treatments-xml/data/03/95/87/039587033921FFE083ADFF232A88197E.xml

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<mods:title id="A8A4AF2CDAA4C7B78459D43DF6081AD1">Two new species of Turleania McLaughlin, 1997 (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, and a redescription of T. balli (McLaughlin &amp; Haig, 1996)</mods:title>
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<taxonomicName id="4C3C4D963921FFE783ADFF2329E91FE6" ID-CoL="962bb1d8-51f2-46e7-bde6-f7bccdc78066" authority="McLaughlin &amp; Haig, 1996" authorityName="McLaughlin &amp; Haig" authorityYear="1996" box="[151,685,151,177]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="balli">
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<emphasis id="B948EA073921FFE783ADFF232A0F1FE6" bold="true" box="[151,331,151,177]" italics="true" pageId="2" pageNumber="51">Turleania balli</emphasis>
(
<bibRefCitation id="EFAD4BE43921FFE78260FF2329E11FE6" author="McLaughlin" box="[346,677,151,177]" pageId="2" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">McLaughlin &amp; Haig, 1996</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
<paragraph id="8B8336153921FFE783ADFF0E2A5C1F83" blockId="2.[151,685,151,212]" box="[151,280,186,212]" pageId="2" pageNumber="51">
(
<figureCitation id="13072A903921FFE783A5FF0E2BB51F83" box="[159,241,186,212]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Figs. 1</figureCitation>
, 
<figureCitation id="13072A903921FFE783C5FF0E2A4B1F83" box="[255,271,186,212]" captionStart="FIGURE 2" captionStartId="4.[151,255,1832,1856]" captionTargetBox="[206,1367,184,1806]" captionTargetId="figure@4.[196,1401,170,1826]" captionTargetPageId="4" captionText="FIGURE 2. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, right cheliped, chela and carpus, dorsal view; B, same, palm and proximal part of dactylus, dorsolateral view (setae omitted); C, same, chela, ventrolateral view (setae omitted); D, same, carpus and distal part of merus, mesial view (setae omitted); E, same, carpus and distal part of merus, lateral view (setae omitted); F, left cheliped, chela and carpus, dorsal view; G, same, carpus and distal part of merus, mesial view (setae omitted); H, same, carpus and distal part of merus, lateral view (setae omitted). Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181759/files/figure.png" pageId="2" pageNumber="51">2</figureCitation>
)
</paragraph>
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<subSubSection id="C326659E3921FFE783ADFEBE292C1E36" pageId="2" pageNumber="51" type="reference_group">
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<taxonomicName id="4C3C4D963921FFE783ADFEBE2A641E76" ID-CoL="9cf6b7f8-4aaa-4e44-a0d4-1b9ba8088b50" box="[151,288,266,289]" class="Malacostraca" family="Paguridae" genus="Anapagrides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073921FFE783ADFEBE2A641E76" box="[151,288,266,289]" italics="true" pageId="2" pageNumber="51">Anapagrides</emphasis>
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sp. — 
<treatmentCitation id="0A9D10043921FFE78256FEBE292D1E76" author="Haig" box="[364,617,266,289]" page="177" pageId="2" pageNumber="51" year="1988">
<bibRefCitation id="EFAD4BE43921FFE78256FEBE29711E76" author="Haig" box="[364,565,266,289]" pageId="2" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">Haig &amp; Ball, 1988</bibRefCitation>
: 177
</treatmentCitation>
, fig.8. 
<taxonomicName id="4C3C4D963921FFE783ADFE9E29C01E16" authority="McLaughlin &amp; Haig, 1996: 84" authorityName="McLaughlin &amp; Haig" authorityPageNumber="84" authorityYear="1996" box="[151,644,298,321]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073921FFE783ADFE9E2A721E16" box="[151,310,298,321]" italics="true" pageId="2" pageNumber="51">Laurentia balli</emphasis>
<treatmentCitation id="0A9D10043921FFE78207FE9E29C01E16" author="McLaughlin" box="[317,644,298,321]" page="84" pageId="2" pageNumber="51" year="1996">
<bibRefCitation id="EFAD4BE43921FFE78207FE9E29191E16" author="McLaughlin" box="[317,605,298,321]" pageId="2" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">McLaughlin &amp; Haig, 1996</bibRefCitation>
: 84
</treatmentCitation>
</taxonomicName>
, fig. 5. 
<taxonomicName id="4C3C4D963921FFE783ADFEFE2A721E36" ID-CoL="962bb1d8-51f2-46e7-bde6-f7bccdc78066" box="[151,310,330,353]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073921FFE783ADFEFE2A721E36" box="[151,310,330,353]" italics="true" pageId="2" pageNumber="51">Turleania balli</emphasis>
</taxonomicName>
. — 
<treatmentCitation id="0A9D10043921FFE7825FFEFE29201E36" author="McLaughlin" box="[357,612,330,353]" page="477" pageId="2" pageNumber="51" year="1997">
<bibRefCitation id="EFAD4BE43921FFE7825FFEFE296B1E36" author="McLaughlin" box="[357,559,330,353]" pageId="2" pageNumber="67" refString="McLaughlin, P. A. (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier, A. &amp; Bouchet, P. (Eds), Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172, 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
: 477
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.
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</paragraph>
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<paragraph id="8B8336153921FFE783ADFE262FB61E83" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
<emphasis id="B948EA073921FFE783ADFE262A0E1EFB" bold="true" box="[151,330,402,428]" pageId="2" pageNumber="51">
<typeStatus id="548788B73921FFE783ADFE262B961EFB" box="[151,210,402,428]" pageId="2" pageNumber="51">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548788B73921FFE78269FE262A811EFB" box="[339,453,402,428]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
: male (sl 
<quantity id="4CC49BF03921FFE78109FE2629D41EFB" box="[563,656,402,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="51" unit="mm" value="2.1">2.1 mm</quantity>
), off 
<collectingCountry id="F32B76853921FFE781F6FE26284B1EFB" box="[716,783,402,428]" name="Iceland" pageId="2" pageNumber="51">Pulau</collectingCountry>
Saparua, 
<collectingCountry id="F32B76853921FFE780B9FE2628BC1EFB" box="[899,1016,402,428]" name="Indonesia" pageId="2" pageNumber="51">Indonesia</collectingCountry>
, 
<geoCoordinate id="EE0850D23921FFE78739FE262F371EFB" box="[1027,1139,402,428]" direction="south" orientation="latitude" pageId="2" pageNumber="51" precision="92" value="-3.6316667">3°37.9’ S</geoCoordinate>
, 128°38.6–39’ E, “Alpha Helix” Saparua St. 3, 
<quantity id="4CC49BF03921FFE782ACFE0E2AB61E83" box="[406,498,442,468]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="2" pageNumber="51" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">0–20 m</quantity>
, cliff face of rock and coral, coll. E. Ball, 
<date id="FF8210D53921FFE780E7FE0E2FD51E83" box="[989,1169,442,468]" pageId="2" pageNumber="51" value="1975-03-29">29 March 1975</date>
(
<collectionCode id="ED2DAED03921FFE7879AFE0E2FA21E83" box="[1184,1254,442,468]" pageId="2" pageNumber="51">NIOJ</collectionCode>
).
</paragraph>
<paragraph id="8B8336153921FFE783FFFE562FF31EAB" blockId="2.[151,1437,402,2028]" box="[197,1207,482,508]" pageId="2" pageNumber="51">
<emphasis id="B948EA073921FFE783FFFE562A3F1EAB" bold="true" box="[197,379,482,508]" pageId="2" pageNumber="51">Redescription.</emphasis>
Nine pairs of quadriserial gills; no arthrobranchs on third maxillipeds.
</paragraph>
<paragraph id="8B8336153921FFE783FFFDBE2E2D1D23" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Shield (
<figureCitation id="13072A903921FFE7821AFDBE2A251D73" box="[288,353,522,548]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
A) 1.1 times longer than broad; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin roundly truncate. Rostrum triangular, well developed, overreaching lateral projections. Lateral projections also well developed, each terminating in small spine.
</paragraph>
<paragraph id="8B8336153921FFE783FFFD362F051D93" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Ocular peduncles (
<figureCitation id="13072A903921FFE78292FD362AA81DCB" box="[424,492,642,668]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
A) 0.8 length of shield, with corneas slightly dilated; lateral surfaces concave medially; ocular acicles narrowly subtriangular, each with small marginal spine.
</paragraph>
<paragraph id="8B8336153921FFE783FFFD662ED21DBB" blockId="2.[151,1437,402,2028]" box="[197,1430,722,748]" pageId="2" pageNumber="51">
Antennular peduncles (
<figureCitation id="13072A903921FFE782EEFD6629501DBB" box="[468,532,722,748]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
A) overreaching distal margins of corneas by 0.6 length of ultimate segment.
</paragraph>
<paragraph id="8B8336153921FFE783FFFD4E29691CDB" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Antennal peduncles (
<figureCitation id="13072A903921FFE78287FD4E2ABA1C43" box="[445,510,762,788]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
A) also overreaching distal margins of corneas by 0.3 length of fifth segments. Second segment with small spine at dorsomesial distal angle; dorsolateral distal angle produced, moderately long, terminating in single or bifid spine. Antennal acicle slightly arcuate, barely reaching proximal margin of cornea, terminating in small spine.
</paragraph>
<paragraph id="8B8336153921FFE783FFFC2E2A991C8B" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">Mouthparts not dissected. Third maxilliped with well developed crista dentata of small teeth but without accessory tooth on ischium.</paragraph>
<paragraph id="8B8336153921FFE783FFFC5E2A611A3B" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Chelipeds (
<figureCitation id="13072A903921FFE78276FC5E2AD71B53" box="[332,403,1002,1028]" captionStart="FIGURE 2" captionStartId="4.[151,255,1832,1856]" captionTargetBox="[206,1367,184,1806]" captionTargetId="figure@4.[196,1401,170,1826]" captionTargetPageId="4" captionText="FIGURE 2. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, right cheliped, chela and carpus, dorsal view; B, same, palm and proximal part of dactylus, dorsolateral view (setae omitted); C, same, chela, ventrolateral view (setae omitted); D, same, carpus and distal part of merus, mesial view (setae omitted); E, same, carpus and distal part of merus, lateral view (setae omitted); F, left cheliped, chela and carpus, dorsal view; G, same, carpus and distal part of merus, mesial view (setae omitted); H, same, carpus and distal part of merus, lateral view (setae omitted). Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181759/files/figure.png" pageId="2" pageNumber="51">Fig. 2</figureCitation>
A–H) subequal in length but right (
<figureCitation id="13072A903921FFE78001FC5E28C61B53" box="[827,898,1002,1028]" captionStart="FIGURE 2" captionStartId="4.[151,255,1832,1856]" captionTargetBox="[206,1367,184,1806]" captionTargetId="figure@4.[196,1401,170,1826]" captionTargetPageId="4" captionText="FIGURE 2. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, right cheliped, chela and carpus, dorsal view; B, same, palm and proximal part of dactylus, dorsolateral view (setae omitted); C, same, chela, ventrolateral view (setae omitted); D, same, carpus and distal part of merus, mesial view (setae omitted); E, same, carpus and distal part of merus, lateral view (setae omitted); F, left cheliped, chela and carpus, dorsal view; G, same, carpus and distal part of merus, mesial view (setae omitted); H, same, carpus and distal part of merus, lateral view (setae omitted). Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181759/files/figure.png" pageId="2" pageNumber="51">Fig. 2</figureCitation>
A–E) distinctly stouter than left (
<figureCitation id="13072A903921FFE78635FC5E2E131B53" box="[1295,1367,1002,1028]" captionStart="FIGURE 2" captionStartId="4.[151,255,1832,1856]" captionTargetBox="[206,1367,184,1806]" captionTargetId="figure@4.[196,1401,170,1826]" captionTargetPageId="4" captionText="FIGURE 2. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, right cheliped, chela and carpus, dorsal view; B, same, palm and proximal part of dactylus, dorsolateral view (setae omitted); C, same, chela, ventrolateral view (setae omitted); D, same, carpus and distal part of merus, mesial view (setae omitted); E, same, carpus and distal part of merus, lateral view (setae omitted); F, left cheliped, chela and carpus, dorsal view; G, same, carpus and distal part of merus, mesial view (setae omitted); H, same, carpus and distal part of merus, lateral view (setae omitted). Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181759/files/figure.png" pageId="2" pageNumber="51">Fig. 2</figureCitation>
F–H). Right chela elongate oval in general outline, 2.6 times as long as broad. Dactylus approximately as long as palm, dorsomesial margin with small proximal spines and short slightly elevated ridges distally, cutting edge with 2 prominent teeth in proximal half and row of small corneous teeth distally. Palm approximately as long as carpus, dorsomesial margin with row of 4 strong spines, dorsolateral margin not delimited, dorsal surface with small but distinct spines laterally, cutting edge of fixed finger with 2 prominent calcareous teeth medially and some smaller teeth on remaining margin. Carpus slightly longer than merus, dorsomesial margin with row of 3 spines decreasing in size proximally, dorsodistal margin with median spine, ventrolateral margin with small distal spine, ventromesial margin unarmed. Merus also with spine at each ventrolateral and ventromesial distal angle.
</paragraph>
<paragraph id="8B8336153921FFE783FFFACE2935190B" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">Left chela elongate, 3.4 times longer than broad. Dactylus 1.2 times longer than palm, unarmed on surfaces, cutting edge with row of short corneous teeth. Palm 0.7 length of carpus, unarmed; cutting edge of fixed finger with row of small calcareous and corneous teeth. Carpus approximately as long as merus, dorsolateral margin with row of 3 spines, dorsomesial margin with only strong distal spine, ventrolateral margin with small distal spine, ventromesial margin unarmed. Merus with small spine at ventromesial distal angle and 2 subdistal spines on ventrolateral margin.</paragraph>
<paragraph id="8B8336153921FFE783FFF9DE2A9A1823" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Ambulatory legs (
<figureCitation id="13072A903921FFE782A0F9DE2A9819D3" box="[410,476,1642,1668]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
E–I) slender [left second pereopod missing, but illustrated by 
<bibRefCitation id="EFAD4BE43921FFE78782F9DE2BBC19FB" author="Haig" pageId="2" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">Haig &amp; Ball (1988, fig. 8E)</bibRefCitation>
]. Dactyli 1.4–1.5 times longer than propodi, slightly curved distally in lateral view, terminating in long corneous claws; dorsal surfaces each with row of small pits; ventral margins each with row of 6 or 7 slen- der corneous spines. Propodi not tapering distally, ventrodistal margins each with slender corneous spine. Carpi each with small subdistal spine on dorsal surface, right second pereopod with additional small spine in proximal half. Meri each with very small subdistal spine (right second) or unarmed (third) on ventral margin. Ischia unarmed on surfaces.
</paragraph>
<paragraph id="8B8336153921FFE783FFF8362F6A18BB" blockId="2.[151,1437,402,2028]" pageId="2" pageNumber="51">
Fourth pereopods [now missing, but right fourth pereopod was illustrated by 
<bibRefCitation id="EFAD4BE43921FFE78768F8362ED818CB" author="Haig" box="[1106,1436,1922,1948]" pageId="2" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">Haig &amp; Ball (1988, fig. 8G)</bibRefCitation>
and described by 
<bibRefCitation id="EFAD4BE43921FFE78259F81E29EE1893" author="McLaughlin" box="[355,682,1962,1988]" pageId="2" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">McLaughlin &amp; Haig (1996)</bibRefCitation>
] semichelate; dactylus without preungual process; propodal rasp composed of single row of corneous scales; carpus unarmed on dorsal surface.
</paragraph>
<caption id="DF43669D3920FFE683ADF8B8280718F9" httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="3" pageNumber="52" targetBox="[158,1412,214,1746]" targetPageId="3">
<paragraph id="8B8336153920FFE683ADF8B8280718F9" blockId="3.[151,1436,1804,1966]" pageId="3" pageNumber="52">
<emphasis id="B948EA073920FFE683ADF8B82A581873" bold="true" box="[151,284,1804,1828]" pageId="3" pageNumber="52">FIGURE 1.</emphasis>
<taxonomicName id="4C3C4D963920FFE6821CF8B9284A1874" authority="McLaughlin &amp; Haig, 1996" authorityName="McLaughlin &amp; Haig" authorityYear="1996" box="[294,782,1804,1828]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073920FFE6821CF8B92A8D1873" box="[294,457,1805,1828]" italics="true" pageId="3" pageNumber="52">Turleania balli</emphasis>
(McLaughlin &amp; Haig, 1996)
</taxonomicName>
. Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm.
</paragraph>
</caption>
<caption id="DF43669D3927FFE183ADF89C2A3F18BA" httpUri="https://zenodo.org/record/181759/files/figure.png" pageId="4" pageNumber="53" targetBox="[206,1367,184,1806]" targetPageId="4">
<paragraph id="8B8336153927FFE183ADF89C2A3F18BA" blockId="4.[151,1437,1832,2029]" pageId="4" pageNumber="53">
<emphasis id="B948EA073927FFE183ADF89C2A5D1817" bold="true" box="[151,281,1832,1856]" pageId="4" pageNumber="53">FIGURE 2.</emphasis>
<taxonomicName id="4C3C4D963927FFE1821AF89D29B31868" authority="McLaughlin &amp; Haig, 1996" authorityName="McLaughlin &amp; Haig" authorityYear="1996" box="[288,759,1832,1856]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073927FFE1821AF89D2AFB1817" box="[288,447,1833,1856]" italics="true" pageId="4" pageNumber="53">Turleania balli</emphasis>
(McLaughlin &amp; Haig, 1996)
</taxonomicName>
. Holotype male (sl 2.1 mm), NIOJ. A, right cheliped, chela and carpus, dorsal view; B, same, palm and proximal part of dactylus, dorsolateral view (setae omitted); C, same, chela, ventrolateral view (setae omitted); D, same, carpus and distal part of merus, mesial view (setae omitted); E, same, carpus and distal part of merus, lateral view (setae omitted); F, left cheliped, chela and carpus, dorsal view; G, same, carpus and distal part of merus, mesial view (setae omitted); H, same, carpus and distal part of merus, lateral view (setae omitted). Scales equal 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B8336153926FFE083FFFF2C2ACC1E7D" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
Coxae of fifth pereopods (
<figureCitation id="13072A903926FFE082CDFF2C297C1FE5" box="[503,568,152,178]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="5" pageNumber="54">Fig. 1</figureCitation>
C) unequal, left coxa larger than right, with well developed, stout and flattish sexual tube directed exteriorly and recurved posteriorly, and then upward, with broad tip obscured by dense fringe of short setae, surfaces also with scattered setae; right coxa with gonopore partially obscured by fringe of short setae.
</paragraph>
<paragraph id="8B8336153926FFE083FFFE8C2F1A1E2D" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
Sixth thoracic sternite with anterior lobe (
<figureCitation id="13072A903926FFE081FFFE8C284F1E05" box="[709,779,312,338]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="5" pageNumber="54">Fig. 1</figureCitation>
B) roundly subquadrate and bearing 4 small marginal spines. Eighth thoracic sternite (
<figureCitation id="13072A903926FFE0812AFED429161E2D" box="[528,594,352,378]" captionStart="FIGURE 1" captionStartId="3.[151,255,1804,1828]" captionTargetBox="[158,1412,214,1746]" captionTargetId="figure@3.[151,1436,194,1780]" captionTargetPageId="3" captionText="FIGURE 1. Turleania balli (McLaughlin &amp; Haig, 1996). Holotype male (sl 2.1 mm), NIOJ. A, shield and cephalic appendages, dorsal view (most setae omitted); B, sixth thoracic sternite, anterior lobe, ventral view; C, coxae of fifth pereopods and eighth thoracic sternite, ventral view; D, left sexual tube, posterior view; E, right second pereopod, lateral view; F, same, dactylus, mesial view; G, same, carpus, mesial view; H, left third pereopod, lateral view; I, same, dactylus, mesial view; J, same, carpus, mesial view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/181758/files/figure.png" pageId="5" pageNumber="54">Fig. 1</figureCitation>
C) developed as single, small subovate lobe.
</paragraph>
<paragraph id="8B8336153926FFE083FFFE3C281B1EA5" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
Telson [missing, but figured by 
<bibRefCitation id="EFAD4BE43926FFE08106FE3C28C11EF5" author="Haig" box="[572,901,392,418]" pageId="5" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">Haig &amp; Ball (1988, fig. 8H)</bibRefCitation>
] with lateral indentations indistinct; posterior lobes markedly asymmetrical, with terminal margins oblique, each with 3 small spines and stronger apical spine; lateral margin of left lobe with narrow chitinous plate.
</paragraph>
<paragraph id="8B8336153926FFE083FFFE4B2FF01D15" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
<emphasis id="B948EA073926FFE083FFFE4B2A691D4E" bold="true" box="[197,301,511,537]" pageId="5" pageNumber="54">Habitat.</emphasis>
<bibRefCitation id="EFAD4BE43926FFE0820FFDB4295B1D4D" author="Haig" box="[309,543,512,538]" pageId="5" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">Haig &amp; Ball (1988)</bibRefCitation>
mentioned that the sole specimen was collected on a cliff face of rock and coral, at a depth of 
<quantity id="4CC49BF03926FFE08243FD9C2AE21D15" box="[377,422,552,578]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="54" unit="m" value="5.0">5 m</quantity>
or less. On the label of the 
<typeStatus id="548788B73926FFE081D2FD9C28161D15" box="[744,850,552,578]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
, the depth is given as 
<quantity id="4CC49BF03926FFE0876EFD9C2FF41D15" box="[1108,1200,552,578]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="5" pageNumber="54" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">
0– 
<specimenCount id="9D3AFD9C3926FFE0874BFD9C2FF41D15" box="[1137,1200,552,578]" pageId="5" pageNumber="54" type="generic">20 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C326659E3926FFE083FFFDFB29B61DED" pageId="5" pageNumber="54" type="distribution">
<paragraph id="8B8336153926FFE083FFFDFB29B61DED" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
<emphasis id="B948EA073926FFE083FFFDFB2A221D3E" bold="true" box="[197,358,591,617]" pageId="5" pageNumber="54">Distribution.</emphasis>
Confirmed records of 
<taxonomicName id="4C3C4D963926FFE0814CFDE429831D3E" box="[630,711,592,617]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE0814CFDE429831D3E" box="[630,711,592,617]" italics="true" pageId="5" pageNumber="54">T. balli</emphasis>
</taxonomicName>
are restricted Saparua Island in 
<collectingCountry id="F32B76853926FFE08771FDE42FEE1D3D" box="[1099,1194,592,618]" name="Indonesia" pageId="5" pageNumber="54">Maluku</collectingCountry>
, 
<collectingCountry id="F32B76853926FFE0878DFDE42E691D3D" box="[1207,1325,592,618]" name="Indonesia" pageId="5" pageNumber="54">Indonesia</collectingCountry>
. A specimen lacking chelipeds and ambulatory legs from “Siboga” station “Banda” was tentatively assigned to 
<taxonomicName id="4C3C4D963926FFE08670FDCC2ED81DC6" box="[1354,1436,632,657]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE08670FDCC2ED81DC6" box="[1354,1436,632,657]" italics="true" pageId="5" pageNumber="54">T. balli</emphasis>
</taxonomicName>
(as 
<taxonomicName id="4C3C4D963926FFE083FAFD142A2B1DEE" box="[192,367,672,697]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE083FAFD142A2B1DEE" box="[192,367,672,697]" italics="true" pageId="5" pageNumber="54">Laurentia balli</emphasis>
</taxonomicName>
) by 
<bibRefCitation id="EFAD4BE43926FFE0829FFD1429AA1DED" author="McLaughlin" box="[421,750,672,698]" pageId="5" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">McLaughlin &amp; Haig (1996)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C326659E3926FFE083FFFD732A88197E" pageId="5" pageNumber="54" type="discussion">
<paragraph id="8B8336153926FFE083FFFD7328281C0D" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
<emphasis id="B948EA073926FFE083FFFD732A7A1DB6" bold="true" box="[197,318,711,737]" pageId="5" pageNumber="54">Remarks.</emphasis>
<bibRefCitation id="EFAD4BE43926FFE08273FD7C29D21DB5" author="McLaughlin" box="[329,662,712,738]" pageId="5" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">McLaughlin &amp; Haig (1996)</bibRefCitation>
described the present specimen as a new species of their genus 
<taxonomicName id="4C3C4D963926FFE083ADFD442A4E1C5E" box="[151,266,752,777]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE083ADFD442A4E1C5E" box="[151,266,752,777]" italics="true" pageId="5" pageNumber="54">Laurentia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D963926FFE0822DFD442A2F1C5E" box="[279,363,752,777]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE0822DFD442A2F1C5E" box="[279,363,752,777]" italics="true" pageId="5" pageNumber="54">L. balli</emphasis>
</taxonomicName>
. Subsequently, 
<bibRefCitation id="EFAD4BE43926FFE08119FD44284E1C5D" author="McLaughlin" box="[547,778,752,778]" pageId="5" pageNumber="67" refString="McLaughlin, P. A. (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier, A. &amp; Bouchet, P. (Eds), Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172, 433 - 572." type="journal article" year="1997">McLaughlin (1997)</bibRefCitation>
proposed the replacement name 
<taxonomicName id="4C3C4D963926FFE087B2FD442FBE1C5E" box="[1160,1274,752,777]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE087B2FD442FBE1C5E" box="[1160,1274,752,777]" italics="true" pageId="5" pageNumber="54">Turleania</emphasis>
</taxonomicName>
for 
<taxonomicName id="4C3C4D963926FFE08613FD442ED81C5E" box="[1321,1436,752,777]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE08613FD442ED81C5E" box="[1321,1436,752,777]" italics="true" pageId="5" pageNumber="54">Laurentia</emphasis>
</taxonomicName>
because the latter proved to be a junior homonym and established an allied genus 
<taxonomicName id="4C3C4D963926FFE08753FCAC2E501C66" box="[1129,1300,792,817]" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE08753FCAC2E501C66" box="[1129,1300,792,817]" italics="true" pageId="5" pageNumber="54">Enneopagurus</emphasis>
</taxonomicName>
for 
<taxonomicName id="4C3C4D963926FFE0867DFCAC2AAB1C0D" authority="McLaughlin, 1997" authorityName="McLaughlin" authorityYear="1997" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="garciagomezi">
<emphasis id="B948EA073926FFE0867DFCAC2A4A1C0E" italics="true" pageId="5" pageNumber="54">E. garciagomezi</emphasis>
<bibRefCitation id="EFAD4BE43926FFE0822FFCF42AAB1C0D" author="McLaughlin" box="[277,495,832,858]" pageId="5" pageNumber="67" refString="McLaughlin, P. A. (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier, A. &amp; Bouchet, P. (Eds), Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172, 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
</taxonomicName>
from deep waters off 
<collectingCountry id="F32B76853926FFE081C8FCF4282C1C0D" box="[754,872,832,858]" name="Indonesia" pageId="5" pageNumber="54">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="8B8336153926FFE083FFFCDC29101AE5" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
The 
<typeStatus id="548788B73926FFE083C0FCDC2A251CD5" box="[250,353,872,898]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C3C4D963926FFE082BDFCDC29731CD6" box="[391,567,872,897]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE082BDFCDC29731CD6" box="[391,567,872,897]" italics="true" pageId="5" pageNumber="54">Turleania balli</emphasis>
</taxonomicName>
is now in poor condition. The specimen is largely decalcified in the integument and has much fewer setae on the chelipeds and ambulatory legs than those illustrated by 
<bibRefCitation id="EFAD4BE43926FFE08630FC2429221C85" author="Haig" pageId="5" pageNumber="67" refString="Haig, J. &amp; Ball, E. E. (1988) Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum, 40, 151 - 196." type="journal article" year="1988">
Haig &amp; Ball (1988, fig. 8 B–G, as 
<taxonomicName id="4C3C4D963926FFE082A6FC0C29761C86" box="[412,562,952,977]" class="Malacostraca" family="Paguridae" genus="Anapagrides" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE082A6FC0C29761C86" box="[412,562,952,977]" italics="true" pageId="5" pageNumber="54">Anapagrides</emphasis>
</taxonomicName>
sp.)
</bibRefCitation>
and 
<bibRefCitation id="EFAD4BE43926FFE0819EFC0C2E241C85" author="McLaughlin" box="[676,1376,952,978]" pageId="5" pageNumber="68" refString="McLaughlin, P. A. &amp; Haig, J. (1996) A new genus for Anapagrides sensu De Saint Laurent-Dechance, 1966 (Decapoda: Anomura: Paguridae) and descriptions of four new species. Proceedings of the Biological Society of Washington, 109, 75 - 90." type="journal article" year="1996">
McLaughlin &amp; Haig (1996, fig. 5B, C, as 
<taxonomicName id="4C3C4D963926FFE08798FC0C2E171C86" box="[1186,1363,952,977]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE08798FC0C2E171C86" box="[1186,1363,952,977]" italics="true" pageId="5" pageNumber="54">Laurentia balli</emphasis>
</taxonomicName>
)
</bibRefCitation>
. The reexamination of the 
<typeStatus id="548788B73926FFE082B5FC542AB11CAD" box="[399,501,992,1018]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
has revealed that it possesses only nine pairs of quadriserial gills (arthrobranchs are absent from the third maxilliped) and the left sexual tube being comparatively broad and flattish. The number of gills is consistent with that of the two new species described below and with the diagnosis of the allied genus 
<taxonomicName id="4C3C4D963926FFE08217FBEC2A9C1B26" box="[301,472,1112,1137]" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE08217FBEC2A9C1B26" box="[301,472,1112,1137]" italics="true" pageId="5" pageNumber="54">Enneopagurus</emphasis>
</taxonomicName>
. However, these three species fit much better in 
<taxonomicName id="4C3C4D963926FFE08726FBEC2FC91B26" box="[1052,1165,1112,1137]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE08726FBEC2FC91B26" box="[1052,1165,1112,1137]" italics="true" pageId="5" pageNumber="54">Turleania</emphasis>
</taxonomicName>
than 
<taxonomicName id="4C3C4D963926FFE087E8FBEC2E391B26" box="[1234,1405,1112,1137]" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE087E8FBEC2E391B26" box="[1234,1405,1112,1137]" italics="true" pageId="5" pageNumber="54">Enneopagurus</emphasis>
</taxonomicName>
in the other characters such as the elongate ocular peduncles with slightly dilated corneas, the right cheliped being much stouter than the left, the right chela with some distinct spines, and the dactyli of the ambulatory legs each armed with a row of well developed corneous spines on the ventral margin. In the sole species of 
<taxonomicName id="4C3C4D963926FFE083ADFB4C2A061A46" box="[151,322,1272,1297]" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE083ADFB4C2A061A46" box="[151,322,1272,1297]" italics="true" pageId="5" pageNumber="54">Enneopagurus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3C4D963926FFE0826AFB4C294B1A46" box="[336,527,1272,1297]" class="Malacostraca" family="Paguridae" genus="Enneopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="garciagomezi">
<emphasis id="B948EA073926FFE0826AFB4C294B1A46" box="[336,527,1272,1297]" italics="true" pageId="5" pageNumber="54">E. garciagomezi</emphasis>
</taxonomicName>
, the ocular peduncles are stout and have strongly dilated corneas, the palm of the right cheliped lacks spines on the dorsomesial margin, and the dactyli of the ambulatory legs are unarmed or have only tiny spinules on the ventral margin. Thus, the generic diagnosis of 
<taxonomicName id="4C3C4D963926FFE0874EFAFC2FA11A36" box="[1140,1253,1352,1377]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B948EA073926FFE0874EFAFC2FA11A36" box="[1140,1253,1352,1377]" italics="true" pageId="5" pageNumber="54">Turleania</emphasis>
</taxonomicName>
is emended to include nine to eleven pairs of quadriserial gills to accommodate McLaughlin &amp; Haig’s species and the two new species from the Ryukyu Islands.
</paragraph>
<paragraph id="8B8336153926FFE083FFFA742A88197E" blockId="5.[151,1436,152,1578]" pageId="5" pageNumber="54">
In the 
<typeStatus id="548788B73926FFE08229FA742A3D1A8D" box="[275,377,1472,1498]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C3C4D963926FFE08299FA742AB01A8E" box="[419,500,1472,1497]" class="Insecta" family="Pyralidae" genus="Laurentia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="balli">
<emphasis id="B948EA073926FFE08299FA742AB01A8E" box="[419,500,1472,1497]" italics="true" pageId="5" pageNumber="54">T. balli</emphasis>
</taxonomicName>
, the dactyli of the ambulatory legs each have a row of small pits on the dorsal surface, which represent setal insertions. The pits show that they had bristles as observed in the 
<taxonomicName id="4C3C4D963926FFE08636FA5C2E3D1956" box="[1292,1401,1512,1537]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="saliens" status="sp. nov.">
<emphasis id="B948EA073926FFE08636FA5C2E3D1956" box="[1292,1401,1512,1537]" italics="true" pageId="5" pageNumber="54">T. saliens</emphasis>
</taxonomicName>
<emphasis id="B948EA073926FFE086BEFA532BFE197E" bold="true" pageId="5" pageNumber="54">
<taxonomicNameLabel id="A27B577C3926FFE086BEFA532BFE197E" pageId="5" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C3C4D963926FFE083C9F9A42AC5197E" box="[243,385,1552,1577]" class="Malacostraca" family="Paguridae" genus="Turleania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp. nov.">
<emphasis id="B948EA073926FFE083C9F9A42AC5197E" box="[243,385,1552,1577]" italics="true" pageId="5" pageNumber="54">T. tenebrosa</emphasis>
</taxonomicName>
<emphasis id="B948EA073926FFE082B3F9BB2A88197E" bold="true" box="[393,460,1551,1577]" pageId="5" pageNumber="54">
<taxonomicNameLabel id="A27B577C3926FFE082B3F9BB2A88197E" box="[393,460,1551,1577]" pageId="5" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>