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<document id="CF056BDDD5AF6D00E103615F8216A747" ID-DOI="10.11646/zootaxa.5506.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747303" ID-ZooBank="B1D3C8F1-DA52-477A-93B9-A0B7F054866D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726054750462" checkinUser="plazi" docAuthor="Zheng, Xinyi &amp; Wu, SanAn" docDate="2024" docId="03958786FFC04550F4AEFC30FBE6F89B" docLanguage="en" docName="zootaxa.5506.4.1.pdf" docOrigin="Zootaxa 5506 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Margarodidae Cockerell 1899" docType="key" docVersion="3" lastPageNumber="497" masterDocId="FFACFFFEFFEE457EF439FFF3FE7AFFAE" masterDocTitle="A review of the Margarodidae sensu stricto (Hemiptera: Coccomorpha) in China, with descriptions of three new species" masterLastPageNumber="500" masterPageNumber="451" pageNumber="497" updateTime="1726080304904" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="856E6806D6103655D4E5B8159F3BE2D9">A review of the Margarodidae sensu stricto (Hemiptera: Coccomorpha) in China, with descriptions of three new species</mods:title>
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<mods:namePart id="182391150F97F25040954A04906833F2">Zheng, Xinyi</mods:namePart>
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<mods:namePart id="88C88AA24C85F1A429E0C0B6A8DB842D">Wu, SanAn</mods:namePart>
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<mods:affiliation id="4FE4594D81993C437B5DCED2DE440BC5">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China.</mods:affiliation>
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<treatment id="03958786FFC04550F4AEFC30FBE6F89B" LSID="urn:lsid:plazi:treatment:03958786FFC04550F4AEFC30FBE6F89B" httpUri="http://treatment.plazi.org/id/03958786FFC04550F4AEFC30FBE6F89B" lastPageNumber="497" pageId="46" pageNumber="497">
<subSubSection id="C326651BFFC04550F4AEFC30FD8BFC73" box="[151,1009,963,989]" pageId="46" pageNumber="497" type="nomenclature">
<paragraph id="8B833690FFC04550F4AEFC30FD8BFC73" blockId="46.[151,1009,963,989]" box="[151,1009,963,989]" pageId="46" pageNumber="497">
<heading id="D0CB81FCFFC04550F4AEFC30FD8BFC73" bold="true" box="[151,1009,963,989]" fontSize="11" level="1" pageId="46" pageNumber="497" reason="1">
<emphasis id="B948EA82FFC04550F4AEFC30FD8BFC73" bold="true" box="[151,1009,963,989]" pageId="46" pageNumber="497">
Key to genera and species of adult female
<taxonomicName id="4C3C4D13FFC04550F69DFC30FD22FC73" ID-CoL="624JZ" authorityName="Cockerell" authorityYear="1899" box="[676,856,963,989]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="family">Margarodidae</taxonomicName>
from
<collectingCountry id="F32B7600FFC04550F79DFC30FD8BFC73" box="[932,1009,963,989]" name="China" pageId="46" pageNumber="497">China</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326651BFFC04550F4AEFBFBFBE6F89B" pageId="46" pageNumber="497" type="key">
<keyStep id="30C82E35FFC04550F4AEFBFBFBE6FB95" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFBFBFBE6FBB3" blockId="46.[151,1437,1032,1846]" box="[151,1436,1032,1054]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFBFBFBE6FBB3" box="[151,1436,1032,1054]" pageId="46" pageNumber="497">
1(0) Abdominal spiracles numbering 8 pairs (genus
<taxonomicName id="4C3C4D13FFC04550F691FBFBFD3BFBB3" ID-CoL="5ZWP" authorityName="Green" authorityYear="1914" box="[680,833,1032,1053]" class="Insecta" family="Margarodidae" genus="Neomargarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA82FFC04550F691FBFBFD3BFBB3" box="[680,833,1032,1053]" italics="true" pageId="46" pageNumber="497">Neomargarodes</emphasis>
</taxonomicName>
)................................................ 2
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFBD5FBE6FB95" blockId="46.[151,1437,1032,1846]" box="[151,1436,1061,1083]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFBD5FBE6FB95" box="[151,1436,1061,1083]" pageId="46" pageNumber="497">- Abdominal spiracles numbering fewer than 8 pairs.......................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEFBB0FBE6FBDB" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFBB0FBE7FBF6" blockId="46.[151,1437,1032,1846]" box="[151,1437,1091,1113]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFBB0FBE7FBF6" box="[151,1437,1091,1113]" pageId="46" pageNumber="497">
2(1) Antenna with segments II and III fused together................................................
<taxonomicName id="4C3C4D13FFC04550F0C4FBB0FBE7FBF6" ID-CoL="46MC2" authority="(Green)" baseAuthorityName="Green" baseAuthorityYear="1912" box="[1277,1437,1091,1112]" class="Insecta" family="Margarodidae" genus="Neomargarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B948EA82FFC04550F0C4FBB0FB36FBF6" box="[1277,1356,1091,1112]" italics="true" pageId="46" pageNumber="497">N. niger</emphasis>
(Green)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFB93FBE6FBDB" blockId="46.[151,1437,1032,1846]" box="[151,1436,1120,1142]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFB93FBE6FBDB" box="[151,1436,1120,1142]" pageId="46" pageNumber="497">- Antenna with segments II and III separate................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEFB8EFBE6FB1E" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFB8EFBE6FB3D" blockId="46.[151,1437,1032,1846]" box="[151,1436,1149,1171]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFB8EFBE6FB3D" box="[151,1436,1149,1171]" pageId="46" pageNumber="497">
3(2) Foreleg claw with basal prominent heel angular..............................
<taxonomicName id="4C3C4D13FFC04550F01AFB8DFB3CFB3D" authority="Zheng &amp; Wu" authorityName="Zheng &amp; Wu" authorityYear="2024" box="[1059,1350,1149,1171]" class="Insecta" family="Margarodidae" genus="Neomargarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="chenopodium" status="sp. nov.">
<emphasis id="B948EA82FFC04550F01AFB8DFABAFB3C" box="[1059,1216,1149,1171]" italics="true" pageId="46" pageNumber="497">N. chenopodium</emphasis>
Zheng &amp; Wu
</taxonomicName>
,
<emphasis id="B948EA82FFC04550F169FB8DFBE6FB3D" bold="true" box="[1360,1436,1150,1171]" pageId="46" pageNumber="497">
<taxonomicNameLabel id="A27B57F9FFC04550F169FB8DFBE6FB3D" box="[1360,1436,1150,1171]" pageId="46" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFB68FBE6FB1E" blockId="46.[151,1437,1032,1846]" box="[151,1436,1179,1201]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFB68FBE6FB1E" box="[151,1436,1179,1201]" pageId="46" pageNumber="497">- Foreleg claw with basal prominent heel swollen and rounded ................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEFB4BFBE6FA88" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFB4BFBE7FB45" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFB4BFBE7FB45" pageId="46" pageNumber="497">
4(3) Multilocular disc-pores each with a bright central loculus; disc-pores on abdominal sternites IIV few, forming transverse segmental bands only 1 or 2 pores wide. Feeding mainly on
<taxonomicName id="4C3C4D13FFC04550F71DFB26FD07FB45" ID-CoL="623QT" box="[804,893,1237,1259]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="46" pageNumber="497" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
............................. ..
<taxonomicName id="4C3C4D13FFC04550F0C3FB25FBE7FB45" ID-CoL="46MBY" authority="Yang" authorityName="Yang" authorityYear="1979" box="[1274,1437,1238,1259]" class="Insecta" family="Margarodidae" genus="Neomargarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="gossypii">
<emphasis id="B948EA82FFC04550F0C3FB25FB1FFB45" box="[1274,1381,1238,1259]" italics="true" pageId="46" pageNumber="497">N. gossypii</emphasis>
Yang
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFB00FBE6FA88" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFB00FBE6FA88" pageId="46" pageNumber="497">
- Multilocular disc-pores each without a bright central loculus; disc-pores on abdominal sternites IIV numerous, forming transverse segmental bands 36 pores wide. Feeding on
<taxonomicName id="4C3C4D13FFC04550F6C7FAE3FDFDFA88" ID-CoL="8SC" box="[766,903,1296,1318]" class="Magnoliopsida" family="Cucurbitaceae" kingdom="Plantae" order="Cucurbitales" pageId="46" pageNumber="497" phylum="Tracheophyta" rank="family">Cucurbitaceae</taxonomicName>
and
<taxonomicName id="4C3C4D13FFC04550F78CFAE3FA30FA88" ID-CoL="68Z" box="[949,1098,1296,1318]" class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" order="Caryophyllales" pageId="46" pageNumber="497" phylum="Tracheophyta" rank="family">Amaranthaceae</taxonomicName>
......
<taxonomicName id="4C3C4D13FFC04550F0A7FAE2FBE6FA88" ID-CoL="46MBS" authority="Tang &amp; Hao" authorityName="Tang &amp; Hao" authorityYear="2000" box="[1182,1436,1296,1318]" class="Insecta" family="Margarodidae" genus="Neomargarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="cucurbitae">
<emphasis id="B948EA82FFC04550F0A7FAE2FB5BFA8B" box="[1182,1313,1296,1318]" italics="true" pageId="46" pageNumber="497">N. cucurbitae</emphasis>
Tang &amp; Hao
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEFADEFBE6FA35" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFADEFBE7FACF" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFADEFBE7FACF" pageId="46" pageNumber="497">
5(1) Abdominal spiracles numbering 6 or 7 pairs (genus
<taxonomicName id="4C3C4D13FFC04550F6F5FADEFD3BFAEC" ID-CoL="62ZP2" authorityName="Guilding" authorityYear="1829" box="[716,833,1325,1346]" class="Insecta" family="Margarodidae" genus="Margarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA82FFC04550F6F5FADEFD3BFAEC" box="[716,833,1325,1346]" italics="true" pageId="46" pageNumber="497">Margarodes</emphasis>
</taxonomicName>
). Body with many spines on thorax and abdomen. Foreleg claw sometimes bifurcate..................................................................
<taxonomicName id="4C3C4D13FFC04550F0F3FAB8FBE7FACF" ID-CoL="3Y49M" authority="(Silvestri)" baseAuthorityName="Silvestri" baseAuthorityYear="1938" box="[1226,1437,1355,1377]" class="Insecta" family="Margarodidae" genus="Margarodes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B948EA82FFC04550F0F3FAB8FB4CFACE" box="[1226,1334,1355,1376]" italics="true" pageId="46" pageNumber="497">M. sinensis</emphasis>
(Silvestri)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFA9BFBE6FA35" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFA9BFBE6FA35" pageId="46" pageNumber="497">
- Abdominal spiracles numbering 2 pairs or absent. Body without spines. Foreleg claw acute, never bifurcate (genus
<taxonomicName id="4C3C4D13FFC04550F4DBFA76FF0CFA34" ID-CoL="63PZQ" authorityName="Ben-Dov" authorityYear="2005" box="[226,374,1413,1434]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA82FFC04550F4DBFA76FF0CFA34" box="[226,374,1413,1434]" italics="true" pageId="46" pageNumber="497">Porphyrophora</emphasis>
</taxonomicName>
)...................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEFA50FBE6F9BE" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEFA50FBE6FA7B" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFA50FBE6FA7B" pageId="46" pageNumber="497">
6(5) Long hair-like setae few and relatively short (each less than 100 μm long); each abdominal tergite with transverse segmental band of setae only 1 or 2 setae wide.....................................
<taxonomicName id="4C3C4D13FFC04550F03EFA32FB3CFA78" authority="Zheng &amp; Wu" authorityName="Zheng &amp; Wu" authorityYear="2024" box="[1031,1350,1472,1494]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="neimonggolensis" status="sp. nov.">
<emphasis id="B948EA82FFC04550F03EFA32FABAFA7B" box="[1031,1216,1472,1494]" italics="true" pageId="46" pageNumber="497">P. neimonggolensis</emphasis>
Zheng &amp; Wu
</taxonomicName>
,
<emphasis id="B948EA82FFC04550F169FA33FBE6FA7B" bold="true" box="[1360,1436,1472,1493]" pageId="46" pageNumber="497">
<taxonomicNameLabel id="A27B57F9FFC04550F169FA33FBE6FA7B" box="[1360,1436,1472,1493]" pageId="46" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEFA2DFBE6F9BE" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEFA2DFBE6F9BE" pageId="46" pageNumber="497">- Long hair-like setae numerous and long (each more than 200 μm long); each abdominal tergite with transverse segmental band of setae 29 setae wide................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEF9EBFBE6F9E5" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEF9EBFBE6F980" blockId="46.[151,1437,1032,1846]" box="[151,1436,1560,1582]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF9EBFBE6F980" box="[151,1436,1560,1582]" pageId="46" pageNumber="497">
7(6) Antenna with 10 segments. Multilocular disc-pores each without a bright central loculus...
<taxonomicName id="4C3C4D13FFC04550F06BF9EAFBE6F980" ID-CoL="4LZBG" authority="Matesova &amp; Jashenko" authorityName="Matesova &amp; Jashenko" authorityYear="1988" box="[1106,1436,1560,1582]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="violaceae">
<emphasis id="B948EA82FFC04550F06BF9EAFABEF983" box="[1106,1220,1560,1582]" italics="true" pageId="46" pageNumber="497">P. violaceae</emphasis>
Matesova &amp; Jashenko
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEF9C5FBE6F9E5" blockId="46.[151,1437,1032,1846]" box="[151,1436,1589,1611]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF9C5FBE6F9E5" box="[151,1436,1589,1611]" pageId="46" pageNumber="497">- Antenna with 9 segments or fewer. Multilocular disc-pores each with a bright central loculus......................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEF9A0FBE6F96E" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEF9A0FBE6F92B" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF9A0FBE6F92B" pageId="46" pageNumber="497">
8(7) Abdominal tergites each with transverse segmental band of setae about 69 setae wide. Thoracic spiracle with 722 spiracular disc-pores....................................................................
<taxonomicName id="4C3C4D13FFC04550F0BDF982FB4FF928" ID-CoL="6VW8W" authority="Yang" authorityName="Yang" authorityYear="1979" box="[1156,1333,1649,1670]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="ningxiana">
<emphasis id="B948EA82FFC04550F0BDF982FA81F928" box="[1156,1275,1649,1670]" italics="true" pageId="46" pageNumber="497">P. ningxiana</emphasis>
Yang
</taxonomicName>
,
<emphasis id="B948EA82FFC04550F106F983FBE6F92B" bold="true" box="[1343,1436,1648,1669]" pageId="46" pageNumber="497">stat. rest.</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEF97DFBE6F96E" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF97DFBE6F96E" pageId="46" pageNumber="497">- Abdominal tergites each with transverse segmental band of setae about 26 setae wide. Thoracic spiracle with 46 spiracular disc-pores........................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C82E35FFC04550F4AEF93BFBE6F89B" pageId="46" pageNumber="497">
<paragraph id="8B833690FFC04550F4AEF93BFBE7F955" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF93BFBE7F955" pageId="46" pageNumber="497">
9(8) Abdominal sternites each with transverse band of setae only 2 or 3 setae wide.Antenna with 79 (mostly 7) segments. Sclerotized plate around eyespot with a reticulated texture............................................
<taxonomicName id="4C3C4D13FFC04550F0FAF915FBE7F955" ID-CoL="78898" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1219,1437,1765,1787]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="polonica">
<emphasis id="B948EA82FFC04550F0FAF915FB54F954" box="[1219,1326,1765,1787]" italics="true" pageId="46" pageNumber="497">P. polonica</emphasis>
(Linnaeus)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833690FFC04550F4AEF8F0FBE6F89B" blockId="46.[151,1437,1032,1846]" pageId="46" pageNumber="497">
<keyLead id="30CD95A5FFC04550F4AEF8F0FBE6F89B" pageId="46" pageNumber="497">
- Abdominal sternites each with transverse band of setae about 46 setae wide. Antenna with 9 segments. Sclerotized plate around eyespot smooth......................................................
<taxonomicName id="4C3C4D13FFC04550F06CF8D2FB3CF898" authority="Zheng &amp; Wu" authorityName="Zheng &amp; Wu" authorityYear="2024" box="[1109,1350,1824,1846]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="ejinensis" status="sp. nov.">
<emphasis id="B948EA82FFC04550F06CF8D2FABAF898" box="[1109,1216,1825,1846]" italics="true" pageId="46" pageNumber="497">P. ejinensis</emphasis>
Zheng &amp; Wu
</taxonomicName>
,
<emphasis id="B948EA82FFC04550F169F8D3FBE6F89B" bold="true" box="[1360,1436,1824,1845]" pageId="46" pageNumber="497">
<taxonomicNameLabel id="A27B57F9FFC04550F169F8D3FBE6F89B" box="[1360,1436,1824,1845]" pageId="46" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>