332 lines
39 KiB
XML
332 lines
39 KiB
XML
<document id="EFFB977CFF9297C0F60FC4F33CFFCC08" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin & Jia, Xinyang" docDate="2023" docId="039587ADBF04FF96FF7F43242191FC50" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Graciliblemus gui Tian & Huang & Jia 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="61" updateTime="1699475196432" updateUser="ExternalLinkService">
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<mods:title id="4A9980A46A3A28A4F18CD137ABC307E0">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="53024BD30CDC15AC554AE1E603E3289E">Tian, Mingyi</mods:namePart>
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<mods:namePart id="EBB0FB8104BBC93E945F868CA0F3E4BA">Huang, Sunbin</mods:namePart>
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<mods:namePart id="47DD9529DD7E9769BE53715A875F6A78">Jia, Xinyang</mods:namePart>
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<mods:title id="B0E633B487F469BCD85F759E0F5E9DA4">Zootaxa</mods:title>
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<mods:date id="9E4164967DD4F236B0EBD9530F49C261">2023</mods:date>
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<mods:number id="374DACB1F3B9E58B0173B69691EADBF6">2023-02-15</mods:number>
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<treatment id="039587ADBF04FF96FF7F43242191FC50" ID-DOI="http://doi.org/10.5281/zenodo.7642396" ID-GBIF-Taxon="209296926" ID-Zenodo-Dep="7642396" LSID="urn:lsid:plazi:treatment:039587ADBF04FF96FF7F43242191FC50" httpUri="http://treatment.plazi.org/id/039587ADBF04FF96FF7F43242191FC50" lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61">
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<subSubSection id="C3266530BF04FF95FF7F432423DBFDAE" box="[151,443,577,604]" pageId="60" pageNumber="61" type="nomenclature">
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<paragraph id="8B8336BBBF04FF95FF7F432423DBFDAE" blockId="60.[151,443,577,639]" box="[151,443,577,604]" pageId="60" pageNumber="61">
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<heading id="D0CB81D7BF04FF95FF7F432423DBFDAE" bold="true" box="[151,443,577,604]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="4">
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<emphasis id="B948EAA9BF04FF95FF7F432423DBFDAE" bold="true" box="[151,443,577,604]" pageId="60" pageNumber="61">
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<taxonomicName id="4C3C4D38BF04FF95FF7F43242311FDAE" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[151,369,577,604]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="gui" status="sp. nov.">
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<emphasis id="B948EAA9BF04FF95FF7F43242311FDAE" bold="true" box="[151,369,577,604]" italics="true" pageId="60" pageNumber="61">Graciliblemus gui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B57D2BF04FF95FE90432423DBFDAE" box="[376,443,577,603]" pageId="60" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3266530BF04FF95FF7F4300230AFD8A" box="[151,362,613,639]" pageId="60" pageNumber="61" type="description">
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<paragraph id="8B8336BBBF04FF95FF7F4300230AFD8A" blockId="60.[151,443,577,639]" box="[151,362,613,639]" pageId="60" pageNumber="61">
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<figureCitation id="13072A3EBF04FF95FF7F43002377FD8A" box="[151,279,613,639]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="60" pageNumber="61">
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Figures
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<quantity id="4CC49B5EBF04FF95FF1A43002377FD8A" box="[242,279,613,639]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="60" pageNumber="61" unit="m" value="1.0">1m</quantity>
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</figureCitation>
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,
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<figureCitation id="13072A3EBF04FF95FECC4300230AFD8A" box="[292,362,613,639]" captionStart-0="FIGURE 41" captionStart-1="FIGURE 42" captionStartId-0="61.[151,250,1943,1967]" captionStartId-1="62.[151,250,1919,1943]" captionTargetBox-0="[276,1326,194,1918]" captionTargetBox-1="[151,1435,181,1894]" captionTargetId-0="figure-20@61.[260,1326,181,1920]" captionTargetId-1="figure-28@62.[151,1435,181,1894]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURE 41. Graciliblemus gui n. sp. A. Habitus of male, holotype; B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." captionText-1="FIGURE 42. Gui Dong, the type locality of Graciliblemus gui n. sp. A. entrance; B. main passage; C. two individuals of G. gui n. sp.; D & F. mould and mushroom growing in cave; E. a cricket of the genus Tachycines." figureDoi-0="http://doi.org/10.5281/zenodo.7642470" figureDoi-1="http://doi.org/10.5281/zenodo.7642472" httpUri-0="https://zenodo.org/record/7642470/files/figure.png" httpUri-1="https://zenodo.org/record/7642472/files/figure.png" pageId="60" pageNumber="61">41–42</figureCitation>
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</paragraph>
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<subSubSection id="C3266530BF04FF95FF7F43CD2171FD13" pageId="60" pageNumber="61" type="materials_examined">
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<paragraph id="8B8336BBBF04FF95FF7F43CD2171FD13" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">
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<emphasis id="B948EAA9BF04FF95FF7F43CD2366FD37" bold="true" box="[151,262,680,706]" pageId="60" pageNumber="61">Material.</emphasis>
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<materialsCitation id="3B543CE6BF04FF95FEE643CD2317FD13" ID-GBIF-Occurrence="4030741307" collectingDate="2020-06-22" collectionCode="SCAU" collectorName="Mingyi Tian" county="Gui Dong" location="Pilin" municipality="Ghost Cave" pageId="60" pageNumber="61" specimenCount="306" specimenCount-female="1" specimenCount-male="306" stateProvince="Guizhou" typeStatus="holotype">
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<typeStatus id="54878819BF04FF95FEE643CD2316FD37" box="[270,374,680,706]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3AFD32BF04FF95FE9543CD23D6FD37" box="[381,438,680,706]" pageId="60" pageNumber="61" type="male">male</specimenCount>
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,
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<collectingCounty id="62E24E37BF04FF95FE2A43CD2051FD36" box="[450,561,680,707]" pageId="60" pageNumber="61">Gui Dong</collectingCounty>
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(
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<collectingMunicipality id="6BE7ACC1BF04FF95FDA843CD20A9FD37" box="[576,713,680,706]" pageId="60" pageNumber="61">Ghost Cave</collectingMunicipality>
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),
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<location id="8EE36060BF04FF95FD3543CD2175FD37" LSID="urn:lsid:plazi:treatment:039587ADBF04FF96FF7F43242191FC50:8EE36060BF04FF95FD3543CD2175FD37" box="[733,789,680,706]" county="Gui Dong" municipality="Ghost Cave" name="Pilin" pageId="60" pageNumber="61" stateProvince="Guizhou">Pilin</location>
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,
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<location id="8EE36060BF04FF95FCC943CD21EFFD37" LSID="urn:lsid:plazi:treatment:039587ADBF04FF96FF7F43242191FC50:8EE36060BF04FF95FCC943CD21EFFD37" box="[801,911,680,706]" county="Gui Dong" municipality="Ghost Cave" name="Zhaoxing" pageId="60" pageNumber="61" stateProvince="Guizhou">Zhaoxing</location>
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,
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<location id="8EE36060BF04FF95FC7243CD2188FD37" LSID="urn:lsid:plazi:treatment:039587ADBF04FF96FF7F43242191FC50:8EE36060BF04FF95FC7243CD2188FD37" box="[922,1000,680,706]" county="Gui Dong" municipality="Ghost Cave" name="Liping" pageId="60" pageNumber="61" stateProvince="Guizhou">Liping</location>
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,
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<collectingRegion id="49F8F859BF04FF95FC1B43CD2636FD37" box="[1011,1110,680,706]" country="China" name="Guizhou" pageId="60" pageNumber="61">Guizhou</collectingRegion>
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,
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<date id="FF82107BBF04FF95FB8A43CD2689FD37" box="[1122,1257,680,706]" pageId="60" pageNumber="61" value="2020-06-22">
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<collectingDate id="EFC6E993BF04FF95FB8A43CD2689FD37" box="[1122,1257,680,706]" pageId="60" pageNumber="61" value="2020-06-22">
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||
2020-VI-
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<quantity id="4CC49B5EBF04FF95FB2443CD2689FD37" box="[1228,1257,680,706]" metricMagnitude="4" metricUnit="m" metricValue="2.2305" pageId="60" pageNumber="61" unit="m" value="22305.0">22</quantity>
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</collectingDate>
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</date>
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,
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<specimenCount id="9D3AFD32BF04FF95FB1C43CD275EFD36" box="[1268,1342,680,707]" pageId="60" pageNumber="61" type="male">305 m</specimenCount>
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,
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<collectorName id="26C9536DBF04FF95FAA243CD22A9FD13" pageId="60" pageNumber="61">Mingyi Tian</collectorName>
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leg., in
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<collectionCode id="ED2DAE7EBF04FF95FECD43A92313FD13" box="[293,371,716,742]" collectionName="SCAU" pageId="60" pageNumber="61">SCAU</collectionCode>
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;
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</materialsCitation>
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<materialsCitation id="3B543CE6BF04FF95FE9643A82171FD13" ID-GBIF-Occurrence="4030741325" box="[382,785,716,743]" collectingDate="2020-06-22" collectionCode="SCAU" collectorName="Mingyi Tian" county="Gui Dong" location="Pilin" municipality="Ghost Cave" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
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<typeStatus id="54878819BF04FF95FE9643A82383FD12" box="[382,483,717,743]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
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:
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<specimenCount id="9D3AFD32BF04FF95FE0543A92031FD13" box="[493,593,716,742]" pageId="60" pageNumber="61" type="female">1 female</specimenCount>
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, idem, in
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<collectionCode id="ED2DAE7EBF04FF95FD5743A9216DFD13" box="[703,781,716,742]" collectionName="SCAU" pageId="60" pageNumber="61">SCAU</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF04FF95FF2F439523D0FCDB" pageId="60" pageNumber="61" type="diagnosis">
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<paragraph id="8B8336BBBF04FF95FF2F439523D0FCDB" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">
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<emphasis id="B948EAA9BF04FF95FF2F43952322FCFF" bold="true" box="[199,322,752,778]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
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Very small and slender trechine beetle, wholly pubescent, anophthalmic, depigmented, fore body much shorter than elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3266530BF04FF96FF2F425D23DAFE00" lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61" type="description">
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<paragraph id="8B8336BBBF04FF95FF2F425D2637FCA7" blockId="60.[151,1437,680,2039]" box="[199,1111,824,851]" pageId="60" pageNumber="61">
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<emphasis id="B948EAA9BF04FF95FF2F425D2339FCA7" bold="true" box="[199,345,824,850]" pageId="60" pageNumber="61">Description.</emphasis>
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Length:
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<quantity id="4CC49B5EBF04FF95FE56425D2028FCA6" box="[446,584,824,851]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.8" metricValueMin="3.1" pageId="60" pageNumber="61" unit="mm" value="3.45" valueMax="3.8" valueMin="3.1">3.1–3.8 mm</quantity>
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; width: 0.9–1.0 mm. Habitus as in
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<figureCitation id="13072A3EBF04FF95FC27425D2633FCA7" box="[975,1107,824,850]" captionStart="FIGURE 41" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[276,1326,194,1918]" captionTargetId="figure-20@61.[260,1326,181,1920]" captionTargetPageId="61" captionText="FIGURE 41. Graciliblemus gui n. sp. A. Habitus of male, holotype; B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642470" httpUri="https://zenodo.org/record/7642470/files/figure.png" pageId="60" pageNumber="61">Figure 41A</figureCitation>
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||
.
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</paragraph>
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<paragraph id="8B8336BBBF04FF95FF2F423920EAFC4A" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">Head with antennae brown, pronotum and elytra yellow, mouthparts, palps and tarsi pale; head slightly shiny, pronotum and elytra dim. Microsculptural engraved meshes indistinct. Head and pronotum smooth, whole elytral covered with coarse and irregular punctures.</paragraph>
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<paragraph id="8B8336BBBF04FF95FF2F42AD23FAFA42" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">
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Head stout though longer than wide excluding mandibles, HLl/HW = 1.0–1.04, HLm/HW = 1.42–1.44, frons rather flat, vertex convex, with longer pubescence; genae broadly expanded, with shorter pubescence; frontal furrows entire, nearly parallel-sided but strongly divergent posteriorly; both anterior and posterior supraorbital pores closely located, the anterior at about middle excluding mandibles, the posterior just before neck constriction which is well marked; clypeus transverse, 4-setose, labrum transverse, frontal margin distinctly bisinuate, making median part bluntly protruding, 6-setose; mandibles rather short, strongly curved apically; right mandibular tooth bidentate; labial suture invisible; mentum bisetose, tooth thin and simple, base largely concaved; submentum 6- setose, with setae very long; 2
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<superScript id="7C499BF3BF04FF95FE0045A12398FB27" attach="left" box="[488,504,1220,1234]" fontSize="6" pageId="60" pageNumber="61">nd</superScript>
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||
labial and 3
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<superScript id="7C499BF3BF04FF95FD6A45A120F0FB27" attach="left" box="[642,656,1220,1234]" fontSize="6" pageId="60" pageNumber="61">rd</superScript>
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maxillary palps strongly tumid, 3
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<superScript id="7C499BF3BF04FF95FBE245A12678FB27" attach="left" box="[1034,1048,1220,1234]" fontSize="6" pageId="60" pageNumber="61">rd</superScript>
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labial and 4
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<superScript id="7C499BF3BF04FF95FB4945A126CEFB27" attach="left" box="[1185,1198,1220,1234]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
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maxillary palps thin; 2
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<superScript id="7C499BF3BF04FF95FF4D458D22D5FB03" attach="left" box="[165,181,1256,1270]" fontSize="6" pageId="60" pageNumber="61">nd</superScript>
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||
labial palp bisetose, with an additional seta near apex on outer margin; penultimate palp slightly longer than the apical ones in both labium and maxillae; antennae filiform, short and stout, extending at about basal 2/3 of elytra; scape stouter and smooth, with several long setae, slightly longer than pedicel which is the shortest; the 11
|
||
<superScript id="7C499BF3BF04FF95FA8C44552711FACB" attach="left" box="[1380,1393,1328,1342]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
the longest; pubescent from pedicel to 11
|
||
<superScript id="7C499BF3BF04FF95FDDD44312022FA97" attach="left" box="[565,578,1364,1378]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
antennomere; the comparative length of each antennomere from the 1
|
||
<superScript id="7C499BF3BF04FF95FAA044312733FA97" attach="left" box="[1352,1363,1364,1378]" fontSize="6" pageId="60" pageNumber="61">st</superScript>
|
||
to 11
|
||
<superScript id="7C499BF3BF04FF95FA78443127FDFA97" attach="none" box="[1424,1437,1364,1378]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
in
|
||
<typeStatus id="54878819BF04FF95FF5C441D2376FA67" box="[180,278,1400,1426]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
as: 1
|
||
<superScript id="7C499BF3BF04FF95FEBA441D233DFA73" attach="left" box="[338,349,1400,1414]" fontSize="6" pageId="60" pageNumber="61">st</superScript>
|
||
(1.16) / 2
|
||
<superScript id="7C499BF3BF04FF95FE24441D23BCFA73" attach="left" box="[460,476,1400,1414]" fontSize="6" pageId="60" pageNumber="61">nd</superScript>
|
||
(1.00) / 3
|
||
<superScript id="7C499BF3BF04FF95FDA4441D203AFA73" attach="left" box="[588,602,1400,1414]" fontSize="6" pageId="60" pageNumber="61">rd</superScript>
|
||
(1.20) / 4
|
||
<superScript id="7C499BF3BF04FF95FD21441D20B6FA73" attach="left" box="[713,726,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.34) / 5
|
||
<superScript id="7C499BF3BF04FF95FCAD441D2132FA73" attach="left" box="[837,850,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.32) / 6
|
||
<superScript id="7C499BF3BF04FF95FC29441D21AEFA73" attach="left" box="[961,974,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.28) / 7
|
||
<superScript id="7C499BF3BF04FF95FBD5441D262AFA73" attach="left" box="[1085,1098,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.24) / 8
|
||
<superScript id="7C499BF3BF04FF95FB51441D26A6FA73" attach="left" box="[1209,1222,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.04) / 9
|
||
<superScript id="7C499BF3BF04FF95FADD441D2722FA73" attach="left" box="[1333,1346,1400,1414]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.16) / 10
|
||
<superScript id="7C499BF3BF04FF95FF5B44F922A0FA5F" attach="left" box="[179,192,1436,1450]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.08) / 11
|
||
<superScript id="7C499BF3BF04FF95FED344F92328FA5F" attach="left" box="[315,328,1436,1450]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
(1.52).
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF04FF95FF2F44A521E4F947" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">Propleura invisible from above; pronotum subcordate, slightly transverse, PnL/PnW = 0.88–0.91, wider than head, PnW/HW = 1.32–1.39, slightly longer than head excluding mandibles, HLl/PnL = 1.02–1.10, HLm/PnL = 1.36–1.44; widest a little before middle, lateral margins finely bordered, strongly sinuate before hind angles which are acute, fore angles broadly obtuse, slightly protruded forwardly; two pairs of latero-marginal setae present, the anterior located at before middle, the posterior a little before hind angle; base almost straight in most part, but shortly oblique forwardly near hind angle on both sides; front nearly straight; median line fine and well-defined; posterior transversal impression faint. Scutellum moderately sized.</paragraph>
|
||
<paragraph id="8B8336BBBF04FF95FF2F47D92072F803" blockId="60.[151,1437,680,2039]" pageId="60" pageNumber="61">
|
||
Elytra oblongus, elongate and slender, moderately convex; much longer than head plus pronotum,EL/(HLm+PnL) = 1.28–1.35; much longer than wide, EL/EW = 1.86–1.93, wider than pronotum, EW/PnW = 1.40–1.44; base unbordered, shoulders rounded, without humeral angle; lateral margins gently expanded, widest at about middle of elytra, sub-serrate and ciliate throughout; disc coarsely and irregularly punctate, striae rather deep, intervals slightly convex; apical striole present in the continuation of stria 6; chaetotaxy (
|
||
<figureCitation id="13072A3EBF04FF95FC5946292655F893" box="[945,1077,1868,1894]" captionStart="FIGURE 41" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[276,1326,194,1918]" captionTargetId="figure-20@61.[260,1326,181,1920]" captionTargetPageId="61" captionText="FIGURE 41. Graciliblemus gui n. sp. A. Habitus of male, holotype; B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642470" httpUri="https://zenodo.org/record/7642470/files/figure.png" pageId="60" pageNumber="61">Figure 41B</figureCitation>
|
||
): basal pore present; both dorsal pores located along 5
|
||
<superScript id="7C499BF3BF04FF95FE60461523F5F88B" attach="left" box="[392,405,1904,1918]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
stria, at about basal 1/5 and a little behind the middle of elytra, respectively, preapical pore located at apical fusion of 2
|
||
<superScript id="7C499BF3BF04FF95FE2F46F123B7F857" attach="left" box="[455,471,1940,1954]" fontSize="6" pageId="60" pageNumber="61">nd</superScript>
|
||
and 3
|
||
<superScript id="7C499BF3BF04FF95FDFF46F12045F857" attach="left" box="[535,549,1940,1954]" fontSize="6" pageId="60" pageNumber="61">rd</superScript>
|
||
striae, much closer to suture than to apical margin of elytra; marginal umbilicate series with 1
|
||
<superScript id="7C499BF3BF04FF95FECA46DD234DF833" attach="none" box="[290,301,1976,1990]" fontSize="6" pageId="60" pageNumber="61">st</superScript>
|
||
, 2
|
||
<superScript id="7C499BF3BF04FF95FEA046DD2338F833" attach="left" box="[328,344,1976,1990]" fontSize="6" pageId="60" pageNumber="61">nd</superScript>
|
||
and anguloapical pore close to the marginal gutter; humeral group equidistantly spaced, while both 5
|
||
<superScript id="7C499BF3BF04FF95FF4D46B922D2F81F" attach="left" box="[165,178,2012,2026]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
and 6
|
||
<superScript id="7C499BF3BF04FF95FF1E46B92363F81F" attach="left" box="[246,259,2012,2026]" fontSize="6" pageId="60" pageNumber="61">th</superScript>
|
||
pores widely separated.
|
||
</paragraph>
|
||
<caption id="DF436633BF05FF94FF7F46F220DAF821" ID-DOI="http://doi.org/10.5281/zenodo.7642470" ID-Zenodo-Dep="7642470" httpUri="https://zenodo.org/record/7642470/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1943,1967]" targetBox="[276,1326,194,1918]" targetPageId="61">
|
||
<paragraph id="8B8336BBBF05FF94FF7F46F220DAF821" blockId="61.[151,1437,1943,2004]" pageId="61" pageNumber="62">
|
||
<emphasis id="B948EAA9BF05FF94FF7F46F22341F85A" bold="true" box="[151,289,1943,1967]" pageId="61" pageNumber="62">FIGURE 41.</emphasis>
|
||
<taxonomicName id="4C3C4D38BF05FF94FEC046FD2380F85A" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[296,480,1944,1968]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="gui" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF05FF94FEC046FD2380F85A" box="[296,480,1944,1968]" italics="true" pageId="61" pageNumber="62">Graciliblemus gui</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF05FF94FE0F46FD2041F845" bold="true" box="[487,545,1944,1968]" pageId="61" pageNumber="62">
|
||
<taxonomicNameLabel id="A27B57D2BF05FF94FE0F46FD2041F845" box="[487,545,1944,1968]" pageId="61" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Habitus of male, holotype; B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF436633BF06FF97FF7F461A264FF84F" ID-DOI="http://doi.org/10.5281/zenodo.7642472" ID-Zenodo-Dep="7642472" httpUri="https://zenodo.org/record/7642472/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,250,1919,1943]" targetBox="[151,1435,181,1894]" targetPageId="62">
|
||
<paragraph id="8B8336BBBF06FF97FF7F461A264FF84F" blockId="62.[151,1436,1919,1979]" pageId="62" pageNumber="63">
|
||
<emphasis id="B948EAA9BF06FF97FF7F461A237FF862" bold="true" box="[151,287,1919,1943]" pageId="62" pageNumber="63">FIGURE 42.</emphasis>
|
||
Gui Dong, the type locality of
|
||
<taxonomicName id="4C3C4D38BF06FF97FDBF461A216EF862" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[599,782,1919,1943]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="gui" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF06FF97FDBF461A216EF862" box="[599,782,1919,1943]" italics="true" pageId="62" pageNumber="63">Graciliblemus gui</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF06FF97FCFC461A212DF862" bold="true" box="[788,845,1919,1943]" pageId="62" pageNumber="63">
|
||
<taxonomicNameLabel id="A27B57D2BF06FF97FCFC461A212DF862" box="[788,845,1919,1943]" pageId="62" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. entrance; B. main passage; C. two individuals of
|
||
<taxonomicName id="4C3C4D38BF06FF97FAB6461A27FCF862" authority="Tian & Huang & Jia, 2023" authorityName="Tian & Huang & Jia" authorityYear="2023" box="[1374,1436,1919,1943]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="gui" status="sp. nov.">
|
||
<emphasis id="B948EAA9BF06FF97FAB6461A27FCF862" box="[1374,1436,1919,1943]" italics="true" pageId="62" pageNumber="63">G. gui</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B948EAA9BF06FF97FF7F46C622B0F84E" bold="true" box="[151,208,1955,1979]" pageId="62" pageNumber="63">
|
||
<taxonomicNameLabel id="A27B57D2BF06FF97FF7F46C622B0F84E" box="[151,208,1955,1979]" pageId="62" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; D & F. mould and mushroom growing in cave; E. a cricket of the genus
|
||
<taxonomicName id="4C3C4D38BF06FF97FC5146C62649F84E" authorityName="Adelung" authorityYear="1902" box="[953,1065,1955,1979]" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" kingdom="Animalia" order="Orthoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EAA9BF06FF97FC5146C62649F84E" box="[953,1065,1955,1979]" italics="true" pageId="62" pageNumber="63">Tachycines</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8336BBBF07FF96FF2F41F2211EFF0C" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
|
||
Legs covered with dense and short hairs; tibiae without longitudinal sulcus; the 1
|
||
<superScript id="7C499BF3BF07FF96FBA441F22637FF50" attach="left" box="[1100,1111,151,165]" fontSize="6" pageId="63" pageNumber="64">st</superScript>
|
||
tarsomere shorter than 2
|
||
<superScript id="7C499BF3BF07FF96FA8041F327E0FF44" attach="none" box="[1384,1408,150,177]" fontSize="6" pageId="63" pageNumber="64">nd-</superScript>
|
||
4
|
||
<superScript id="7C499BF3BF07FF96FA7841F327FDFF51" attach="right" box="[1424,1437,150,164]" fontSize="6" pageId="63" pageNumber="64">th</superScript>
|
||
combined in fore leg, while as long as 2
|
||
<superScript id="7C499BF3BF07FF96FDA341DF2004FF20" attach="none" box="[587,612,186,213]" fontSize="6" pageId="63" pageNumber="64">nd-</superScript>
|
||
4
|
||
<superScript id="7C499BF3BF07FF96FD9B41DF20E0FF3D" attach="left" box="[627,640,186,200]" fontSize="6" pageId="63" pageNumber="64">th</superScript>
|
||
combined in middle and hind legs. In male, the 1
|
||
<superScript id="7C499BF3BF07FF96FB7341DE26C6FF3C" attach="left" box="[1179,1190,187,201]" fontSize="6" pageId="63" pageNumber="64">st</superScript>
|
||
and 2
|
||
<superScript id="7C499BF3BF07FF96FB0041DF2698FF3D" attach="left" box="[1256,1272,186,200]" fontSize="6" pageId="63" pageNumber="64">nd</superScript>
|
||
protarsomeres feebly more widened, both distinctly denticulate inwardly at apex.
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF07FF96FF2F40662380FEB4" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">Abdominal ventrites pubescent, ventrites IV–VI each with a pair of paramedial setae; VII bisetose in male, while quadrisetose in female.</paragraph>
|
||
<paragraph id="8B8336BBBF07FF96FF2F402E23DAFE00" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
|
||
Male genitalia (
|
||
<figureCitation id="13072A3EBF07FF96FE90402E2041FE90" box="[376,545,331,357]" captionStart="FIGURE 41" captionStartId="61.[151,250,1943,1967]" captionTargetBox="[276,1326,194,1918]" captionTargetId="figure-20@61.[260,1326,181,1920]" captionTargetPageId="61" captionText="FIGURE 41. Graciliblemus gui n. sp. A. Habitus of male, holotype; B. left elytron (white dots indicating the chaetotaxy); C. median lobe, lateral view; D. apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7642470" httpUri="https://zenodo.org/record/7642470/files/figure.png" pageId="63" pageNumber="64">Figure 41C, D</figureCitation>
|
||
): Weakly sclerotized, very small but elongate, evidently arcuate medially, base large, with a small sagittal aileron; apical dorsal membranous open fairly wide, inner sac without distinct copulatory piece, apical lobe evenly narrowed, blunt at apex in lateral view; in dorsal view, apical lobe wide, gradually narrowed toward the broad apex; both parameres fairly developed, slightly curved, right one with 4, while the left with 3 long setae at apex respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF07FF96FF2F409A201CFD94" pageId="63" pageNumber="64" type="discussion">
|
||
<paragraph id="8B8336BBBF07FF96FF2F409A201CFD94" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
|
||
<emphasis id="B948EAA9BF07FF96FF2F409A235BFDEC" bold="true" box="[199,315,511,537]" pageId="63" pageNumber="64">Remarks.</emphasis>
|
||
This is the second species of the genus
|
||
<taxonomicName id="4C3C4D38BF07FF96FD1F409A21F7FDEC" authorityName="Deuve & Tian" authorityYear="2016" box="[759,919,511,537]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EAA9BF07FF96FD1F409A21F7FDEC" box="[759,919,511,537]" italics="true" pageId="63" pageNumber="64">Graciliblemus</emphasis>
|
||
</taxonomicName>
|
||
, differing from the
|
||
<typeStatus id="54878819BF07FF96FB9B409A26C3FDEC" box="[1139,1187,511,537]" pageId="63" pageNumber="64">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C3C4D38BF07FF96FAE9409A231BFDC8" authority="Deuve & Tian, 2016" authorityName="Deuve & Tian" authorityYear="2016" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="lipingensis">
|
||
<emphasis id="B948EAA9BF07FF96FAE9409A27FCFDEC" box="[1281,1436,511,537]" italics="true" pageId="63" pageNumber="64">G. lipingensis</emphasis>
|
||
<bibRefCitation id="EFAD4B4ABF07FF96FF7F4346231BFDC8" author="Deuve, T. & Tian, M. Y." box="[151,379,547,573]" pageId="63" pageNumber="64" pagination="343 - 354" refId="ref27810" refString="Deuve, T. & Tian, M. Y. (2016) Descriptions de sept nouveaux Trechini cavernicoles de la Chine continentale (Coleoptera, Caraboidea). Bulletin de la Societe entomologique de France, 121 (3), 343 - 354. https: // doi. org / 10.3406 / bsef. 2016.2174" type="journal article" year="2016">Deuve & Tian, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
by its larger body size, pronotum with lateral margins strongly sinuate before hind angles, and elytra more elongate (
|
||
<bibRefCitation id="EFAD4B4ABF07FF96FE6443222010FD94" author="Deuve, T. & Tian, M. Y." box="[396,624,583,609]" pageId="63" pageNumber="64" pagination="343 - 354" refId="ref27810" refString="Deuve, T. & Tian, M. Y. (2016) Descriptions de sept nouveaux Trechini cavernicoles de la Chine continentale (Coleoptera, Caraboidea). Bulletin de la Societe entomologique de France, 121 (3), 343 - 354. https: // doi. org / 10.3406 / bsef. 2016.2174" type="journal article" year="2016">Deuve & Tian 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF07FF96FF2F430E26C9FD70" box="[199,1193,619,645]" pageId="63" pageNumber="64" type="etymology">
|
||
<paragraph id="8B8336BBBF07FF96FF2F430E26C9FD70" blockId="63.[151,1437,150,933]" box="[199,1193,619,645]" pageId="63" pageNumber="64">
|
||
<emphasis id="B948EAA9BF07FF96FF2F430E232EFD70" bold="true" box="[199,334,619,645]" pageId="63" pageNumber="64">Etymology.</emphasis>
|
||
“Gui” means ghost in Chinese. It refers to the
|
||
<typeStatus id="54878819BF07FF96FCB3430E21EBFD70" box="[859,907,619,645]" pageId="63" pageNumber="64">type</typeStatus>
|
||
locality, Gui Dong Cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3266530BF07FF96FF2F43EB2191FC50" pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph id="8B8336BBBF07FF96FF2F43EB26C6FD38" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
|
||
<emphasis id="B948EAA9BF07FF96FF2F43EB2301FD5D" bold="true" box="[199,353,654,680]" pageId="63" pageNumber="64">Distribution.</emphasis>
|
||
<collectingCountry id="F32B762BBF07FF96FE8243EA23CDFD5C" box="[362,429,655,681]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F8F859BF07FF96FE5643EA2045FD5C" box="[446,549,655,681]" country="China" name="Guizhou" pageId="63" pageNumber="64">Guizhou</collectingRegion>
|
||
). Known only from the limestone cave Gui Dong in southern Liping County, Qiandongnan Miao & Dong Autonomous Prefecture, eastern
|
||
<collectingRegion id="49F8F859BF07FF96FCAF43D62672FD38" box="[839,1042,691,717]" country="China" name="Guizhou" pageId="63" pageNumber="64">Guizhou Province</collectingRegion>
|
||
(
|
||
<figureCitation id="13072A3EBF07FF96FBC843D626F8FD38" box="[1056,1176,691,717]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="63" pageNumber="64">
|
||
Figure
|
||
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</figureCitation>
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).
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||
</paragraph>
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||
<paragraph id="8B8336BBBF07FF96FF2F43B2237DFCA8" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
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||
Gui Dong is near Pilin Village, and opens just along a road. There is a monument near the entrance. It is a short cave, about 30 metres long, connecting to an underground creek in the inner most part. As it was muddy and wet, moulds and mushrooms were growing inside. Apart from the trechine beetles, cave crickets also occur in Gui Dong (
|
||
<figureCitation id="13072A3EBF07FF96FF7742262371FCA8" box="[159,273,835,861]" captionStart="FIGURE 42" captionStartId="62.[151,250,1919,1943]" captionTargetBox="[151,1435,181,1894]" captionTargetId="figure-28@62.[151,1435,181,1894]" captionTargetPageId="62" captionText="FIGURE 42. Gui Dong, the type locality of Graciliblemus gui n. sp. A. entrance; B. main passage; C. two individuals of G. gui n. sp.; D & F. mould and mushroom growing in cave; E. a cricket of the genus Tachycines." figureDoi="http://doi.org/10.5281/zenodo.7642472" httpUri="https://zenodo.org/record/7642472/files/figure.png" pageId="63" pageNumber="64">Figure 42</figureCitation>
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||
).
|
||
</paragraph>
|
||
<paragraph id="8B8336BBBF07FF96FF2F42032191FC50" blockId="63.[151,1437,150,933]" pageId="63" pageNumber="64">
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<emphasis id="B948EAA9BF07FF96FF2F4203265BFC75" bold="true" box="[199,1083,870,897]" pageId="63" pageNumber="64">
|
||
Additional generic characteristics of
|
||
<taxonomicName id="4C3C4D38BF07FF96FD6642032656FC75" authority="Deuve & Tian, 2016" authorityName="Deuve & Tian" authorityYear="2016" box="[654,1078,870,897]" class="Insecta" family="Carabidae" genus="Graciliblemus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B948EAA9BF07FF96FD6642032151FC74" bold="true" box="[654,817,870,897]" italics="true" pageId="63" pageNumber="64">Graciliblemus</emphasis>
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<bibRefCitation id="EFAD4B4ABF07FF96FCD442022656FC75" author="Deuve, T. & Tian, M. Y." box="[828,1078,870,897]" pageId="63" pageNumber="64" pagination="343 - 354" refId="ref27810" refString="Deuve, T. & Tian, M. Y. (2016) Descriptions de sept nouveaux Trechini cavernicoles de la Chine continentale (Coleoptera, Caraboidea). Bulletin de la Societe entomologique de France, 121 (3), 343 - 354. https: // doi. org / 10.3406 / bsef. 2016.2174" type="journal article" year="2016">Deuve & Tian, 2016</bibRefCitation>
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||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Ventrite VII bisetose in male, while quadrisetose in female; the 1
|
||
<superScript id="7C499BF3BF07FF96FDF642EE2049FC6C" attach="left" box="[542,553,907,921]" fontSize="6" pageId="63" pageNumber="64">st</superScript>
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and 2
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<superScript id="7C499BF3BF07FF96FD8642EF201EFC6D" attach="left" box="[622,638,906,920]" fontSize="6" pageId="63" pageNumber="64">nd</superScript>
|
||
protarsomeres modified in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |