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<document id="103397AFC41AE6A45D2A4D25336E40EA" ID-CLB-Dataset="56069" ID-DOI="10.11646/zootaxa.5125.3.5" ID-GBIF-Dataset="da274829-479f-4ea1-8f2e-1e598de786b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6443876" ID-ZooBank="4645A035-17E6-488C-8F10-DA66E146487D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649661642485" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2022" docId="039587B4FFB3FFA6FF5CFF71D95DFCC8" docLanguage="en" docName="zootaxa.5125.3.5.pdf" docOrigin="Zootaxa 5125 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudomicracis tomicoides" docType="treatment" docVersion="7" lastPageNumber="338" masterDocId="FFACFFCCFFBEFFABFFCBFFAED841FF85" masterDocTitle="The strongly dimorphic bark beetle genus Pseudomicracis (Coleoptera, Scolytinae) in Madagascar-an integrated taxonomic revision" masterLastPageNumber="343" masterPageNumber="325" pageNumber="338" updateTime="1699319771094" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F8472CD89176057BEFBCE81DE2FF14C7">The strongly dimorphic bark beetle genus Pseudomicracis (Coleoptera, Scolytinae) in Madagascar-an integrated taxonomic revision</mods:title>
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<mods:namePart id="5615E43E81F8BAB6B215C2D2A87E1D7A">Jordal, Bjarte H.</mods:namePart>
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<mods:date id="D0EC7B308CC7DA89B1C3659A274E1C0C">2022</mods:date>
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<treatment id="039587B4FFB3FFA6FF5CFF71D95DFCC8" ID-DOI="http://doi.org/10.5281/zenodo.6448753" ID-GBIF-Taxon="194565927" ID-Zenodo-Dep="6448753" LSID="urn:lsid:plazi:treatment:039587B4FFB3FFA6FF5CFF71D95DFCC8" httpUri="http://treatment.plazi.org/id/039587B4FFB3FFA6FF5CFF71D95DFCC8" lastPageNumber="338" pageId="13" pageNumber="338">
<subSubSection id="C3266529FFB3FFA6FF5CFF71DAD4FF7C" box="[151,661,223,250]" pageId="13" pageNumber="338" type="nomenclature">
<paragraph id="8B8336A2FFB3FFA6FF5CFF71DAD4FF7C" blockId="13.[151,661,223,286]" box="[151,661,223,250]" pageId="13" pageNumber="338">
<heading id="D0CB81CEFFB3FFA6FF5CFF71DAD4FF7C" bold="true" box="[151,661,223,250]" fontSize="11" level="1" pageId="13" pageNumber="338" reason="1">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CFF71DAD4FF7C" authority="(Schedl, 1961)" baseAuthorityName="Schedl" baseAuthorityYear="1961" box="[151,661,223,250]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB3FFA6FF5CFF71DAD4FF7C" bold="true" box="[151,661,223,250]" pageId="13" pageNumber="338">
<emphasis id="B948EAB0FFB3FFA6FF5CFF71D99FFF7F" bold="true" box="[151,478,223,250]" italics="true" pageId="13" pageNumber="338">Pseudomicracis tomicoides</emphasis>
(
<bibRefCitation id="EFAD4B53FFB3FFA6FE26FF71DACCFF7C" author="Schedl, K. E." box="[493,653,223,249]" pageId="13" pageNumber="338" pagination="127 - 170" refId="ref10492" refString="Schedl, K. E. (1961) Fauna Madagascariensis, IV. 188 contribution. Institut Scientifique de Madagascar, Memoires, Series E, 12, 127 - 170." type="journal article" year="1961">Schedl, 1961</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF5CFEADD991FE9B" box="[151,464,259,286]" pageId="13" pageNumber="338" type="description">
<paragraph id="8B8336A2FFB3FFA6FF5CFEADD991FE9B" blockId="13.[151,661,223,286]" box="[151,464,259,286]" pageId="13" pageNumber="338">
(
<figureCitation id="13072A27FFB3FFA6FF54FEADD90AFE98" box="[159,331,259,285]" captionStart="FIGURES 3644" captionStartId="14.[151,264,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-25@14.[151,1436,181,1890]" captionTargetPageId="14" captionText="FIGURES 3644. Dorsal, lateral and front view of Pseudomicracis tomicoides male holotype (36, 39, 43); Pseudomicracis dispar male (37, 40, 43); Pseudomicracis pennata male holotype (38, 41, 44)." figureDoi="http://doi.org/10.5281/zenodo.6443900" httpUri="https://zenodo.org/record/6443900/files/figure.png" pageId="13" pageNumber="338">Figs 36, 39, 42</figureCitation>
,
<figureCitation id="13072A27FFB3FFA6FE9DFEADD986FE9B" box="[342,455,259,286]" captionStart="FIGURES 4553" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[178,1418,185,1919]" captionTargetId="figure-15@15.[170,1418,181,1919]" captionTargetPageId="15" captionText="FIGURES 4553. Dorsal, lateral and front view of Pseudomicracis tomicoides female allotype (45, 48, 51); Pseudomicracis dispar female (46, 49, 52); Pseudomicracis pennata female allotype (47, 50, 53)." figureDoi="http://doi.org/10.5281/zenodo.6443904" httpUri="https://zenodo.org/record/6443904/files/figure.png" pageId="13" pageNumber="338">45, 48, 51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF5CFEE6DBCCFE18" pageId="13" pageNumber="338" type="reference_group">
<paragraph id="8B8336A2FFB3FFA6FF5CFEE6DA2CFEE5" blockId="13.[151,909,327,413]" box="[151,621,327,352]" pageId="13" pageNumber="338">
<treatmentCitationGroup id="AB2C118CFFB3FFA6FF5CFEE6DA2CFEE5" box="[151,621,327,352]" pageId="13" pageNumber="338">
<treatmentCitation id="0A9D10B3FFB3FFA6FF5CFEE6DA2CFEE5" author="Schedl, K. E." box="[151,621,327,352]" page="141" pageId="13" pageNumber="338" year="1961">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CFEE6DA2CFEE5" ID-CoL="42PW5" authority="Schedl, 1961: 141" authorityName="Schedl" authorityPageNumber="141" authorityYear="1961" box="[151,621,327,352]" class="Insecta" family="Curculionidae" genus="Micracidendron" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB3FFA6FF5CFEE6D9EEFEE5" box="[151,431,328,352]" italics="true" pageId="13" pageNumber="338">Micracidendron tomicoides</emphasis>
<bibRefCitation id="EFAD4B53FFB3FFA6FE7EFEE6DA2CFEE5" author="Schedl, K. E." box="[437,621,327,352]" pageId="13" pageNumber="338" pagination="127 - 170" refId="ref10492" refString="Schedl, K. E. (1961) Fauna Madagascariensis, IV. 188 contribution. Institut Scientifique de Madagascar, Memoires, Series E, 12, 127 - 170." type="journal article" year="1961">Schedl, 1961: 141</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8B8336A2FFB3FFA6FF5CFEC9DB08FEFB" blockId="13.[151,909,327,413]" box="[151,841,358,382]" pageId="13" pageNumber="338">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CFEC9DA54FEFB" ID-CoL="6XRNX" authority="(Schedl, 1961)" baseAuthorityName="Schedl" baseAuthorityYear="1961" box="[151,533,358,382]" class="Insecta" family="Curculionidae" genus="Saurotocis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB3FFA6FF5CFEC9D936FEFB" box="[151,375,358,382]" italics="true" pageId="13" pageNumber="338">Saurotocis tomicoides</emphasis>
(
<bibRefCitation id="EFAD4B53FFB3FFA6FE4FFEC8DA4CFEFB" author="Schedl, K. E." box="[388,525,358,382]" pageId="13" pageNumber="338" pagination="127 - 170" refId="ref10492" refString="Schedl, K. E. (1961) Fauna Madagascariensis, IV. 188 contribution. Institut Scientifique de Madagascar, Memoires, Series E, 12, 127 - 170." type="journal article" year="1961">Schedl, 1961</bibRefCitation>
)
</taxonomicName>
, combination by
<bibRefCitation id="EFAD4B53FFB3FFA6FD08FEC8DB08FEFB" author="Wood, S. L." box="[707,841,358,382]" pageId="13" pageNumber="338" pagination="223 - 230" refId="ref10686" refString="Wood, S. L. (1984) New generic synonomy and new genera of Scolytidae (Coleoptera). Great Basin Naturalist, 44, 223 - 230." type="journal article" year="1984">Wood (1984)</bibRefCitation>
</paragraph>
<paragraph id="8B8336A2FFB3FFA6FF5CFE2BDBCCFE18" blockId="13.[151,909,327,413]" box="[151,909,389,413]" pageId="13" pageNumber="338">
<taxonomicName id="4C3C4D21FFB3FFA6FF5CFE2BDA0AFE18" authority="(Schedl, 1961)" baseAuthorityName="Schedl" baseAuthorityYear="1961" box="[151,587,389,413]" class="Insecta" family="Curculionidae" genus="Pseudomicracis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="tomicoides">
<emphasis id="B948EAB0FFB3FFA6FF5CFE2BD9ECFE18" box="[151,429,389,413]" italics="true" pageId="13" pageNumber="338">Pseudomicracis tomicoides</emphasis>
(
<bibRefCitation id="EFAD4B53FFB3FFA6FE70FE2BDA02FE18" author="Schedl, K. E." box="[443,579,389,413]" pageId="13" pageNumber="338" pagination="127 - 170" refId="ref10492" refString="Schedl, K. E. (1961) Fauna Madagascariensis, IV. 188 contribution. Institut Scientifique de Madagascar, Memoires, Series E, 12, 127 - 170." type="journal article" year="1961">Schedl, 1961</bibRefCitation>
)
</taxonomicName>
, combination by
<bibRefCitation id="EFAD4B53FFB3FFA6FD31FE2BDBCCFE18" author="Jordal, B. H." box="[762,909,389,413]" pageId="13" pageNumber="338" pagination="245 - 278" refId="ref10239" refString="Jordal, B. H. (2021 c) A phylogenetic and taxonomic assessment of Afrotropical Micracidini (Coleoptera, Scolytinae) reveals a strong diversifying role for Madagascar. Organisms Diversity &amp; Evolution, 21, 245 - 278. https: // doi. org / 10.1007 / s 13127 - 021 - 00481 - 4" type="journal article" year="2021">Jordal (2021c)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF5CFE65DAA9FD8C" pageId="13" pageNumber="338" type="materials_examined">
<paragraph id="8B8336A2FFB3FFA6FF5CFE65DAA9FD8C" blockId="13.[151,1437,458,845]" pageId="13" pageNumber="338">
<emphasis id="B948EAB0FFB3FFA6FF5CFE65D907FE60" bold="true" box="[151,326,459,485]" pageId="13" pageNumber="338">Type material.</emphasis>
<materialsCitation id="3B543CFFFFB3FFA6FE84FE65DC80FE60" ID-GBIF-Occurrence="3750353311" box="[335,1217,458,486]" collectingDate="1952-11-23" collectionCode="MNHN" collectorName="Dr K. E. Schedl" country="Madagascar" location="Perinet" pageId="13" pageNumber="338" specimenCount="1" typeStatus="holotype">
<typeStatus id="54878800FFB3FFA6FE84FE65D9FBFE60" box="[335,442,459,485]" pageId="13" pageNumber="338" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="F32B7632FFB3FFA6FDE8FE65DAEDFE60" box="[547,684,459,485]" name="Madagascar" pageId="13" pageNumber="338">Madagascar</collectingCountry>
,
<location id="8EE36079FFB3FFA6FD73FE65DB4BFE60" LSID="urn:lsid:plazi:treatment:039587B4FFB3FFA6FF5CFF71D95DFCC8:8EE36079FFB3FFA6FD73FE65DB4BFE60" box="[696,778,459,485]" country="Madagascar" name="Perinet" pageId="13" pageNumber="338">Perinet</location>
,
<date id="FF821062FFB3FFA6FCDCFE65DBD0FE63" box="[791,913,459,486]" pageId="13" pageNumber="338" value="1952-11-23">
<collectingDate id="EFC6E98AFFB3FFA6FCDCFE65DBD0FE63" box="[791,913,459,486]" pageId="13" pageNumber="338" value="1952-11-23">23.xi.1952</collectingDate>
</date>
,
<collectorName id="26C95374FFB3FFA6FC55FE62DC0DFE60" box="[926,1100,459,486]" pageId="13" pageNumber="338">Dr K.E. Schedl</collectorName>
[
<collectionCode id="ED2DAE67FFB3FFA6FB90FE64DCFBFE60" box="[1115,1210,458,485]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="338">MNHN</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B543CFFFFB3FFA6FB00FE65DAA5FD8C" ID-GBIF-Occurrence="3750353315" collectingDate="1952-11-23" collectionCode="MNHN, NHMW" collectorName="Dr K. E. Schedl" country="Madagascar" location="Perinet" pageId="13" pageNumber="338" specimenCount="2" typeStatus="allotype">
<typeStatus id="54878800FFB3FFA6FB00FE65DD6CFE60" box="[1227,1325,459,485]" pageId="13" pageNumber="338" type="allotype">Allotype</typeStatus>
male and
<typeStatus id="54878800FFB3FFA6FF5CFE5ED946FD8F" box="[151,263,496,522]" pageId="13" pageNumber="338" type="paratype">paratypes</typeStatus>
: same data as holotype [
<collectionCode id="ED2DAE67FFB3FFA6FDDEFE5EDA31FD8F" box="[533,624,496,522]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="338">MNHN</collectionCode>
,
<collectionCode id="ED2DAE67FFB3FFA6FDB7FE40DA9FFD8C" box="[636,734,494,521]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="338" type="Museum">NHMW</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF0CFDBDD996FD38" pageId="13" pageNumber="338" type="diagnosis">
<paragraph id="8B8336A2FFB3FFA6FF0CFDBDD996FD38" blockId="13.[151,1437,458,845]" pageId="13" pageNumber="338">
<emphasis id="B948EAB0FFB3FFA6FF0CFDBDD903FDA8" bold="true" box="[199,322,531,557]" pageId="13" pageNumber="338">Diagnosis.</emphasis>
Length
<quantity id="4CC49B47FFB3FFA6FE6AFDBDDA6AFDAB" box="[417,555,531,558]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="13" pageNumber="338" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.62.9 mm</quantity>
, 2.32.5 × as long as wide. Antennal club almost three times as long as wide, sutures not evident, basal segment shiny, very short;
<emphasis id="B948EAB0FFB3FFA6FD32FD99DB72FDD4" bold="true" box="[761,819,567,593]" pageId="13" pageNumber="338">male</emphasis>
declivity impressed, demarcated on each side by 57 large tubercles; posterior margin of elytra with a distinct rim; interstriae with densely placed and strongly confused setae;
<emphasis id="B948EAB0FFB3FFA6FF14FDD1D96FFD1C" bold="true" box="[223,302,639,665]" pageId="13" pageNumber="338">female</emphasis>
frons with dense, long, soft setae; antennal scapus short, with a very long and thin dorsal spike with long setae longer than spike.
</paragraph>
</subSubSection>
<subSubSection id="C3266529FFB3FFA6FF0CFD69D95DFCC8" pageId="13" pageNumber="338" type="distribution">
<paragraph id="8B8336A2FFB3FFA6FF0CFD69D95DFCC8" blockId="13.[151,1437,458,845]" pageId="13" pageNumber="338">
<emphasis id="B948EAB0FFB3FFA6FF0CFD69D9B1FD64" bold="true" box="[199,496,711,737]" pageId="13" pageNumber="338">Distribution and biology.</emphasis>
<collectingCountry id="F32B7632FFB3FFA6FE3DFD69DA3EFD64" box="[502,639,711,737]" name="Madagascar" pageId="13" pageNumber="338">Madagascar</collectingCountry>
. Only known from the
<typeStatus id="54878800FFB3FFA6FCB4FD66DBEEFD67" box="[895,943,712,738]" pageId="13" pageNumber="338">type</typeStatus>
locality where it was dissected from
<taxonomicName id="4C3C4D21FFB3FFA6FA80FD66D8AAFC80" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="338" phylum="Tracheophyta" rank="species" species="cymosa">
<emphasis id="B948EAB0FFB3FFA6FA80FD66D8AAFC80" italics="true" pageId="13" pageNumber="338">Ocotea cymosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4D21FFB3FFA6FEEFFD42DA51FC80" box="[292,528,747,774]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="338" phylum="Tracheophyta" rank="species" species="laens">
<emphasis id="B948EAB0FFB3FFA6FEEFFD42D9C3FC80" box="[292,386,747,774]" italics="true" pageId="13" pageNumber="338">O. laens</emphasis>
(Lauraceae)
</taxonomicName>
branches about
<quantity id="4CC49B47FFB3FFA6FD01FD45DB5FFC83" box="[714,798,747,774]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="13" pageNumber="338" unit="cm" value="4.5" valueMax="5.0" valueMin="4.0">45 cm</quantity>
diameter (
<bibRefCitation id="EFAD4B53FFB3FFA6FC5CFD45DC68FC80" author="Schedl, K. E." box="[919,1065,747,773]" pageId="13" pageNumber="338" pagination="1 - 326" refId="ref10622" refString="Schedl, K. E. (1977) Die Scolytidae und Platypodidae Madagaskars und einiger naheliegender Inselgruppen. Mitteilungen der Forstlichen Bundes-Versuchsanstalt Wien, 119, 1 - 326." type="journal article" year="1977">Schedl 1977</bibRefCitation>
). One male may accept one, two or
<specimenCount id="9D3AFD2BFFB3FFA6FF7DFCA1D90EFCAC" box="[182,335,783,809]" pageId="13" pageNumber="338" type="female">three females</specimenCount>
, copulate with these just inside the entrance, and each female excavate longitudinal egg tunnels in the phloem.
</paragraph>
</subSubSection>
</treatment>
</document>