treatments-xml/data/03/95/87/039587A153275430FF14DED4E512FAE9.xml
2024-06-21 12:22:17 +02:00

531 lines
56 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="ECA0CDB218BD1A7953AC3E41DF6D9A60" ID-DOI="10.5281/zenodo.196425" ID-GBIF-Dataset="e18a85f3-43a2-4c80-8a1f-698148f15a9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="196425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460243284416" checkinUser="plazi" docAuthor="Mironov, Sergey V. &amp; Dabert, Jacek" docDate="2010" docId="039587A153275430FF14DED4E512FAE9" docLanguage="en" docName="zt02526p036.pdf" docOrigin="Zootaxa 2526" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protolichus strangulatus Favette et Trouessart 1904" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FFACFFD9532C543EFF83DF16E753FFBD" masterDocTitle="Systematic revision of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="12" updateTime="1698241288831" updateUser="plazi">
<mods:mods id="1CA8277F247C20372D596EA0166E1E55" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6B4CC4B1D0F456E1B96F46F6D7AF071A">
<mods:title id="C59B9C78BF9E0A49BFB7BCEDB83A5A04">Systematic revision of the feather mite genus Protolichus Trouessart, 1884 (Astigmata, Pterolichidae)</mods:title>
</mods:titleInfo>
<mods:name id="26A1E2BD4676FFE8075205B9EF5DF591" type="personal">
<mods:role id="623A37F87A6233B51AD24DC2E054B94D">
<mods:roleTerm id="8EDE54FC1E4EF0CE3B1CC46223A0C2F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43A3B710492236083BE08206FFB36BF1">Mironov, Sergey V.</mods:namePart>
</mods:name>
<mods:name id="A674FF712D538CBAAD59DAC07C32F16C" type="personal">
<mods:role id="EFF7498957DF2271B45103E09A60C03A">
<mods:roleTerm id="9BA21364CFF4A86456A68F2C4476F317">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3E9744CBB9B35D2CFC122B234F2B0CE">Dabert, Jacek</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B95458391222CADB5CA006C6E2CBDB0">text</mods:typeOfResource>
<mods:relatedItem id="7EA69185681FEA2C557B6518D9B9C21C" type="host">
<mods:titleInfo id="86551650B03CDF29AFE5820F4E5D7E20">
<mods:title id="0510E7926F3E8676F28FD9ED8AA96074">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="308D45C92AE4917E17B01459A979BD4B">
<mods:date id="694D8FAACE8A8DD89980F15034EF2FC4">2010</mods:date>
<mods:detail id="A023CB5A2FCD38B6C8D4E0A643A886F9" type="volume">
<mods:number id="63A7A5575076AF466485E6B3FB3CFDF0">2526</mods:number>
</mods:detail>
<mods:extent id="50A1EB141F7EAD6C8455E35D6A5263E9" unit="page">
<mods:start id="29C977AA45F41680C75804EAEBCB48D9">1</mods:start>
<mods:end id="4C7D4121800615DEFB0B2C5D2BDA3803">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9142B5331D873F7B1DA24D3B770D8902">journal article</mods:classification>
<mods:identifier id="E827DC8588A73AE2B33A5EF7328A55B3" type="DOI">10.5281/zenodo.196425</mods:identifier>
<mods:identifier id="2A6ACBC41CC2AD27E2F64A793C18B1D8" type="GBIF-Dataset">e18a85f3-43a2-4c80-8a1f-698148f15a9b</mods:identifier>
<mods:identifier id="ABF6C21B2A6F48C8B6355A6477B55760" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1092F4F0C140249A2161191FFFC198A3" type="Zenodo-Dep">196425</mods:identifier>
</mods:mods>
<treatment id="039587A153275430FF14DED4E512FAE9" ID-DOI="http://doi.org/10.5281/zenodo.5686093" ID-GBIF-Taxon="119390087" ID-Zenodo-Dep="5686093" LSID="urn:lsid:plazi:treatment:039587A153275430FF14DED4E512FAE9" httpUri="http://treatment.plazi.org/id/039587A153275430FF14DED4E512FAE9" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C326653C53275435FF14DED4E6DBFE42" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B8336B753275435FF14DED4E448FE61" blockId="11.[151,795,450,511]" box="[151,795,450,476]" pageId="11" pageNumber="12">
<heading id="D0CB81DB53275435FF14DED4E448FE61" bold="true" box="[151,795,450,476]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C3C4D3453275435FF14DED4E448FE61" authority="Favette et Trouessart, 1904" authorityName="Favette et Trouessart" authorityYear="1904" box="[151,795,450,476]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553275435FF14DED4E448FE61" bold="true" box="[151,795,450,476]" pageId="11" pageNumber="12">
<emphasis id="B948EAA553275435FF14DED4E692FE61" bold="true" box="[151,449,450,476]" italics="true" pageId="11" pageNumber="12">Protolichus strangulatus</emphasis>
<bibRefCitation id="EFAD4B4653275435FE4ADED4E448FE61" author="Favette" box="[457,795,450,476]" pageId="11" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette et Trouessart, 1904</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8336B753275435FF14DEF3E6DBFE42" blockId="11.[151,795,450,511]" box="[151,392,485,511]" pageId="11" pageNumber="12">
(
<figureCitation id="13072A3253275435FF1CDEF3E7BAFE42" box="[159,233,485,511]" captionStart="FIGURE 5" captionStartId="12.[151,258,1961,1985]" captionTargetBox="[213,1394,197,1906]" captionTargetId="figure@12.[183,1399,197,1934]" captionTargetPageId="12" captionText="FIGURE 5. Dorsal view of male opisthosoma of Protolichus brachiatus (A, B) and P. strangulatus (C, D). A, Cheteromorph male, B, D—homeomorph male." httpUri="https://zenodo.org/record/196430/files/figure.png" pageId="11" pageNumber="12">Figs 5</figureCitation>
C, D, 6, 11A)
</paragraph>
</subSubSection>
<paragraph id="8B8336B753275435FF14DD26E585FD3A" pageId="11" pageNumber="12">
<table id="F93CC4175327ABC1FF14DD26E2CEFD3A" box="[151,1437,560,647]" gridcols="12" gridrows="3" pageId="11" pageNumber="12">
<tr id="350C34F55327ABC1FF14DD26E2CEFDFA" box="[151,1437,560,583]" gridrow="0" pageId="11" pageNumber="12">
<th id="76DD5D895327ABC1FF14DD26E5ADFDFA" box="[151,766,560,583]" gridcol="0" gridrow="0" pageId="11" pageNumber="12">
<emphasis id="B948EAA553275435FF14DD26E527FDFA" box="[151,628,560,583]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C3C4D3453275435FF14DD26E640FDFA" box="[151,275,560,583]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Protolichus</taxonomicName>
(s. str.)
<taxonomicName id="4C3C4D3453275435FEF0DD26E6B6FDFA" box="[371,485,560,583]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brachiatus">brachiatus</taxonomicName>
<taxonomicName id="4C3C4D3453275435FE6CDD26E527FDFA" authority="Favette and" authorityName="Favette and" box="[495,628,560,583]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="strangulatus">strangulatus</taxonomicName>
</emphasis>
Favette and
</th>
<th id="76DD5D895327ABC1FC8BDD26E42BFDFA" box="[776,888,560,583]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">Trouessart</th>
<th id="76DD5D895327ABC1FC01DD26E4ECFDFA" box="[898,959,560,583]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">1904:</th>
<th id="76DD5D895327ABC1FC4ADD26E4A4FDFA" box="[969,1015,560,583]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">136,</th>
<th id="76DD5D895327ABC1FB81DD26E34CFDFA" box="[1026,1055,560,583]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">Pl.</th>
<th id="76DD5D895327ABC1FBAADD26E36CFDFA" box="[1065,1087,560,583]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">V,</th>
<th id="76DD5D895327ABC1FBCADD26E33DFDFA" box="[1097,1134,560,583]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">fig.</th>
<th id="76DD5D895327ABC1FBFBDD26E3DEFDFA" box="[1144,1165,560,583]" gridcol="7" gridrow="0" pageId="11" pageNumber="12">4;</th>
<th id="76DD5D895327ABC1FB14DD26E3BDFDFA" box="[1175,1262,560,583]" gridcol="8" gridrow="0" pageId="11" pageNumber="12">Dubinin</th>
<th id="76DD5D895327ABC1FB7BDD26E266FDFA" box="[1272,1333,560,583]" gridcol="9" gridrow="0" pageId="11" pageNumber="12">1956:</th>
<th id="76DD5D895327ABC1FABCDD26E23EFDFA" box="[1343,1389,560,583]" gridcol="10" gridrow="0" pageId="11" pageNumber="12">295,</th>
<th id="76DD5D895327ABC1FAFBDD26E2CEFDFA" box="[1400,1437,560,583]" gridcol="11" gridrow="0" pageId="11" pageNumber="12">fig.</th>
</tr>
<tr id="350C34F55327ABC1FF14DD46E2CEFDDA" box="[151,1437,592,615]" gridrow="1" pageId="11" pageNumber="12" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76DD5D895327ABC1FF14DD46E5ADFDDA" box="[151,766,592,615]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">131A.</th>
</tr>
<tr id="350C34F55327ABC1FF14DD66E2CEFD3A" box="[151,1437,624,647]" gridrow="2" pageId="11" pageNumber="12" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76DD5D895327ABC1FF14DD66E5ADFD3A" box="[151,766,624,647]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">
<taxonomicName id="4C3C4D3453275435FF14DD66E582FD3A" authority="Gaud and Atyeo 1996: 132" authorityName="Gaud and Atyeo" authorityPageNumber="132" authorityYear="1996" box="[151,721,624,647]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553275435FF14DD66E6F4FD3A" box="[151,423,624,647]" italics="true" pageId="11" pageNumber="12">Protolichus strangulatus:</emphasis>
Gaud and Atyeo 1996: 132
</taxonomicName>
.
</th>
</tr>
</table>
</paragraph>
<subSubSection id="C326653C53275435FF14DDA3E5D3FC82" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8B8336B753275435FF14DDA3E5D3FC82" blockId="11.[151,1437,693,1988]" pageId="11" pageNumber="12">
<emphasis id="B948EAA553275435FF14DDA3E6DDFD72" bold="true" box="[151,398,693,719]" pageId="11" pageNumber="12">Material examined.</emphasis>
Heteromorph male
<typeStatus id="5487881553275435FDFEDDA3E5BEFD72" box="[637,749,693,719]" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
, homeomorph male and
<specimenCount id="9D3AFD3E53275435FB97DDA3E32FFD72" box="[1044,1148,693,719]" pageId="11" pageNumber="12" type="female">2 female</specimenCount>
<typeStatus id="5487881553275435FB07DDA3E27CFD72" box="[1156,1327,693,719]" pageId="11" pageNumber="12" type="paralectotype">paralectotypes</typeStatus>
(3 slides with number
<collectionCode id="ED2DAE7253275435FEB6DDCCE638FD49" LSID="urn:lsid:biocol.org:col:15277" box="[309,363,730,756]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15277" name="Royal Ontario Museum - Herbarium" pageId="11" pageNumber="12">TRT</collectionCode>
34F1) ex
<taxonomicName id="4C3C4D3453275435FE66DDCCE4E8FD49" authority="Gray G.R., 1859" authorityName="Gray G.R." authorityYear="1859" box="[485,955,730,756]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553275435FE66DDCCE5A2FD4E" box="[485,753,730,755]" italics="true" pageId="11" pageNumber="12">Charmosyna pulchella</emphasis>
Gray G.R., 1859
</taxonomicName>
(
<taxonomicName id="4C3C4D3453275435FC4FDDCCE301FD49" box="[972,1106,730,756]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="family">Psittacidae</taxonomicName>
:
<taxonomicName id="4C3C4D3453275435FBDEDDCCE395FD49" box="[1117,1222,730,756]" class="Aves" family="Psittacidae" kingdom="Animalia" order="Psittaciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="subFamily" subFamily="Loriinae">Loriinae</taxonomicName>
), New
<collectingCountry id="F32B762753275435FA99DDCCE221FD49" box="[1306,1394,730,756]" name="Guinea-Bissau" pageId="11" pageNumber="12">Guinea</collectingCountry>
, no other data;
<typeStatus id="5487881553275435FE9CDC16E6DFFCA7" box="[287,396,768,794]" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
designated here.
<specimenCount id="9D3AFD3E53275435FDDCDC16E594FCA7" box="[607,711,768,794]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
(
<collectionCode id="ED2DAE7253275435FD54DC16E45CFCA7" LSID="urn:lsid:biocol.org:col:15277" box="[727,783,768,794]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15277" name="Royal Ontario Museum - Herbarium" pageId="11" pageNumber="12">TRT</collectionCode>
34F14, labeled as
<taxonomicName id="4C3C4D3453275435FC77DC16E3C1FCA4" box="[1012,1170,768,793]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553275435FC77DC16E3C1FCA4" box="[1012,1170,768,793]" italics="true" pageId="11" pageNumber="12">P. brachiatus</emphasis>
</taxonomicName>
) ex
<taxonomicName id="4C3C4D3453275435FB4BDC16E613FC82" authority="Wallace, 1863" authorityName="Wallace" authorityYear="1863" class="Aves" family="Psittacidae" genus="Loriculus" kingdom="Animalia" order="Psittaciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="sclateri">
<emphasis id="B948EAA553275435FB4BDC16E2CFFCA4" box="[1224,1436,768,793]" italics="true" pageId="11" pageNumber="12">Loriculus sclateri</emphasis>
Wallace, 1863
</taxonomicName>
, Sulawesi (contamination).
</paragraph>
</subSubSection>
<subSubSection id="C326653C53275430FF45DC5CE4CAFD09" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8B8336B753275435FF45DC5CE58FFA62" blockId="11.[151,1437,693,1988]" pageId="11" pageNumber="12">
<emphasis id="B948EAA553275435FF45DC5CE60DFCD9" bold="true" box="[198,350,842,868]" pageId="11" pageNumber="12">Description.</emphasis>
Heteromorph male (
<typeStatus id="5487881553275435FDDDDC5CE583FCD9" box="[606,720,842,868]" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
). Idiosoma, length x width, 590 x 305. Subcapitulum with posterior margin strongly convex, widest part crossed by single transverse fold, area with setae
<emphasis id="B948EAA553275435FB7DDC66E267FC34" box="[1278,1332,880,905]" italics="true" pageId="11" pageNumber="12">subc</emphasis>
outlined by short folds forming lateral and posterior sides (
<figureCitation id="13072A3253275435FD61DC83E47BFC12" box="[738,808,917,943]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
G). Prodorsal shield: posterior margin straight, length along midline 150, surface monotonously punctate, transverse bar across scapular setae poorly sclerotized. Setae
<emphasis id="B948EAA553275435FF5EDCF6E7A2FC44" box="[221,241,992,1017]" italics="true" pageId="11" pageNumber="12">si</emphasis>
missing (probably spiculiform). Distance between scapular setae:
<emphasis id="B948EAA553275435FC7FDCF6E364FC44" box="[1020,1079,992,1017]" italics="true" pageId="11" pageNumber="12">se-se</emphasis>
65,
<emphasis id="B948EAA553275435FBE9DCF6E3C8FC44" box="[1130,1179,992,1017]" italics="true" pageId="11" pageNumber="12">si-si</emphasis>
28. Hysterosoma 420 long. Length of hysteronotal shield 405, width 240, anterior margin slightly convex, most of surface monotonously punctate, posterior part near bases of opisthosomal lobes with numerous poorly expressed pits (
<figureCitation id="13072A3253275435FF23DB46E7BFFBD7" box="[160,236,1104,1130]" captionStart="FIGURE 5" captionStartId="12.[151,258,1961,1985]" captionTargetBox="[213,1394,197,1906]" captionTargetId="figure@12.[183,1399,197,1934]" captionTargetPageId="12" captionText="FIGURE 5. Dorsal view of male opisthosoma of Protolichus brachiatus (A, B) and P. strangulatus (C, D). A, Cheteromorph male, B, D—homeomorph male." httpUri="https://zenodo.org/record/196430/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
C). Opisthosomal lobes with acute apices, without transverse crest posterior to base of setae
<emphasis id="B948EAA553275435FAF5DB46E2C7FBD4" box="[1398,1428,1104,1129]" italics="true" pageId="11" pageNumber="12">h3</emphasis>
. Terminal cleft as a narrow inverted U, slightly enlarged in anterior third, length 82, width at level of setae
<geoCoordinate id="EE08507053275435FA03DB63E7EAFB09" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="1.25">
<emphasis id="B948EAA553275435FA03DB63E2C8FB33" box="[1408,1435,1141,1166]" italics="true" pageId="11" pageNumber="12">e1</emphasis>
15
</geoCoordinate>
, width in anterior part 22. Setae
<emphasis id="B948EAA553275435FDD8DB8DE52BFB09" box="[603,632,1179,1204]" italics="true" pageId="11" pageNumber="12">c2</emphasis>
spiculiform, 42 long; setae
<emphasis id="B948EAA553275435FC5EDB8DE4AFFB09" box="[989,1020,1179,1204]" italics="true" pageId="11" pageNumber="12">d2</emphasis>
, setiform, about 5 long, setae
<emphasis id="B948EAA553275435FAFCDB8DE2CEFB09" box="[1407,1437,1179,1204]" italics="true" pageId="11" pageNumber="12">e2</emphasis>
represented by macrosetae extending far beyond lobar apices, terminal part thick without filiform end, 255 260 long; setae
<emphasis id="B948EAA553275435FED1DBF3E63EFB43" box="[338,365,1253,1278]" italics="true" pageId="11" pageNumber="12">e1</emphasis>
setiform, situated on margin of terminal cleft, slightly posterior to level of macrosetae
<emphasis id="B948EAA553275435FAF5DBF3E2C0FB43" box="[1398,1427,1253,1278]" italics="true" pageId="11" pageNumber="12">h2</emphasis>
; setae
<emphasis id="B948EAA553275435FF5FDA1DE7A0FA99" box="[220,243,1291,1316]" italics="true" pageId="11" pageNumber="12">f2</emphasis>
flattened, spatuliform with bidentate apex. Distance between dorsal setae:
<emphasis id="B948EAA553275435FB0BDA1DE39EFA99" box="[1160,1229,1291,1316]" italics="true" pageId="11" pageNumber="12">c2:d2</emphasis>
, 135,
<emphasis id="B948EAA553275435FA99DA1DE20CFA99" box="[1306,1375,1291,1316]" italics="true" pageId="11" pageNumber="12">d2:e2</emphasis>
150,
<emphasis id="B948EAA553275435FF14DA26E78FFAF4" box="[151,220,1328,1353]" italics="true" pageId="11" pageNumber="12">e2:h3</emphasis>
125,
<emphasis id="B948EAA553275435FEA1DA26E636FAF4" box="[290,357,1328,1353]" italics="true" pageId="11" pageNumber="12">
e2:
<geoCoordinate id="EE08507053275435FECADA26E636FAF4" box="[329,357,1328,1353]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="2.1666665">e1</geoCoordinate>
</emphasis>
70,
<emphasis id="B948EAA553275435FE1FDA26E68CFAF4" box="[412,479,1328,1353]" italics="true" pageId="11" pageNumber="12">e2:e2</emphasis>
140,
<emphasis id="B948EAA553275435FDA6DA26E53EFAF4" box="[549,621,1328,1353]" italics="true" pageId="11" pageNumber="12">h2:h2</emphasis>
90,
<emphasis id="B948EAA553275435FD20DA26E5BAFAF4" box="[675,745,1328,1353]" italics="true" pageId="11" pageNumber="12">h3:h3</emphasis>
58,
<emphasis id="B948EAA553275435FCA3DA26E42DFAF4" box="[800,894,1328,1353]" italics="true" pageId="11" pageNumber="12">ps1:ps1</emphasis>
48,
<emphasis id="B948EAA553275435FC36DA26E4A8FAF4" box="[949,1019,1328,1353]" italics="true" pageId="11" pageNumber="12">e2:h2</emphasis>
62,
<emphasis id="B948EAA553275435FBB2DA26E32BFAF4" box="[1073,1144,1328,1353]" italics="true" pageId="11" pageNumber="12">h2:h3</emphasis>
32,
<emphasis id="B948EAA553275435FB2CDA26E3A0FAF4" box="[1199,1267,1328,1353]" italics="true" pageId="11" pageNumber="12">
h2:
<geoCoordinate id="EE08507053275435FB5BDA26E3A0FAF4" box="[1240,1267,1328,1353]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="1.1333333">e1</geoCoordinate>
</emphasis>
8,
<emphasis id="B948EAA553275435FA9FDA26E23DFAF4" box="[1308,1390,1328,1353]" italics="true" pageId="11" pageNumber="12">ps1:h3</emphasis>
17. Genital apparatus
<date id="FF82107753275435FEE8DA43E696FAD2" box="[363,453,1365,1391]" pageId="11" pageNumber="12" value="1922-10-27">27 x 22</date>
, its base at level of trochanters IV. Anterior ends of paragenital apodemes connected with inner tips of epimerites IIIa, middle parts of apodemes connected by large and poorly sclerotized transverse bridge (
<figureCitation id="13072A3253275435FEEDDAB6E6E7FA07" box="[366,436,1440,1466]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
A). Distance between ventral setae:
<emphasis id="B948EAA553275435FCD4DAB6E4C8FA04" box="[855,923,1440,1465]" italics="true" pageId="11" pageNumber="12">3b:3a</emphasis>
8,
<emphasis id="B948EAA553275435FC3CDAB6E4A6FA04" box="[959,1013,1440,1465]" italics="true" pageId="11" pageNumber="12">3a:g</emphasis>
30,
<emphasis id="B948EAA553275435FBA4DAB6E30FFA04" box="[1063,1116,1440,1465]" italics="true" pageId="11" pageNumber="12">g:4a</emphasis>
68,
<emphasis id="B948EAA553275435FB0CDAB6E38DFA04" box="[1167,1246,1440,1465]" italics="true" pageId="11" pageNumber="12">4a:ps3</emphasis>
118. Cupules
<emphasis id="B948EAA553275435FA06DAB6E2C8FA04" box="[1413,1435,1440,1465]" italics="true" pageId="11" pageNumber="12">ih</emphasis>
at level of setae
<emphasis id="B948EAA553275435FED6DAD3E62EFA63" box="[341,381,1477,1502]" italics="true" pageId="11" pageNumber="12">ps2</emphasis>
. Diameter of anal suckers 21.
</paragraph>
<paragraph id="8B8336B753275435FF47DAFCE484F902" blockId="11.[151,1437,693,1988]" pageId="11" pageNumber="12">
Legs II approximately 1.5 times longer than legs I. Tarsus I with crest-like ventral extension stretching along all segment, proximal end of this extension widely rounded (
<figureCitation id="13072A3253275435FC37D906E4AFF997" box="[948,1020,1552,1578]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
B). Tibia I, II with small verrucous ventral tubercle, solenidia φ of these segments with verrucae (
<figureCitation id="13072A3253275435FCE4D923E4E1F9F2" box="[871,946,1589,1615]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Figs 6</figureCitation>
B, C). Genu I and femora I, II without any apophyses. Tarsus II with small ventral extensions bearing setae
<emphasis id="B948EAA553275435FC0FD94DE4C4F9C9" box="[908,919,1627,1652]" italics="true" pageId="11" pageNumber="12">s</emphasis>
and
<emphasis id="B948EAA553275435FC53D94DE4A1F9C9" box="[976,1010,1627,1652]" italics="true" pageId="11" pageNumber="12">wa</emphasis>
. Genu II with dorso-apical tubercle; seta
<emphasis id="B948EAA553275435FF4FD996E7BDF924" box="[204,238,1664,1689]" italics="true" pageId="11" pageNumber="12">cG</emphasis>
II strongly thickened along its entire length, smooth, nearly 3 times longer than segment. Setae
<emphasis id="B948EAA553275435FAD1D996E22FF924" box="[1362,1404,1664,1689]" italics="true" pageId="11" pageNumber="12">d, e</emphasis>
of tarsus IV cone-shaped, with inflated base, poorly sclerotized (
<figureCitation id="13072A3253275435FCE8D9B3E4E1F902" box="[875,946,1701,1727]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
D).
</paragraph>
<paragraph id="8B8336B753275435FF47D9DCE442F8E9" blockId="11.[151,1437,693,1988]" pageId="11" pageNumber="12">
Homeomorph male (1
<typeStatus id="5487881553275435FE51D9DCE524F959" box="[466,631,1738,1764]" pageId="11" pageNumber="12" type="paralectotype">paralectotype</typeStatus>
). Subcapitulum with posterior margin strongly convex, median part with several transverse striae (
<figureCitation id="13072A3253275435FE78D9E6E512F8B7" box="[507,577,1776,1802]" captionStart="FIGURE 6" captionStartId="13.[151,257,1741,1765]" captionTargetBox="[155,1431,204,1717]" captionTargetId="figure@13.[151,1436,194,1720]" captionTargetPageId="13" captionText="FIGURE 6. Protolichus strangulatus, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view." httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
H). Terminal cleft as a very narrow U, approximately 6 times longer than wide. Anterior ends of paragenital apodemes not connected with epimerites IIa, IIIa; transverse bridge between middle parts of paragenital apodemes absent.
</paragraph>
<paragraph id="8B8336B753275435FF47D876E653F879" blockId="11.[151,1437,693,1988]" pageId="11" pageNumber="12">
Legs II about 1.3 times longer than legs I. Tarsi I, II with blunt-angular ventral margin. Tibiae II with scarcely distinct ventral tubercles. Setae
<emphasis id="B948EAA553275435FD1AD893E5EEF823" box="[665,701,1925,1950]" italics="true" pageId="11" pageNumber="12">cG</emphasis>
of genu II thickened, approximately 1.4 times longer than segment.
</paragraph>
<caption id="DF43663F53205432FF14D8BFE5D7F85D" httpUri="https://zenodo.org/record/196430/files/figure.png" pageId="12" pageNumber="13" targetBox="[213,1394,197,1906]" targetPageId="12">
<paragraph id="8B8336B753205432FF14D8BFE5D7F85D" blockId="12.[151,1436,1961,2016]" pageId="12" pageNumber="13">
<emphasis id="B948EAA553205432FF14D8BFE645F87C" bold="true" box="[151,278,1961,1985]" pageId="12" pageNumber="13">FIGURE 5</emphasis>
. Dorsal view of male opisthosoma of
<taxonomicName id="4C3C4D3453205432FD45D8BFE495F87D" box="[710,966,1961,1984]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553205432FD45D8BFE495F87D" box="[710,966,1961,1984]" italics="true" pageId="12" pageNumber="13">Protolichus brachiatus</emphasis>
</taxonomicName>
(A, B) and
<taxonomicName id="4C3C4D3453205432FBCED8BFE3A5F87D" box="[1101,1270,1961,1984]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553205432FBCED8BFE3A5F87D" box="[1101,1270,1961,1984]" italics="true" pageId="12" pageNumber="13">P. strangulatus</emphasis>
</taxonomicName>
(C,D). A, Cheteromorph male, B, D—homeomorph male.
</paragraph>
</caption>
<caption id="DF43663F53215433FF14D9DBE403F8F8" httpUri="https://zenodo.org/record/196431/files/figure.png" pageId="13" pageNumber="14" targetBox="[155,1431,204,1717]" targetPageId="13">
<paragraph id="8B8336B753215433FF14D9DBE403F8F8" blockId="13.[151,1436,1741,1861]" pageId="13" pageNumber="14">
<emphasis id="B948EAA553215433FF14D9DBE645F958" bold="true" box="[151,278,1741,1765]" pageId="13" pageNumber="14">FIGURE 6</emphasis>
.
<taxonomicName id="4C3C4D3453215433FEA6D9D8E564F958" box="[293,567,1742,1765]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553215433FEA6D9D8E564F958" box="[293,567,1742,1765]" italics="true" pageId="13" pageNumber="14">Protolichus strangulatus</emphasis>
</taxonomicName>
, male. A—ventral view of heteromorph male, B—leg I of heteromorph male, dorsal view, C—leg II of heteromorph male, dorsal view, D—tarsus IV of heteromorph male, dorsal view, E—leg I of homeomorph male, dorsal view, F—leg II of homeomorph male, G—gnathosoma of heteromorph male, ventral view, H—gnathosoma of homeomorph male, dorsal view, ventral view.
</paragraph>
</caption>
<paragraph id="8B8336B753215430FF45D878E5B1FF4A" blockId="13.[151,1437,1902,2032]" lastBlockId="14.[151,1437,152,1364]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Measurements: Idiosoma, length x width, 490 x 260. Prodorsal shield 135 long. Setae
<emphasis id="B948EAA553215433FB37D878E394F83A" box="[1204,1223,1902,1927]" italics="true" pageId="13" pageNumber="14">si</emphasis>
22 long. Distance between scapular setae:
<emphasis id="B948EAA553215433FE39D887E6A5F817" box="[442,502,1937,1962]" italics="true" pageId="13" pageNumber="14">se:se</emphasis>
65,
<emphasis id="B948EAA553215433FDAFD887E50EF817" box="[556,605,1937,1962]" italics="true" pageId="13" pageNumber="14">si:si</emphasis>
35. Hysterosoma 350 long. Length of hysteronotal shield 330, width 200. Terminal cleft
<date id="FF82107753215433FEFDD8A5E69CF870" box="[382,463,1971,1997]" pageId="13" pageNumber="14" value="1952-10-05">52 x 5.</date>
Lateral setae
<emphasis id="B948EAA553215433FDF1D8A5E5DEF871" box="[626,653,1971,1996]" italics="true" pageId="13" pageNumber="14">c2</emphasis>
28 long; setae
<emphasis id="B948EAA553215433FCC3D8A5E40FF871" box="[832,860,1971,1996]" italics="true" pageId="13" pageNumber="14">d2</emphasis>
5 long; setae
<emphasis id="B948EAA553215433FB83D8A5E34FF871" box="[1024,1052,1971,1996]" italics="true" pageId="13" pageNumber="14">e2</emphasis>
thick represented by macrosetae extending beyond lobar apices, terminal part thick without filiform tip, about 180 long. Distances between dorsal setae:
<emphasis id="B948EAA553225430FEB2DF8EE627FF0C" box="[305,372,152,177]" italics="true" pageId="14" pageNumber="15">c2:d2</emphasis>
120,
<emphasis id="B948EAA553225430FE35DF8EE6AAFF0C" box="[438,505,152,177]" italics="true" pageId="14" pageNumber="15">d2:e2</emphasis>
130,
<emphasis id="B948EAA553225430FDBFDF8EE52CFF0C" box="[572,639,152,177]" italics="true" pageId="14" pageNumber="15">e2:h3</emphasis>
75,
<emphasis id="B948EAA553225430FD30DF8EE5A7FF0C" box="[691,756,152,177]" italics="true" pageId="14" pageNumber="15">
e2:
<geoCoordinate id="EE08507053225430FD5ADF8EE5A7FF0C" box="[729,756,152,177]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="1.9166667">e1</geoCoordinate>
</emphasis>
55,
<emphasis id="B948EAA553225430FCABDF8EE43BFF0C" box="[808,872,152,177]" italics="true" pageId="14" pageNumber="15">e2:e2</emphasis>
122,
<emphasis id="B948EAA553225430FC2FDF8EE4A2FF0C" box="[940,1009,152,177]" italics="true" pageId="14" pageNumber="15">h2:h2</emphasis>
55,
<emphasis id="B948EAA553225430FBA6DF8EE33AFF0C" box="[1061,1129,152,177]" italics="true" pageId="14" pageNumber="15">h3:h3</emphasis>
32,
<emphasis id="B948EAA553225430FB1EDF8EE3AAFF0C" box="[1181,1273,152,177]" italics="true" pageId="14" pageNumber="15">ps1:ps1</emphasis>
28,
<emphasis id="B948EAA553225430FAAEDF8EE23CFF0C" box="[1325,1391,152,177]" italics="true" pageId="14" pageNumber="15">e2:h2</emphasis>
50,
<emphasis id="B948EAA553225430FF14DFACE788FF6E" box="[151,219,186,211]" italics="true" pageId="14" pageNumber="15">h2:h3</emphasis>
22,
<emphasis id="B948EAA553225430FE92DFACE600FF6E" box="[273,339,186,211]" italics="true" pageId="14" pageNumber="15">
h2:
<geoCoordinate id="EE08507053225430FEBADFACE600FF6E" box="[313,339,186,211]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="1.1333333">e1</geoCoordinate>
</emphasis>
8,
<emphasis id="B948EAA553225430FEF9DFACE699FF6E" box="[378,458,186,211]" italics="true" pageId="14" pageNumber="15">ps1:h3</emphasis>
17. Genital apparatus
<date id="FF82107753225430FD54DFACE46BFF69" box="[727,824,186,212]" pageId="14" pageNumber="15" value="1928-10-18">28 x 18.</date>
Distance between ventral setae:
<emphasis id="B948EAA553225430FB3CDFACE257FF6E" box="[1215,1284,186,211]" italics="true" pageId="14" pageNumber="15">3b:3a</emphasis>
12,
<emphasis id="B948EAA553225430FABADFACE23CFF6E" box="[1337,1391,186,211]" italics="true" pageId="14" pageNumber="15">3a:g</emphasis>
18,
<emphasis id="B948EAA553225430FF14DFCBE79FFF4B" box="[151,204,221,246]" italics="true" pageId="14" pageNumber="15">g:4a</emphasis>
58,
<emphasis id="B948EAA553225430FE83DFCBE61CFF4B" box="[256,335,221,246]" italics="true" pageId="14" pageNumber="15">4a:ps3</emphasis>
100. Diameter of anal suckers 20.
</paragraph>
<paragraph id="8B8336B753225430FF47DE16E4CAFD09" blockId="14.[151,1437,152,1364]" pageId="14" pageNumber="15">
Female (2
<typeStatus id="5487881553225430FEC5DE16E6AAFEA7" box="[326,505,256,282]" pageId="14" pageNumber="15" type="paralectotype">paralectotypes</typeStatus>
). Gnathosoma as in homeomorph male. Idiosoma, length x width, 440450 x 242250. Prodorsal shield with posterior margin slightly concave; length along midline 118128. Setae
<emphasis id="B948EAA553225430FA0BDE33E2CFFE83" box="[1416,1436,293,318]" italics="true" pageId="14" pageNumber="15">si</emphasis>
setiform, 1012 long. Distance between scapular setae:
<emphasis id="B948EAA553225430FCA6DE5CE433FEDE" box="[805,864,330,355]" italics="true" pageId="14" pageNumber="15">se:se</emphasis>
7578,
<emphasis id="B948EAA553225430FC43DE5CE4A2FEDE" box="[960,1009,330,355]" italics="true" pageId="14" pageNumber="15">si:si</emphasis>
4550. Hysterosoma 320325 long. Opisthosoma with short bluntly rounded opisthosomal lobes, terminal cleft as a small inverted U (
<figureCitation id="13072A3253225430FAA2DE66E226FE37" box="[1313,1397,368,394]" captionStart="FIGURE 11" captionStartId="21.[151,255,1560,1584]" captionTargetBox="[158,1420,460,1559]" captionTargetId="figure@21.[151,1436,450,1574]" captionTargetPageId="21" captionText="FIGURE 11. Protolichus females, dorsal view. A— Protolichus strangulatus, B— P. magnificus." httpUri="https://zenodo.org/record/196436/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
A). Length of hysteronotal shield 300315, width 205210, anterior margin slightly concave, most of surface monotonously punctate with faint net-like striations, medial area posterior to level of setae
<emphasis id="B948EAA553225430FB3EDEACE38AFE6E" box="[1213,1241,442,467]" italics="true" pageId="14" pageNumber="15">e2</emphasis>
with several (6 10) lacunae; subtegumental sclerotized bars in postero-lateral parts of opisthosoma narrow. Setae
<emphasis id="B948EAA553225430FA93DEF6E278FE44" box="[1296,1323,480,505]" italics="true" pageId="14" pageNumber="15">c3</emphasis>
setiform, 1518 long. Setae
<emphasis id="B948EAA553225430FEF0DD13E6DCFDA3" box="[371,399,517,542]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
,
<emphasis id="B948EAA553225430FE1DDD13E690FDA3" box="[414,451,517,542]" italics="true" pageId="14" pageNumber="15">d2,</emphasis>
setiform, short, setae
<emphasis id="B948EAA553225430FD4BDD13E5B0FDA3" box="[712,739,517,542]" italics="true" pageId="14" pageNumber="15">e2</emphasis>
flattened, saber-shaped; setae
<emphasis id="B948EAA553225430FBCFDD13E3C9FDA3" box="[1100,1178,517,542]" italics="true" pageId="14" pageNumber="15">f2, ps2</emphasis>
slightly flattened and enlarged in middle part,
<emphasis id="B948EAA553225430FE3ADD3CE683FDFE" box="[441,464,554,579]" italics="true" pageId="14" pageNumber="15">f2</emphasis>
usually with additional spine on outer margin; setae
<emphasis id="B948EAA553225430FBC6DD3CE331FDFE" box="[1093,1122,554,579]" italics="true" pageId="14" pageNumber="15">e1</emphasis>
situated between levels of openings
<emphasis id="B948EAA553225430FE8BDD46E64CFDD4" box="[264,287,592,617]" italics="true" pageId="14" pageNumber="15">gl</emphasis>
and cupules
<emphasis id="B948EAA553225430FE38DD46E68BFDD4" box="[443,472,592,617]" italics="true" pageId="14" pageNumber="15">im</emphasis>
. Length of hysteronotal setae:
<emphasis id="B948EAA553225430FCC5DD46E432FDD4" box="[838,865,592,617]" italics="true" pageId="14" pageNumber="15">
c
<date id="FF82107753225430FCD0DD46E432FDD4" box="[851,865,592,617]" pageId="14" pageNumber="15" value="1910-02-12">2</date>
</emphasis>
1012,
<emphasis id="B948EAA553225430FC41DD46E48CFDD4" box="[962,991,592,617]" italics="true" pageId="14" pageNumber="15">d2</emphasis>
5,
<geoCoordinate id="EE08507053225430FB86DD46E315FDD7" box="[1029,1094,592,618]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="3.0833335">
<emphasis id="B948EAA553225430FB86DD46E372FDD4" box="[1029,1057,592,617]" italics="true" pageId="14" pageNumber="15">e2</emphasis>
65
</geoCoordinate>
68,
<emphasis id="B948EAA553225430FB02DD46E3CBFDD4" box="[1153,1176,592,617]" italics="true" pageId="14" pageNumber="15">f2</emphasis>
40. Distance between setae:
<emphasis id="B948EAA553225430FF62DD63E677FD33" box="[225,292,629,654]" italics="true" pageId="14" pageNumber="15">c2:d2</emphasis>
115125,
<emphasis id="B948EAA553225430FE1CDD63E6B1FD33" box="[415,482,629,654]" italics="true" pageId="14" pageNumber="15">d2:e2</emphasis>
128130,
<emphasis id="B948EAA553225430FDDDDD63E5F2FD33" box="[606,673,629,654]" italics="true" pageId="14" pageNumber="15">e2:h3</emphasis>
6870,
<emphasis id="B948EAA553225430FD7CDD63E412FD33" box="[767,833,629,654]" italics="true" pageId="14" pageNumber="15">e2:e2</emphasis>
, 140145,
<emphasis id="B948EAA553225430FC47DD63E35AFD33" box="[964,1033,629,654]" italics="true" pageId="14" pageNumber="15">h2:h2</emphasis>
7580,
<emphasis id="B948EAA553225430FBEBDD63E390FD33" box="[1128,1219,629,654]" italics="true" pageId="14" pageNumber="15">
ps1:ps
<date id="FF82107753225430FB34DD63E390FD33" box="[1207,1219,629,654]" pageId="14" pageNumber="15" value="1920-01-22">1</date>
</emphasis>
2022. Epigynum bow-shaped,
<date id="FF82107753225430FEB0DD8CE6EBFD09" box="[307,440,666,692]" pageId="14" pageNumber="15" value="1940-10-13" valueMax="1940-10-15" valueMin="1940-10-13">1315 x 40</date>
42. Seta
<emphasis id="B948EAA553225430FDA8DD8CE506FD0E" box="[555,597,666,691]" italics="true" pageId="14" pageNumber="15">mG</emphasis>
of genu I thin, spiculiform.
</paragraph>
</subSubSection>
<subSubSection id="C326653C53225430FF45DDA9E338FD67" box="[198,1131,703,730]" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8B8336B753225430FF45DDA9E338FD67" blockId="14.[151,1437,152,1364]" box="[198,1131,703,730]" pageId="14" pageNumber="15">
<emphasis id="B948EAA553225430FF45DDA9E68DFD64" bold="true" box="[198,478,703,729]" pageId="14" pageNumber="15">
<typeStatus id="5487881553225430FF45DDA9E650FD64" box="[198,259,703,729]" pageId="14" pageNumber="15">Type</typeStatus>
host and locality.
</emphasis>
<taxonomicName id="4C3C4D3453225430FE66DDD6E5BCFD64" box="[485,751,704,729]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553225430FE66DDD6E5BCFD64" box="[485,751,704,729]" italics="true" pageId="14" pageNumber="15">Charmosyna pulchella</emphasis>
</taxonomicName>
(designated here), New
<collectingCountry id="F32B762753225430FB8DDDD6E335FD67" box="[1038,1126,704,730]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C326653C53225430FF45DDF3E512FAE9" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B8336B753225430FF45DDF3E565FB59" blockId="14.[151,1437,152,1364]" pageId="14" pageNumber="15">
<emphasis id="B948EAA553225430FF45DDF3E667FD42" bold="true" box="[198,308,741,767]" pageId="14" pageNumber="15">Remark.</emphasis>
<bibRefCitation id="EFAD4B4653225430FEC2DDF3E5E2FD42" author="Favette" box="[321,689,741,767]" pageId="14" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">Favette and Trouessart (1904)</bibRefCitation>
described this species as a subspecies of
<taxonomicName id="4C3C4D3453225430FB47DDF3E237FD43" box="[1220,1380,741,766]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553225430FB47DDF3E237FD43" box="[1220,1380,741,766]" italics="true" pageId="14" pageNumber="15">P. brachiatus</emphasis>
</taxonomicName>
and mentioned two hosts,
<taxonomicName id="4C3C4D3453225430FE1EDC1CE50BFC9E" box="[413,600,778,803]" class="Aves" family="Psittacidae" genus="Lorius" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="garrulus">
<emphasis id="B948EAA553225430FE1EDC1CE50BFC9E" box="[413,600,778,803]" italics="true" pageId="14" pageNumber="15">Lorius garrulus</emphasis>
</taxonomicName>
and
<emphasis id="B948EAA553225430FD10DC1CE4FBFC9E" box="[659,936,778,803]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3C4D3453225430FD10DC1CE4F7FC9E" box="[659,932,778,803]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">Charmosyna pulchella</taxonomicName>
,
</emphasis>
from New
<collectingCountry id="F32B762753225430FBB1DC1CE3D8FC99" box="[1074,1163,778,804]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
. There are formally no specimens labeled as “
<emphasis id="B948EAA553225430FE21DC26E516FCF4" box="[418,581,816,841]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3C4D3453225430FE21DC26E569FCF4" box="[418,570,816,841]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">strangulatus</taxonomicName>
</emphasis>
in the Trouessart collection. Nevertheless, this collection contains slides with a specimen from
<taxonomicName id="4C3C4D3453225430FE1EDC43E587FCD3" box="[413,724,853,878]" class="Aves" family="Psittacidae" genus="Charmosynopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553225430FE1EDC43E587FCD3" box="[413,724,853,878]" italics="true" pageId="14" pageNumber="15">Charmosynopsis pulchella</emphasis>
</taxonomicName>
(recent name
<taxonomicName id="4C3C4D3453225430FCFFDC43E3D5FCD3" box="[892,1158,853,878]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553225430FCFFDC43E3D5FCD3" box="[892,1158,853,878]" italics="true" pageId="14" pageNumber="15">Charmosyna pulchella</emphasis>
</taxonomicName>
) from New
<collectingCountry id="F32B762753225430FA95DC43E239FCD2" box="[1302,1386,853,879]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
and identified by Trouessart as “
<emphasis id="B948EAA553225430FE64DC6CE5C7FC2E" box="[487,660,890,915]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3C4D3453225430FE64DC6CE5DAFC2E" box="[487,649,890,915]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">P. brachiatus</taxonomicName>
</emphasis>
. Probably it was a preliminary identification, because neither the original description of
<taxonomicName id="4C3C4D3453225430FE26DCB6E512FC04" box="[421,577,928,953]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553225430FE26DCB6E512FC04" box="[421,577,928,953]" italics="true" pageId="14" pageNumber="15">P. brachiatus</emphasis>
</taxonomicName>
nor the subsequent revision of
<taxonomicName id="4C3C4D3453225430FC35DCB6E36DFC04" box="[950,1086,928,953]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAA553225430FC35DCB6E36DFC04" box="[950,1086,928,953]" italics="true" pageId="14" pageNumber="15">Protolichus</emphasis>
</taxonomicName>
listed
<taxonomicName id="4C3C4D3453225430FB0DDCB6E27EFC04" box="[1166,1325,928,953]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553225430FB0DDCB6E27EFC04" box="[1166,1325,928,953]" italics="true" pageId="14" pageNumber="15">Ch. pulchella</emphasis>
</taxonomicName>
as a host of
<taxonomicName id="4C3C4D3453225430FF39DCD3E618FC63" box="[186,331,965,990]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B948EAA553225430FF39DCD3E618FC63" box="[186,331,965,990]" italics="true" pageId="14" pageNumber="15">Protolichus</emphasis>
</taxonomicName>
species other than
<emphasis id="B948EAA553225430FDC3DCD3E44DFC63" box="[576,798,965,990]" italics="true" pageId="14" pageNumber="15">
P. b.
<taxonomicName id="4C3C4D3453225430FD02DCD3E44DFC63" box="[641,798,965,990]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">strangulatus</taxonomicName>
</emphasis>
. Based on this indirect evidence, we consider that specimens in three slides (34F1) from
<taxonomicName id="4C3C4D3453225430FD03DCFCE479FBBE" box="[640,810,1002,1027]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B948EAA553225430FD03DCFCE479FBBE" box="[640,810,1002,1027]" italics="true" pageId="14" pageNumber="15">Ch. pulchella</emphasis>
</taxonomicName>
represent a part of
<typeStatus id="5487881553225430FBA7DCFCE309FBB9" box="[1060,1114,1002,1028]" pageId="14" pageNumber="15">type</typeStatus>
series of
<taxonomicName id="4C3C4D3453225430FB54DCFCE2C7FBBE" box="[1239,1428,1002,1027]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553225430FB54DCFCE2C7FBBE" box="[1239,1428,1002,1027]" italics="true" pageId="14" pageNumber="15">P. strangulatus</emphasis>
</taxonomicName>
. Additional evidence is a very clear correspondence of the homeomorph male in this slides to the photo of
<emphasis id="B948EAA553225430FAE9DB06E67FFBF3" italics="true" pageId="14" pageNumber="15">
P. b.
<taxonomicName id="4C3C4D3453225430FF14DB23E67FFBF3" box="[151,300,1077,1102]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">strangulatus</taxonomicName>
</emphasis>
given by
<bibRefCitation id="EFAD4B4653225430FE26DB23E4EAFBF2" author="Favette" box="[421,953,1077,1103]" pageId="14" pageNumber="36" refString="Favette, J. &amp; Trouessart, E. L. (1904) Monographie du genre Protolichus (Trt) et revision des Sarcoptides plumicoles (Analgesinae) qui vivent sur les perroquets. Memoires de la Societe zoologique de France, 17, 120 - 166 + pls. V - XV." type="journal article" year="1904">
Favette and Trouessart (1904: Pl. V,
<figureCitation id="13072A3253225430FCDDDB23E4FAFBF2" box="[862,937,1077,1103]" captionStart="FIGURE 4" captionStartId="10.[151,255,1275,1299]" captionTargetBox="[160,1422,223,1234]" captionTargetId="figure@10.[151,1436,194,1254]" captionTargetPageId="10" captionText="FIGURE 4. Protolichus brachiatus. A—leg I of homeomorph male, dorsal view, B—leg II of homeomorph male, dorsal view, C—gnathosoma of homeomorph male, ventral view, D—leg I of female, dorsal view, E—leg II of female, dorsal view, F—tarsus III of female, dorsal view, G—tarsus IV of female, dorsal view." httpUri="https://zenodo.org/record/196429/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
.)
</bibRefCitation>
, particularly in respect to a very narrow terminal cleft (compare
<figureCitation id="13072A3253225430FE32DB4CE6A7FBC9" box="[433,500,1114,1140]" captionStart="FIGURE 5" captionStartId="12.[151,258,1961,1985]" captionTargetBox="[213,1394,197,1906]" captionTargetId="figure@12.[183,1399,197,1934]" captionTargetPageId="12" captionText="FIGURE 5. Dorsal view of male opisthosoma of Protolichus brachiatus (A, B) and P. strangulatus (C, D). A, Cheteromorph male, B, D—homeomorph male." httpUri="https://zenodo.org/record/196430/files/figure.png" pageId="14" pageNumber="15">Fig. 5</figureCitation>
B with 5D), a feature that was specifically stressed in the original description of this species as the character discriminating it from
<taxonomicName id="4C3C4D3453225430FD7BDB96E4C6FB24" box="[760,917,1152,1177]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553225430FD7BDB96E4C6FB24" box="[760,917,1152,1177]" italics="true" pageId="14" pageNumber="15">P. brachiatus</emphasis>
</taxonomicName>
. It is necessary to add that one male in this slide is actually a heteromorph one. Based on the facts above, we mark the
<typeStatus id="5487881553225430FB8ADBB3E327FB02" box="[1033,1140,1189,1215]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C3C4D3453225430FB1FDBB3E218FB03" box="[1180,1355,1189,1214]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553225430FB1FDBB3E218FB03" box="[1180,1355,1189,1214]" italics="true" pageId="14" pageNumber="15">P. strangulatus</emphasis>
</taxonomicName>
, in the specimen 34F1 from
<emphasis id="B948EAA553225430FE12DBDDE565FB59" box="[401,566,1227,1252]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3C4D3453225430FE12DBDDE561FB59" box="[401,562,1227,1252]" class="Aves" family="Psittacidae" genus="Charmosyna" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulchella">Ch. pulchella</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B8336B753225430FF47DBE6E512FAE9" blockId="14.[151,1437,152,1364]" pageId="14" pageNumber="15">
Trouessarts (1884) report of
<taxonomicName id="4C3C4D3453225430FDB9DBE6E5B2FAB4" box="[570,737,1264,1289]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">
<emphasis id="B948EAA553225430FDB9DBE6E5B2FAB4" box="[570,737,1264,1289]" italics="true" pageId="14" pageNumber="15">P. brachiatus</emphasis>
</taxonomicName>
from
<taxonomicName id="4C3C4D3453225430FCACDBE6E35CFAB4" box="[815,1039,1264,1289]" class="Aves" family="Psittacidae" genus="Loriculus" kingdom="Animalia" order="Psittaciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="sclateri">
<emphasis id="B948EAA553225430FCACDBE6E35CFAB4" box="[815,1039,1264,1289]" italics="true" pageId="14" pageNumber="15">Loriculus sclateri</emphasis>
</taxonomicName>
is quite obviously the result of contamination. Examination of the slide 34F14 determined as
<emphasis id="B948EAA553225430FC26DA03E300FA93" box="[933,1107,1301,1326]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3C4D3453225430FC26DA03E31EFA93" box="[933,1101,1301,1326]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brachiatus">P. brachiatus</taxonomicName>
,
</emphasis>
has shown that it actually contains a female of
<taxonomicName id="4C3C4D3453225430FE08DA2DE569FAE9" box="[395,570,1339,1364]" class="Arachnida" family="Pterolichidae" genus="Protolichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="strangulatus">
<emphasis id="B948EAA553225430FE08DA2DE569FAE9" box="[395,570,1339,1364]" italics="true" pageId="14" pageNumber="15">P. strangulatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>