439 lines
56 KiB
XML
439 lines
56 KiB
XML
<document id="27B29721DC1EFD6ADA470C3324C3A2D3" ID-CLB-Dataset="45643" ID-DOI="10.5281/zenodo.215499" ID-GBIF-Dataset="3d66a055-e64c-4ff7-9f9a-e4340ffd9105" ID-ISSN="1175-5326" ID-Zenodo-Dep="215499" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460428309642" checkinUser="plazi" docAuthor="Richard, Jasmine, Grave, Sammy De & Clark, Paul F." docDate="2012" docId="03958796FFC4FFE2FF7EFE95252AFBA3" docLanguage="en" docName="zt03162p038.pdf" docOrigin="Zootaxa 3162" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Monsamnis Richard, Grave & Clark, 2012, gen. nov." docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FFACFFEEFFC5FFE0FFE9FFF2263CFFFB" masterDocTitle="A new atyid genus and species from Madagascar (Crustacea: Decapoda: Caridea)" masterLastPageNumber="38" masterPageNumber="31" pageNumber="32" updateTime="1698330938830" updateUser="ExternalLinkService">
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<mods:title id="AF8911DF981DB10917E766127DD926ED">A new atyid genus and species from Madagascar (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="53AF0E20BAA83DD8AD6E456110601676">Richard, Jasmine</mods:namePart>
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<mods:namePart id="20090AA1262C1F185B5A44DA6D275C3A">Grave, Sammy De</mods:namePart>
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<mods:namePart id="03E61E036BBAD79FF44FF3511A519F27">Clark, Paul F.</mods:namePart>
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<mods:date id="6B4E746D9EEA4E5CA7A9E54B825065F3">2012</mods:date>
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<mods:number id="23097C4AF91FFCF391D35581F2780A6E">3162</mods:number>
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<treatment id="03958796FFC4FFE2FF7EFE95252AFBA3" ID-DOI="http://doi.org/10.5281/zenodo.6177496" ID-GBIF-Taxon="119429194" ID-Zenodo-Dep="6177496" LSID="urn:lsid:plazi:treatment:03958796FFC4FFE2FF7EFE95252AFBA3" httpUri="http://treatment.plazi.org/id/03958796FFC4FFE2FF7EFE95252AFBA3" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32">
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<subSubSection id="C326650BFFC4FFE1FF7EFE9527A5FE7A" box="[151,409,359,385]" pageId="1" pageNumber="32" type="nomenclature">
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<paragraph id="8B833680FFC4FFE1FF7EFE9527A5FE7A" blockId="1.[151,409,359,385]" box="[151,409,359,385]" pageId="1" pageNumber="32">
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<taxonomicName id="4C3C4D03FFC4FFE1FF7EFE952719FE7A" ID-CoL="8VJXW" box="[151,293,359,385]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B948EA92FFC4FFE1FF7EFE952719FE7A" bold="true" box="[151,293,359,385]" italics="true" pageId="1" pageNumber="32">Monsamnis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B57E9FFC4FFE1FEC5FE9527A5FE7A" box="[300,409,359,385]" pageId="1" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326650BFFC4FFE1FF7EFE5D2259FE3C" box="[151,1125,430,455]" pageId="1" pageNumber="32" type="materials_examined">
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<paragraph id="8B833680FFC4FFE1FF7EFE5D2259FE3C" blockId="1.[151,1437,430,2003]" box="[151,1125,430,455]" pageId="1" pageNumber="32">
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<emphasis id="B948EA92FFC4FFE1FF7EFE5D270EFE3C" bold="true" box="[151,306,430,455]" pageId="1" pageNumber="32">
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<typeStatus id="54878822FFC4FFE1FF7EFE5D26EEFE3C" box="[151,210,431,455]" pageId="1" pageNumber="32">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3C4D03FFC4FFE1FED0FE5D2471FE3D" box="[313,589,430,454]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="carpolongus" status="sp. nov.">
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<emphasis id="B948EA92FFC4FFE1FED0FE5D2471FE3D" box="[313,589,430,454]" italics="true" pageId="1" pageNumber="32">Monsamnis carpolongus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B57E9FFC4FFE1FDBDFE5D2492FE3C" box="[596,686,431,455]" pageId="1" pageNumber="32" rank="species">
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<emphasis id="B948EA92FFC4FFE1FDBDFE5D2492FE3C" bold="true" box="[596,686,431,455]" pageId="1" pageNumber="32">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, by present designation and monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326650BFFC4FFE1FF2EFE2022A1FDAD" pageId="1" pageNumber="32" type="diagnosis">
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<paragraph id="8B833680FFC4FFE1FF2EFE2022A1FDAD" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="32">
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<emphasis id="B948EA92FFC4FFE1FF2EFE20277DFE10" bold="true" box="[199,321,466,491]" pageId="1" pageNumber="32">Diagnosis.</emphasis>
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Carapace without supraorbital tooth, suborbital margin not produced, rostrum short and unarmed, exopods present on all pereiopods (fifth vestigial or reduced), carpus of second pereiopod elongated 7–9 × long as broad with anterior excavation, uropod diaeresis with a single spine, appendix masculina long and armed with numerous spiniform setae, female carrying ca. 14 large eggs, 1.4–1.5 × 1.00–
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<quantity id="4CC49B65FFC4FFE1FC10FDCC227AFDAD" box="[1017,1094,574,598]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="32" unit="mm" value="1.1">1.1mm</quantity>
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in size.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326650BFFC4FFE1FF2EFD912318FD65" pageId="1" pageNumber="32" type="etymology">
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<paragraph id="8B833680FFC4FFE1FF2EFD912318FD65" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="32">
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<emphasis id="B948EA92FFC4FFE1FF2EFD912772FD80" bold="true" box="[199,334,611,635]" pageId="1" pageNumber="32">Etymology.</emphasis>
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The genus name is formed by an arbitrary combination from the Latin
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<emphasis id="B948EA92FFC4FFE1FB6CFD9622FCFD80" box="[1157,1216,612,635]" italics="true" pageId="1" pageNumber="32">mons</emphasis>
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meaning mountain and
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<emphasis id="B948EA92FFC4FFE1FF2FFD752735FD65" box="[198,265,647,670]" italics="true" pageId="1" pageNumber="32">amnis</emphasis>
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meaning stream; for the habitat in which these crustaceans were collected; gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326650BFFC4FFE2FF2EFD5924A5FE7C" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32" type="discussion">
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<paragraph id="8B833680FFC4FFE1FF2EFD592436FB6D" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="32">
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<emphasis id="B948EA92FFC4FFE1FF2EFD592707FD38" bold="true" box="[199,315,683,707]" pageId="1" pageNumber="32">Remarks.</emphasis>
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The traditional characters used to distinguish atyid subfamilies (série sensu
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<bibRefCitation id="EFAD4B71FFC4FFE1FB77FD582303FD39" author="Bouvier" box="[1182,1343,682,706]" pageId="1" pageNumber="32" refString="Bouvier, E. - L. (1925) Recherches sur la morphologie, les variations et la distribution systematique des crevettes d'eau douce de la famille des Atyides. Encyclopedie Entomologique, 4, 1 - 365." type="journal article" year="1925">Bouvier, 1925</bibRefCitation>
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) are the relative development of the exopods on the pereiopods, branchial formulae, pigment presence and reduction in eyes, number of spines on the uropodal diaresis, as well as the shape and proportions of the chelipeds (see
|
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<bibRefCitation id="EFAD4B71FFC4FFE1FADCFD0026EEFCD5" author="Holthuis" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1986) A new genus and species of subterranean shrimp from Western Australia (Crustacea: Decapoda: Atyidae). Zoologische Mededelingen, 60, 103 - 111." type="journal article" year="1986">Holthuis, 1986</bibRefCitation>
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,
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<bibRefCitation id="EFAD4B71FFC4FFE1FF35FCE42724FCD5" author="Holthuis" box="[220,280,790,814]" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1993) The recent genera of the Caridean and Stenopodidean shrimps (Crustacea, Decapoda) with an appendix on the order Amphionidacea. Leiden, Nationaal Natuurhistorisch Museum. 1 - 328." type="book chapter" year="1993">1993</bibRefCitation>
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||
). Four subfamilies are recognised in the traditional classification, three of which were originally termed “série” in
|
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<bibRefCitation id="EFAD4B71FFC4FFE1FEE1FCC82789FCA9" author="Bouvier" box="[264,437,826,850]" pageId="1" pageNumber="32" refString="Bouvier, E. - L. (1925) Recherches sur la morphologie, les variations et la distribution systematique des crevettes d'eau douce de la famille des Atyides. Encyclopedie Entomologique, 4, 1 - 365." type="journal article" year="1925">Bouvier (1925)</bibRefCitation>
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, viz. caridienne, caridellienne, paratyienne; with the série typhlatienne added by
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<bibRefCitation id="EFAD4B71FFC4FFE1FAD5FCC826DDFC8D" author="Holthuis" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1965) The Atyidae of Madagascar. Memoires du Museum National D'Histoire Naturelle, nouvelle serie, Serie A. Zoologie, 33 (1), 1 - 48." type="journal article" year="1965">Holthuis (1965)</bibRefCitation>
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(see also
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<bibRefCitation id="EFAD4B71FFC4FFE1FEB1FCAC245FFC8D" author="Monod" box="[344,611,862,886]" pageId="1" pageNumber="32" refString="Monod, T. & Cals, P. (1970) Sur une espece nouvelle de crevette cavernicole: Typhlatya galapagensis (Decapoda, Natantia; Atyidae). In: Mission Zoologique Belge aux iles Galapagos et en Ecuador (N. et J. Leleup, 1964 - 1965) Resu 1 tats Scientifiques. Musee Royal de l'Afrique Centrale, Institut Royal des sciences naturelles de Belgique, Charles Darwin Foundation for the Galapagos Isles, Brussels, 2, 57 - 103, 67 figs." type="journal article" year="1970">Monod and Cals, 1970</bibRefCitation>
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). All four were formally elevated to subfamily level in
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<bibRefCitation id="EFAD4B71FFC4FFE1FB01FCAC23A0FC8D" author="Holthuis" box="[1256,1436,862,886]" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1986) A new genus and species of subterranean shrimp from Western Australia (Crustacea: Decapoda: Atyidae). Zoologische Mededelingen, 60, 103 - 111." type="journal article" year="1986">Holthuis (1986)</bibRefCitation>
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(série caridienne as
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<taxonomicName id="4C3C4D03FFC4FFE1FE91FC7027E9FC61" box="[376,469,898,922]" class="Malacostraca" family="Atyinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="family">Atyinae</taxonomicName>
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), although their usage since then has not been consistent in systematic literature (e.g.,
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<bibRefCitation id="EFAD4B71FFC4FFE1FF7EFC542794FC45" author="Sanz" box="[151,424,934,958]" pageId="1" pageNumber="32" refString="Sanz, S. & Platvoet, D. (1995) New perspectives on the evolution of the genus Typhlatya (Crustacea, Decapoda): first record of a cavernicolous atyid in the Iberian Pensinsula, Typhlatya miravetensis n. sp. Contributions to Zoology, 65, 79 - 99." type="journal article" year="1995">Sanz and Platvoet, 1995</bibRefCitation>
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;
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<bibRefCitation id="EFAD4B71FFC4FFE1FE5AFC542489FC45" author="Sket" box="[435,693,934,958]" pageId="1" pageNumber="32" refString="Sket, B. & Zaksek, V. (2009) European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society, 155, 786 - 818." type="journal article" year="2009">Sket and Zakšek, 2009</bibRefCitation>
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), resulting in them not being recognised in
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<bibRefCitation id="EFAD4B71FFC4FFE1FB71FC5523A4FC45" box="[1176,1432,934,959]" pageId="1" pageNumber="32" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D., & Wetzer, R. (2009) A Classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, Supplement 21, 1 - 109." type="book chapter">
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<collectingCountry id="F32B7610FFC4FFE1FB71FC552285FC45" box="[1176,1209,935,958]" name="South Georgia and the South Sandwich Islands" pageId="1" pageNumber="32">De</collectingCountry>
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Grave
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<emphasis id="B948EA92FFC4FFE1FAE5FC5A2300FC45" box="[1292,1340,934,959]" italics="true" pageId="1" pageNumber="32">et al</emphasis>
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. (2009)
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</bibRefCitation>
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. A growing body of literature, primarily molecular based (e.g.,
|
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<bibRefCitation id="EFAD4B71FFC4FFE1FCBCFC392228FC19" author="Page" box="[853,1044,970,995]" pageId="1" pageNumber="32" refString="Page, T. J., Rintelen K. von & Hughes, J. M. (2007) Phylogenetic and biogeographic relationships of subterranean and surface genera of Australian Atyidae (Crustacea: Decapoda; Atyidae) inferred with mitochondrial DNA. Invertebrate Systematics, 21, 137 - 145." type="journal article" year="2007">
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Page
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||
<emphasis id="B948EA92FFC4FFE1FC7AFC3E25F8FC18" box="[915,964,971,995]" italics="true" pageId="1" pageNumber="32">et al</emphasis>
|
||
., 2007
|
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</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FBF6FC382266FC19" author="Page" box="[1055,1114,970,994]" pageId="1" pageNumber="32" refString="Page, T. J., Humphreys, W. F. & Hughes, J. M. (2008) Shrimps down under: evolutionary relationships of subterranean crustaceans from Western Australia (Decapoda: Atyidae: Stygiocaris). Plos ONE 3 (2): e 1618." type="journal article" year="2008">2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FB8FFC382354FC19" author="Sket" box="[1126,1384,970,994]" pageId="1" pageNumber="32" refString="Sket, B. & Zaksek, V. (2009) European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society, 155, 786 - 818." type="journal article" year="2009">Sket and Zakšek, 2009</bibRefCitation>
|
||
) has shed considerable doubt on some of the characters used by
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FCC6FC1C25DFFBFD" author="Holthuis" box="[815,995,1006,1030]" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1965) The Atyidae of Madagascar. Memoires du Museum National D'Histoire Naturelle, nouvelle serie, Serie A. Zoologie, 33 (1), 1 - 48." type="journal article" year="1965">Holthuis (1965)</bibRefCitation>
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, especially the presence and number of exopods as well as the branchial formulae, both of which are perhaps better interpreted as environmental adaptations, such as low oxygen environments and a troglobitic lifestyle.
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||
<collectingCountry id="F32B7610FFC4FFE1FC7DFBC5258FFBB4" box="[916,947,1079,1103]" name="American Samoa" pageId="1" pageNumber="32">As</collectingCountry>
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the true phylogenetic significance of these characters will not be resolved for some time, the relationship of
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<taxonomicName id="4C3C4D03FFC4FFE1FC64FBAE2232FB88" box="[909,1038,1116,1139]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
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||
<emphasis id="B948EA92FFC4FFE1FC64FBAE2232FB88" box="[909,1038,1116,1139]" italics="true" pageId="1" pageNumber="32">Monsamnis</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A27B57E9FFC4FFE1FBF1FBA922BFFB88" box="[1048,1155,1115,1139]" pageId="1" pageNumber="32" rank="genus">
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||
<emphasis id="B948EA92FFC4FFE1FBF1FBA922BFFB88" bold="true" box="[1048,1155,1115,1139]" pageId="1" pageNumber="32">gen. nov.</emphasis>
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</taxonomicNameLabel>
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||
to the majority of other atyid genera is relatively unclear.
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</paragraph>
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<paragraph id="8B833680FFC4FFE1FF2EFB502429FA69" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="32">
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Based on characters (presence of exopods, branchial formula, absence of carapacial teeth) examined for the present study, the new genus is related to the troglobitic and anchialine genera, traditionally included in the subfamily Typhlatyinae.
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FE6FFB192470FAF9" box="[390,588,1258,1283]" pageId="1" pageNumber="32" refString="Page, T. J., Humphreys, W. F. & Hughes, J. M. (2008) Shrimps down under: evolutionary relationships of subterranean crustaceans from Western Australia (Decapoda: Atyidae: Stygiocaris). Plos ONE 3 (2): e 1618." type="journal article">
|
||
Page
|
||
<emphasis id="B948EA92FFC4FFE1FE2AFB1E27C8FAF8" box="[451,500,1259,1283]" italics="true" pageId="1" pageNumber="32">et al</emphasis>
|
||
. (2008)
|
||
</bibRefCitation>
|
||
already provided support to recognise a série typhlatienne, if perhaps not in a formal Linnean hierarchy, but justifying contrasting the new genus to those genera currently included in that subfamily. Recent molecular phylogenetic work (Von Rintelen
|
||
<emphasis id="B948EA92FFC4FFE1FCDBFAC6255FFAB0" box="[818,867,1331,1355]" italics="true" pageId="1" pageNumber="32">et al</emphasis>
|
||
. in review) has re-emphasised that the “série” proposed by
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FEEDFAA4278CFA95" author="Bouvier" box="[260,432,1366,1390]" pageId="1" pageNumber="32" refString="Bouvier, E. - L. (1925) Recherches sur la morphologie, les variations et la distribution systematique des crevettes d'eau douce de la famille des Atyides. Encyclopedie Entomologique, 4, 1 - 365." type="journal article" year="1925">Bouvier (1925)</bibRefCitation>
|
||
are indeed natural groupings, although in a somewhat modified composition, and a série typhlatienne is indeed recognised.
|
||
</paragraph>
|
||
<paragraph id="8B833680FFC4FFE1FF2EFA6C234EF855" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="32">
|
||
According to
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FE83FA6C241FFA4D" author="Holthuis" box="[362,547,1438,1462]" pageId="1" pageNumber="32" refString="Holthuis, L. B. (1986) A new genus and species of subterranean shrimp from Western Australia (Crustacea: Decapoda: Atyidae). Zoologische Mededelingen, 60, 103 - 111." type="journal article" year="1986">Holthuis (1986)</bibRefCitation>
|
||
, this subfamily (série) comprises of
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FC31FA6C2256FA4D" box="[984,1130,1438,1462]" class="Malacostraca" family="Atyidae" genus="Antecaridina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FC31FA6C2256FA4D" box="[984,1130,1438,1462]" italics="true" pageId="1" pageNumber="32">Antecaridina</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FB95FA6D22C5FA4D" box="[1148,1273,1439,1462]" class="Malacostraca" family="Atyidae" genus="Stygiocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FB95FA6D22C5FA4D" box="[1148,1273,1439,1462]" italics="true" pageId="1" pageNumber="32">Stygiocaris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FAE5FA6C23A8FA4D" box="[1292,1428,1438,1462]" class="Malacostraca" family="Atyidae" genus="Typhlopatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FAE5FA6C23A8FA4D" box="[1292,1428,1438,1462]" italics="true" pageId="1" pageNumber="32">Typhlopatsa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FF7EFA312739FA20" box="[151,261,1475,1499]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FF7EFA312739FA20" box="[151,261,1475,1499]" italics="true" pageId="1" pageNumber="32">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FED6FA3127ECFA20" box="[319,464,1475,1499]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FED6FA3127ECFA20" box="[319,464,1475,1499]" italics="true" pageId="1" pageNumber="32">Spelaeocaris</emphasis>
|
||
</taxonomicName>
|
||
. The status of the monotypic genus
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FC9AFA312238FA20" box="[883,1028,1475,1499]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FC9AFA312238FA20" box="[883,1028,1475,1499]" italics="true" pageId="1" pageNumber="32">Spelaeocaris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54878822FFC4FFE1FBFCFA30227BFA21" box="[1045,1095,1474,1498]" pageId="1" pageNumber="32">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FB40FA312703FA05" authority="Matjasic, 1956" authorityName="Matjasic" authorityYear="1956" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B948EA92FFC4FFE1FB40FA3123A7FA20" box="[1193,1435,1475,1499]" italics="true" pageId="1" pageNumber="32">Spelaeocaris pretneri</emphasis>
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FF7EFA142703FA05" author="Matjasic" box="[151,319,1510,1534]" pageId="1" pageNumber="32" refString="Matjasic, J. (1956) Ein neuer Hohlendecapode aus Jugoslawien. Zoologischer Anzeiger 157, 65 - 68." type="journal article" year="1956">Matjašič, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
) remains problematic.
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FDAAFA14256AFA05" box="[579,854,1510,1534]" pageId="1" pageNumber="32" refString="Monod, T. & Cals, P. (1970) Sur une espece nouvelle de crevette cavernicole: Typhlatya galapagensis (Decapoda, Natantia; Atyidae). In: Mission Zoologique Belge aux iles Galapagos et en Ecuador (N. et J. Leleup, 1964 - 1965) Resu 1 tats Scientifiques. Musee Royal de l'Afrique Centrale, Institut Royal des sciences naturelles de Belgique, Charles Darwin Foundation for the Galapagos Isles, Brussels, 2, 57 - 103, 67 figs." type="journal article">Monod and Cals, (1970)</bibRefCitation>
|
||
continued to separate this genus from
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FAE2FA142345FA05" box="[1291,1401,1510,1534]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FAE2FA142345FA05" box="[1291,1401,1510,1534]" italics="true" pageId="1" pageNumber="32">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
on the basis that the third and fourth pereiopodal exopodites are reduced, with the fifth being absent. This contrast to
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FF7EF9DC2739F9BD" box="[151,261,1582,1606]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FF7EF9DC2739F9BD" box="[151,261,1582,1606]" italics="true" pageId="1" pageNumber="32">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
in which the exopodites are well developed on the third and fourth pereiopod, and well developed or absent on the fifth (see
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FE77F9A1248AF991" author="Jaume" box="[414,694,1618,1642]" pageId="1" pageNumber="32" refString="Jaume, D. & Brehier, F. (2005) A new species of Typhlatya (Crustacea: Decapoda: Atyidae) from anchialine caves on the French Mediterranean coast. Zoological Journal of the Linnean Society, 144, 387 - 414." type="journal article" year="2005">Jaume and Bréhier, 2005</bibRefCitation>
|
||
for an overview). However,
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FC13F9A02329F991" author="Sanz" box="[1018,1301,1618,1642]" pageId="1" pageNumber="32" refString="Sanz, S. & Platvoet, D. (1995) New perspectives on the evolution of the genus Typhlatya (Crustacea, Decapoda): first record of a cavernicolous atyid in the Iberian Pensinsula, Typhlatya miravetensis n. sp. Contributions to Zoology, 65, 79 - 99." type="journal article" year="1995">Sanz and Platvoet (1995)</bibRefCitation>
|
||
re-assigned
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FF7EF9852732F975" box="[151,270,1655,1678]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B948EA92FFC4FFE1FF7EF9852732F975" box="[151,270,1655,1678]" italics="true" pageId="1" pageNumber="32">S. pretneri</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FEDFF9842798F975" box="[310,420,1654,1678]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FEDFF9842798F975" box="[310,420,1654,1678]" italics="true" pageId="1" pageNumber="32">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
, as they interpreted the reduction in exopod size from P1 through to P5 as a convergent character, given that similar reduced exopods on P3–4 and absence on P5 are also present in
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FB30F969271FF92D" authority="Chace, 1954" authorityName="Chace" authorityYear="1954" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monae">
|
||
<emphasis id="B948EA92FFC4FFE1FB30F96923A0F948" box="[1241,1436,1691,1715]" italics="true" pageId="1" pageNumber="32">Typhlatya monae</emphasis>
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FF7EF94C271FF92D" author="Chace" box="[151,291,1726,1750]" pageId="1" pageNumber="32" refString="Chace, F. A. Jr. (1954) Two new subterranean shrimps (Decapoda: Caridea) from Florida and the West Indies, with a revised key to the American species. Journal of the Washington Academy of Sciences, 44, 318 - 324." type="journal article" year="1954">Chace, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FEB0F94C24F9F92D" authority="Creaser, 1936" authorityName="Creaser" authorityYear="1936" box="[345,709,1726,1750]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="pearsei">
|
||
<emphasis id="B948EA92FFC4FFE1FEB0F94C241DF92D" box="[345,545,1726,1750]" italics="true" pageId="1" pageNumber="32">Typhlatya pearsei</emphasis>
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FDCEF94C24F9F92D" author="Creaser" box="[551,709,1726,1750]" pageId="1" pageNumber="32" refString="Creaser, E. P. (1936) Crustaceans from Yucatan. In: Pearse, A. S. Creaser E. P. & Hall F. G. (eds.) Cenotes of Yucatan: A Zoological and Hydrographic Survey, Washington, Carnegie Institution, 117 - 132." type="book chapter" year="1936">Creaser, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FD26F94D25CFF92D" author="Jaume" box="[719,1011,1726,1750]" pageId="1" pageNumber="32" refString="Jaume, D. & Brehier, F. (2005) A new species of Typhlatya (Crustacea: Decapoda: Atyidae) from anchialine caves on the French Mediterranean coast. Zoological Journal of the Linnean Society, 144, 387 - 414." type="journal article" year="2005">Jaume and Bréhier (2005)</bibRefCitation>
|
||
agreed and maintained the synonymy of both genera. However,
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FE52F91024FAF901" author="Sket" box="[443,710,1762,1786]" pageId="1" pageNumber="32" refString="Sket, B. & Zaksek, V. (2009) European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society, 155, 786 - 818." type="journal article" year="2009">Sket and Zakšek (2009)</bibRefCitation>
|
||
using molecular data from previous studies combined with morphological information of European cave dwelling shrimp taxa, recognised that
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FBF2F8F42290F8E5" box="[1051,1196,1798,1822]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FBF2F8F42290F8E5" box="[1051,1196,1798,1822]" italics="true" pageId="1" pageNumber="32">Spelaeocaris</emphasis>
|
||
</taxonomicName>
|
||
was allied to
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FAA0F8F426F3F8B8" class="Malacostraca" family="Atyidae" genus="Troglocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FAA0F8F426F3F8B8" italics="true" pageId="1" pageNumber="32">Troglocaris</emphasis>
|
||
</taxonomicName>
|
||
(série paratyienne). Further, in that study
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FD4EF8D92504F8B8" box="[679,824,1835,1859]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FD4EF8D92504F8B8" box="[679,824,1835,1859]" italics="true" pageId="1" pageNumber="32">Spelaeocaris</emphasis>
|
||
</taxonomicName>
|
||
was formally recognised as a subgenus of
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FAF3F8D923A6F8B8" box="[1306,1434,1835,1859]" class="Malacostraca" family="Atyidae" genus="Troglocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FAF3F8D923A6F8B8" box="[1306,1434,1835,1859]" italics="true" pageId="1" pageNumber="32">Troglocaris</emphasis>
|
||
</taxonomicName>
|
||
and three new species described in it, as well as a single unassigned specimen, indicative of higher species richness than previously believed (
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FE5EF8802487F871" author="Sket" box="[439,699,1906,1930]" pageId="1" pageNumber="32" refString="Sket, B. & Zaksek, V. (2009) European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society, 155, 786 - 818." type="journal article" year="2009">Sket and Zakšek, 2009</bibRefCitation>
|
||
). Although this may not be the final position of
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FB27F8812363F870" box="[1230,1375,1907,1931]" class="Malacostraca" family="Atyidae" genus="Spelaeocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FB27F8812363F870" box="[1230,1375,1907,1931]" italics="true" pageId="1" pageNumber="32">Spelaeocaris</emphasis>
|
||
</taxonomicName>
|
||
, here
|
||
<bibRefCitation id="EFAD4B71FFC4FFE1FF7EF864279EF855" author="Sket" box="[151,418,1942,1966]" pageId="1" pageNumber="32" refString="Sket, B. & Zaksek, V. (2009) European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society, 155, 786 - 818." type="journal article" year="2009">Sket and Zakšek (2009)</bibRefCitation>
|
||
are followed and this genus is not considered part of Typhlatyinae (série typhlatienne).
|
||
</paragraph>
|
||
<paragraph id="8B833680FFC4FFE2FF2EF84824A5FE7C" blockId="1.[151,1437,430,2003]" lastBlockId="2.[151,1437,151,1112]" lastPageId="2" lastPageNumber="33" pageId="1" pageNumber="32">
|
||
Considering the remaining genera in Typhlatyinae (série typhlatienne),
|
||
<taxonomicName id="4C3C4D03FFC4FFE1FBF1F84E22A5F828" box="[1048,1177,1980,2003]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC4FFE1FBF1F84E22A5F828" box="[1048,1177,1980,2003]" italics="true" pageId="1" pageNumber="32">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC4FFE1FB4EF849232AF828" box="[1191,1302,1979,2003]" pageId="1" pageNumber="32" rank="genus">
|
||
<emphasis id="B948EA92FFC4FFE1FB4EF849232AF828" bold="true" box="[1191,1302,1979,2003]" pageId="1" pageNumber="32">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
appears to occupy an intermediate position between those with a well-developed exopod on the fifth pereiopod and a spiniform, well-developed sub-orbital angle (viz.
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FD7AFF4E2519FF2F" box="[659,805,188,212]" class="Malacostraca" family="Atyidae" genus="Antecaridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD7AFF4E2519FF2F" box="[659,805,188,212]" italics="true" pageId="2" pageNumber="33">Antecaridina</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FCDDFF4F258DFF2F" box="[820,945,189,212]" class="Malacostraca" family="Atyidae" genus="Stygiocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FCDDFF4F258DFF2F" box="[820,945,189,212]" italics="true" pageId="2" pageNumber="33">Stygiocaris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FC29FF4E2274FF2F" box="[960,1096,188,212]" class="Malacostraca" family="Atyidae" genus="Typhlopatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FC29FF4E2274FF2F" box="[960,1096,188,212]" italics="true" pageId="2" pageNumber="33">Typhlopatsa</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FB62FF4E22C5FF2F" box="[1163,1273,188,212]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FB62FF4E22C5FF2F" box="[1163,1273,188,212]" italics="true" pageId="2" pageNumber="33">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
which lacks a produced sub-orbital angle (exopods on fifth pereiopod present, reduced or absent).
|
||
<collectingCountry id="F32B7610FFC7FFE2FB8EFF1222BAFF03" box="[1127,1158,224,248]" name="American Samoa" pageId="2" pageNumber="33">As</collectingCountry>
|
||
the exopod on the fifth pereiopod varies from present to absent in
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FD78FEF624C3FEE7" box="[657,767,260,284]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD78FEF624C3FEE7" box="[657,767,260,284]" italics="true" pageId="2" pageNumber="33">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FCABFEF72255FEE7" author="Jaume" box="[834,1129,260,284]" pageId="2" pageNumber="33" refString="Jaume, D. & Brehier, F. (2005) A new species of Typhlatya (Crustacea: Decapoda: Atyidae) from anchialine caves on the French Mediterranean coast. Zoological Journal of the Linnean Society, 144, 387 - 414." type="journal article" year="2005">Jaume and Bréhier, 2005</bibRefCitation>
|
||
), it is tempting to suggest
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF7EFEDA2739FEBB" box="[151,261,296,320]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FF7EFEDA2739FEBB" box="[151,261,296,320]" italics="true" pageId="2" pageNumber="33">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
could be the sister taxon to
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FDA4FEDB24F2FEBB" box="[589,718,297,320]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FDA4FEDB24F2FEBB" box="[589,718,297,320]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FD3EFEDA257EFEBB" box="[727,834,296,320]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD3EFEDA257EFEBB" bold="true" box="[727,834,296,320]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, a hypothesis refrained upon in the present study, as current biogeographical understanding of this genus would counter argue. Instead the morphological differences between all genera are simply outlined below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326650BFFC7FFE2FF2EFE67252AFBA3" pageId="2" pageNumber="33" type="reference_group">
|
||
<paragraph id="8B833680FFC7FFE2FF2EFE6722D5FD5C" blockId="2.[151,1437,151,1112]" pageId="2" pageNumber="33">
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF2EFE672774FE57" ID-CoL="8VJXW" box="[199,328,405,428]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FF2EFE672774FE57" box="[199,328,405,428]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FEBBFE662783FE57" box="[338,447,404,428]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FEBBFE662783FE57" bold="true" box="[338,447,404,428]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to the anchialine genus,
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FCB4FE6625D3FE57" ID-CoL="8V549" box="[861,1007,404,428]" class="Malacostraca" family="Atyidae" genus="Antecaridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FCB4FE6625D3FE57" box="[861,1007,404,428]" italics="true" pageId="2" pageNumber="33">Antecaridina</emphasis>
|
||
</taxonomicName>
|
||
, in having a short unarmed rostrum, absence of carapacial teeth and exopods present on all pereiopods. The new genus differs from
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FB57FE4A236CFE2B" ID-CoL="8V549" box="[1214,1360,440,464]" class="Malacostraca" family="Atyidae" genus="Antecaridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FB57FE4A236CFE2B" box="[1214,1360,440,464]" italics="true" pageId="2" pageNumber="33">Antecaridina</emphasis>
|
||
</taxonomicName>
|
||
by the possession of well developed eyes (degenerate in
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FD2CFE2E256BFE0F" ID-CoL="8V549" box="[709,855,476,500]" class="Malacostraca" family="Atyidae" genus="Antecaridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD2CFE2E256BFE0F" box="[709,855,476,500]" italics="true" pageId="2" pageNumber="33">Antecaridina</emphasis>
|
||
</taxonomicName>
|
||
) and the long carpus of the second pereiopod (vs. 4 × long as broad, see
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FE60FE0D2413FDEC" author="Holthuis" box="[393,559,511,535]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1965) The Atyidae of Madagascar. Memoires du Museum National D'Histoire Naturelle, nouvelle serie, Serie A. Zoologie, 33 (1), 1 - 48." type="journal article" year="1965">Holthuis 1965</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FDACFDF324FAFDE3" ID-CoL="8VJXW" box="[581,710,513,536]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FDACFDF324FAFDE3" box="[581,710,513,536]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FD26FDF22507FDE3" box="[719,827,512,536]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD26FDF22507FDE3" bold="true" box="[719,827,512,536]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<emphasis id="B948EA92FFC7FFE2FC3FFDF22262FDE3" box="[982,1118,512,536]" italics="true" pageId="2" pageNumber="33">Typhlopasta</emphasis>
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FB81FE0D2331FDEC" author="Holthuis" box="[1128,1293,511,535]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1956) The troglobic Atyidae of Madagascar (Crustacea, Decapoda, Natantia). Memoires de l'Institut Scientifique de Madagascar, series A, 11, 97 - 110." type="journal article" year="1956">Holthuis 1956</bibRefCitation>
|
||
, in having a short unarmed rostrum and no supraorbital teeth, but differs conspicuously from
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FBA3FDD622EEFDC7" ID-CoL="8VJYD" box="[1098,1234,548,572]" class="Malacostraca" family="Atyidae" genus="Typhlopatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FBA3FDD622EEFDC7" box="[1098,1234,548,572]" italics="true" pageId="2" pageNumber="33">Typhlopatsa</emphasis>
|
||
</taxonomicName>
|
||
in having a well developed eye (vs. degenerate) and in having a long carpus on the second pereiopod (vs. 5 as long as broad).
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF7EFD9E2739FD7F" ID-CoL="8VJQZ" box="[151,261,620,644]" class="Malacostraca" family="Atyidae" genus="Typhlatya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FF7EFD9E2739FD7F" box="[151,261,620,644]" italics="true" pageId="2" pageNumber="33">Typhlatya</emphasis>
|
||
</taxonomicName>
|
||
and the Australian genus,
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FDC6FD9F2497FD7F" ID-CoL="8VJY9" box="[559,683,621,644]" class="Malacostraca" family="Atyidae" genus="Stygiocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FDC6FD9F2497FD7F" box="[559,683,621,644]" italics="true" pageId="2" pageNumber="33">Stygiocaris</emphasis>
|
||
</taxonomicName>
|
||
are relatively closely related to each other (
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FB7CFD9E236BFD7F" author="Page" box="[1173,1367,620,644]" pageId="2" pageNumber="33" refString="Page, T. J., Humphreys, W. F. & Hughes, J. M. (2008) Shrimps down under: evolutionary relationships of subterranean crustaceans from Western Australia (Decapoda: Atyidae: Stygiocaris). Plos ONE 3 (2): e 1618." type="journal article" year="2008">
|
||
Page
|
||
<emphasis id="B948EA92FFC7FFE2FB3DFD9F2339FD7F" box="[1236,1285,620,644]" italics="true" pageId="2" pageNumber="33">et al</emphasis>
|
||
., 2008
|
||
</bibRefCitation>
|
||
); both can easily be separated from
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FE35FD632461FD53" ID-CoL="8VJXW" box="[476,605,657,680]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FE35FD632461FD53" box="[476,605,657,680]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FD8DFD6224F2FD53" box="[612,718,656,680]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD8DFD6224F2FD53" bold="true" box="[612,718,656,680]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in having degenerate eyes (vs. well developed).
|
||
</paragraph>
|
||
<paragraph id="8B833680FFC7FFE2FF2EFD4727B4FCC3" blockId="2.[151,1437,151,1112]" pageId="2" pageNumber="33">
|
||
Two further, perhaps distantly related genera are
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FD05FD4625BFFD37" ID-CoL="8V5J7" box="[748,899,692,716]" class="Malacostraca" family="Atyidae" genus="Halocaridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FD05FD4625BFFD37" box="[748,899,692,716]" italics="true" pageId="2" pageNumber="33">Halocaridina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FC53FD46224CFD37" ID-CoL="8VJXF" box="[954,1136,692,716]" class="Malacostraca" family="Atyidae" genus="Halocaridinides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FC53FD46224CFD37" box="[954,1136,692,716]" italics="true" pageId="2" pageNumber="33">Halocaridinides</emphasis>
|
||
</taxonomicName>
|
||
, which are anchialine taxa with eyes lacking any clear cornea, although they are relatively large. Additionally, their diaresis harbours numerous spines and no exopods are present (
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FDB1FD0E2547FCEF" author="Fujino" box="[600,891,764,788]" pageId="2" pageNumber="33" refString="Fujino, T. & Shokita, S. (1975) Report on some new atyid shrimps (Crustacea, Decapoda, Caridea) from the Rykyu Islands. Bulletin of the Sciences and Engineering Division, University of the Ryukyus, 18, 93 - 113." type="journal article" year="1975">Fujino and Shokita, 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FC61FD0E220CFCEF" author="Holthuis" box="[904,1072,764,788]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1973) Caridean shrimps found in land-locked saltwater pools at four Indo-West Pacific localities (Sinai Peninsula, Funafuti Atoll, Maui and Hawaii Islands), with the description of one new genus and four new species. Zoologische Verhandelingen, 128, 1 - 48, Plates 1 - 7." type="journal article" year="1973">Holthuis, 1973</bibRefCitation>
|
||
) facilitating the distinction with
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF7EFCD32724FCC3" ID-CoL="8VJXW" box="[151,280,801,824]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FF7EFCD32724FCC3" box="[151,280,801,824]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FEF6FCD227B4FCC3" box="[287,392,800,824]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FEF6FCD227B4FCC3" bold="true" box="[287,392,800,824]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B833680FFC7FFE2FF2EFCB6252AFBA3" blockId="2.[151,1437,151,1112]" pageId="2" pageNumber="33">
|
||
Another two, quite possibly unrelated, genera are known from
|
||
<collectingCountry id="F32B7610FFC7FFE2FC6FFCB6222DFCA7" box="[902,1041,836,860]" name="Madagascar" pageId="2" pageNumber="33">Madagascar</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FBF5FCB623A0FCA7" ID-CoL="8V57Q" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[1052,1436,836,860]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FBF5FCB622BEFCA7" box="[1052,1154,836,860]" italics="true" pageId="2" pageNumber="33">Caridina</emphasis>
|
||
H.
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FB43FCB623A0FCA7" author="Milne" box="[1194,1436,836,860]" pageId="2" pageNumber="33" refString="Milne Edwards, H. (1837) Histoire naturelle naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux. Paris, Librairie de Roret, 2, 1 - 532." type="journal article" year="1837">Milne Edwards, 1837</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF2CFC9B27F8FC84" ID-CoL="6HVP" authority="Holthuis, 1956" authorityName="Holthuis" authorityYear="1956" box="[197,452,871,896]" class="Bryopsida" family="Dicranaceae" genus="Parisia" kingdom="Plantae" order="Dicranales" pageId="2" pageNumber="33" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FF2CFC9B272BFC7B" box="[197,279,873,896]" italics="true" pageId="2" pageNumber="33">Parisia</emphasis>
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FEF7FC9527F8FC84" author="Holthuis" box="[286,452,871,895]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1956) The troglobic Atyidae of Madagascar (Crustacea, Decapoda, Natantia). Memoires de l'Institut Scientifique de Madagascar, series A, 11, 97 - 110." type="journal article" year="1956">Holthuis, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FE3BFC952441FC84" author="Holthuis" box="[466,637,871,895]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1965) The Atyidae of Madagascar. Memoires du Museum National D'Histoire Naturelle, nouvelle serie, Serie A. Zoologie, 33 (1), 1 - 48." type="journal article" year="1965">Holthuis, 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FD61FC95251FFC84" author="Gurney" box="[648,803,871,895]" pageId="2" pageNumber="33" refString="Gurney, A. R. (1984) Freshwater shrimp genera Caridina and Parisia (Decapoda: Caridea: Atyidae) of Madagascar, with descriptions of four new species. Journal of Natural History 18, 567 - 590." type="journal article" year="1984">Gurney, 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FCC7FC9525A3FC84" author="Cai" box="[814,927,871,895]" pageId="2" pageNumber="33" refString="Cai, Y. (2005) On a small collection of atyid shrimps from cave Lakata Zafera, western Madagascar, with a description of a new species (Decapoda: Atyidae). Proceedings of the Biological Society of Washington, 118 (2), 312 - 318." type="journal article" year="2005">Cai, 2005</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FC59FC9B220DFC7B" ID-CoL="8VJXW" box="[944,1073,873,896]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FC59FC9B220DFC7B" box="[944,1073,873,896]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FBD1FC9A229CFC7B" box="[1080,1184,872,896]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FBD1FC9A229CFC7B" bold="true" box="[1080,1184,872,896]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FAD9FC9A23A9FC7B" ID-CoL="8V57Q" box="[1328,1429,872,896]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FAD9FC9A23A9FC7B" box="[1328,1429,872,896]" italics="true" pageId="2" pageNumber="33">Caridina</emphasis>
|
||
</taxonomicName>
|
||
, in not exhibiting a supra-orbital tooth and possessing slender pereiopods. However, the new genus differs from
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FF7EFC4226C0FC33" ID-CoL="8V57Q" box="[151,252,944,968]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FF7EFC4226C0FC33" box="[151,252,944,968]" italics="true" pageId="2" pageNumber="33">Caridina</emphasis>
|
||
</taxonomicName>
|
||
by possessing exopods (vs. absent) on all pereiopods and a long carpus on the second pereiopod, being 7– 9 × long as broad (vs. 3.5–7.1 × long as broad in the Madagascan species) and being excavated (vs. absent) anteriorly. Further, in
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FEBCFC0A2786FBEB" ID-CoL="8V57Q" box="[341,442,1016,1040]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FEBCFC0A2786FBEB" box="[341,442,1016,1040]" italics="true" pageId="2" pageNumber="33">Caridina</emphasis>
|
||
</taxonomicName>
|
||
the uropod diaeresis is armed with more than one spine (
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FBB2FC05233EFBF4" author="Holthuis" box="[1115,1282,1015,1039]" pageId="2" pageNumber="33" refString="Holthuis, L. B. (1965) The Atyidae of Madagascar. Memoires du Museum National D'Histoire Naturelle, nouvelle serie, Serie A. Zoologie, 33 (1), 1 - 48." type="journal article" year="1965">Holthuis 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAD4B71FFC7FFE2FAF8FC052727FBCF" author="Richard" pageId="2" pageNumber="33" refString="Richard, J. & Clark, P. F. (2009) African Caridina (Crustacea: Decapoda: Caridea: Atyidae): redescriptions of C. africana Kingsley, 1882, C. togoensis Hilgendorf, 1893, C. natalensis (Bouvier, 1925) and C. roubaudi (Bouvier, 1925) with descriptions of 14 new species. Zootaxa, 1995, 1 - 75." type="journal article" year="2009">Richard and Clark 2009</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FED9FBEF278DFBCF" ID-CoL="8VJXW" box="[304,433,1053,1076]" class="Malacostraca" family="Atyidae" genus="Monsamnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B948EA92FFC7FFE2FED9FBEF278DFBCF" box="[304,433,1053,1076]" italics="true" pageId="2" pageNumber="33">Monsamnis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B57E9FFC7FFE2FE52FBEE241AFBCF" box="[443,550,1052,1076]" pageId="2" pageNumber="33" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FE52FBEE241AFBCF" bold="true" box="[443,550,1052,1076]" pageId="2" pageNumber="33">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be separated from the troglobic genus
|
||
<taxonomicName id="4C3C4D03FFC7FFE2FBF5FBEF2252FBCF" ID-CoL="6HVP" box="[1052,1134,1053,1076]" class="Bryopsida" family="Dicranaceae" genus="Parisia" kingdom="Plantae" order="Dicranales" pageId="2" pageNumber="33" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="B948EA92FFC7FFE2FBF5FBEF2252FBCF" box="[1052,1134,1053,1076]" italics="true" pageId="2" pageNumber="33">Parisia</emphasis>
|
||
</taxonomicName>
|
||
, in having exopods on all pereiopods (vs. absent) as well a well-developed, cornea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |