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<document id="F07014A116E69CFFEA63F5B6FD0CF682" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin &amp; Jia, Xinyang" docDate="2023" docId="039587ADBF14FF84FF7F47A42752F800" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dongodytes mingruoae Tian &amp; Huang &amp; Jia 2023, n. sp." docType="treatment" docVersion="5" lastPageNumber="46" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="45" updateTime="1699475196432" updateUser="ExternalLinkService">
<mods:mods id="30564E643A5BAF92409988CD1ADFEE52" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="465D367945A2CF3EB02FB1F3BFEC9F2B">
<mods:title id="D284A3798B586D6F43F3D87C3B976218">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
</mods:titleInfo>
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<mods:role id="73A68E9CC1F120F7CDFA17F4BB97D051">
<mods:roleTerm id="B9DB1B69A4046D43A995CC25F90D7148">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F0F3B4EE6BF0030F6326D948318511C">Tian, Mingyi</mods:namePart>
<mods:nameIdentifier id="5184196FFD968568D6BF8FE60A410D9E" type="ORCID">0000-0003-2823-7619</mods:nameIdentifier>
<mods:nameIdentifier id="7837683AEB55AAC431A8ED4C0D2C0E4F" type="email">mytian@scau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="73D56F15806AA21A7F5ABA32402AB36D">Huang, Sunbin</mods:namePart>
<mods:nameIdentifier id="EE9040A594D32D6C0A57BE9CA85C7586" type="ORCID">0000-0001-8357-6651</mods:nameIdentifier>
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<mods:roleTerm id="7534D88AB7113ED782212DACA475ED54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="84E526736C5CE9A1F2F5CC9DC67491EF">Jia, Xinyang</mods:namePart>
<mods:nameIdentifier id="C94913B9B7589E7FF048BE19AFF49843" type="ORCID">0000-0003-1936-5926</mods:nameIdentifier>
<mods:nameIdentifier id="A57061C7976AD65AFC9E077006C5B95A" type="email">527748207@qq.com</mods:nameIdentifier>
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<mods:typeOfResource id="566D88D4BC79998415073D6257C2BEBC">text</mods:typeOfResource>
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<mods:titleInfo id="6850FAB9B705CDDD85438F307665C1EC">
<mods:title id="5DD4C5A339E1A07B3FAEAA7AC814ED1F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1885D5645177E63D74B2739C14FFBEF9">
<mods:date id="6FF8C26D0C95B3C218CACD1BB3FFD3DF">2023</mods:date>
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<mods:number id="909AC9CA87029D729FAB807FE154B5F0">2023-02-15</mods:number>
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<mods:number id="1DBA73D128E2A445D528FCFA29D88271">5243</mods:number>
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<mods:number id="37112A50A6F0A0CB9699F3F59BBE8A1E">1</mods:number>
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<mods:url id="F37A1AD04B62A0A9652D0AC4ACC19961">http://dx.doi.org/10.11646/zootaxa.5243.1.1</mods:url>
</mods:location>
<mods:classification id="F769C0B4D1DE23ACA8FDB4A44D80CEF9">journal article</mods:classification>
<mods:identifier id="1AEA97690CBB7291F39DDDA0FB0FA2AD" type="CLB-Dataset">54139</mods:identifier>
<mods:identifier id="765F13BE56BC22198BC7F6DEBA8C86FA" type="DOI">10.11646/zootaxa.5243.1.1</mods:identifier>
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<mods:identifier id="66DFE67355F8AB5417F0B9CB03B38594" type="Zenodo-Dep">7642400</mods:identifier>
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<treatment id="039587ADBF14FF84FF7F47A42752F800" ID-DOI="http://doi.org/10.5281/zenodo.7647044" ID-GBIF-Taxon="209296919" ID-Zenodo-Dep="7647044" LSID="urn:lsid:plazi:treatment:039587ADBF14FF84FF7F47A42752F800" httpUri="http://treatment.plazi.org/id/039587ADBF14FF84FF7F47A42752F800" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C3266530BF14FF85FF7F47A42034F92E" box="[151,596,1729,1756]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8B8336BBBF14FF85FF7F47A42034F92E" blockId="44.[151,596,1729,1791]" box="[151,596,1729,1756]" pageId="44" pageNumber="45">
<heading id="D0CB81D7BF14FF85FF7F47A42034F92E" bold="true" box="[151,596,1729,1756]" fontSize="11" level="2" pageId="44" pageNumber="45" reason="4">
<emphasis id="B948EAA9BF14FF85FF7F47A42034F92E" bold="true" box="[151,596,1729,1756]" pageId="44" pageNumber="45">
<taxonomicName id="4C3C4D38BF14FF85FF7F47A4206AF92E" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[151,522,1729,1756]" class="Insecta" family="Carabidae" genus="Dongodytes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mingruoae" status="sp. nov.">
<emphasis id="B948EAA9BF14FF85FF7F47A4206AF92E" bold="true" box="[151,522,1729,1756]" italics="true" pageId="44" pageNumber="45">Dongodytes (s. str.) mingruoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57D2BF14FF85FDF947A42034F92E" box="[529,596,1729,1755]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF14FF85FF7F478023F9F90A" box="[151,409,1765,1791]" pageId="44" pageNumber="45" type="description">
<paragraph id="8B8336BBBF14FF85FF7F478023F9F90A" blockId="44.[151,596,1729,1791]" box="[151,409,1765,1791]" pageId="44" pageNumber="45">
<figureCitation id="13072A3EBF14FF85FF7F47802369F90A" box="[151,265,1765,1791]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="44" pageNumber="45">Figures 1j</figureCitation>
,
<figureCitation id="13072A3EBF14FF85FEFE47802327F90A" box="[278,327,1765,1791]" captionStart="FIGURE 27" captionStartId="42.[151,250,1965,1989]" captionTargetBox="[280,1306,453,1927]" captionTargetId="figure-129@42.[280,1307,453,1942]" captionTargetPageId="42" captionText="FIGURE 27. Left elytra (white dots indicating the chaetotaxy) of Dongodytes species (white dots indicating the chaetotaxy). A. Dongodytes (Dongodytodes) pingjingae sp. n.; B. Dongodytes (s. str.) mingruoae n. sp." figureDoi="http://doi.org/10.5281/zenodo.7642446" httpUri="https://zenodo.org/record/7642446/files/figure.png" pageId="44" pageNumber="45">27B</figureCitation>
,
<figureCitation id="13072A3EBF14FF85FEBB478023F9F90A" box="[339,409,1765,1791]" captionStart-0="FIGURE 29" captionStart-1="FIGURE 30" captionStart-2="FIGURE 31" captionStartId-0="46.[151,250,1943,1967]" captionStartId-1="47.[151,250,1867,1891]" captionStartId-2="48.[151,250,1869,1893]" captionTargetBox-1="[161,1422,193,1831]" captionTargetBox-2="[151,1435,181,1845]" captionTargetId-0="figure-28@46.[346,1241,181,1920]" captionTargetId-1="figure-20@47.[160,1428,181,1844]" captionTargetId-2="figure-28@48.[151,1435,181,1845]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionTargetPageId-2="48" captionText-0="FIGURE 29. Habitus of Dongodytes (s. str.) mingruoae n. sp., male, holotype." captionText-1="FIGURE 30. Dongodytes (s. str.) mingruoae n. sp. A. median lobe, lateral view; B. median lobe, dorsal view. Scale for both." captionText-2="FIGURE 31. Jiudu Cave I, the type locality of Dongodytes (s. str.) mingruoae n. sp. A. entrance; B. several plants of Begonia edulis Lévl., 1909; C. a running specimen of Dongodytes (s. str.) mingruoae n. sp. in cave." figureDoi-0="http://doi.org/10.5281/zenodo.7647034" figureDoi-1="http://doi.org/10.5281/zenodo.7642450" figureDoi-2="http://doi.org/10.5281/zenodo.7642452" httpUri-0="https://zenodo.org/record/7647034/files/figure.png" httpUri-1="https://zenodo.org/record/7642450/files/figure.png" httpUri-2="https://zenodo.org/record/7642452/files/figure.png" pageId="44" pageNumber="45">29–31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF14FF85FF7F4648263DF822" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8B8336BBBF14FF85FF7F4648263DF822" blockId="44.[151,1437,1837,2008]" pageId="44" pageNumber="45">
<emphasis id="B948EAA9BF14FF85FF7F46482366F8B2" bold="true" box="[151,262,1837,1863]" pageId="44" pageNumber="45">Material.</emphasis>
<materialsCitation id="3B543CE6BF14FF85FEF946482020F87A" ID-GBIF-Occurrence="4030741306" collectingDate="2018-09-25" collectingDateMax="2018-09-25" collectingDateMin="2015-07-27" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Sunbin Huang &amp; Mengzhen Chen" country="China" county="Du'an Yao Autonomous County" elevation="333" latitude="24.27" location="Cave I" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="10" specimenCount-male="1" stateProvince="Guangxi Zhuang Autonomous Region" typeStatus="holotype">
<typeStatus id="54878819BF14FF85FEF94648231EF8B2" box="[273,382,1837,1863]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D3AFD32BF14FF85FE64464823A5F8B2" box="[396,453,1837,1863]" pageId="44" pageNumber="45" type="male">male</specimenCount>
,
<location id="8EE36060BF14FF85FE3D46482044F8BD" LSID="urn:lsid:plazi:treatment:039587ADBF14FF84FF7F47A42752F800:8EE36060BF14FF85FE3D46482044F8BD" box="[469,548,1837,1864]" country="China" county="Du'an Yao Autonomous County" latitude="24.27" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" name="Cave I" pageId="44" pageNumber="45" stateProvince="Guangxi Zhuang Autonomous Region">Cave I</location>
,
<location id="8EE36060BF14FF85FDDD464820A3F8B2" LSID="urn:lsid:plazi:treatment:039587ADBF14FF84FF7F47A42752F800:8EE36060BF14FF85FDDD464820A3F8B2" box="[565,707,1837,1863]" country="China" county="Du'an Yao Autonomous County" latitude="24.27" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" name="Jiudu Xiang" pageId="44" pageNumber="45" stateProvince="Guangxi Zhuang Autonomous Region">Jiudu Xiang</location>
,
<collectingCounty id="62E24E37BF14FF85FD3B46482627F8B2" box="[723,1095,1837,1864]" pageId="44" pageNumber="45">Du’an Yao Autonomous County</collectingCounty>
,
<location id="8EE36060BF14FF85FBBE464826ACF8B2" LSID="urn:lsid:plazi:treatment:039587ADBF14FF84FF7F47A42752F800:8EE36060BF14FF85FBBE464826ACF8B2" box="[1110,1228,1837,1863]" country="China" county="Du'an Yao Autonomous County" latitude="24.27" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" name="Hechi Shi" pageId="44" pageNumber="45" stateProvince="Guangxi Zhuang Autonomous Region">Hechi Shi</location>
,
<collectingRegion id="49F8F859BF14FF85FB33464823E2F89E" pageId="44" pageNumber="45">Guangxi Zhuang Autonomous Region</collectingRegion>
, southern
<collectingCountry id="F32B762BBF14FF85FE1B46342059F89E" box="[499,569,1873,1899]" name="China" pageId="44" pageNumber="45">China</collectingCountry>
,
<geoCoordinate id="EE08507CBF14FF85FDAB463420C1F89E" box="[579,673,1873,1899]" degrees="24.27" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="555" value="24.27">24.27°N</geoCoordinate>
/
<geoCoordinate id="EE08507CBF14FF85FD5D46342141F89E" box="[693,801,1873,1899]" degrees="108.23" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="555" value="108.23">108.23°E</geoCoordinate>
,
<quantity id="4CC49B5EBF14FF85FCC446342112F899" box="[812,882,1873,1900]" metricMagnitude="2" metricUnit="m" metricValue="3.3299999999999996" pageId="44" pageNumber="45" unit="m" value="333.0">
<elevation id="0011D188BF14FF85FCC446342112F899" box="[812,882,1873,1900]" metricMagnitude="2" metricUnit="m" metricValue="3.3299999999999996" pageId="44" pageNumber="45" unit="m" value="333.0">333 m</elevation>
</quantity>
in altitude,
<date id="FF82107BBF14FF85FC1D4634261CF899" box="[1013,1148,1873,1900]" pageId="44" pageNumber="45" value="2018-09-25">
<collectingDate id="EFC6E993BF14FF85FC1D4634261CF899" box="[1013,1148,1873,1900]" pageId="44" pageNumber="45" value="2018-09-25">2018-IX-25</collectingDate>
</date>
, leg.
<collectorName id="26C9536DBF14FF85FB5D46342722F89E" box="[1205,1346,1873,1899]" pageId="44" pageNumber="45">Mingyi Tian</collectorName>
,
<collectorName id="26C9536DBF14FF85FAA546342282F865" pageId="44" pageNumber="45">Sunbin Huang</collectorName>
&amp;
<collectorName id="26C9536DBF14FF85FEE0461023A4F87A" box="[264,452,1909,1935]" pageId="44" pageNumber="45">Mengzhen Chen</collectorName>
, in
<collectionCode id="ED2DAE7EBF14FF85FE064610205CF87A" box="[494,572,1909,1935]" pageId="44" pageNumber="45">SCAU</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B543CE6BF14FF85FDA046132723F87A" ID-GBIF-Occurrence="4030741324" box="[584,1347,1909,1936]" collectingDate="2018-09-25" collectingDateMax="2018-09-25" collectingDateMin="2015-07-27" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Sunbin Huang &amp; Mengzhen Chen" country="China" county="Du'an Yao Autonomous County" elevation="333" latitude="24.27" location="Cave I" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" pageId="44" pageNumber="45" specimenCount="14" specimenCount-female="8" specimenCount-male="6" stateProvince="Guangxi Zhuang Autonomous Region" typeStatus="paratype">
<typeStatus id="54878819BF14FF85FDA0461320D8F865" box="[584,696,1910,1936]" pageId="44" pageNumber="45" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D3AFD32BF14FF85FD2B4610217AF87A" box="[707,794,1909,1935]" pageId="44" pageNumber="45" type="male">6 males</specimenCount>
and
<specimenCount id="9D3AFD32BF14FF85FCBB461021A3F87A" box="[851,963,1909,1935]" pageId="44" pageNumber="45" type="female">8 females</specimenCount>
, same data as holotype, in
<collectionCode id="ED2DAE7EBF14FF85FB1D46102723F87A" box="[1269,1347,1909,1935]" collectionName="SCAU" pageId="44" pageNumber="45">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B543CE6BF14FF85FAB846102776F846" ID-GBIF-Occurrence="4030741316" collectingDate="2015-07-27" collectingDateMax="2018-09-25" collectingDateMin="2015-07-27" collectionCode="SCAU" collectorName="Xinhui Wang &amp; Mingruo Tang &amp; Jujian Chen" country="China" county="Du'an Yao Autonomous County" elevation="333" latitude="24.27" location="Cave I" longLatPrecision="751" longitude="108.23" municipality="Autonomous Region" pageId="44" pageNumber="45" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Guangxi Zhuang Autonomous Region">
<specimenCount id="9D3AFD32BF14FF85FAB8461027FCF87A" box="[1360,1436,1909,1935]" pageId="44" pageNumber="45" type="male">2 male</specimenCount>
and
<specimenCount id="9D3AFD32BF14FF85FF2F46FC234AF846" box="[199,298,1945,1971]" pageId="44" pageNumber="45" type="female">1 female</specimenCount>
, same cave,
<date id="FF82107BBF14FF85FE5F46FC2027F846" box="[439,583,1945,1972]" pageId="44" pageNumber="45" value="2015-07-27">
<collectingDate id="EFC6E993BF14FF85FE5F46FC2027F846" box="[439,583,1945,1972]" pageId="44" pageNumber="45" value="2015-07-27">2015-VII-27</collectingDate>
</date>
, leg.
<collectorName id="26C9536DBF14FF85FD6B46FC217BF841" box="[643,795,1945,1972]" pageId="44" pageNumber="45">Xinhui Wang</collectorName>
,
<collectorName id="26C9536DBF14FF85FCCD46FC21A6F841" box="[805,966,1945,1972]" pageId="44" pageNumber="45">Mingruo Tang</collectorName>
&amp;
<collectorName id="26C9536DBF14FF85FC0146FC260EF846" box="[1001,1134,1945,1971]" pageId="44" pageNumber="45">Jujian Chen</collectorName>
leg., in
<collectionCode id="ED2DAE7EBF14FF85FB2046FC2776F846" box="[1224,1302,1945,1971]" collectionName="SCAU" pageId="44" pageNumber="45">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B543CE6BF14FF85FACA46FC2639F822" ID-GBIF-Occurrence="4030741317" collectingDate="2017-03-17" collectionCode="SCAU" collectorName="Sunbin Huang &amp; Pingjing Yang &amp; Dianmei Wang" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D3AFD32BF14FF85FACA46FC270DF846" box="[1314,1389,1945,1971]" pageId="44" pageNumber="45" type="male">1 male</specimenCount>
and
<specimenCount id="9D3AFD32BF14FF85FF7F46D8229BF822" box="[151,251,1981,2007]" pageId="44" pageNumber="45" type="female">1 female</specimenCount>
,
<date id="FF82107BBF14FF85FEEE46D823EBF822" box="[262,395,1981,2007]" pageId="44" pageNumber="45" value="2017-03-17">
<collectingDate id="EFC6E993BF14FF85FEEE46D823EBF822" box="[262,395,1981,2007]" pageId="44" pageNumber="45" value="2017-03-17">2017-III-17</collectingDate>
</date>
, leg.
<collectorName id="26C9536DBF14FF85FE2F46D8200BF82D" box="[455,619,1981,2008]" pageId="44" pageNumber="45">Sunbin Huang</collectorName>
,
<collectorName id="26C9536DBF14FF85FD9E46D82175F82D" box="[630,789,1981,2008]" pageId="44" pageNumber="45">Pingjing Yang</collectorName>
&amp;
<collectorName id="26C9536DBF14FF85FCD146D82184F82D" box="[825,996,1981,2008]" pageId="44" pageNumber="45">Dianmei Wang</collectorName>
, in
<collectionCode id="ED2DAE7EBF14FF85FBE346D82639F822" box="[1035,1113,1981,2007]" collectionName="SCAU" pageId="44" pageNumber="45">SCAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF15FF84FF2F41F32739FF20" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8B8336BBBF15FF84FF2F41F32739FF20" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
<emphasis id="B948EAA9BF15FF84FF2F41F32322FF45" bold="true" box="[199,322,150,176]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
Large sized cavernicolous trechines, not pubescent, head and pronotum with a few sparse setae, elytra glabrous, antennae very long, with the apical part (9
<superScript id="7C499BF3BF15FF84FD3641DF208BFF3D" attach="left" box="[734,747,186,200]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
to 11
<superScript id="7C499BF3BF15FF84FCC241DF2157FF3D" attach="left" box="[810,823,186,200]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
antennomeres) extended over the apex of elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF15FF84FF2F41BB21A3F97B" pageId="45" pageNumber="46" type="description">
<paragraph id="8B8336BBBF15FF84FF2F41BB263FFF0C" blockId="45.[151,1437,150,2037]" box="[199,1119,222,249]" pageId="45" pageNumber="46">
<emphasis id="B948EAA9BF15FF84FF2F41BB2339FF0D" bold="true" box="[199,345,222,248]" pageId="45" pageNumber="46">Description.</emphasis>
Length:
<quantity id="4CC49B5EBF15FF84FE5641BA2028FF0C" box="[446,584,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.6" metricValueMin="8.1" pageId="45" pageNumber="46" unit="mm" value="8.35" valueMax="8.6" valueMin="8.1">8.1–8.6 mm</quantity>
; width:
<quantity id="4CC49B5EBF15FF84FD4A41BA2128FF0C" box="[674,840,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" metricValueMax="2.13" metricValueMin="1.89" pageId="45" pageNumber="46" unit="mm" value="2.01" valueMax="2.13" valueMin="1.89">1.89–2.13 mm</quantity>
. Habitus as in
<figureCitation id="13072A3EBF15FF84FC0341BA263BFF0C" box="[1003,1115,223,249]" captionStart="FIGURE 29" captionStartId="46.[151,250,1943,1967]" captionTargetId="figure-28@46.[346,1241,181,1920]" captionTargetPageId="46" captionText="FIGURE 29. Habitus of Dongodytes (s. str.) mingruoae n. sp., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7647034" httpUri="https://zenodo.org/record/7647034/files/figure.png" pageId="45" pageNumber="46">Figure 29</figureCitation>
.
</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F406623AEFE7D" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">Brown or light yellowish brown, apical half of antennae, mouthparts palps and tarsi pale. Underside of head, pro- and mesothorax covered sparsely with rather long setae; ventrites densely pubescent. Microsculpture: transverse striate on head and pronotum, dense isodiametric engraved meshes on elytra. Moderately shiny. Fore body including mandibles as long as elytra.</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F40F72043FC89" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
Head elongate, much longer than wide, HLm/HW=3.22–3.47, HLl/HW= 2.54–2.68; widest at about frontal 1/3 of head excluding mandibles, much longer than pronotum, HLl/PnL = 1.48–1.65; genae hardly expanded, gradually narrowed posteriad, but strongly narrowed from basal 1/3, neck constriction fairly long, about 1/3 as wide as head; frons flat, vertex convex; clypeus transverse, 6-setose; labrum moderately sinuate at frontal margin, 6-setose; two pairs of supraorbital setae present, the anterior two at about the widest point, while the posterior two much shorter, only slightly longer than the secondary setae and so not easy to be noticed, at about 3/8 of head from base; a pair of suborbital setae present at about 1/4 from base; mentum and submentum well separated by a clear labial suture, mentum bisetose, mental tooth small and short, simple at tip; submentum 9-setose; mouthparts palps very thin and slender, glabrous except 2
<superScript id="7C499BF3BF15FF84FE5443CA23ACFD48" attach="left" box="[444,460,687,701]" fontSize="6" pageId="45" pageNumber="46">nd</superScript>
labial palpomere which is bisetose on inner margin; the 2
<superScript id="7C499BF3BF15FF84FBB243CA260AFD48" attach="left" box="[1114,1130,687,701]" fontSize="6" pageId="45" pageNumber="46">nd</superScript>
labial palpomere 1.4 times longer than 3
<superScript id="7C499BF3BF15FF84FEC543B6235BFD14" attach="none" box="[301,315,723,737]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
; the 3
<superScript id="7C499BF3BF15FF84FE6C43B623F2FD14" attach="left" box="[388,402,723,737]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
maxillary palpomere 1.1 times longer than 4
<superScript id="7C499BF3BF15FF84FC7A43B621FFFD14" attach="none" box="[914,927,723,737]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
; ligula 6 or 8-setose; antennae filiform, very slender and long, apical three antennomeres extended at over elytral apices; pubescent from pedicel, scape stouter and shorter than other, with several long setae; the 4
<superScript id="7C499BF3BF15FF84FD30427E2085FCDC" attach="left" box="[728,741,795,809]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
the longest; the comparative length of each antennomere from the 1
<superScript id="7C499BF3BF15FF84FF26425A22B9FCB8" attach="left" box="[206,217,831,845]" fontSize="6" pageId="45" pageNumber="46">st</superScript>
to 11
<superScript id="7C499BF3BF15FF84FEF0425B2345FCB9" attach="left" box="[280,293,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
in
<typeStatus id="54878819BF15FF84FEA0425A23CAFCAC" box="[328,426,831,857]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
as: 1
<superScript id="7C499BF3BF15FF84FE0D425A2390FCB8" attach="left" box="[485,496,831,845]" fontSize="6" pageId="45" pageNumber="46">st</superScript>
(1.00) / 2
<superScript id="7C499BF3BF15FF84FDB6425B200EFCB9" attach="left" box="[606,622,830,844]" fontSize="6" pageId="45" pageNumber="46">nd</superScript>
(1.23) / 3
<superScript id="7C499BF3BF15FF84FD35425B208BFCB9" attach="left" box="[733,747,830,844]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
(2.51) / 4
<superScript id="7C499BF3BF15FF84FCB1425B2106FCB9" attach="left" box="[857,870,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(2.60) / 5
<superScript id="7C499BF3BF15FF84FC3C425B2181FCB9" attach="left" box="[980,993,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.70) / 6
<superScript id="7C499BF3BF15FF84FBA7425B263CFCB9" attach="left" box="[1103,1116,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.97) / 7
<superScript id="7C499BF3BF15FF84FB22425B26B7FCB9" attach="left" box="[1226,1239,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.77) / 8
<superScript id="7C499BF3BF15FF84FAAD425B2732FCB9" attach="left" box="[1349,1362,830,844]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.49) / 9
<superScript id="7C499BF3BF15FF84FF5C420722A1FC85" attach="left" box="[180,193,866,880]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.23) / 10
<superScript id="7C499BF3BF15FF84FED54207232AFC85" attach="left" box="[317,330,866,880]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.11) / 11
<superScript id="7C499BF3BF15FF84FE2C420723B1FC85" attach="left" box="[452,465,866,880]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
(1.23).
</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F42E326ABFB83" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">Prothorax elongate, propleura strongly expanded and visible from above except base and front, longer than wide, PrL/PrW =1.41–1.70; slightly wider than head, PrW/HW= 1.10–1.21; wider than pronotum, PrW/PnW = 1.27–1.34; widest at about 1/3 from base. Pronotum elongate, much longer than wide, PnL/PnW = 1.90–2.22; slightly narrower than head, PnW/HW= 0.85–0.89; widest at about basal 1/3, slightly sinuate just before hind angles which are bluntly rectangular, front angles obtuse; base distinctly wider than front, PbW/PfW= 1.23–1.35; both anterior and posterior latero-marginal setae present, located at about 4/7 from base and a little before hind angles respectively; posterior transversal impression well-marked. Scutellum large, in inverse triangle, nearly as long as wide.</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F45E523DDFA84" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
Elytra very elongate, much longer than wide, EL/EW=2.05–2.24, much wider than prothorax; apex wider than base; lateral margins finely bordered, smooth and glabrous throughout; base slightly narrower than base of pronotum, humeral parts nearly straight, humeral angles widely rounded off; widest at about 2/3 from base; disc moderately convex except the basal area which is evidently depressed; striae indistinct, only the 1
<superScript id="7C499BF3BF15FF84FB02458E2695FB0C" attach="left" box="[1258,1269,1259,1273]" fontSize="6" pageId="45" pageNumber="46">st</superScript>
and 2
<superScript id="7C499BF3BF15FF84FAD3458E272BFB0C" attach="left" box="[1339,1355,1259,1273]" fontSize="6" pageId="45" pageNumber="46">nd</superScript>
faintly traceable; chaetotaxy (
<figureCitation id="13072A3EBF15FF84FE7D446A207CFADC" box="[405,540,1295,1321]" captionStart="FIGURE 27" captionStartId="42.[151,250,1965,1989]" captionTargetBox="[280,1306,453,1927]" captionTargetId="figure-129@42.[280,1307,453,1942]" captionTargetPageId="42" captionText="FIGURE 27. Left elytra (white dots indicating the chaetotaxy) of Dongodytes species (white dots indicating the chaetotaxy). A. Dongodytes (Dongodytodes) pingjingae sp. n.; B. Dongodytes (s. str.) mingruoae n. sp." figureDoi="http://doi.org/10.5281/zenodo.7642446" httpUri="https://zenodo.org/record/7642446/files/figure.png" pageId="45" pageNumber="46">Figure 27B</figureCitation>
): two dorsal pores present on the site of the 3
<superScript id="7C499BF3BF15FF84FBCA446A2650FAE8" attach="left" box="[1058,1072,1295,1309]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
stria, at about basal 2/7 and 4/7 respectively; preapical pore present at apical 1/7 of elytron, pattern of the marginal umbilicate pores similar in
<taxonomicName id="4C3C4D38BF15FF84FA69445123D8FA84" authority="Tian, 2011" authorityName="Tian" authorityYear="2011" class="Insecta" family="Carabidae" genus="Dongodytes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="baxian">
<emphasis id="B948EAA9BF15FF84FA6944512357FA84" italics="true" pageId="45" pageNumber="46">D. (s. str.) baxian</emphasis>
Tian, 2011
</taxonomicName>
.
</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F441E21ADFA4D" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
Legs thin and very slender, tibiae not longitudinally sulcate; the 1
<superScript id="7C499BF3BF15FF84FC58441F21DBFA7D" attach="left" box="[944,955,1402,1416]" fontSize="6" pageId="45" pageNumber="46">st</superScript>
tarsomere subequal to 2
<superScript id="7C499BF3BF15FF84FB39441F2681FA7D" attach="left" box="[1233,1249,1402,1416]" fontSize="6" pageId="45" pageNumber="46">nd</superScript>
– 4
<superScript id="7C499BF3BF15FF84FAE6441F277BFA7D" attach="left" box="[1294,1307,1402,1416]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
tarsomeres combined in fore leg, while slightly longer those in middle and hind legs.
</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F44A72087FA29" blockId="45.[151,1437,150,2037]" box="[199,743,1474,1500]" pageId="45" pageNumber="46">Ventrite IV–VII each quadrisetose in both sexes.</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F448021A3F97B" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
Male genitalia (
<figureCitation id="13072A3EBF15FF84FE934480238FF9F5" box="[379,495,1509,1536]" captionStart="FIGURE 30" captionStartId="47.[151,250,1867,1891]" captionTargetBox="[161,1422,193,1831]" captionTargetId="figure-20@47.[160,1428,181,1844]" captionTargetPageId="47" captionText="FIGURE 30. Dongodytes (s. str.) mingruoae n. sp. A. median lobe, lateral view; B. median lobe, dorsal view. Scale for both." figureDoi="http://doi.org/10.5281/zenodo.7642450" httpUri="https://zenodo.org/record/7642450/files/figure.png" pageId="45" pageNumber="46">Figure 30</figureCitation>
): Median lobe of aedeagus moderately elongate, nearly straight ventrally in apical half though shortly sinuate just before apex, which is blunt; basal orifice large, with a very small sagittal aileron; inner sac armed with a large copulatory piece which covered with scale structures on surface, as long as 1/4 of the median lobe; in dorsal, apical lobe short and narrow, rounded at apex; parameres very elongate and clavoid, evenly developed, each with 5 long and one or two short setae at apical portion.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF15FF84FF2F47FD208BF90F" pageId="45" pageNumber="46" type="discussion">
<paragraph id="8B8336BBBF15FF84FF2F47FD208BF90F" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
<emphasis id="B948EAA9BF15FF84FF2F47FD235BF947" bold="true" box="[199,315,1688,1714]" pageId="45" pageNumber="46">Remarks.</emphasis>
Similar to
<taxonomicName id="4C3C4D38BF15FF84FE5F47FD2116F947" authority="(Tian, 2010)" baseAuthorityName="Tian" baseAuthorityYear="2010" box="[439,886,1688,1714]" class="Insecta" family="Carabidae" genus="Dongodytes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="baxian">
<emphasis id="B948EAA9BF15FF84FE5F47FD205DF947" box="[439,573,1688,1714]" italics="true" pageId="45" pageNumber="46">Dongodytes</emphasis>
(
<emphasis id="B948EAA9BF15FF84FDA547FC20E7F947" box="[589,647,1689,1714]" italics="true" pageId="45" pageNumber="46">s. str.</emphasis>
)
<emphasis id="B948EAA9BF15FF84FD7E47FD2082F947" box="[662,738,1688,1714]" italics="true" pageId="45" pageNumber="46">baxian</emphasis>
(Tian, 2010)
</taxonomicName>
, but easily separated from the latter by its thinner head with longer and narrower neck constriction, the median lobe of aedeagus thinner, and narrower at apical part, and ventrite VII 4-setose in male, instead of bisetose.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF15FF84FF2F4661265DF8EB" box="[199,1085,1795,1822]" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8B8336BBBF15FF84FF2F4661265DF8EB" blockId="45.[151,1437,150,2037]" box="[199,1085,1795,1822]" pageId="45" pageNumber="46">
<emphasis id="B948EAA9BF15FF84FF2F4661232EF8EB" bold="true" box="[199,334,1796,1822]" pageId="45" pageNumber="46">Etymology.</emphasis>
Named after Ms. Mingruo Tang, a member of SCAU caving team.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF15FF84FF2F46422752F800" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8B8336BBBF15FF84FF2F464226CBF8B4" blockId="45.[151,1437,150,2037]" box="[199,1195,1831,1858]" pageId="45" pageNumber="46">
<emphasis id="B948EAA9BF15FF84FF2F46422301F8B4" bold="true" box="[199,353,1831,1857]" pageId="45" pageNumber="46">Distribution.</emphasis>
<collectingRegion id="49F8F859BF15FF84FE8F464223A7F8B4" box="[359,455,1831,1857]" country="China" name="Guangxi" pageId="45" pageNumber="46">Guangxi</collectingRegion>
(Du’an). Known only from the Cave I in Jiudu Xiang (
<figureCitation id="13072A3EBF15FF84FBDB464226FBF8B4" box="[1075,1179,1831,1857]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="45" pageNumber="46">Figure 1j</figureCitation>
).
</paragraph>
<paragraph id="8B8336BBBF15FF84FF2F462E2752F800" blockId="45.[151,1437,150,2037]" pageId="45" pageNumber="46">
Cave I opens on a hillside not far from a village. It is about 200 metres long, from the large opening to its bottom, through a steep slope. It is surrounded by big trees and dense bushes but sun light can reach most parts of the cave. There is a large population of
<taxonomicName id="4C3C4D38BF15FF84FE9E46F620F2F859" authority="Levl., 1909" authorityName="Levl." authorityYear="1909" box="[374,658,1938,1964]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="edulis">
<emphasis id="B948EAA9BF15FF84FE9E46F62074F859" box="[374,532,1938,1964]" italics="true" pageId="45" pageNumber="46">Begonia edulis</emphasis>
Lévl., 1909
</taxonomicName>
(
<figureCitation id="13072A3EBF15FF84FD7646F72159F859" box="[670,825,1938,1964]" captionStart="FIGURE 31" captionStartId="48.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-28@48.[151,1435,181,1845]" captionTargetPageId="48" captionText="FIGURE 31. Jiudu Cave I, the type locality of Dongodytes (s. str.) mingruoae n. sp. A. entrance; B. several plants of Begonia edulis Lévl., 1909; C. a running specimen of Dongodytes (s. str.) mingruoae n. sp. in cave." figureDoi="http://doi.org/10.5281/zenodo.7642452" httpUri="https://zenodo.org/record/7642452/files/figure.png" pageId="45" pageNumber="46">Figure 31A, B</figureCitation>
). There are droplets of water in some places at half way or near the bottom. The blind beetles were discovered lower half of the cave (
<figureCitation id="13072A3EBF15FF84FC4746D3264CF825" box="[943,1068,1974,2000]" captionStart="FIGURE 31" captionStartId="48.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-28@48.[151,1435,181,1845]" captionTargetPageId="48" captionText="FIGURE 31. Jiudu Cave I, the type locality of Dongodytes (s. str.) mingruoae n. sp. A. entrance; B. several plants of Begonia edulis Lévl., 1909; C. a running specimen of Dongodytes (s. str.) mingruoae n. sp. in cave." figureDoi="http://doi.org/10.5281/zenodo.7642452" httpUri="https://zenodo.org/record/7642452/files/figure.png" pageId="45" pageNumber="46">Figure 31C</figureCitation>
). Apart from
<taxonomicName id="4C3C4D38BF15FF84FB5246D227FCF825" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[1210,1436,1974,2000]" class="Insecta" family="Carabidae" genus="Dongodytes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="mingruoae" status="sp. nov.">
<emphasis id="B948EAA9BF15FF84FB5246D227FCF825" box="[1210,1436,1974,2000]" italics="true" pageId="45" pageNumber="46">D. (s. str.) mingruoae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF15FF84FF7F46BF22B3F801" bold="true" box="[151,211,2010,2036]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A27B57D2BF15FF84FF7F46BF22B3F801" box="[151,211,2010,2036]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C3C4D38BF15FF84FF0846BF2072F801" authorityName="Tian" authorityYear="2011" box="[224,530,2010,2036]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Dongodytodes">
<emphasis id="B948EAA9BF15FF84FF0846BF2072F801" box="[224,530,2010,2036]" italics="true" pageId="45" pageNumber="46">Dongodytes (Dongodytodes)</emphasis>
</taxonomicName>
sp. and
<emphasis id="B948EAA9BF15FF84FD8246BF216BF801" box="[618,779,2010,2036]" italics="true" pageId="45" pageNumber="46">Eutrichodemus</emphasis>
millipede species also occur in this limestone cave.
</paragraph>
</subSubSection>
</treatment>
</document>
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