,717,223,248]" italics="true" pageId="6" pageNumber="582">M. batak</emphasis>
</taxonomicName>
, is strongly granular and rugose (see Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FB4363DFFC2719F9" author="Wowor" box="[1209,1358,224,248]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Wowor, D. (1991) A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from northern Sumatra. Treubia, 30 (2), 165 - 170." type="journal article" year="1991">Wowor 1991</bibRefCitation>
: pl. 1). The G1 of 
<taxonomicName id="4C3C4D559E2EF745FEED623AF815181C" box="[279,380,260,285]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2EF745FEED623AF815181C" box="[279,380,260,285]" italics="true" pageId="6" pageNumber="582">M. batak</emphasis>
</taxonomicName>
has a proportionately shorter and more quadrate terminal segment (Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FB28623AFC00181D" author="Wowor" box="[1234,1385,260,284]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Wowor, D. (1991) A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from northern Sumatra. Treubia, 30 (2), 165 - 170." type="journal article" year="1991">Wowor 1991</bibRefCitation>
: fig. 1B–E; Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FED16217F8FE1841" author="Tan" box="[299,407,296,320]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
: fig. 2a, b) than in 
<taxonomicName id="4C3C4D559E2EF745FD936216FBD21841" box="[617,699,295,320]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FD936216FBD21841" box="[617,699,295,320]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<emphasis id="B948EAC49E2EF745FD386218FA751841" bold="true" box="[706,796,295,320]" pageId="6" pageNumber="582">
<taxonomicNameLabel id="A27B57BF9E2EF745FD386218FA751841" box="[706,796,295,320]" pageId="6" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, where the terminal segment is long and cylindrical (
<figureCitation id="13072A539E2EF745FA946218F9CF1865" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="6" pageNumber="582">Fig. 4</figureCitation>
A–C). 
<taxonomicName id="4C3C4D559E2EF745FF0B6273FB7D1864" box="[241,532,332,357]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="tobaense">
<emphasis id="B948EAC49E2EF745FF0B6273FB7D1864" box="[241,532,332,357]" italics="true" pageId="6" pageNumber="582">Malayopotamon tobaense</emphasis>
</taxonomicName>
can be separated from 
<taxonomicName id="4C3C4D559E2EF745FCE26272FA031864" box="[792,874,332,357]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FCE26272FA031864" box="[792,874,332,357]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FC8B6273FAA21864" box="[881,971,332,357]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FC8B6273FAA21864" bold="true" box="[881,971,332,357]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
by the postorbital cristae being confluent with the epibranchial tooth, and the postorbital and epibranchial cristae being more distinct and sharp (see 
<bibRefCitation id="EFAD4B279E2EF745FA96624FF9BA18AD" author="Bott" pageId="6" pageNumber="582" refString="Bott, R. (1970) Die Sußwasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft, 526, 1 - 338, figs. 1 - 8, pls. 1 - 58, Map 1." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 58, fig. 89) (postorbital cristae not joining the epibranchial teeth and the postorbital and epigastric cristae not sharp in 
<taxonomicName id="4C3C4D559E2EF745FEDF6286F81118D1" box="[293,376,439,464]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FEDF6286F81118D1" box="[293,376,439,464]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FE7A6288F8A918D1" box="[384,448,439,464]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FE7A6288F8A918D1" bold="true" box="[384,448,439,464]" pageId="6" pageNumber="582">n. sp.</emphasis>
</taxonomicNameLabel>
; 
<figureCitation id="13072A539E2EF745FE2A6288FB7B18D1" box="[464,530,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="6" pageNumber="582">Fig. 2</figureCitation>
A, B), and the distal part of the subterminal segment of the G1 is distinctly bent with the terminal segment short and quadrate (Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FD1862E2FA2718F5" author="Tan" box="[738,846,476,500]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
: fig. 6) (distal part of the subterminal segment gently curving and the terminal segment elongated and cylindrical in 
<taxonomicName id="4C3C4D559E2EF745FC81613EFABB1B19" box="[891,978,511,536]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FC81613EFABB1B19" box="[891,978,511,536]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FC2762C0FD551B19" box="[989,1084,511,536]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FC2762C0FD551B19" bold="true" box="[989,1084,511,536]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<figureCitation id="13072A539E2EF745FBB462C0FDFC1B19" box="[1102,1173,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="6" pageNumber="582">Fig. 4</figureCitation>
A–C). 
<taxonomicName id="4C3C4D559E2EF745FB1E62C0F9891B3D" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="turgeo">
<emphasis id="B948EAC49E2EF745FB1E62C0F9891B3D" italics="true" pageId="6" pageNumber="582">Malayopotamon turgeo</emphasis>
</taxonomicName>
is distinctive in having the dorsal surface of the carapace convex, giving it a more inflated physiognomy; and its postorbital cristae is also confluent with and joins the epibranchial tooth (Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FB3B6177FC5D1B61" author="Tan" box="[1217,1332,584,608]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
: fig. 7a) (submarginally interrupted in 
<taxonomicName id="4C3C4D559E2EF745FE1F6152FB511B84" box="[485,568,620,645]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FE1F6152FB511B84" box="[485,568,620,645]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FDC56153FBF11B84" box="[575,664,620,645]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FDC56153FBF11B84" bold="true" box="[575,664,620,645]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<figureCitation id="13072A539E2EF745FD5C6153FB801B85" box="[678,745,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1923]" captionTargetId="figure@3.[151,1436,193,1924]" captionTargetPageId="3" captionText="FIGURE 2. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace." httpUri="https://zenodo.org/record/229085/files/figure.png" pageId="6" pageNumber="582">Fig. 2</figureCitation>
A). The structures of the G1 are also very different. While the terminal segments are both cylindrical, that of 
<taxonomicName id="4C3C4D559E2EF745FD4561AEFA581BA9" box="[703,817,657,680]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="turgeo">
<emphasis id="B948EAC49E2EF745FD4561AEFA581BA9" box="[703,817,657,680]" italics="true" pageId="6" pageNumber="582">M. turgeo</emphasis>
</taxonomicName>
is gently curving upwards and the distal part of the subdistal segment is strongly bent (Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FD9C618AFBBB1BCD" author="Tan" box="[614,722,692,716]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
: fig. 7c, e), whereas the distal part of the subterminal segment is gently curving and the terminal segment evenly cylindrical in 
<taxonomicName id="4C3C4D559E2EF745FCB161E6FAF41BF1" box="[843,925,727,752]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FCB161E6FAF41BF1" box="[843,925,727,752]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FC5E61E8FA941BF1" box="[932,1021,727,752]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FC5E61E8FA941BF1" bold="true" box="[932,1021,727,752]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13072A539E2EF745FBF661E8FD261BF1" box="[1036,1103,727,752]" captionStart="FIGURE 4" captionStartId="5.[151,250,1227,1249]" captionTargetBox="[174,1423,203,1214]" captionTargetId="figure@5.[151,1436,193,1224]" captionTargetPageId="5" captionText="FIGURE 4. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left G 1, ventral view; B, terminal segment of left G 1, ventral view; C, terminal segment of left G 1, dorsal view; D, left G 2. Scales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/229087/files/figure.png" pageId="6" pageNumber="582">Fig. 4</figureCitation>
A–C). In addition, while male thoracic sternites 3 and 4 are completely fused in 
<taxonomicName id="4C3C4D559E2EF745FD3661C2FA761A14" box="[716,799,764,789]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">
<emphasis id="B948EAC49E2EF745FD3661C2FA761A14" box="[716,799,764,789]" italics="true" pageId="6" pageNumber="582">M. weh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57BF9E2EF745FCDD61C3FAEB1A14" box="[807,898,764,789]" pageId="6" pageNumber="582" rank="species">
<emphasis id="B948EAC49E2EF745FCDD61C3FAEB1A14" bold="true" box="[807,898,764,789]" pageId="6" pageNumber="582">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13072A539E2EF745FC6B61C3FABD1A15" box="[913,980,764,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1901,1923]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure@4.[151,1436,193,1879]" captionTargetPageId="4" captionText="FIGURE 3. Malayopotamon weh sp. nov., holotype male (15.5 × 12.1 mm) (MZB), Pulau Weh, Sumatra. A, left third maxilliped; B, outer view of right major chela; C, outer view of left minor chela; D, anterior thoracic sternum and male abdomen; E, male abdomen and thoracic sternum; F, posterior part of carapace showing abdomen (first somite detached from carapace)." httpUri="https://zenodo.org/record/229086/files/figure.png" pageId="6" pageNumber="582">Fig. 3</figureCitation>
D), there is a shallow but visible groove present between these sternites in 
<taxonomicName id="4C3C4D559E2EF745FDE9601EFB1F1A39" box="[531,630,799,824]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="batak">
<emphasis id="B948EAC49E2EF745FDE9601EFB1F1A39" box="[531,630,799,824]" italics="true" pageId="6" pageNumber="582">M. batak</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D559E2EF745FD7E601EFA641A39" box="[644,781,799,824]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="tobaense">
<emphasis id="B948EAC49E2EF745FD7E601EFA641A39" box="[644,781,799,824]" italics="true" pageId="6" pageNumber="582">M. tobaense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3C4D559E2EF745FCB8601EFAC61A39" box="[834,943,801,824]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="turgeo">
<emphasis id="B948EAC49E2EF745FCB8601EFAC61A39" box="[834,943,801,824]" italics="true" pageId="6" pageNumber="582">M. turgeo</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFAD4B279E2EF745FC10601FFD341A39" author="Bott" box="[1002,1117,800,824]" pageId="6" pageNumber="582" refString="Bott, R. (1970) Die Sußwasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft, 526, 1 - 338, figs. 1 - 8, pls. 1 - 58, Map 1." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 58 fig. 89; Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FAB0601FF9BA1A5D" author="Wowor" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Wowor, D. (1991) A new species of Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae) from northern Sumatra. Treubia, 30 (2), 165 - 170." type="journal article" year="1991">Wowor 1991</bibRefCitation>
: pl. 1; Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FE9E607AF8B81A5D" author="Tan" box="[356,465,836,860]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Tan, S. H. (1999) Revision of the Southeast Asian potamid crabs of the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 33 (2), 207 - 231. http: // dx. doi. org / 10.1080 / 002229399300380" type="journal article" year="1999">Tan 1999</bibRefCitation>
: fig. 7b).
</paragraph>
<paragraph id="8B8336D69E2EF745FF3D6055FC381AEE" blockId="6.[151,1436,151,1007]" pageId="6" pageNumber="582">
The small island of 
<collectingCountry id="F32B76469E2EF745FE586055F88B1A82" box="[418,482,874,899]" name="Iceland" pageId="6" pageNumber="582">Pulau</collectingCountry>
<taxonomicName id="4C3C4D559E2EF745FE126055FB721A82" box="[488,539,874,899]" class="Malacostraca" family="Potamidae" genus="Malayopotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="weh" status="sp. nov.">Weh</taxonomicName>
(ca. 
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2) is relatively isolated, being about 
<quantity id="4CC49B339E2EF745FBC96055FD101A82" box="[1075,1145,874,899]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="582" unit="km" value="15.0">15 km</quantity>
north of the northernmost tip of Sumatra. Relatively high (about 
<quantity id="4CC49B339E2EF745FDAD60B1FBC81AA6" box="[599,673,910,935]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="582" unit="m" value="600.0">600 m</quantity>
in altitude), it has several waterfalls and streams. De 
<bibRefCitation id="EFAD4B279E2EF745FAEE60AFFCF51AA6" author="Man" box="[1300,1436,910,935]" pageId="6" pageNumber="582" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 9 (1), 75 - 218." type="journal article" year="1895">Man (1895)</bibRefCitation>
reported on some intertidal crabs from the island (see also Ng &amp; 
<bibRefCitation id="EFAD4B279E2EF745FC54608DFD741ACA" author="Liu" box="[942,1053,946,971]" pageId="6" pageNumber="582" refString="Ng, P. K. L. &amp; Liu, H. - C. (1999) The taxonomy of Sesarma tangi Rathbun, 1931 and S. stormi De Man, 1895 (Crustacea: Decapoda: Brachyura: Grapsidae: Sesarminae), with establishment of a new genus for S. stormi. Zoological Studies, 38 (2), 228 - 237." type="journal article" year="1999">Liu 1999</bibRefCitation>
), but surprisingly, no freshwater decapods have ever been recorded. The present record of a freshwater potamid crab is therefore noteworthy.
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