326 lines
39 KiB
XML
326 lines
39 KiB
XML
<document id="DDEF27215E1E1A7CF87B56DC982D1B7B" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin & Jia, Xinyang" docDate="2023" docId="039587ADBF0DFF91FF7F41BA21F5FC3C" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Shilinotrechus zhangfani Tian & Huang & Jia 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="57" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="54" updateTime="1731937328377" updateUser="felipe">
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<mods:title id="D134BF1783E75F9DBFF7DD86BD5B8C84">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="CE57BC7FD7ACC8DD76A8CCD70A31E7BB">Tian, Mingyi</mods:namePart>
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<mods:namePart id="9842E217A3FBEA53722931C67ABB9CBA">Huang, Sunbin</mods:namePart>
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<mods:namePart id="3683F1CAEC1A8E7EB6AB10D91293ECC5">Jia, Xinyang</mods:namePart>
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<mods:date id="4F4F723494B6BFA191D73BFCB5F47E48">2023</mods:date>
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<mods:number id="E1AED30B592BC9B6A3887BCA95A2EAAA">2023-02-15</mods:number>
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<treatment id="039587ADBF0DFF91FF7F41BA21F5FC3C" ID-DOI="http://doi.org/10.5281/zenodo.7642392" ID-GBIF-Taxon="209296922" ID-Zenodo-Dep="7642392" LSID="urn:lsid:plazi:treatment:039587ADBF0DFF91FF7F41BA21F5FC3C" httpUri="http://treatment.plazi.org/id/039587ADBF0DFF91FF7F41BA21F5FC3C" lastPageId="56" lastPageNumber="57" pageId="53" pageNumber="54">
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<subSubSection id="C3266530BF0DFF9CFF7F41BA2074FF0C" box="[151,532,223,250]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="8B8336BBBF0DFF9CFF7F41BA2074FF0C" blockId="53.[151,532,223,286]" box="[151,532,223,250]" pageId="53" pageNumber="54">
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<heading id="D0CB81D7BF0DFF9CFF7F41BA2074FF0C" bold="true" box="[151,532,223,250]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
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<emphasis id="B948EAA9BF0DFF9CFF7F41BA2074FF0C" bold="true" box="[151,532,223,250]" pageId="53" pageNumber="54">
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<taxonomicName id="4C3C4D38BF0DFF9CFF7F41BA23AAFF0F" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[151,458,223,250]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="zhangfani" status="sp. nov.">
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<emphasis id="B948EAA9BF0DFF9CFF7F41BA23AAFF0F" bold="true" box="[151,458,223,250]" italics="true" pageId="53" pageNumber="54">Shilinotrechus zhangfani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B57D2BF0DFF9CFE3A41BA2074FF0C" box="[466,532,223,249]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3266530BF0DFF9CFF7F40662302FEEB" box="[151,354,259,286]" pageId="53" pageNumber="54" type="description">
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<paragraph id="8B8336BBBF0DFF9CFF7F40662302FEEB" blockId="53.[151,532,223,286]" box="[151,354,259,286]" pageId="53" pageNumber="54">
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<figureCitation id="13072A3EBF0DFF9CFF7F4066236EFEE8" box="[151,270,259,285]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="53" pageNumber="54">Figures 1a</figureCitation>
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,
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<figureCitation id="13072A3EBF0DFF9CFEF340662302FEEB" box="[283,354,259,286]" captionStart-0="FIGURE 35" captionStart-1="FIGURE 36" captionStart-2="FIGURE 37" captionStartId-0="53.[151,250,2007,2031]" captionStartId-1="54.[151,250,1974,1998]" captionStartId-2="55.[151,250,1919,1943]" captionTargetBox-0="[240,1347,644,1970]" captionTargetBox-1="[153,1433,1066,1950]" captionTargetBox-2="[151,1435,181,1894]" captionTargetId-0="figure-137@53.[225,1362,633,1983]" captionTargetId-1="figure-503@54.[153,1434,1065,1950]" captionTargetId-2="figure-20@55.[151,1435,181,1894]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionTargetPageId-2="55" captionText-0="FIGURE 35. Habitus of Shilinotrechus zhangfani n. sp., male, holotype." captionText-1="FIGURE 36. Shilinotrechus zhangfani n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." captionText-2="FIGURE 37. Xianren Dong, the type locality of Shilinotrechus zhangfani n. sp. A. entrance; B. speleothems in cave; C. a running specimen of Shilinotrechus zhangfani n. sp.; D & E. millipedes of the genera Eutrichodesmus and Glyphiulus; F. mould growing in cave." figureDoi-0="http://doi.org/10.5281/zenodo.7642460" figureDoi-1="http://doi.org/10.5281/zenodo.7642462" figureDoi-2="http://doi.org/10.5281/zenodo.7642464" httpUri-0="https://zenodo.org/record/7642460/files/figure.png" httpUri-1="https://zenodo.org/record/7642462/files/figure.png" httpUri-2="https://zenodo.org/record/7642464/files/figure.png" pageId="53" pageNumber="54">35−37</figureCitation>
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</paragraph>
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<subSubSection id="C3266530BF0DFF9CFF7F402E228AFE58" pageId="53" pageNumber="54" type="materials_examined">
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<paragraph id="8B8336BBBF0DFF9CFF7F402E228AFE58" blockId="53.[151,1436,331,610]" pageId="53" pageNumber="54">
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<emphasis id="B948EAA9BF0DFF9CFF7F402E2366FE90" bold="true" box="[151,262,331,357]" pageId="53" pageNumber="54">Material.</emphasis>
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<materialsCitation id="3B543CE6BF0DFF9CFEE3402E228AFE58" ID-GBIF-Occurrence="4030741321" collectingDate="2020-08-17" collectionCode="SCAU" collectorName="Mingyi Tian & Jingli Cheng" county="cave Xianren Dong" elevation="1901" latitude="24.48" location="Mile Shi" longLatPrecision="750" longitude="103.2" municipality="Xi'er Township" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
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<typeStatus id="54878819BF0DFF9CFEE3402E2313FE90" box="[267,371,331,357]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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male,
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<collectingCounty id="62E24E37BF0DFF9CFE52402920F2FE93" box="[442,658,331,358]" pageId="53" pageNumber="54">cave Xianren Dong</collectingCounty>
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,
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<collectingMunicipality id="6BE7ACC1BF0DFF9CFD72402E212BFE90" box="[666,843,331,357]" pageId="53" pageNumber="54">Xi’er Township</collectingMunicipality>
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,
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<location id="8EE36060BF0DFF9CFCBB402E21D5FE90" LSID="urn:lsid:plazi:treatment:039587ADBF0DFF91FF7F41BA21F5FC3C:8EE36060BF0DFF9CFCBB402E21D5FE90" box="[851,949,331,357]" county="cave Xianren Dong" latitude="24.48" longLatPrecision="750" longitude="103.2" municipality="Xi'er Township" name="Mile Shi" pageId="53" pageNumber="54" stateProvince="Yunnan Province">Mile Shi</location>
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,
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<collectingCounty id="62E24E37BF0DFF9CFC56402E27F8FE90" box="[958,1432,331,357]" pageId="53" pageNumber="54">Honghe Hani &Yi Autonomous Prefecture</collectingCounty>
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,
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<collectingRegion id="49F8F859BF0DFF9CFF7F4015233BFE7C" box="[151,347,367,394]" country="China" name="Yunnan" pageId="53" pageNumber="54">Yunnan Province</collectingRegion>
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,
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<geoCoordinate id="EE08507CBF0DFF9CFE80400A23AFFE7C" box="[360,463,367,393]" degrees="103.20" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="103.2">103.20 E</geoCoordinate>
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,
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<geoCoordinate id="EE08507CBF0DFF9CFE3B400A2054FE7F" box="[467,564,367,394]" degrees="24.48" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="24.48">24.48 N</geoCoordinate>
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,
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<elevation id="0011D188BF0DFF9CFDA9400A212FFE7C" box="[577,847,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.9009999999999998" pageId="53" pageNumber="54" unit="m" value="1901.0">
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<quantity id="4CC49B5EBF0DFF9CFDA9400A20F7FE7F" box="[577,663,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.9009999999999998" pageId="53" pageNumber="54" unit="m" value="1901.0">1901 m</quantity>
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above sea level
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</elevation>
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,
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<date id="FF82107BBF0DFF9CFCB4400A2195FE7C" box="[860,1013,367,393]" pageId="53" pageNumber="54" value="2020-08-17">
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<collectingDate id="EFC6E993BF0DFF9CFCB4400A2195FE7C" box="[860,1013,367,393]" pageId="53" pageNumber="54" value="2020-08-17">2020-VIII-17</collectingDate>
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</date>
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,
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<collectorName id="26C9536DBF0DFF9CFBEA400A26EFFE7C" box="[1026,1167,367,393]" pageId="53" pageNumber="54">Mingyi Tian</collectorName>
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&
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<collectorName id="26C9536DBF0DFF9CFB5E400A2725FE7C" box="[1206,1349,367,393]" pageId="53" pageNumber="54">Jingli Cheng</collectorName>
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leg., in
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<collectionCode id="ED2DAE7EBF0DFF9CFF7F40F62285FE58" box="[151,229,403,429]" collectionName="SCAU" pageId="53" pageNumber="54">SCAU</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF0DFF9CFF2F40D22732FE24" box="[199,1362,439,466]" pageId="53" pageNumber="54" type="diagnosis">
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<paragraph id="8B8336BBBF0DFF9CFF2F40D22732FE24" blockId="53.[151,1436,331,610]" box="[199,1362,439,466]" pageId="53" pageNumber="54">
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<emphasis id="B948EAA9BF0DFF9CFF2F40D22322FE24" bold="true" box="[199,322,439,465]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
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Small-sized and stout trechine species, eyeless and depigmented, body concolourous brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF0DFF91FF2F40BE23FFFDC8" lastPageId="56" lastPageNumber="57" pageId="53" pageNumber="54" type="description">
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<paragraph id="8B8336BBBF0DFF9CFF2F40BE2181FE03" blockId="53.[151,1436,331,610]" box="[199,993,475,502]" pageId="53" pageNumber="54">
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<emphasis id="B948EAA9BF0DFF9CFF2F40BE2339FE00" bold="true" box="[199,345,475,501]" pageId="53" pageNumber="54">Description.</emphasis>
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Length:
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<quantity id="4CC49B5EBF0DFF9CFE5640BE2077FE03" box="[446,535,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="53" pageNumber="54" unit="mm" value="4.9">4.9 mm</quantity>
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; width:
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<quantity id="4CC49B5EBF0DFF9CFD9940BE20AAFE03" box="[625,714,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="53" pageNumber="54" unit="mm" value="1.8">1.8 mm</quantity>
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. Habitus as in
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<figureCitation id="13072A3EBF0DFF9CFC8540BE21BDFE03" box="[877,989,475,502]" captionStart="FIGURE 35" captionStartId="53.[151,250,2007,2031]" captionTargetBox="[240,1347,644,1970]" captionTargetId="figure-137@53.[225,1362,633,1983]" captionTargetPageId="53" captionText="FIGURE 35. Habitus of Shilinotrechus zhangfani n. sp., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7642460" httpUri="https://zenodo.org/record/7642460/files/figure.png" pageId="53" pageNumber="54">Figure 35</figureCitation>
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.
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</paragraph>
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<paragraph id="8B8336BBBF0DFF9CFF2F409A2602FD94" blockId="53.[151,1436,331,610]" pageId="53" pageNumber="54">Body stout and strongly convex on dorsum; concolourous brown, but antennae, palps and tarsi slightly lighter; surface with fine and sparse pubescence; microsculpture engraved meshes irregularly isodiametric or polygonal on head, pronotum and elytra. For body much shorter than elytra, (HLm+PL) / EL = 0.71.</paragraph>
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<caption id="DF436633BF0DFF9CFF7F46B2211CF81A" ID-DOI="http://doi.org/10.5281/zenodo.7642460" ID-Zenodo-Dep="7642460" box="[151,892,2007,2031]" httpUri="https://zenodo.org/record/7642460/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,250,2007,2031]" targetBox="[240,1347,644,1970]" targetPageId="53">
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<paragraph id="8B8336BBBF0DFF9CFF7F46B2211CF81A" blockId="53.[151,892,2007,2031]" box="[151,892,2007,2031]" pageId="53" pageNumber="54">
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<emphasis id="B948EAA9BF0DFF9CFF7F46B22340F81A" bold="true" box="[151,288,2007,2031]" pageId="53" pageNumber="54">FIGURE 35.</emphasis>
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Habitus of
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<taxonomicName id="4C3C4D38BF0DFF9CFE7F46B220F2F81A" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[407,658,2007,2031]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="zhangfani" status="sp. nov.">
|
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<emphasis id="B948EAA9BF0DFF9CFE7F46B220F2F81A" box="[407,658,2007,2031]" italics="true" pageId="53" pageNumber="54">Shilinotrechus zhangfani</emphasis>
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</taxonomicName>
|
||
<emphasis id="B948EAA9BF0DFF9CFD7146B220B2F81A" bold="true" box="[665,722,2007,2031]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="A27B57D2BF0DFF9CFD7146B220B2F81A" box="[665,722,2007,2031]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
, male, holotype.
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||
</paragraph>
|
||
</caption>
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<paragraph id="8B8336BBBF0EFF9FFF2F41F2263BFDC8" blockId="54.[151,1437,151,1041]" pageId="54" pageNumber="55">
|
||
Head slightly longer than wide, HLm/HW = 1.67, HLl/HW = 1.22; widest at about middle excluding mandibles, frontal furrows widened, nearly parallel-sided; two pairs of supraorbital pores present; clypeus 4-setose; labrum distinctly emarginate, 6-setose; right mandibular tridentate; mentum and submentum separated; mentum bisetose, base bi-foveate, tooth thick and short, bluntly bifid at tip; submentum sharply spined medially, 6-setose; mouthparts palps very thin and elongated; apical one in maxillae widened at base, narrowed towards apex, reverse in penultimate one; apical one in labium similar to that in maxillae, while penultimate one gently narrowed towards base; the 2
|
||
<superScript id="7C499BF3BF0EFF9FFF4D400B22D5FE89" attach="left" box="[165,181,366,380]" fontSize="6" pageId="54" pageNumber="55">nd</superScript>
|
||
labial palpomere bisetose on inner margin, with three additional setae near apical portion, the 3
|
||
<superScript id="7C499BF3BF0EFF9FFAF1400B2747FE89" attach="left" box="[1305,1319,366,380]" fontSize="6" pageId="54" pageNumber="55">rd</superScript>
|
||
maxillary palpomere with four short set ae on apical half, other palps smooth and glabrous; the 2
|
||
<superScript id="7C499BF3BF0EFF9FFBB540F7260DFE55" attach="left" box="[1117,1133,402,416]" fontSize="6" pageId="54" pageNumber="55">nd</superScript>
|
||
labial palpomere 1.2 times longer than 3
|
||
<superScript id="7C499BF3BF0EFF9FFEC340D32359FE31" attach="none" box="[299,313,438,452]" fontSize="6" pageId="54" pageNumber="55">rd</superScript>
|
||
; the 3
|
||
<superScript id="7C499BF3BF0EFF9FFE9740D323EDFE31" attach="left" box="[383,397,438,452]" fontSize="6" pageId="54" pageNumber="55">rd</superScript>
|
||
maxillary palpomere as long as 4
|
||
<superScript id="7C499BF3BF0EFF9FFCEE40D32173FE31" attach="none" box="[774,787,438,452]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
; suborbital pores present. Antennae filiform, extending to 1/3 of elytra from base; scape stout, with several setae, as long as pedicel; pedicel to 11
|
||
<superScript id="7C499BF3BF0EFF9FFB8640BF261BFE1D" attach="left" box="[1134,1147,474,488]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
antennomeres pubescent; the comparative length of each antennomere from the 1
|
||
<superScript id="7C499BF3BF0EFF9FFCEC409A216FFDF8" attach="left" box="[772,783,511,525]" fontSize="6" pageId="54" pageNumber="55">st</superScript>
|
||
to 11
|
||
<superScript id="7C499BF3BF0EFF9FFCA5409B213AFDF9" attach="left" box="[845,858,510,524]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
in
|
||
<typeStatus id="54878819BF0EFF9FFC96409A2180FDEC" box="[894,992,511,537]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
as: 1
|
||
<superScript id="7C499BF3BF0EFF9FFBF3409A2646FDF8" attach="left" box="[1051,1062,511,525]" fontSize="6" pageId="54" pageNumber="55">st</superScript>
|
||
(1,00) / 2
|
||
<superScript id="7C499BF3BF0EFF9FFB7D409B26C5FDF9" attach="left" box="[1173,1189,510,524]" fontSize="6" pageId="54" pageNumber="55">nd</superScript>
|
||
(1.00) / 3
|
||
<superScript id="7C499BF3BF0EFF9FFAFB409B2741FDF9" attach="left" box="[1299,1313,510,524]" fontSize="6" pageId="54" pageNumber="55">rd</superScript>
|
||
(1.36) / 4
|
||
<superScript id="7C499BF3BF0EFF9FFA78409B27FDFDF9" attach="none" box="[1424,1437,510,524]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.41) / 5
|
||
<superScript id="7C499BF3BF0EFF9FFF174347236CFDC5" attach="left" box="[255,268,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.41) / 6
|
||
<superScript id="7C499BF3BF0EFF9FFE92434723E7FDC5" attach="left" box="[378,391,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.41) / 7
|
||
<superScript id="7C499BF3BF0EFF9FFE1D43472062FDC5" attach="left" box="[501,514,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.27) / 8
|
||
<superScript id="7C499BF3BF0EFF9FFD984347201DFDC5" attach="left" box="[624,637,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.09) / 9
|
||
<superScript id="7C499BF3BF0EFF9FFD0343472098FDC5" attach="left" box="[747,760,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.09) / 10
|
||
<superScript id="7C499BF3BF0EFF9FFC9C434721E1FDC5" attach="left" box="[884,897,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.00) / 11
|
||
<superScript id="7C499BF3BF0EFF9FFC1443472669FDC5" attach="left" box="[1020,1033,546,560]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
(1.23).
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF0EFF9FFF2F432221D6FD04" blockId="54.[151,1437,151,1041]" pageId="54" pageNumber="55">Pronotum subtrapezoidal, wider than head, PW/HW = 1.33, clearly wider than long, PW/PL = 1.26; lateral margins finely bordered, widely and upwardly reflexed throughout, more so near base; hind angles obtuse, fore angles protruding; base much wider than front, PbW/PfW = 1.42; base shallowly bisinuate, slightly convex backwardly in median portion, front straight; two marginal setae present, the anterior located at about 1/6 from front, and a little after the front angles, the posterior at hind angle. Scutellum very small.</paragraph>
|
||
<paragraph id="8B8336BBBF0EFF9FFF2F439E21EEFBE4" blockId="54.[151,1437,151,1041]" pageId="54" pageNumber="55">
|
||
Elytra elongate, amygdaloidal, base more widened than apices; longer than wide, EL/EW = 1.53, much wider than pronotum, EW/PW = 1.72, more than two times longer than pronotum, EL/PL = 3.26, widest at about 1/4 from base; shoulders broadly protruding, lateral margins ciliate throughout; striae deep, the 8
|
||
<superScript id="7C499BF3BF0EFF9FFB49422726CEFCA5" attach="left" box="[1185,1198,834,848]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
stria invisible; apical recurrent striole short; chaetotaxy (
|
||
<figureCitation id="13072A3EBF0EFF9FFDC9420220C9FC74" box="[545,681,871,897]" captionStart="FIGURE 36" captionStartId="54.[151,250,1974,1998]" captionTargetBox="[153,1433,1066,1950]" captionTargetId="figure-503@54.[153,1434,1065,1950]" captionTargetPageId="54" captionText="FIGURE 36. Shilinotrechus zhangfani n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642462" httpUri="https://zenodo.org/record/7642462/files/figure.png" pageId="54" pageNumber="55">Figure 36A</figureCitation>
|
||
): basal pore present; two dorsal setiferous pores present along stria 3 before and behind middle of elytra respectively; the preapical dorsal pore present, located in the anastomosis of 3
|
||
<superScript id="7C499BF3BF0EFF9FFF4D42CB22D3FC49" attach="left" box="[165,179,942,956]" fontSize="6" pageId="54" pageNumber="55">rd</superScript>
|
||
and 4
|
||
<superScript id="7C499BF3BF0EFF9FFF1242CB2367FC49" attach="left" box="[250,263,942,956]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
striae, closer to elytral suture; the humeral group of marginal umbilicate pores not aggregated, 1
|
||
<superScript id="7C499BF3BF0EFF9FFABF42CA2702FC48" attach="left" box="[1367,1378,943,957]" fontSize="6" pageId="54" pageNumber="55">st</superScript>
|
||
pore forwardly shifted; 1
|
||
<superScript id="7C499BF3BF0EFF9FFE9C42B6231FFC14" attach="left" box="[372,383,979,993]" fontSize="6" pageId="54" pageNumber="55">st</superScript>
|
||
and 2
|
||
<superScript id="7C499BF3BF0EFF9FFE2942B723B1FC15" attach="left" box="[449,465,978,992]" fontSize="6" pageId="54" pageNumber="55">nd</superScript>
|
||
pores close to the marginal gutter; middle group (5
|
||
<superScript id="7C499BF3BF0EFF9FFBE242B72677FC15" attach="left" box="[1034,1047,978,992]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
and 6
|
||
<superScript id="7C499BF3BF0EFF9FFBB142B72606FC15" attach="left" box="[1113,1126,978,992]" fontSize="6" pageId="54" pageNumber="55">th</superScript>
|
||
pores) locate before middle of elytra, not very close to each other; the anguloapical pore absent.
|
||
</paragraph>
|
||
<caption id="DF436633BF0EFF9FFF7F46D323CBF807" ID-DOI="http://doi.org/10.5281/zenodo.7642462" ID-Zenodo-Dep="7642462" httpUri="https://zenodo.org/record/7642462/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1974,1998]" targetBox="[153,1433,1066,1950]" targetPageId="54">
|
||
<paragraph id="8B8336BBBF0EFF9FFF7F46D323CBF807" blockId="54.[151,1436,1974,2034]" pageId="54" pageNumber="55">
|
||
<emphasis id="B948EAA9BF0EFF9FFF7F46D3237FF83B" bold="true" box="[151,287,1974,1998]" pageId="54" pageNumber="55">FIGURE 36.</emphasis>
|
||
<taxonomicName id="4C3C4D38BF0EFF9FFECC46D3207FF83B" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[292,543,1974,1998]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="zhangfani" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF0EFF9FFECC46D3207FF83B" box="[292,543,1974,1998]" italics="true" pageId="54" pageNumber="55">Shilinotrechus zhangfani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF0EFF9FFDCC46D3203DF83B" bold="true" box="[548,605,1974,1998]" pageId="54" pageNumber="55">
|
||
<taxonomicNameLabel id="A27B57D2BF0EFF9FFDCC46D3203DF83B" box="[548,605,1974,1998]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF436633BF0FFF9EFF7F461A2322F82B" ID-DOI="http://doi.org/10.5281/zenodo.7642464" ID-Zenodo-Dep="7642464" httpUri="https://zenodo.org/record/7642464/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1919,1943]" targetBox="[151,1435,181,1894]" targetPageId="55">
|
||
<paragraph id="8B8336BBBF0FFF9EFF7F461A2322F82B" blockId="55.[151,1436,1919,2015]" pageId="55" pageNumber="56">
|
||
<emphasis id="B948EAA9BF0FFF9EFF7F461A2343F862" bold="true" box="[151,291,1919,1943]" pageId="55" pageNumber="56">FIGURE 37.</emphasis>
|
||
Xianren Dong, the type locality of
|
||
<taxonomicName id="4C3C4D38BF0FFF9EFD75461A21FBF862" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[669,923,1919,1943]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="zhangfani" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF0FFF9EFD75461A21FBF862" box="[669,923,1919,1943]" italics="true" pageId="55" pageNumber="56">Shilinotrechus zhangfani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF0FFF9EFC4C461A2180F862" bold="true" box="[932,992,1919,1943]" pageId="55" pageNumber="56">
|
||
<taxonomicNameLabel id="A27B57D2BF0FFF9EFC4C461A2180F862" box="[932,992,1919,1943]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. entrance; B. speleothems in cave; C. a running specimen of
|
||
<taxonomicName id="4C3C4D38BF0FFF9EFE8346C62005F84E" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[363,613,1955,1979]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="zhangfani" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF0FFF9EFE8346C62005F84E" box="[363,613,1955,1979]" italics="true" pageId="55" pageNumber="56">Shilinotrechus zhangfani</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF0FFF9EFD8346C620C3F84E" bold="true" box="[619,675,1955,1979]" pageId="55" pageNumber="56">
|
||
<taxonomicNameLabel id="A27B57D2BF0FFF9EFD8346C620C3F84E" box="[619,675,1955,1979]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; D & E. millipedes of the genera
|
||
<taxonomicName id="4C3C4D38BF0FFF9EFC1946C626F4F84E" authorityName="Silvestri" authorityYear="1910" box="[1009,1172,1955,1979]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EAA9BF0FFF9EFC1946C626F4F84E" box="[1009,1172,1955,1979]" italics="true" pageId="55" pageNumber="56">Eutrichodesmus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D38BF0FFF9EFB2B46C62752F84E" authorityName="Gervais" authorityYear="1847" box="[1219,1330,1955,1979]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EAA9BF0FFF9EFB2B46C62752F84E" box="[1219,1330,1955,1979]" italics="true" pageId="55" pageNumber="56">Glyphiulus</emphasis>
|
||
</taxonomicName>
|
||
; F. mould growing in cave.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8336BBBF00FF91FF2F41F22103FF0C" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">
|
||
Legs slender, protibiae without longitudinal sulcus.the 1
|
||
<superScript id="7C499BF3BF00FF91FCC641F22159FF50" attach="left" box="[814,825,151,165]" fontSize="6" pageId="56" pageNumber="57">st</superScript>
|
||
tarsomere much shorter than 2
|
||
<superScript id="7C499BF3BF00FF91FB6E41F326F6FF51" attach="none" box="[1158,1174,150,164]" fontSize="6" pageId="56" pageNumber="57">nd</superScript>
|
||
–4
|
||
<superScript id="7C499BF3BF00FF91FB5A41F326DFFF51" attach="left" box="[1202,1215,150,164]" fontSize="6" pageId="56" pageNumber="57">th</superScript>
|
||
tarsomeres together in fore leg, whereas as long as and longer than those in middle and hind legs respectively. The 1
|
||
<superScript id="7C499BF3BF00FF91FB1C41DE269FFF3C" attach="left" box="[1268,1279,187,201]" fontSize="6" pageId="56" pageNumber="57">st</superScript>
|
||
protarsomere enlarged and widened, not denticulate inwardly at apex in male.
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF00FF91FF2F40662197FEB4" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">Underside smooth and mostly glabrous in head and thorax; abdominal ventrites shortly and sparsely pubescent sparsely setose; IV−VI each with two paramedial setae; VII bisetose in male.</paragraph>
|
||
<paragraph id="8B8336BBBF00FF91FF2F402E23BEFDEC" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">
|
||
Male genitalia (
|
||
<figureCitation id="13072A3EBF00FF91FE9C402E2077FE90" box="[372,535,331,357]" captionStart="FIGURE 36" captionStartId="54.[151,250,1974,1998]" captionTargetBox="[153,1433,1066,1950]" captionTargetId="figure-503@54.[153,1434,1065,1950]" captionTargetPageId="54" captionText="FIGURE 36. Shilinotrechus zhangfani n. sp. A. left elytron (white dots indicating the chaetotaxy); B. median lobe, lateral view; C. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642462" httpUri="https://zenodo.org/record/7642462/files/figure.png" pageId="56" pageNumber="57">Figure 36B, C</figureCitation>
|
||
): The median lobe of aedeagus very short and stout, moderately sclerotised, basal portion strongly bent ventrally; apical part nearly straight, parallel-sided until apex, which suddenly contracted and widely broad; in dorsal view, apical lobe gradually widened toward apex, which is broadly rounded, copulatory piece extending over apex; basal orifice small, with a small sagittal aileron; copulatory piece large and long, about 2/7 as long as the median lobe; parameres thin and elongated, slightly shorter than the median lobe, each bearing with three long setae at apex.
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF00FF91FF2F434623FFFDC8" blockId="56.[151,1437,150,969]" box="[199,415,547,573]" pageId="56" pageNumber="57">Female: Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF00FF91FF2F4322228EFD38" pageId="56" pageNumber="57" type="discussion">
|
||
<paragraph id="8B8336BBBF00FF91FF2F4322228EFD38" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">
|
||
<emphasis id="B948EAA9BF00FF91FF2F4322235BFD94" bold="true" box="[199,315,583,609]" pageId="56" pageNumber="57">Remarks.</emphasis>
|
||
It is the fourth species of the Yunnan’s endemic genus
|
||
<taxonomicName id="4C3C4D38BF00FF91FC534322268EFD94" authority="Ueno, 2003" authorityName="Ueno" authorityYear="2003" box="[955,1262,583,609]" class="Insecta" family="Carabidae" genus="Shilinotrechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EAA9BF00FF91FC534322263CFD94" box="[955,1116,583,609]" italics="true" pageId="56" pageNumber="57">Shilinotrechus</emphasis>
|
||
<bibRefCitation id="EFAD4B4ABF00FF91FB8D4322268EFD94" author="Ueno, S. I." box="[1125,1262,583,609]" pageId="56" pageNumber="57" pagination="245 - 252" refId="ref29062" refString="Ueno, S. I. (2003) A new genus and species of extraordinary cave trechine (Coleoptera, Trechinae) from eastern Yunnan, Southwest China. Elytra, 31, 245 - 252." type="journal article" year="2003">Uéno, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
. It differs from other congeners by thinner body, shoulders of elytra strongly protruded, the 1
|
||
<superScript id="7C499BF3BF00FF91FBE8430E266BFD8C" attach="left" box="[1024,1035,619,633]" fontSize="6" pageId="56" pageNumber="57">st</superScript>
|
||
marginal umbilicate pore evidently shifted forwardly, and in particular the strongly bent male genitalia (
|
||
<bibRefCitation id="EFAD4B4ABF00FF91FC7F43EA2678FD5C" author="Ueno, S. I." box="[919,1048,655,681]" pageId="56" pageNumber="57" pagination="245 - 252" refId="ref29062" refString="Ueno, S. I. (2003) A new genus and species of extraordinary cave trechine (Coleoptera, Trechinae) from eastern Yunnan, Southwest China. Elytra, 31, 245 - 252." type="journal article" year="2003">Uéno 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAD4B4ABF00FF91FBCB43EA2766FD5C" author="Huang, S. B. & Tian, M. Y." box="[1059,1286,654,681]" pageId="56" pageNumber="57" pagination="727 - 733" refId="ref28116" refString="Huang, S. B. & Tian, M. Y. (2015) New species and new record of subterranean trechine beetles (Coleoptera: Carabidae: Trechinae) from southwestern China. The Coleopterists Bulletin, 49 (4), 727 - 733. https: // doi. org / 10.1649 / 0010 - 065 X- 69.4.727" type="journal article" year="2015">Huang & Tian 2015</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAD4B4ABF00FF91FAFA43EA2282FD38" author="Huang, S. B. & Tian, M. Y. & Faille, A." pageId="56" pageNumber="57" pagination="22 - 31" refId="ref28063" refString="Huang, S. B., Tian, M. Y. & Faille, A. (2020 b) Two new species of subterranean ground beetles from China (Coleoptera: Carabidae: Trechinae). Raffles Bulletin of Zoology, Supplement No. 35, 22 - 31." type="book chapter" year="2020">
|
||
Huang
|
||
<emphasis id="B948EAA9BF00FF91FA8C43EA27FDFD5C" box="[1380,1437,655,681]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
||
2020b
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF00FF91FF2F43B2209DFCE0" pageId="56" pageNumber="57" type="etymology">
|
||
<paragraph id="8B8336BBBF00FF91FF2F43B2209DFCE0" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">
|
||
<emphasis id="B948EAA9BF00FF91FF2F43B2232EFD04" bold="true" box="[199,334,727,753]" pageId="56" pageNumber="57">Etymology.</emphasis>
|
||
This new species is named in honour of Mr. Fan Zhang (Institute of Geology,
|
||
<collectingRegion id="49F8F859BF00FF91FB2C43B2277AFD04" box="[1220,1306,727,753]" country="China" name="Yunnan" pageId="56" pageNumber="57">Yunnan</collectingRegion>
|
||
University, Kunming), who is a leading caver in
|
||
<collectingRegion id="49F8F859BF00FF91FDDD439E2099FCE0" box="[565,761,763,789]" country="China" name="Yunnan" pageId="56" pageNumber="57">Yunnan Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF00FF91FF2F427B21F5FC3C" pageId="56" pageNumber="57" type="distribution">
|
||
<paragraph id="8B8336BBBF00FF91FF2F427B27FCFCCC" blockId="56.[151,1437,150,969]" box="[199,1436,798,825]" pageId="56" pageNumber="57">
|
||
<emphasis id="B948EAA9BF00FF91FF2F427B2301FCCD" bold="true" box="[199,353,798,824]" pageId="56" pageNumber="57">Distribution.</emphasis>
|
||
<collectingCountry id="F32B762BBF00FF91FE8D427A23C8FCCC" box="[357,424,799,825]" name="China" pageId="56" pageNumber="57">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F8F859BF00FF91FE5C427A2070FCCC" box="[436,528,799,825]" country="China" name="Yunnan" pageId="56" pageNumber="57">Yunnan</collectingRegion>
|
||
).Known only from the limestone cave called Xianren Dong in Mile Shi (
|
||
<figureCitation id="13072A3EBF00FF91FACA427A27EDFCCC" box="[1314,1421,799,825]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="56" pageNumber="57">Figure 1a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF00FF91FF2F422621F5FC3C" blockId="56.[151,1437,150,969]" pageId="56" pageNumber="57">
|
||
Xianren Dong is located on the left side of Road X016 about 7 kilometres from Xi’er Town. The length remains unknown, with a small entrance. Inside the cave, the humidity was rather high. Bats, two species of millipedes and growing mould were also observed inside the cave in addition to the unique trechine beetle, which was discovered under a stone in the inner most part of the main passage (
|
||
<figureCitation id="13072A3EBF00FF91FCFE42CA21E8FC3C" box="[790,904,943,969]" captionStart="FIGURE 37" captionStartId="55.[151,250,1919,1943]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-20@55.[151,1435,181,1894]" captionTargetPageId="55" captionText="FIGURE 37. Xianren Dong, the type locality of Shilinotrechus zhangfani n. sp. A. entrance; B. speleothems in cave; C. a running specimen of Shilinotrechus zhangfani n. sp.; D & E. millipedes of the genera Eutrichodesmus and Glyphiulus; F. mould growing in cave." figureDoi="http://doi.org/10.5281/zenodo.7642464" httpUri="https://zenodo.org/record/7642464/files/figure.png" pageId="56" pageNumber="57">Figure 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |